treatments-xml/data/03/9A/87/039A878CFFFDFFC19005F94CF24F012D.xml

<document id="7023F2EFEF364445E7616BAC5A992F29" ID-DOI="10.11646/zootaxa.1218.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5064858" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625283180981" checkinUser="felipe" docAuthor="Nolan, Matthew J. &amp; Cribb, Thomas H." docDate="2006" docId="039A878CFFFDFFC19005F94CF24F012D" docLanguage="en" docName="zt01218p080.pdf" docOrigin="Zootaxa 1218 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1218.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Phthinomita robertsthomsoni Nolan &amp; Cribb 2006, n. sp." docType="treatment" docUuid="01CBBB79-E84F-4CA1-80AC-1AD82FEED152" docUuidSource="ZooBank" docVersion="5" lastPageNumber="55" masterDocId="FFA3FFF4FFCEFFF7910DFF99F011030B" masterDocTitle="An exceptionally rich complex of Sanguinicolidae von Graff, 1907 (Platyhelminthes: Trematoda) from Siganidae, Labridae and Mullidae (Teleostei: Perciformes) from the Indo-west Pacific Region" masterLastPageNumber="80" masterPageNumber="1" pageNumber="52" updateTime="1698986292376" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="96DC8FAAF400A9854A7A10E41BE13E50">An exceptionally rich complex of Sanguinicolidae von Graff, 1907 (Platyhelminthes: Trematoda) from Siganidae, Labridae and Mullidae (Teleostei: Perciformes) from the Indo-west Pacific Region</mods:title>
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<mods:namePart id="4771BB5C6C9974F35132372845BBCC1E">Nolan, Matthew J.</mods:namePart>
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<mods:namePart id="35D93AB38A5C8CCFE75BF882A1DDE800">Cribb, Thomas H.</mods:namePart>
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<subSubSection id="C3296511FFFDFFC49005F94CF34A05E5" box="[264,859,1748,1775]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8B8C369AFFFDFFC49005F94CF34A05E5" blockId="51.[264,859,1748,1775]" box="[264,859,1748,1775]" pageId="51" pageNumber="52">
<heading id="D0C481F6FFFDFFC49005F94CF34A05E5" bold="true" box="[264,859,1748,1775]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B947EA88FFFDFFC49005F94CF34A05E5" bold="true" box="[264,859,1748,1775]" pageId="51" pageNumber="52">
<taxonomicName id="4C334D19FFFDFFC49005F94CF27B05E4" ID-CoL="8PHH2" authority="Nolan &amp; Cribb, 2006" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[264,618,1749,1775]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="51" pageNumber="52" phylum="Platyhelminthes" rank="species" species="robertsthomsoni" status="sp. nov.">
<emphasis id="B947EA88FFFDFFC49005F94CF27B05E4" bold="true" box="[264,618,1749,1775]" italics="true" pageId="51" pageNumber="52">Phthinomita robertsthomsoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27457F3FFFDFFC4937FF94DF2A405E5" box="[626,693,1748,1774]" pageId="51" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13082A1FFFFDFFC493C9F94DF34205E5" box="[708,851,1748,1774]" captionStart="FIGURES 41–43" captionStartId="52.[264,383,1749,1773]" captionTargetBox="[346,1188,323,1730]" captionTargetId="figure-63@52.[292,1240,295,1744]" captionTargetPageId="52" captionText="FIGURES 41–43. Phthinomita robertsthomsoni n. sp. from the intertrabecular spaces of ventricle (heart) of Siganus argenteus off Lizard Island. 41. Adult. Lateral view, whole mount. 42. Male terminal genitalia, lateral view. 43. Female terminal genitalia, lateral view. Scale­bars: 41,43, 250" figureDoi="http://doi.org/10.5281/zenodo.5064908" httpUri="https://zenodo.org/record/5064908/files/figure.png" pageId="51" pageNumber="52">Figs. 41–43</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<paragraph id="8B8C369AFFFDFFC49005F886F19C046A" blockId="51.[264,1322,1823,1889]" pageId="51" pageNumber="52">
<typeStatus id="54888838FFFDFFC49005F886F1530432" box="[264,322,1823,1849]" pageId="51" pageNumber="52">Type</typeStatus>
host: 
<taxonomicName id="4C334D19FFFDFFC49096F8B9F36E0432" authority="(Quoy &amp; Gaimard)" baseAuthorityName="Quoy &amp; Gaimard" box="[411,895,1823,1849]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B947EA88FFFDFFC49096F8B9F26B0432" box="[411,634,1824,1849]" italics="true" pageId="51" pageNumber="52">Siganus argenteus</emphasis>
(Quoy &amp; Gaimard)
</taxonomicName>
, Forktail Rabbitfish (
<taxonomicName id="4C334D19FFFDFFC4959FF886F5370432" box="[1170,1318,1823,1849]" class="Actinopterygii" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="order">Perciformes</taxonomicName>
: 
<taxonomicName id="4C334D19FFFDFFC49005F8DEF191046A" authorityName="Richardson" authorityYear="1837" box="[264,384,1863,1889]" class="Actinopterygii" family="Siganidae" kingdom="Animalia" order="Perciformes" pageId="51" pageNumber="52" phylum="Chordata" rank="family">Siganidae</taxonomicName>
).
