290 lines
40 KiB
XML
290 lines
40 KiB
XML
<document id="DF768E7C0B222343FF5969DCA249F961" ID-DOI="10.11646/zootaxa.4763.1.6" ID-GBIF-Dataset="181f12d1-3c7c-4f39-bc5c-eb42a9bb4913" ID-ISSN="1175-5326" ID-Zenodo-Dep="3744071" ID-ZooBank="9941C5F0-5EEA-4303-AB04-230BCED735CB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586325599653" checkinUser="plazi" docAuthor="Kataev, Boris M. & Muilwijk, Jan" docDate="2020" docId="03821331FFD3FFADFF691EC6FB3E15E8" docLanguage="en" docName="zootaxa.4763.1.6.pdf" docOrigin="Zootaxa 4763 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Disciferella Kataev & Muilwijk 2020, gen. n." docType="treatment" docUuid="07580209-929F-4C1B-B63A-2AC864923E7B" docUuidSource="ZooBank" docVersion="6" lastPageNumber="75" masterDocId="FFBB6B49FFD2FFAFFFFE1A74FFD11237" masterDocTitle="A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini)" masterLastPageNumber="84" masterPageNumber="73" pageNumber="74" updateTime="1698736403035" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C40E4489BB1C15BEBFEC4420DA08EF79">
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<mods:title id="68D88533C87DD0948894D91703A0F38C">A remarkable new lebiine genus with two new species from Pakistan, Afghanistan and Iran (Coleoptera: Carabidae: Lebiini)</mods:title>
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<mods:namePart id="E55E7B24B83517D6D626C28ED81B822F">Kataev, Boris M.</mods:namePart>
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<mods:namePart id="EF2C488A29EDE39F88F2150159FD8C55">Muilwijk, Jan</mods:namePart>
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<mods:title id="F640F73034332DCAA2A91B46445142E4">Zootaxa</mods:title>
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<mods:part id="6183F7FD3BCCE220F36BC9E41EDAB236">
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<mods:date id="D8633957D9C2FEFB3E6187105AF0168D">2020</mods:date>
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<treatment id="03821331FFD3FFADFF691EC6FB3E15E8" ID-DOI="http://doi.org/10.5281/zenodo.3809629" ID-GBIF-Taxon="163538457" ID-Zenodo-Dep="3809629" LSID="urn:lsid:zoobank.org:act:07580209-929F-4C1B-B63A-2AC864923E7B" httpUri="http://treatment.plazi.org/id/03821331FFD3FFADFF691EC6FB3E15E8" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74">
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<paragraph id="8B94A227FFD3FFAEFF691EC6FEAC16FA" blockId="1.[151,919,1202,1264]" box="[151,381,1202,1229]" pageId="1" pageNumber="74">
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<heading id="D0DC154BFFD3FFAEFF691EC6FEAC16FA" bold="true" box="[151,381,1202,1229]" fontSize="11" level="1" pageId="1" pageNumber="74" reason="1">
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<emphasis id="B95F7E35FFD3FFAEFF691EC6FEAC16FA" bold="true" box="[151,381,1202,1229]" pageId="1" pageNumber="74">
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<taxonomicName id="4C2BD9A4FFD3FFAEFF691EC6FEF216FA" ID-CoL="8LYP6" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[151,291,1202,1229]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="74" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95F7E35FFD3FFAEFF691EC6FEF216FA" bold="true" box="[151,291,1202,1229]" italics="true" pageId="1" pageNumber="74">Disciferella</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26CC34EFFD3FFAEFED51EC7FEAC16FA" box="[299,381,1203,1229]" pageId="1" pageNumber="74" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8B94A227FFD3FFAEFF691EACFC4616C7" blockId="1.[151,919,1202,1264]" box="[151,919,1240,1264]" pageId="1" pageNumber="74">
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urn:lsid:zoobank.org:act:
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<uuid id="FF8D98F2FFD3FFAEFE621EACFC4616C7" box="[412,919,1240,1264]" pageId="1" pageNumber="74">07580209-929F-4C1B-B63A-2AC864923E7B</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331F1ACFFD3FFAEFF691F6FFDEF1704" box="[151,574,1306,1331]" pageId="1" pageNumber="74" type="materials_examined">
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<paragraph id="8B94A227FFD3FFAEFF691F6FFDEF1704" blockId="1.[151,574,1306,1331]" box="[151,574,1306,1331]" pageId="1" pageNumber="74">
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<typeStatus id="54901C85FFD3FFAEFF691F6FFF181704" box="[151,201,1307,1331]" pageId="1" pageNumber="74">Type</typeStatus>
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species:
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<taxonomicName id="4C2BD9A4FFD3FFAEFED81F6EFE2F1705" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[294,510,1306,1330]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="74" phylum="Arthropoda" rank="species" species="kabakovi" status="sp. nov.">