</paragraph>
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<paragraph id="8B8C369AFFFAFFC39005F94CF53A0439" blockId="52.[264,1323,1749,1877]" pageId="52" pageNumber="53">
<emphasis id="B947EA88FFFAFFC39005F94CF1C105E6" bold="true" box="[264,464,1749,1773]" pageId="52" pageNumber="53">FIGURES 41–43.</emphasis>
<taxonomicName id="4C334D19FFFAFFC390DAF94FF30105E6" authority="Nolan &amp; Cribb, 2006" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[471,784,1750,1773]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="52" pageNumber="53" phylum="Platyhelminthes" rank="species" species="robertsthomsoni" status="sp. nov.">
<emphasis id="B947EA88FFFAFFC390DAF94FF30105E6" box="[471,784,1750,1773]" italics="true" pageId="52" pageNumber="53">Phthinomita robertsthomsoni</emphasis>
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<emphasis id="B947EA88FFFAFFC39215F94CF34505E6" bold="true" box="[792,852,1749,1773]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="A27457F3FFFAFFC39215F94CF34505E6" box="[792,852,1749,1773]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from the intertrabecular spaces of ventricle (heart) of 
<taxonomicName id="4C334D19FFFAFFC39077F960F250041B" box="[378,577,1785,1808]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B947EA88FFFAFFC39077F960F250041B" box="[378,577,1785,1808]" italics="true" pageId="52" pageNumber="53">Siganus argenteus</emphasis>
</taxonomicName>
off Lizard Island. 41. Adult. Lateral view, whole mount. 42. Male terminal genitalia, lateral view. 43. Female terminal genitalia, lateral view. 
<emphasis id="B947EA88FFFAFFC39538F882F4A10439" box="[1077,1200,1819,1842]" italics="true" pageId="52" pageNumber="53">Scale­bars:</emphasis>
41,43, 250
</paragraph>
</caption>
<paragraph id="8B8C369AFFFAFFC3901AF8A7F1AD045E" blockId="52.[264,1323,1749,1877]" box="[279,444,1854,1877]" pageId="52" pageNumber="53">
m; 
<quantity id="4CCB9B7FFFFAFFC39037F8A7F1A9045E" box="[314,440,1854,1877]" metricMagnitude="4" metricUnit="m" metricValue="4.21" pageId="52" pageNumber="53" unit="m" value="42100.0">42, 100 m</quantity>
.