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<emphasis id="B95F7E35FFD3FFAEFED81F6EFE2F1705" box="[294,510,1306,1330]" italics="true" pageId="1" pageNumber="74">Disciferella kabakovi</emphasis>
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</taxonomicName>
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<emphasis id="B95F7E35FFD3FFAEFDFB1F6FFDEF1704" bold="true" box="[517,574,1307,1331]" pageId="1" pageNumber="74">
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<taxonomicNameLabel id="A26CC34EFFD3FFAEFDFB1F6FFDEF1704" box="[517,574,1307,1331]" pageId="1" pageNumber="74" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331F1ACFFD3FFAEFF691F14FB211578" pageId="1" pageNumber="74" type="diagnosis">
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<paragraph id="8B94A227FFD3FFAEFF691F14FB211578" blockId="1.[151,1437,1376,2015]" pageId="1" pageNumber="74">
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<emphasis id="B95F7E35FFD3FFAEFF691F14FEC3174C" bold="true" box="[151,274,1376,1403]" pageId="1" pageNumber="74">Diagnosis.</emphasis>
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Recognizable by the combination of the following distinctive characters: size rather small (body length
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<quantity id="4CD30FC2FFD3FFAEFF691FF0FEF717A8" box="[151,294,1412,1439]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="5.2" metricValueMin="4.4" pageId="1" pageNumber="74" unit="mm" value="4.8" valueMax="5.2" valueMin="4.4">4.4–5.2 mm</quantity>
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); body, including eyes, covered with very fine, short and rather dense inclined setae (hairs); head with moderately deep neck constriction and without suborbital setae; mandibles elongate; mentum and submentum completely fused; ligular sclerite with two ventroapical setae; apical palpomeres not widened apically; antennae densely pubescent from antennomere 4, with sparser setae on antennomere 3 and with a few fine setae on two basal antennomeres; pronotal base projecting distally of basal angles; pronotal sides each with a long seta at apex of basal angle and with a short seta at apical angle, with or without anterior lateral seta; proepipleura (prohypomera) basally visible from above; elytra very wide, discoidal, with apices truncate and with striae strongly obliterate, not evident; epipleura of elytra incomplete, not reaching apical angle and not visible from above; abdominal sternites (ventrites) and metasternum normal (three basal abdominal sternites not fused, metasternum not shortened, and metepisternum much longer than wide); protibia without spines along outer margin and with only anterior spur apically (posterior spur absent); apical spurs of meso- and metatibia not longer than half of tarsomere 1; tarsi dorsally setose, slen- der; tarsomere 4 not bilobe; tarsal claws smooth; male protarsomeres 1–4 slightly dilated and covered with sparse spongy adhesive vestiture; gonosubcoxite without spines, and gonocoxite short and wide, glabrous.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331F1ACFFD3FFADFF391D2DFB3E15E8" lastPageId="2" lastPageNumber="75" pageId="1" pageNumber="74" type="etymology">
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<paragraph id="8B94A227FFD3FFAEFF391D2DFE3D158C" blockId="1.[151,1437,1376,2015]" pageId="1" pageNumber="74">
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<emphasis id="B95F7E35FFD3FFAEFF391D2DFE9F1544" bold="true" box="[199,334,1881,1907]" pageId="1" pageNumber="74">Etymology.</emphasis>
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The generic name is a feminine Latin compound noun based on the words
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<emphasis id="B95F7E35FFD3FFAEFB711D2DFB041544" box="[1167,1237,1881,1907]" italics="true" pageId="1" pageNumber="74">discus</emphasis>
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(disk or disc) and
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<emphasis id="B95F7E35FFD3FFAEFF691D09FF1215A0" box="[151,195,1917,1943]" italics="true" pageId="1" pageNumber="74">fĕro</emphasis>
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(to bring, to bear), plus the diminutive suffix
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<emphasis id="B95F7E35FFD3FFAEFD3A1D09FD2615A0" box="[708,759,1917,1943]" italics="true" pageId="1" pageNumber="74">-ella</emphasis>
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, and referring to the small body size and the discoidal shape of the elytra of the new genus.
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</paragraph>
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<paragraph id="8B94A227FFD3FFAEFF391DB0FADD15E8" blockId="1.[151,1437,1376,2015]" box="[199,1292,1988,2015]" pageId="1" pageNumber="74">
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<emphasis id="B95F7E35FFD3FFAEFF391DB0FEB615E8" bold="true" box="[199,359,1988,2015]" pageId="1" pageNumber="74">Composition.</emphasis>
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This new genus includes two new allopatric species described in the present paper.
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</paragraph>
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<paragraph id="8B94A227FFD0FFADFF391AECFAEA1704" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="75">
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<emphasis id="B95F7E35FFD0FFADFF391AECFE061284" bold="true" box="[199,471,152,179]" pageId="2" pageNumber="75">Comparative remarks.</emphasis>
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The following characteristics of the new genus suggest that it belongs to the tribe
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<taxonomicName id="4C2BD9A4FFD0FFADFA8B1AEDFF1E12E0" authorityName="Bonelli" authorityYear="1810" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="tribe" tribe="Lebiini">Lebiini</taxonomicName>
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: 1) antennae densely pubescent from antennomere 4; 2) head not pedunculate posteriorly; 3) protibia without spines along outer margin; 4) meso- and metatibial apical spurs short; and 5) gonosubcoxite lacking spines and setae. Within this tribe,
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<taxonomicName id="4C2BD9A4FFD0FFADFE561B71FDFA1328" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[424,555,261,287]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95F7E35FFD0FFADFE561B71FDFA1328" box="[424,555,261,287]" italics="true" pageId="2" pageNumber="75">Disciferella</emphasis>
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</taxonomicName>
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<emphasis id="B95F7E35FFD0FFADFDCA1B71FD541328" bold="true" box="[564,645,261,287]" pageId="2" pageNumber="75">
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<taxonomicNameLabel id="A26CC34EFFD0FFADFDCA1B71FD541328" box="[564,645,261,287]" pageId="2" pageNumber="75" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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should be treated as a member of
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<taxonomicName id="4C2BD9A4FFD0FFADFBEC1B71FF1F1374" authority="sensu Bouchard et al., 2011" authorityName="Bouchard" authorityYear="2011" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">
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Dromiusina
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<emphasis id="B95F7E35FFD0FFADFB611B72FB0C1328" box="[1183,1245,262,287]" italics="true" pageId="2" pageNumber="75">sensu</emphasis>
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<bibRefCitation id="EFBADFD6FFD0FFADFB181B71FF1F1374" author="Bouchard, P. & Bousquet, Y. & Davies, A. E. & Alonso-Zarazaga, M. A. & Lawrence, J. F. & Lyal, Ch. H. C. & Newton, A. F. & Reid, Ch. A. M. & Schmitt, M. & Slipinski, S. A. & Smith, A. B. T." pageId="2" pageNumber="75" pagination="1 - 972" refId="ref7104" refString="Bouchard, P., Bousquet, Y., Davies, A. E., Alonso-Zarazaga, M. A., Lawrence, J. F., Lyal, Ch. H. C., Newton, A. F., Reid, Ch. A. M., Schmitt, M., Slipinski, S. A. & Smith, A. B. T. (2011) Family-group names in Coleoptera (Insecta). ZooKeys, 88, 1 - 972. https: // doi. org / 10.3897 / zookeys. 88.807" type="journal article" year="2011">
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Bouchard
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<emphasis id="B95F7E35FFD0FFADFAA21B72FA471328" box="[1372,1430,261,287]" italics="true" pageId="2" pageNumber="75">et al.</emphasis>
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, 2011
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</bibRefCitation>
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</taxonomicName>
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(including Singilina Jeannel 1949, Lionychina Jeannel 1948, and Metadromiina
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<bibRefCitation id="EFBADFD6FFD0FFADFB8E1B5DFAE91374" author="Basilewsky, P." box="[1136,1336,297,323]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
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) since it possesses the following distinctive characters of this subtribe (e. g.