</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FE85F3A7023D" blockId="53.[264,1323,284,550]" box="[312,950,284,310]" pageId="53" pageNumber="54">Site in host: Intertrabecular spaces of ventricle (heart).</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FEDDF18B028D" blockId="53.[264,1323,284,550]" pageId="53" pageNumber="54">
<materialsCitation id="3B5B3CC7FFFBFFC29035FEDDF187028D" latitude="-14.666667" location="northern Great Barrier Reef" longLatPrecision="1287" longitude="145.45" municipality="Lizard Island" pageId="53" pageNumber="54" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="54888838FFFBFFC29035FEDDF1630255" box="[312,370,324,350]" pageId="53" pageNumber="54">Type</typeStatus>
locality: 
<collectingMunicipality id="6BE8ACE0FFFBFFC290FBFEDDF28F0255" box="[502,670,324,350]" pageId="53" pageNumber="54">Lizard Island</collectingMunicipality>
, 
<location id="8EEC6041FFFBFFC293B8FEDDF40B0255" LSID="urn:lsid:plazi:treatment:039A878CFFFDFFC19005F94CF24F012D:8EEC6041FFFBFFC293B8FEDDF40B0255" box="[693,1050,324,350]" latitude="-14.666667" longLatPrecision="1287" longitude="145.45" municipality="Lizard Island" name="northern Great Barrier Reef" pageId="53" pageNumber="54" stateProvince="Queensland">northern Great Barrier Reef</location>
(
<geoCoordinate id="EE07505DFFFBFFC2953BFEDDF4860255" box="[1078,1175,324,350]" degrees="14" direction="south" minutes="40" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="-14.666667">14°40’S</geoCoordinate>
<geoCoordinate id="EE07505DFFFBFFC295A4FEDDF50C0255" box="[1193,1309,324,350]" degrees="145" direction="east" minutes="27" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="145.45">145°27’E</geoCoordinate>
), 
<collectingRegion id="49F7F878FFFBFFC29005FEF5F187028D" box="[264,406,364,390]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FE0DF1C902F5" blockId="53.[264,1323,284,550]" pageId="53" pageNumber="54">
<materialsCitation id="3B5B3CC7FFFBFFC29035FE0DF1C502F5" collectingDate="Aug. 2002, Jan. 2003" collectionCode="QM" pageId="53" pageNumber="54" specimenCode="G 225613, G 225614-225622" specimenCount="2" typeStatus="holotype">
Material examined: ex 
<taxonomicName id="4C334D19FFFBFFC29354FE0DF35E02A5" authority=", Lizard" authorityName="Lizard" box="[601,847,404,430]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B947EA88FFFBFFC29354FE0DF27902A6" box="[601,616,404,429]" italics="true" pageId="53" pageNumber="54">S</emphasis>
. 
<emphasis id="B947EA88FFFBFFC29371FE0DF2FF02A6" box="[636,750,404,429]" italics="true" pageId="53" pageNumber="54">argenteus</emphasis>
, Lizard
</taxonomicName>
Island (QLD), 
<collectingDate id="EFC9E9B2FFFBFFC2951AFE0DF53702A5" box="[1047,1318,404,430]" isEnumeration="true" pageId="53" pageNumber="54">
<date id="FF8D105AFFFBFFC2951AFE0DF48A02A5" box="[1047,1179,404,430]" pageId="53" pageNumber="54" value="2002-08">Aug. 2002</date>
, 
<date id="FF8D105AFFFBFFC295A0FE0DF53702A5" box="[1197,1318,404,430]" pageId="53" pageNumber="54" value="2003-01">Jan. 2003</date>
</collectingDate>
, twenty­two specimens (
<typeStatus id="54888838FFFBFFC29349FE25F2A202DD" box="[580,691,444,470]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
no. 
<collectionCode id="ED22AE5FFFFBFFC29201FE25F32A02DD" box="[780,827,444,470]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="53" pageNumber="54" type="Museum">QM</collectionCode>
<specimenCode id="DB959EE1FFFBFFC2925BFE25F3F002DD" box="[854,993,444,470]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="53" pageNumber="54" type="Museum">G 225613</specimenCode>
; 
<typeStatus id="54888838FFFBFFC2950DFE25F47602DD" box="[1024,1127,444,470]" pageId="53" pageNumber="54" type="paratype">Paratype</typeStatus>
nos. 
<collectionCode id="ED22AE5FFFFBFFC295C1FE25F4EA02DD" box="[1228,1275,444,470]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="53" pageNumber="54" type="Museum">QM</collectionCode>
<specimenCode id="DB959EE1FFFBFFC29418FE25F1DD02F5" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="53" pageNumber="54" type="Museum">G 225614–225622</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FD95F22E012D" blockId="53.[264,1323,284,550]" box="[312,575,524,550]" pageId="53" pageNumber="54">Collector: M.J. Nolan.</paragraph>
<paragraph id="8B8C369AFFFBFFC29005FDC5F194017E" blockId="53.[264,1322,604,710]" box="[264,389,604,629]" pageId="53" pageNumber="54">
<emphasis id="B947EA88FFFBFFC29005FDC5F194017E" box="[264,389,604,629]" italics="true" pageId="53" pageNumber="54">Etymology</emphasis>
</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FD1DF31801CD" blockId="53.[264,1322,604,710]" pageId="53" pageNumber="54">
Specific name for our good friend and colleague Mr Ashley Roberts­Thomson who collected the first 
<taxonomicName id="4C334D19FFFBFFC290D6FD35F27B01CE" box="[475,618,684,710]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="53" pageNumber="54" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B947EA88FFFBFFC290D6FD35F1FB01CE" box="[475,490,684,709]" italics="true" pageId="53" pageNumber="54">S</emphasis>
. 