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<bibRefCitation id="EFBADFD6FFD0FFADFC661B39FBCE1350" author="Habu, A." box="[920,1055,333,359]" pageId="2" pageNumber="75" refId="ref7334" refString="Habu, A. (1967) Carabidae Truncatipennes group (Insecta: Coleoptera). Fauna Japonica. Biogeographical Society of Japan, Tokyo, xiv + 338 pp., 27 pls." type="book" year="1967">Habu, 1967</bibRefCitation>
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,
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<bibRefCitation id="EFBADFD6FFD0FFADFBD21B39FBB81350" author="Habu, A." box="[1068,1129,333,359]" pageId="2" pageNumber="75" pagination="105 - 146" refId="ref7369" refString="Habu, A. (1983) Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (3) (Coleoptera, Carabidae). The Entomological Review of Japan, 38, 105 - 146." type="journal article" year="1983">1983</bibRefCitation>
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): 1) head ventrally without suborbital setae; 2) three basal abdominal sternites normal, not fused; 3) tarsomere 4 not bilobe; and 4) gonocoxite not slender nor tapering. Based on the mentum and submentum completely fused, the new genus should formally be included in the
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<taxonomicName id="4C2BD9A4FFD0FFADFE9B1BCEFE1713E4" box="[357,454,442,467]" class="Insecta" family="Carabidae" genus="Dromius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
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<emphasis id="B95F7E35FFD0FFADFE9B1BCEFE1713E4" box="[357,454,442,467]" italics="true" pageId="2" pageNumber="75">Dromius</emphasis>
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</taxonomicName>
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genus-group
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<emphasis id="B95F7E35FFD0FFADFDA11BCEFD4C13E4" box="[607,669,442,467]" italics="true" pageId="2" pageNumber="75">sensu</emphasis>
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||
<bibRefCitation id="EFBADFD6FFD0FFADFD5A1BCDFC6013E4" author="Ball, G. E. & Bousquet, Y." box="[676,945,441,467]" pageId="2" pageNumber="75" pagination="32 - 132" refId="ref6839" refString="Ball, G. E. & Bousquet, Y. (2000) Carabidae Latreille, 1810. In: Arnett, R. H. Jr. & Thomas, M. C. (Eds.), American beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 32 - 132." type="book chapter" year="2000">Ball & Bousquet (2000)</bibRefCitation>
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||
(=
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<taxonomicName id="4C2BD9A4FFD0FFADFC291BCDFB8A13E4" box="[983,1115,441,467]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
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in narrow sense; = Dromiini
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<emphasis id="B95F7E35FFD0FFADFF691BAAFF0413C0" box="[151,213,478,503]" italics="true" pageId="2" pageNumber="75">sensu</emphasis>
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFF231BA9FE7613C0" author="Basilewsky, P." box="[221,423,477,503]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
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). Additional characteristics of this group, which are also present in the new genus, are: 1) mandibles not widened and not arcuately curved basally, with narrow scrobe; 2) apical palpomeres not widened apically, fusiform; 3) epipleura incomplete, not reaching apical angle and not visible from above; and 4) tarsi slender. However, the very characteristic body shape and the peculiar set of the other morphological character states make it difficult to specify the position of this genus and indicate its closely related genera. Within
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB511819FAE710B0" box="[1199,1334,621,647]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
|
||
, it is distinctly recognized already by the very wide, discoidal elytra lacking striae and with widely explanate margins. The peculiar set of its distinctive features also includes: 1) dorsal and ventral surfaces of body covered with very fine, short setae; 2) labium with acute median tooth and with two ventroapical setae on ligular sclerite; 3) three basal antennomeres with sparse setation; 4) pronotum with a short seta at apical angle; 5) proepipleura basally visible in dorsal view; 6) tarsal claws smooth; and 7) male protarsomeres 1–4 slightly dilated and covered with ventral adhesive vestiture. In body pubescence,