<emphasis id="B947EA88FFFBFFC290F5FD35F27B01CE" box="[504,618,684,709]" italics="true" pageId="53" pageNumber="54">argenteus</emphasis>
</taxonomicName>
of this study.
</paragraph>
<paragraph id="8B8C369AFFFBFFC29005FD65F183001E" blockId="53.[264,1323,764,1430]" box="[264,402,764,789]" pageId="53" pageNumber="54">
<emphasis id="B947EA88FFFBFFC29005FD65F183001E" box="[264,402,764,789]" italics="true" pageId="53" pageNumber="54">Description</emphasis>
</paragraph>
<paragraph id="8B8C369AFFFBFFC29035FCBDF2E1069D" blockId="53.[264,1323,764,1430]" pageId="53" pageNumber="54">Based on 22 whole mounts. With features of genus. Body notched at male genital pore. Intestine; posterior caeca sinuous. Anterior testis originating at posterior end of intercaecal field, margins lobed. Posterior testis tear­shaped. Cirrus­sac tear­shaped. Internal seminal vesicle spherical, occupying most of cirrus­sac, positioned ventrally; ejaculatory duct straight; prostatic cells large. Ovary oblanceolate, overlapping posterior margin of anterior testis. Oviduct originating at centre of posterior margin of ovary, sinuous, dorsal to vas deferens, entering oötype postero­dorsally. Vitelline duct forming anterior to cirrus­sac, passing posteriorly sinuously, dextral to vas deferens; vitelline reservoir forming anterior to anterior margin of cirrus­sac, entering oötype posterodextrally. Oötype ovoid. Mehlis’ gland extending anteriorly to posterior margin of cirrussac, extending posteriorly to mid­section of posterior testis. Uterus extending from oötype, slightly sinuous, sinistral to oviduct. Uterine chamber forming anterior to posterior margin of ovary and anterior testis, ventral to ovary, sinuous, curving dorsally posteriorly to female pore. Vitelline follicles extending anteriorly past intestinal bifurcation, extending posteriorly to level female genital pore, sinistral and dextral to oesophagus, filling intercaecal field, dextral to anterior testis.</paragraph>
<paragraph id="8B8C369AFFFBFFC29005FA55F17F06EE" blockId="53.[264,1323,1484,1870]" box="[264,366,1484,1509]" pageId="53" pageNumber="54">
<emphasis id="B947EA88FFFBFFC29005FA55F17F06EE" box="[264,366,1484,1509]" italics="true" pageId="53" pageNumber="54">Remarks</emphasis>
</paragraph>
<paragraph id="8B8C369AFFFBFFC19035FA6DF4EE0255" blockId="53.[264,1323,1484,1870]" lastBlockId="54.[264,1323,284,550]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
This species possesses the characters differentially diagnostic of the genus 
<taxonomicName id="4C334D19FFFBFFC29005F985F188053E" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[264,409,1564,1589]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="53" pageNumber="54" phylum="Platyhelminthes" rank="genus">
<emphasis id="B947EA88FFFBFFC29005F985F188053E" box="[264,409,1564,1589]" italics="true" pageId="53" pageNumber="54">Phthinomita</emphasis>
</taxonomicName>
and is here identified as a new species in that genus. 