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFDBB1931FD191168" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[581,712,837,863]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B95F7E35FFD0FFADFDBB1931FD191168" box="[581,712,837,863]" italics="true" pageId="2" pageNumber="75">Disciferella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F7E35FFD0FFADFD2E1931FCF11168" bold="true" box="[720,800,837,863]" pageId="2" pageNumber="75">
|
||
<taxonomicNameLabel id="A26CC34EFFD0FFADFD2E1931FCF11168" box="[720,800,837,863]" pageId="2" pageNumber="75" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to the enigmatic monotypic dromiusine genus
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFF69191DFE0A11B4" authority="Bedel, 1898" authorityName="Bedel" authorityYear="1898" box="[151,475,873,899]" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFF69191DFE9B11B4" box="[151,330,873,899]" italics="true" pageId="2" pageNumber="75">Brachynopterus</emphasis>
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFEAF191DFE0A11B4" author="Bedel, L." box="[337,475,873,899]" pageId="2" pageNumber="75" pagination="241 - 242" refId="ref7014" refString="Bedel, L. (1898) Diagnoses de deux genres nouveaux de carabiques du Sahara septentrional. Bulletin de la Societe Entomologique de France, 1898, 241 - 242." type="journal article" year="1898">Bedel, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
, described from one female specimen collected in
|
||
<collectingCountry id="F33CE2B7FFD0FFADFBF0191DFBB411B4" box="[1038,1125,873,899]" name="Algeria" pageId="2" pageNumber="75">Algeria</collectingCountry>
|
||
, and known to us only from the rather brief original and subsequent additional, also very brief, descriptions, probably of the same specimen (
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFF6119C5FEFA11FC" author="Bedel, L." box="[159,299,944,971]" pageId="2" pageNumber="75" pagination="1895 - 1925" refId="ref7046" refString="Bedel, L. (1905) Catalogue raisonne des coleopteres du nord de l'Afrique. (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, pp. 229 - 252. [1895 - 1925]" type="book chapter" year="1905">Bedel, 1905</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFEC619C5FE0C11FC" author="Jedlicka, A." box="[312,477,945,971]" pageId="2" pageNumber="75" refId="ref7480" refString="Jedlicka, A. (1941) Versuch einer Monographie der pal. Carabiden-Gattungen mit abgestutzten Flugeldecken (Truncatipennen) mit Berucksichtigung der indischen Fauna. Mit 2 Tabellen. Praha, 27 pp. + 2 pls." type="book" year="1941">Jedlička, 1941</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFE1719C5FD6211FC" author="Basilewsky, P." box="[489,691,945,971]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
|
||
), and from two photographs (in dorsal view) of the
|
||
<typeStatus id="54901C85FFD0FFADFB0219C5FA8F11FC" box="[1276,1374,945,971]" pageId="2" pageNumber="75" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFA7A19C6FEA811D8" authority="Bedel, 1898" authorityName="Bedel" authorityYear="1898" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="B95F7E35FFD0FFADFA7A19C6FF3711D8" italics="true" pageId="2" pageNumber="75">B. rufulus</emphasis>
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFF1019A1FEA811D8" author="Bedel, L." box="[238,377,981,1007]" pageId="2" pageNumber="75" pagination="241 - 242" refId="ref7014" refString="Bedel, L. (1898) Diagnoses de deux genres nouveaux de carabiques du Sahara septentrional. Bulletin de la Societe Entomologique de France, 1898, 241 - 242." type="journal article" year="1898">Bedel, 1898</bibRefCitation>
|
||
</taxonomicName>
|
||
. Although
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFE0A19A1FD7611D8" authorityName="Bedel" authorityYear="1898" box="[500,679,981,1007]" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFE0A19A1FD7611D8" box="[500,679,981,1007]" italics="true" pageId="2" pageNumber="75">Brachynopterus</emphasis>
|
||
</taxonomicName>
|
||
is treated by all authors as a member of
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB8F19A1FB2911D8" box="[1137,1272,981,1007]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
|
||
, its systematic position within this subtribe is obscure. Like the new genus,
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFCAD198DFBD71624" authorityName="Bedel" authorityYear="1898" box="[851,1030,1017,1043]" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFCAD198DFBD71624" box="[851,1030,1017,1043]" italics="true" pageId="2" pageNumber="75">Brachynopterus</emphasis>
|
||
</taxonomicName>
|
||
has a seta in apical pronotal angle, elytra not striate, tarsal claws smooth and antennomeres with a narrow glabrous strip on outer margin, but it is distinctly differing at least in having pronotum markedly narrowed behind basal angles, proepipleura basally not visible in dorsal view, elytra much narrower, more coarsely punctate and more narrowly explanate along sides, and gonocoxite elongate, slightly arcuate and narrowed apically. By the latter character,
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFBB61EFDFB2A1694" authorityName="Bedel" authorityYear="1898" box="[1096,1275,1161,1187]" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFBB61EFDFB2A1694" box="[1096,1275,1161,1187]" italics="true" pageId="2" pageNumber="75">Brachynopterus</emphasis>
|
||
</taxonomicName>
|
||
markedly differs from other
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFEB41ED9FD6316F0" authority="(Basilewsky, 1984)" baseAuthorityName="Basilewsky" baseAuthorityYear="1984" box="[330,690,1197,1223]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">
|
||
Dromiusina (
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFE201ED9FD7B16F0" author="Basilewsky, P." box="[478,682,1197,1223]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. It is also worthy of note that according to the habitus drawing of
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFF691EA6FED716DC" authorityName="Bedel" authorityYear="1898" box="[151,262,1233,1259]" class="Insecta" family="Carabidae" genus="Brachynopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="species" species="rufulus">
|
||
<emphasis id="B95F7E35FFD0FFADFF691EA6FED716DC" box="[151,262,1233,1259]" italics="true" pageId="2" pageNumber="75">B. rufulus</emphasis>
|
||
</taxonomicName>
|
||
included in the original description of this taxon (
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFCBE1EA5FBD716DC" author="Bedel, L." box="[832,1030,1233,1259]" pageId="2" pageNumber="75" pagination="241 - 242" refId="ref7014" refString="Bedel, L. (1898) Diagnoses de deux genres nouveaux de carabiques du Sahara septentrional. Bulletin de la Societe Entomologique de France, 1898, 241 - 242." type="journal article" year="1898">Bedel, 1898: 241</bibRefCitation>
|
||
), its mandibles are more rounded at sides than those in the new genus. Unfortunately, there is no available information about the characters visible from ventral side (structure of the labium, presence or absence of the suborbital setae, abdominal sternites, etc).
|
||
</paragraph>
|
||
<paragraph id="8B94A227FFD0FFADFF391F49FCB614AC" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="75">
|
||
The new genus also somewhat reminds the monotypic genus
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFC811F49FB241760" authority="Rosenhauer, 1856" authorityName="Rosenhauer" authorityYear="1856" box="[895,1269,1340,1367]" class="Insecta" family="Carabidae" genus="Pseudotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFC811F49FBF21760" box="[895,1059,1341,1367]" italics="true" pageId="2" pageNumber="75">Pseudotrechus</emphasis>
|
||
Rosenhauer, 1856
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F33CE2B7FFD0FFADFF691F15FF0B174C" box="[151,218,1377,1403]" name="Spain" pageId="2" pageNumber="75">Spain</collectingCountry>
|
||
,
|
||
<collectingCountry id="F33CE2B7FFD0FFADFF1B1F15FE9B174C" box="[229,330,1377,1403]" name="Morocco" pageId="2" pageNumber="75">Morocco</collectingCountry>
|
||
and
|
||
<collectingCountry id="F33CE2B7FFD0FFADFE801F15FE04174C" box="[382,469,1377,1403]" name="Algeria" pageId="2" pageNumber="75">Algeria</collectingCountry>
|
||
, which also has wide, discoidal elytra without striae and with very fine and short, scattered setae on disc. Some authors (e. g.