<taxonomicName id="4C334D19FFFBFFC29597F985F1D60556" authorityName="Nolan &amp; Cribb" authorityYear="2006" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="53" pageNumber="54" phylum="Platyhelminthes" rank="species" species="robertsthomsoni">
<emphasis id="B947EA88FFFBFFC29597F985F1D60556" italics="true" pageId="53" pageNumber="54">Phthinomita robertsthomsoni</emphasis>
</taxonomicName>
is immediately distinguished by the combined possession of a body 3784–5706 (4851) long, a body 192 wide on average, an anterior testis 38–62% of the body length and 14.8–31.3 times longer than the posterior one, a posterior testis 15–46% of the body width, a cirrus­sac 64–96 (78) x 45–80 (64), an ovary 87–183 (138) x 64–144 (103) and a uterine chamber 257–385 (319) x 39–80 (59). The species differs further in having an anterior testis with the anterior margin in the posterior end of the intercaecal field, an ovary with an anterior margin lateral to the posterior margin of the anterior testis, a uterine chamber originating anteriorly to the posterior margin of both the anterior testis and the ovary and vitelline follicles extending anteriorly past the intestinal bifurcation.
</paragraph>
<paragraph id="8B8C369AFFF8FFC19035FEF5F24F012D" blockId="54.[264,1323,284,550]" pageId="54" pageNumber="55">
There are in addition, 20–29 base differences (5.8–8.6% sequence divergence) between the ITS2 rDNA sequence of 
<taxonomicName id="4C334D19FFF8FFC193DDFE0DF3BF02A6" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[720,942,404,429]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="54" pageNumber="55" phylum="Platyhelminthes" rank="species" species="robertsthomsoni">
<emphasis id="B947EA88FFF8FFC193DDFE0DF3BF02A6" box="[720,942,404,429]" italics="true" pageId="54" pageNumber="55">P. robertsthomsoni</emphasis>
</taxonomicName>
and the remaining 
<taxonomicName id="4C334D19FFF8FFC19597FE0DF14D02DD" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="54" pageNumber="55" phylum="Platyhelminthes" rank="species" species="undetermined">
<emphasis id="B947EA88FFF8FFC19597FE0DF53A02A6" box="[1178,1323,404,429]" italics="true" pageId="54" pageNumber="55">Phthinomita</emphasis>
species
</taxonomicName>
sequenced here. Between the sequences from 
<taxonomicName id="4C334D19FFF8FFC19286FE25F47902DE" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[907,1128,444,469]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="54" pageNumber="55" phylum="Platyhelminthes" rank="species" species="robertsthomsoni">
<emphasis id="B947EA88FFF8FFC19286FE25F47902DE" box="[907,1128,444,469]" italics="true" pageId="54" pageNumber="55">P. robertsthomsoni</emphasis>
</taxonomicName>
(two replicates) from 
<taxonomicName id="4C334D19FFF8FFC19044FE7DF1C802F6" box="[329,473,484,509]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B947EA88FFF8FFC19044FE7DF1C802F6" box="[329,473,484,509]" italics="true" pageId="54" pageNumber="55">S. argenteus</emphasis>
</taxonomicName>
off Lizard Island and 
<taxonomicName id="4C334D19FFF8FFC193EDFE7DF35002F6" authorityName="Nolan &amp; Cribb" authorityYear="2006" box="[736,833,484,509]" class="Trematoda" family="Aporocotylidae" genus="Phthinomita" kingdom="Animalia" order="Diplostomida" pageId="54" pageNumber="55" phylum="Platyhelminthes" rank="species" species="sasali">
<emphasis id="B947EA88FFF8FFC193EDFE7DF35002F6" box="[736,833,484,509]" italics="true" pageId="54" pageNumber="55">P. sasali</emphasis>
</taxonomicName>
(one replicate) from 
<taxonomicName id="4C334D19FFF8FFC19531FE7DF4A902F6" box="[1084,1208,484,509]" class="Actinopterygii" family="Siganidae" genus="Siganus" kingdom="Animalia" order="Perciformes" pageId="54" pageNumber="55" phylum="Chordata" rank="species" species="doliatus">
<emphasis id="B947EA88FFF8FFC19531FE7DF4A902F6" box="[1084,1208,484,509]" italics="true" pageId="54" pageNumber="55">S. doliatus</emphasis>
</taxonomicName>
off 
<collectingCountry id="F324760AFFF8FFC195E4FE7DF53B02F5" box="[1257,1322,484,510]" name="Palau" pageId="54" pageNumber="55">Palau</collectingCountry>
there are 20 base differences.
</paragraph>
</subSubSection>
</treatment>
</document>