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFDA51FF1FCB317A8" author="Ball, G. E. & Bousquet, Y." box="[603,866,1413,1439]" pageId="2" pageNumber="75" pagination="32 - 132" refId="ref6839" refString="Ball, G. E. & Bousquet, Y. (2000) Carabidae Latreille, 1810. In: Arnett, R. H. Jr. & Thomas, M. C. (Eds.), American beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, Florida, pp. 32 - 132." type="book chapter" year="2000">Ball & Bousquet, 2000</bibRefCitation>
|
||
) include this genus in
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB9D1FF1FB3B17A8" baseAuthorityName="Basilewsky" baseAuthorityYear="1984" box="[1123,1258,1413,1439]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
|
||
, but it is distinguished from members of this subtribe by many peculiar characters, which are also absent in
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB451FDDFAEF17F4" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[1211,1342,1449,1475]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B95F7E35FFD0FFADFB451FDDFAEF17F4" box="[1211,1342,1449,1475]" italics="true" pageId="2" pageNumber="75">Disciferella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F7E35FFD0FFADFAB81FDDFA4717F4" bold="true" box="[1350,1430,1449,1475]" pageId="2" pageNumber="75">
|
||
<taxonomicNameLabel id="A26CC34EFFD0FFADFAB81FDDFA4717F4" box="[1350,1430,1449,1475]" pageId="2" pageNumber="75" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, among them the metasternum very short, the metepisternum wider than long and three basal abdominal sternites fused; for this reason
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFE641F85FDEF143C" authorityName="Rosenhauer" authorityYear="1856" box="[410,574,1521,1547]" class="Insecta" family="Carabidae" genus="Pseudotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFE641F85FDEF143C" box="[410,574,1521,1547]" italics="true" pageId="2" pageNumber="75">Pseudotrechus</emphasis>
|
||
</taxonomicName>
|
||
is usually treated as a member of the monotypic subtribe
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB0F1F85FE8C1418" authority="Basilewsky, 1984" authorityName="Basilewsky" authorityYear="1984" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Pseudotrechina">
|
||
Pseudotrechina
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFF691C61FE8C1418" author="Basilewsky, P." box="[151,349,1557,1583]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
(e. g.
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFE5A1C61FDBD1418" author="Basilewsky, P." box="[420,620,1557,1583]" pageId="2" pageNumber="75" pagination="525 - 559" refId="ref6978" refString="Basilewsky, P. (1984) Essai d'une classification supragenerique naturelle des carabides lebiens d'Afrique et de Madagascar (Coleoptera Carabidae Lebiinae). Revue de Zoologie Africaine, 98, 525 - 559." type="journal article" year="1984">Basilewsky, 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFD861C61FCC01418" author="Lorenz, W." box="[632,785,1556,1583]" pageId="2" pageNumber="75" refId="ref7578" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the world (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Author, Titzing, (ii) + 530 pp." type="book" year="2005">Lorenz, 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFCE31C61FBC01418" author="Bouchard, P. & Bousquet, Y. & Davies, A. E. & Alonso-Zarazaga, M. A. & Lawrence, J. F. & Lyal, Ch. H. C. & Newton, A. F. & Reid, Ch. A. M. & Schmitt, M. & Slipinski, S. A. & Smith, A. B. T." box="[797,1041,1557,1583]" pageId="2" pageNumber="75" pagination="1 - 972" refId="ref7104" refString="Bouchard, P., Bousquet, Y., Davies, A. E., Alonso-Zarazaga, M. A., Lawrence, J. F., Lyal, Ch. H. C., Newton, A. F., Reid, Ch. A. M., Schmitt, M., Slipinski, S. A. & Smith, A. B. T. (2011) Family-group names in Coleoptera (Insecta). ZooKeys, 88, 1 - 972. https: // doi. org / 10.3897 / zookeys. 88.807" type="journal article" year="2011">
|
||
Bouchard
|
||
<emphasis id="B95F7E35FFD0FFADFC6F1C62FC131418" box="[913,962,1557,1583]" italics="true" pageId="2" pageNumber="75">et al</emphasis>
|
||
., 2011
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFBADFD6FFD0FFADFBE31C61FB631418" author="Kabak, I. I." box="[1053,1202,1557,1583]" pageId="2" pageNumber="75" pagination="579 - 625" refId="ref7519" refString="Kabak, I. I. (2017) Tribe Lebiini Bonelli, 1810. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Revised and Updated Edition. Archostemata-Myxophaga-Adephaga. Brill, Leiden / Boston, pp. 579 - 625." type="book chapter" year="2017">Kabak, 2017</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB3A1C61FAB91418" authorityName="Rosenhauer" authorityYear="1856" box="[1220,1384,1557,1583]" class="Insecta" family="Carabidae" genus="Pseudotrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFB3A1C61FAB91418" box="[1220,1384,1557,1583]" italics="true" pageId="2" pageNumber="75">Pseudotrechus</emphasis>
|
||
</taxonomicName>
|
||
also differs from the new genus in many other sufficient characters including head without neck constriction, mandibles short, arcuately curved, pronotum very wide, with very prominent acute apical angles lacking seta, protibia with posterior spur, tarsal claws basally with two small denticles, etc.
|
||
</paragraph>
|
||
<paragraph id="8B94A227FFD0FFADFF391CD1FB3E15E8" blockId="2.[151,1437,152,2015]" pageId="2" pageNumber="75">
|
||
In discoidal elytra,
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFE561CD1FDFA1488" authority="Kataev & Muilwijk, 2020" authorityName="Kataev & Muilwijk" authorityYear="2020" box="[424,555,1701,1727]" class="Insecta" family="Carabidae" genus="Disciferella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B95F7E35FFD0FFADFE561CD1FDFA1488" box="[424,555,1701,1727]" italics="true" pageId="2" pageNumber="75">Disciferella</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95F7E35FFD0FFADFDC81CD1FD591488" bold="true" box="[566,648,1701,1727]" pageId="2" pageNumber="75">
|
||
<taxonomicNameLabel id="A26CC34EFFD0FFADFDC81CD1FD591488" box="[566,648,1701,1727]" pageId="2" pageNumber="75" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also somewhat similar to the South Palaearctic genus
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFADE1CD2FE9214D4" authority="Semenov, 1889" authorityName="Semenov" authorityYear="1889" class="Insecta" family="Carabidae" genus="Discoptera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFADE1CD2FA4D1488" box="[1312,1436,1702,1727]" italics="true" pageId="2" pageNumber="75">Discoptera</emphasis>
|
||
Semenov, 1889
|
||
</taxonomicName>
|
||
and to some species of the predominantly African genus
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFC3E1CBDFAD414D4" authority="Latreille, 1802" authorityName="Latreille" authorityYear="1802" box="[960,1285,1737,1763]" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFC3E1CBDFB8614D4" box="[960,1111,1737,1763]" italics="true" pageId="2" pageNumber="75">Graphipterus</emphasis>
|
||
Latreille, 1802
|
||
</taxonomicName>
|
||
, for example, the species of the
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFEA51C99FD421530" authority="(Forsskal, 1775)" baseAuthorityName="Forsskal" baseAuthorityYear="1775" box="[347,659,1772,1799]" class="Insecta" family="Carabidae" genus="Graphipterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="species" species="serrator">
|
||
<emphasis id="B95F7E35FFD0FFADFEA51C99FE091530" box="[347,472,1773,1799]" italics="true" pageId="2" pageNumber="75">G. serrator</emphasis>
|
||
(Forsskål, 1775)
|
||
</taxonomicName>
|
||
group, but these two genera belong to the tribes Cyclosomini Laporte, 1834 and Graphipterini Latreille, 1802, respectively. Both tribes differ from the new genus in numerous characters, including the longer meso- and metatibial apical spurs (distinctly longer than half of the tarsomere 1) and the presence of the strong spines on the outer margin of the protibia. In these two characters, Cyclosomini and Graphipterini markedly differ from all
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFE461D09FDD815A0" authorityName="Bonelli" authorityYear="1810" box="[440,521,1917,1943]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="tribe" tribe="Lebiini">Lebiini</taxonomicName>
|
||
including
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFD781D09FCDF15A0" baseAuthorityName="Basilewsky" baseAuthorityYear="1984" box="[646,782,1917,1943]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
|
||
. Although meso- and metatibial spurs in
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFB121D09FAA115A0" baseAuthorityName="Basilewsky" baseAuthorityYear="1984" box="[1260,1392,1917,1943]" class="Insecta" family="Carabidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="subTribe" subTribe="Dromiusina">Dromiusina</taxonomicName>
|
||
are variable in length, they are at most half as long as the tarsomere 1. In the new genus, the meso- and metatibial spurs are not longer than those, for example, in
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFD951DB1FC7115E8" authority="Wissmann, 1846" authorityName="Wissmann" authorityYear="1846" box="[619,928,1989,2015]" class="Insecta" family="Carabidae" genus="Lionychus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFD951DB1FD0F15E8" box="[619,734,1989,2015]" italics="true" pageId="2" pageNumber="75">Lionychus</emphasis>
|
||
Wissmann, 1846
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2BD9A4FFD0FFADFC281DB1FB3A15E8" authority="Chaudoir, 1846" authorityName="Chaudoir" authorityYear="1846" box="[982,1259,1989,2015]" class="Insecta" family="Carabidae" genus="Apristus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="75" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95F7E35FFD0FFADFC281DB1FBE215E8" box="[982,1075,1989,2015]" italics="true" pageId="2" pageNumber="75">Apristus</emphasis>
|
||
Chaudoir, 1846
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |