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<mods:title id="8720E53CBDCC9C4800C6B7FDF7C18DFE">Rediscovered syntypes of Procrangonyx japonicus, with nomenclatural consideration of some crangonyctoidean subterranean amphipods (Crustacea: Amphipoda: Allocrangonyctidae, Niphargidae, Pseudocrangonyctidae)</mods:title>
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218.—
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<bibRefCitation id="EFEA4B3EFF8CFFAD3198F9563B052212" author="Bousfield, E. L." box="[645,817,1110,1132]" pageId="1" pagination="282 - 316" refId="ref4338" refString="Bousfield, E. L. (1977) A new look at the systematics of gammaroidean amphipods of the world. Crustaceana Supplement, 4, 282 - 316." type="journal article" year="1977">Bousfield, 1977</bibRefCitation>
: 306
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.—
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<bibRefCitation id="EFEA4B3EFF8CFFAD3099F9563C042212" author="Holsinger, J. R." box="[900,1072,1110,1132]" pageId="1" pagination="244 - 281" refId="ref4555" refString="Holsinger, J. R. (1977) A review of the systematics of the Holarctic amphipod family Crangonyctidae. Crustaceana Supplement, 4, 244 - 281." type="journal article" year="1977">Holsinger, 1977</bibRefCitation>
: 248
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(in discussion).—
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: 444
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445.—
<treatmentCitation id="0ADA10DEFF8CFFAD32C6F9753A8622F5" author="Holsinger, J. R." box="[475,690,1141,1163]" page="542" pageId="1" year="1986">
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: 542
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.—
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: 956
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.—
<treatmentCitation id="0ADA10DEFF8CFFAD30DFF9753C9022F5" author="Stock, J. H. &amp; Jo, Y. W." box="[962,1188,1141,1163]" page="119" pageId="1" year="1990">
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: 119
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(as an unavailable name without
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: 584
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585 (as an unavailable name).
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: 37
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(
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species:
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, fixed by monotypy).—
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: 444
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(as a junior synonym of
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).—
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: 272
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.—
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: 121
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.
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<emphasis id="B90FEADDFF8CFFAD338AF9F1392A2378" box="[151,286,1265,1286]" italics="true" pageId="1">Eocrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD323BF9F039DF2378" author="Schellenberg, A." box="[294,491,1264,1286]" pageId="1" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg, 1934</bibRefCitation>
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(sic).—
<treatmentCitation id="0ADA10DEFF8CFFAD3126F9F03B362378" author="Ishimaru, S." box="[571,770,1264,1286]" page="41" pageId="1" year="1994">
<bibRefCitation id="EFEA4B3EFF8CFFAD3126F9F03AE92378" author="Ishimaru, S." box="[571,733,1264,1286]" pageId="1" pagination="29 - 86" refId="ref5966" refString="Ishimaru, S. - i. (1994) A catalogue of gammaridean and ingolfiellidean Amphipoda recorded from the vicinity of Japan. Report of the Sado Marine Biological Station Niigata University, 24, 29 - 86." type="journal article" year="1994">Ishimaru, 1994</bibRefCitation>
: 41
</treatmentCitation>
(attribution only, without corresponding reference).—
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: 41
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(attribution only, without corresponding reference).
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<paragraph id="8BC436CFFF8CFFAD338AF82E3D5C23FE" blockId="1.[151,1436,1110,1408]" pageId="1">
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<taxonomicName id="4C7B4D4CFF8CFFAD338AF82E39C6233D" ID-CoL="6fa36d80-f8d9-49c4-a543-56b559503260" authority="Schellenberg, 1936" authorityName="Schellenberg" authorityYear="1936" box="[151,498,1325,1347]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD338AF82E392A233D" box="[151,286,1326,1347]" italics="true" pageId="1">Eocrangonyx</emphasis>
Schellenberg, 1936
</taxonomicName>
(sic).—
<treatmentCitation id="0ADA10DEFF8CFFAD315BF82D3B1F233D" author="Holsinger, J. R." box="[582,811,1325,1347]" page="248" pageId="1" year="1977">
<bibRefCitation id="EFEA4B3EFF8CFFAD315BF82D3AC1233D" author="Holsinger, J. R." box="[582,757,1325,1347]" pageId="1" pagination="244 - 281" refId="ref4555" refString="Holsinger, J. R. (1977) A review of the systematics of the Holarctic amphipod family Crangonyctidae. Crustaceana Supplement, 4, 244 - 281." type="journal article" year="1977">Holsinger, 1977</bibRefCitation>
: 248
</treatmentCitation>
(in discussion, as a junior synonym of
<taxonomicName id="4C7B4D4CFF8CFFAD37F9F82E3D40233D" ID-CoL="7PVNZ" authorityName=", Schellenberg" authorityYear="1934" box="[1252,1396,1326,1347]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD37F9F82E3D40233D" box="[1252,1396,1326,1347]" italics="true" pageId="1">Procrangonyx</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADA10DEFF8CFFAD33DAF84C39AA231C" author="Holsinger, J. R." box="[199,414,1356,1378]" page="956" pageId="1" year="1989">
<bibRefCitation id="EFEA4B3EFF8CFFAD33DAF84C3959231C" author="Holsinger, J. R." box="[199,365,1356,1378]" pageId="1" pagination="947 - 959" refId="ref4648" refString="Holsinger, J. R. (1989) Allocrangonyctidae and Pseudocrangonyctidae, two new families of Holarctic subterranean amphipod crustaceans (Gammaridea), with comments on their phylogenetic and zoogeographic relationships. Proceedings of the Biological Society of Washington, 102, 947 - 959." type="journal article" year="1989">Holsinger, 1989</bibRefCitation>
: 956
</treatmentCitation>
(as a junior synonym of
<taxonomicName id="4C7B4D4CFF8CFFAD31BDF84D3B1B231C" ID-CoL="7PVNZ" authorityName=", Schellenberg" authorityYear="1934" box="[672,815,1357,1378]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD31BDF84D3B1B231C" box="[672,815,1357,1378]" italics="true" pageId="1">Procrangonyx</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADA10DEFF8CFFAD3048F84C3CE6231C" author="Sidorov, D. A. &amp; Barabanschikov, E. I." box="[853,1234,1356,1378]" page="70" pageId="1" year="2010">
<bibRefCitation id="EFEA4B3EFF8CFFAD3048F84C3C84231C" author="Sidorov, D. A. &amp; Barabanschikov, E. I." box="[853,1200,1356,1378]" pageId="1" pagination="70 - 75" refId="ref6394" refString="Sidorov, D. A. &amp; Barabanschikov, E. I. (2010) Findings of stygobiotic and amphibiotic Peracarida (Amphipoda) in subsurface waters of the Samarga R. Basin (Northern Primorye) and some considerations about the taxonomic position of &quot; Orchestia &quot; solifuga Iwasa. Bulletin of the North-East Scientific Center, Russian Academy of Sciences Far East Branch, 2010, 70 - 75." type="journal article" year="2010">Sidorov &amp; Barabanschikov, 2010</bibRefCitation>
: 70
</treatmentCitation>
(in discussion, as a junior synonym of
<taxonomicName id="4C7B4D4CFF8CFFAD3295F86B3A2C23FE" ID-CoL="7PVNZ" authorityName=", Schellenberg" authorityYear="1934" box="[392,536,1387,1408]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD3295F86B3A2C23FE" box="[392,536,1387,1408]" italics="true" pageId="1">Procrangonyx</emphasis>
</taxonomicName>
).—
<treatmentCitation id="0ADA10DEFF8CFFAD3121F86A3BA423FE" author="Tomikawa, K. &amp; Shinoda, S." box="[572,912,1386,1408]" page="584" pageId="1" year="2016">
<bibRefCitation id="EFEA4B3EFF8CFFAD3121F86A3B5423FE" author="Tomikawa, K. &amp; Shinoda, S." box="[572,864,1386,1408]" pageId="1" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda, 2016</bibRefCitation>
: 584
</treatmentCitation>
.—Tomikawa
<emphasis id="B90FEADDFF8CFFAD373CF86B3C6023FE" box="[1057,1108,1386,1408]" italics="true" pageId="1">et al.</emphasis>
, 2016: 200 (in discussion).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3616544FF8CFFAF338AF8A839F524B2" lastPageId="3" pageId="1" type="materials_examined">
<paragraph id="8BC436CFFF8CFFAD338AF8A83B5E23C0" blockId="1.[151,874,1448,1470]" box="[151,874,1448,1470]" pageId="1">
<emphasis id="B90FEADDFF8CFFAD338AF8A8391123C0" bold="true" box="[151,293,1448,1470]" pageId="1">
<typeStatus id="54C0886DFF8CFFAD338AF8A838F823C0" box="[151,204,1448,1470]" pageId="1">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7B4D4CFF8CFFAD3236F8A93AAD23C0" authority="Ueno, 1930" authorityName="Ueno" authorityYear="1930" box="[299,665,1448,1470]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8CFFAD3236F8A93A2823C0" box="[299,540,1449,1470]" italics="true" pageId="1">Eucrangonyx japonicus</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD313CF8A83AAD23C0" author="Ueno, M." box="[545,665,1448,1470]" pageId="1" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
</taxonomicName>
, fixed by monotypy.
</paragraph>
<paragraph id="8BC436CFFF8CFFAD338AF8EA3CB72081" blockId="1.[151,1437,1514,2007]" pageId="1">
<emphasis id="B90FEADDFF8CFFAD338AF8EA399A207D" bold="true" box="[151,430,1514,1539]" pageId="1">Nomenclatural history.</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD32A5F8EB3A4C207D" author="Bate, C. S." box="[440,632,1514,1539]" pageId="1" pagination="237 - 240" refId="ref4305" refString="Bate, C. S. (1859) On the genus Niphargus (Schiodte). Proceedings of the Dublin University Zoology &amp; Botanical Association, 1, 237 - 240." type="journal article" year="1859">C.S. Bate (1859)</bibRefCitation>
—the citation of Charles Spence Bateherein follows
<bibRefCitation id="EFEA4B3EFF8CFFAD37FFF8EA3D42207D" author="Clark, P. F." box="[1250,1398,1514,1539]" pageId="1" pagination="429 - 438" refId="ref4454" refString="Clark, P. F. (2018) Charles Spence Bate: what's in a name? Zootaxa, 4497 (3), 429 - 438. https: // doi. org / 10.11646 / zootaxa. 4497.3.7" type="journal article" year="2018">Clark (2018)</bibRefCitation>
— established a new genus,
<taxonomicName id="4C7B4D4CFF8CFFAD32ABFB0F3ADB2059" authority="C. S. Bate, 1859" authorityName="C. S. Bate" authorityYear="1859" box="[438,751,1550,1575]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD32ABFB0F3A072058" box="[438,563,1551,1574]" italics="true" pageId="1">Crangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD3127FB0F3ADB2059" author="Bate, C. S." box="[570,751,1550,1575]" pageId="1" pagination="237 - 240" refId="ref4305" refString="Bate, C. S. (1859) On the genus Niphargus (Schiodte). Proceedings of the Dublin University Zoology &amp; Botanical Association, 1, 237 - 240." type="journal article" year="1859">C.S. Bate, 1859</bibRefCitation>
</taxonomicName>
, for a single new subterranean species,
<taxonomicName id="4C7B4D4CFF8CFFAD37ADFB0F39212035" authority="C. S. Bate, 1859" authorityName="C. S. Bate" authorityYear="1859" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis id="B90FEADDFF8CFFAD37ADFB0F3CF72058" box="[1200,1219,1551,1574]" italics="true" pageId="1">C</emphasis>
.
<emphasis id="B90FEADDFF8CFFAD37CFFB0E3D502059" box="[1234,1380,1550,1575]" italics="true" pageId="1">subterraneus</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD3671FB0F39212035" author="Bate, C. S." pageId="1" pagination="237 - 240" refId="ref4305" refString="Bate, C. S. (1859) On the genus Niphargus (Schiodte). Proceedings of the Dublin University Zoology &amp; Botanical Association, 1, 237 - 240." type="journal article" year="1859">C.S. Bate, 1859</bibRefCitation>
</taxonomicName>
, which was collected from Ringwood,
<collectingCountry id="F36C765FFF8CFFAD31C6FB333B252034" box="[731,785,1587,1610]" name="United Kingdom" pageId="1">U.K.</collectingCountry>
The
<typeStatus id="54C0886DFF8CFFAD304DFB333BB42035" box="[848,896,1587,1611]" pageId="1">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8CFFAD3718FB333CB62034" authorityName="C. S. Bate" authorityYear="1859" box="[1029,1154,1587,1610]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD3718FB333CB62034" box="[1029,1154,1587,1610]" italics="true" pageId="1">Crangonyx</emphasis>
</taxonomicName>
was thereby fixed as
<taxonomicName id="4C7B4D4CFF8CFFAD369FFB33391D2011" authorityName="C. S. Bate" authorityYear="1859" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis id="B90FEADDFF8CFFAD369FFB33391D2011" italics="true" pageId="1">C. subterraneus</emphasis>
</taxonomicName>
by monotypy, but C.S. Bate provided no morphological details of the uropods. Later,
<bibRefCitation id="EFEA4B3EFF8CFFAD37F5FB563DA82011" author="Stebbing, T. R. R." box="[1256,1436,1622,1647]" pageId="1" pagination="395 - 432" refId="ref6616" refString="Stebbing, T. R. R. (1899) Amphipoda from the Copenhagen Museum and other sources. - Part II. Transactions of the Linnean Society, Series 2 (Zoology), 7, 395 - 432. https: // doi. org / 10.1111 / j. 1096 - 3642.1899. tb 00202. x" type="journal article" year="1899">Stebbing (1899)</bibRefCitation>
erected a new
<taxonomicName id="4C7B4D4CFF8CFFAD3220FB7B398D20EC" authorityName="C. S. Bate" authorityYear="1859" box="[317,441,1659,1682]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD3220FB7B398D20EC" box="[317,441,1659,1682]" italics="true" pageId="1">Crangonyx</emphasis>
</taxonomicName>
-like genus,
<taxonomicName id="4C7B4D4CFF8CFFAD315BFB7C3BA620ED" authority="Stebbing, 1899" authorityName="Stebbing" authorityYear="1899" box="[582,914,1658,1683]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD315BFB7C3AEF20ED" box="[582,731,1660,1683]" italics="true" pageId="1">Eucrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD31F9FB7A3BA620ED" author="Stebbing, T. R. R." box="[740,914,1658,1683]" pageId="1" pagination="395 - 432" refId="ref6616" refString="Stebbing, T. R. R. (1899) Amphipoda from the Copenhagen Museum and other sources. - Part II. Transactions of the Linnean Society, Series 2 (Zoology), 7, 395 - 432. https: // doi. org / 10.1111 / j. 1096 - 3642.1899. tb 00202. x" type="journal article" year="1899">Stebbing, 1899</bibRefCitation>
</taxonomicName>
, for subterranean amphipods with a biramous uropod 3, i.e., with both inner and outer rami. He included five species in this new genus but did not designate its
<typeStatus id="54C0886DFF8CFFAD338AFBC338F320A5" box="[151,199,1731,1755]" pageId="1">type</typeStatus>
species; one of the originally included five species,
<taxonomicName id="4C7B4D4CFF8CFFAD3000FBC33D5720A5" authority="Smith in Smith &amp; Verrill, 1871" authorityName="Smith in Smith &amp; Verrill" authorityYear="1871" box="[797,1379,1730,1755]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B90FEADDFF8CFFAD3000FBC33BCC20A5" box="[797,1016,1730,1755]" italics="true" pageId="1">Crangonyx gracilis</emphasis>
Smith in
<bibRefCitation id="EFEA4B3EFF8CFFAD3774FBC23D5720A5" author="Smith, S. I. &amp; Verrill, A. E." box="[1129,1379,1730,1755]" pageId="1" pagination="448 - 454" refId="ref6528" refString="Smith, S. I. &amp; Verrill, A. E. (1871) Notice of the Invertebrata dredged in Lake Superior in 1871, by the U. S. Lake Survey, under the direction of Gen. C. B. Comstock, S. I. Smith, naturalist. The American Journal of Science and Arts, Series 3, 2, 448 - 454. https: // doi. org / 10.2475 / ajs. s 3 - 2.12.448" type="journal article" year="1871">Smith &amp; Verrill, 1871</bibRefCitation>
</taxonomicName>
, was subsequently designated as the
<typeStatus id="54C0886DFF8CFFAD32EAFBE73A132081" box="[503,551,1767,1791]" pageId="1">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8CFFAD31B9FBE73B0D2080" authorityName="Stebbing" authorityYear="1899" box="[676,825,1767,1790]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD31B9FBE73B0D2080" box="[676,825,1767,1790]" italics="true" pageId="1">Eucrangonyx</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFEA4B3EFF8CFFAD307EFBE63C4B2081" author="Chevreux, E. &amp; Fage, L." box="[867,1151,1766,1791]" pageId="1" refId="ref4424" refString="Chevreux, E. &amp; Fage, L. (1925) Faune de France. 9. Amphipodes. Paul Lechevalier, Paris, xix, 488 pp." type="book" year="1925">Chevreux &amp; Fage (1925)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC436CFFF8CFFAD33DAFA0A3CB721F1" blockId="1.[151,1437,1514,2007]" pageId="1">
Much later,
<bibRefCitation id="EFEA4B3EFF8CFFAD324DFA0B39D4215D" author="Ueno, M." box="[336,480,1803,1827]" pageId="1" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno (1930)</bibRefCitation>
described a Japanese subterranean amphipod collected from
<collectingRegion id="49BFF82DFF8CFFAD3785FA0A3CEB215D" box="[1176,1247,1802,1827]" country="Japan" name="Tokyo" pageId="1">Tokyo</collectingRegion>
as
<taxonomicName id="4C7B4D4CFF8CFFAD361BFA0C39952139" authority="Ueno, 1930" authorityName="Ueno" authorityYear="1930" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8CFFAD361BFA0C39302138" italics="true" pageId="1">Eucrangonyx japonicus</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD320EFA2F39952139" author="Ueno, M." box="[275,417,1839,1863]" pageId="1" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
</taxonomicName>
. He highlighted the fact that its uniramous (with outer ramus only) uropod 3 was uniarticulate, contrary to the condition in the Asian subterranean amphipod genus
<taxonomicName id="4C7B4D4CFF8CFFAD375FFA52391821F1" authority="Akatsuka &amp; Komai, 1922" authorityName="Akatsuka &amp; Komai" authorityYear="1922" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8CFFAD375FFA523D3F2115" box="[1090,1291,1874,1899]" italics="true" pageId="1">Pseudocrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8CFFAD3609FA52391821F1" author="Akatsuka, K. &amp; Komai, T." pageId="1" pagination="119 - 126" refId="ref4227" refString="Akatsuka, K. &amp; Komai, T. (1922) Pseudocrangonyx, a new genus of subterranean amphipods from Japan. Annotationes Zoologicae Japonenses, 10, 119 - 126." type="journal article" year="1922">Akatsuka &amp; Komai, 1922</bibRefCitation>
</taxonomicName>
, whose authors had diagnosed it by the uniramous but biarticulate uropod 3.
</paragraph>
<paragraph id="8BC436CFFF8CFFAE33DAFA9A3BC32444" blockId="1.[151,1437,1514,2007]" lastBlockId="2.[151,1437,151,2038]" lastPageId="2" pageId="1">
<bibRefCitation id="EFEA4B3EFF8CFFAD33DAFA9A399F21CD" author="Schellenberg, A." box="[199,427,1946,1971]" pageId="1" pagination="49 - 58" refId="ref6241" refString="Schellenberg, A. (1932) Bemerkungen uber subterrane Amphipoden Grossbritanniens. Zoologischer Anzeiger, 99, 49 - 58." type="journal article" year="1932">Schellenberg (1932)</bibRefCitation>
examined non-type material of the British
<taxonomicName id="4C7B4D4CFF8CFFAD308BFA9B3C7E21CD" authorityName="C. S. Bate" authorityYear="1859" box="[918,1098,1946,1971]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="1" phylum="Arthropoda" rank="species" species="subterraneus">
<emphasis id="B90FEADDFF8CFFAD308BFA9B3B9D21CC" box="[918,937,1947,1970]" italics="true" pageId="1">C</emphasis>
.
<emphasis id="B90FEADDFF8CFFAD30A5FA9A3C7E21CD" box="[952,1098,1946,1971]" italics="true" pageId="1">subterraneus</emphasis>
</taxonomicName>
and reported that this species has a rudimentary inner ramus on uropod 3, which is therefore biramous. Referring to this work again two years later,
<bibRefCitation id="EFEA4B3EFF8FFFAE33C1FD9739F126CE" author="Schellenberg, A." box="[220,453,151,176]" pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg (1934)</bibRefCitation>
concluded that the generic name
<taxonomicName id="4C7B4D4CFF8FFFAE3079FD993BCD26CE" authorityName="Stebbing" authorityYear="1899" box="[868,1017,153,176]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3079FD993BCD26CE" box="[868,1017,153,176]" italics="true" pageId="2">Eucrangonyx</emphasis>
</taxonomicName>
is a junior subjective synonym of
<taxonomicName id="4C7B4D4CFF8FFFAE338AFDBC392726AD" authorityName="C. S. Bate" authorityYear="1859" box="[151,275,188,211]" class="Malacostraca" family="Crangonyctidae" genus="Crangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AFDBC392726AD" box="[151,275,188,211]" italics="true" pageId="2">Crangonyx</emphasis>
</taxonomicName>
, the latter being diagnosed by a biramous uropod 3. At the same time he proposed a new generic name,
<taxonomicName id="4C7B4D4CFF8FFFAE338AFDE03A2E2686" authority="Schellenberg, 1934" authorityName=", Schellenberg" authorityYear="1934" box="[151,538,223,248]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AFDE039022689" box="[151,310,224,247]" italics="true" pageId="2">Procrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE325DFDDF3A2E2686" author="Schellenberg, A." box="[320,538,223,248]" pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg, 1934</bibRefCitation>
</taxonomicName>
(p. 217), for the species with a uniramous uropod 3 that had been previously classified under
<taxonomicName id="4C7B4D4CFF8FFFAE3250FC0439D62765" authorityName="Stebbing" authorityYear="1899" box="[333,482,260,283]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3250FC0439D62765" box="[333,482,260,283]" italics="true" pageId="2">Eucrangonyx</emphasis>
</taxonomicName>
. Throughout the (German) text, he referred to these uniramous “species” in the plural (i.e., as “Arten”), which indicates that his concept of the genus comprised more than one species.
<bibRefCitation id="EFEA4B3EFF8FFFAE3616FC2738D5271D" author="Schellenberg, A." pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg (1934)</bibRefCitation>
did not designate a
<typeStatus id="54C0886DFF8FFFAE32C1FC4B3A38271D" box="[476,524,331,355]" pageId="2">type</typeStatus>
species for
<taxonomicName id="4C7B4D4CFF8FFFAE31BFFC4C3B75271D" authorityName=", Schellenberg" authorityYear="1934" box="[674,833,332,355]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE31BFFC4C3B75271D" box="[674,833,332,355]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
, but he did explicitly mention a single species,
<taxonomicName id="4C7B4D4CFF8FFFAE3699FC4C393027F8" authorityName="Ueno" authorityYear="1930" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE3699FC4C3DA1271D" box="[1412,1429,332,355]" italics="true" pageId="2">E</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE338AFC6F393027F8" box="[151,260,367,390]" italics="true" pageId="2">japonicus</emphasis>
</taxonomicName>
(cited as
<taxonomicName id="4C7B4D4CFF8FFFAE326FFC6F3AB227F8" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[370,646,367,390]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE326FFC6F3AB227F8" box="[370,646,367,390]" italics="true" pageId="2">Procrangonyx japonicus</emphasis>
</taxonomicName>
), as “an example” of such a uniramous species (p. 218):
<emphasis id="B90FEADDFF8FFFAE360AFC6F39B127D4" italics="true" pageId="2">… ist er bei anderen Arten, z. B.</emphasis>
<taxonomicName id="4C7B4D4CFF8FFFAE328CFC933A9227D5" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[401,678,402,427]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">Procrangonyx japonicus</taxonomicName>
<emphasis id="B90FEADDFF8FFFAE31AFFC933B6D27D5" box="[690,857,403,427]" italics="true" pageId="2">(Uéno, p. 22),</emphasis>
<taxonomicName id="4C7B4D4CFF8FFFAE3079FC923C1F27D5" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[868,1067,402,427]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">Pseudocrangonyx</taxonomicName>
<emphasis id="B90FEADDFF8FFFAE3725FC923D3E27D4" box="[1080,1290,402,427]" italics="true" pageId="2">(l. c.) einästig, …</emphasis>
. Although Schellenberg did not cite
<taxonomicName id="4C7B4D4CFF8FFFAE32D8FCB73AE627B0" authorityName="Ueno" authorityYear="1930" box="[453,722,439,462]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE32D8FCB73AE627B0" box="[453,722,439,462]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
</taxonomicName>
in its original combination, there can be no doubt about the identity of the species involved. He explicitly attributed
<taxonomicName id="4C7B4D4CFF8FFFAE3036FCDB3C75278C" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[811,1089,475,498]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE3036FCDB3C75278C" box="[811,1089,475,498]" italics="true" pageId="2">Procrangonyx japonicus</emphasis>
</taxonomicName>
to
<bibRefCitation id="EFEA4B3EFF8FFFAE3771FCDA3D35278C" author="Ueno, M." box="[1132,1281,474,498]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno (1930)</bibRefCitation>
, whose work includes on page 22 a figure showing the uniramous uropod 3 of
<taxonomicName id="4C7B4D4CFF8FFFAE309DFCFF3C392468" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[896,1037,511,534]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE309DFCFF3BA52468" box="[896,913,511,534]" italics="true" pageId="2">E</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE30BDFCFF3C392468" box="[928,1037,511,534]" italics="true" pageId="2">japonicus</emphasis>
</taxonomicName>
and the description of this uropod.
<bibRefCitation id="EFEA4B3EFF8FFFAE338AFF21394E2444" author="Schellenberg, A." box="[151,378,545,570]" pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg (1934)</bibRefCitation>
also listed
<bibRefCitation id="EFEA4B3EFF8FFFAE32E4FF223AAE2444" author="Ueno, M." box="[505,666,546,570]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uénos (1930)</bibRefCitation>
work in his references section.
</paragraph>
<paragraph id="8BC436CFFF8FFFAE33DAFF4539092492" blockId="2.[151,1437,151,2038]" pageId="2">
Three years after the initial proposal of
<taxonomicName id="4C7B4D4CFF8FFFAE3198FF463BF12420" authority=", Schellenberg" authorityName=", Schellenberg" authorityYear="1934" box="[645,965,581,606]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3198FF463B102423" box="[645,804,582,605]" italics="true" pageId="2">Procrangonyx</emphasis>
, Schellenberg
</taxonomicName>
himself (1937a) established a new genus,
<taxonomicName id="4C7B4D4CFF8FFFAE338AFF6A3A2D24FF" authority="Schellenberg, 1937 a" authorityName="Schellenberg" authorityYear="1937" box="[151,537,617,642]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AFF6A391824FF" box="[151,300,618,641]" italics="true" pageId="2">Eocrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE322EFF693A2D24FF" author="Schellenberg, A." box="[307,537,617,642]" pageId="2" pagination="31 - 44" refId="ref6294" refString="Schellenberg, A. (1937 a) Die Amphipodengattungen um Crangonyx, ihre Verbreitung und ihre Arten. Mitteilungen aus dem Zoologischen Museum in Berlin, 22, 31 - 44. https: // doi. org / 10.1002 / mmnz. 19370220103" type="journal article" year="1937">Schellenberg, 1937a</bibRefCitation>
</taxonomicName>
, with a detailed diagnosis. He included one species in it,
<taxonomicName id="4C7B4D4CFF8FFFAE378EFF6A391824DB" authority="(Ueno, 1930)" baseAuthorityName="Ueno" baseAuthorityYear="1930" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE378EFF6A3DA824FF" box="[1171,1436,618,641]" italics="true" pageId="2">Eocrangonyx japonicus</emphasis>
(
<bibRefCitation id="EFEA4B3EFF8FFFAE3382FF8D391024DB" author="Ueno, M." box="[159,292,653,677]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
)
</taxonomicName>
cited as follows (p. 37):
<taxonomicName id="4C7B4D4CFF8FFFAE315BFF8D3B1E24DA" authority="(Ueno)" authorityName="Ueno" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[582,810,653,678]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
E. japonicus
<emphasis id="B90FEADDFF8FFFAE31C6FF8D3B1E24DA" box="[731,810,653,676]" italics="true" pageId="2">(Uéno)</emphasis>
</taxonomicName>
<emphasis id="B90FEADDFF8FFFAE302CFF8D3C7E24D8" box="[817,1098,653,678]" italics="true" pageId="2">
<quantity id="4C839B2AFF8FFFAE302CFF8D3B4424DB" box="[817,880,653,677]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="2" unit="mm" value="9.0">9 mm</quantity>
, Wasserwerk
<collectingRegion id="49BFF82DFF8FFFAE3715FF8D3C7E24D8" box="[1032,1098,653,678]" country="Japan" name="Tokyo" pageId="2">Tokyo</collectingRegion>
</emphasis>
.
<taxonomicName id="4C7B4D4CFF8FFFAE377CFF8E3D5E24DB" authorityName="Ueno" authorityYear="1930" box="[1121,1386,654,677]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE377CFF8E3D5E24DB" box="[1121,1386,654,677]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
</taxonomicName>
was thereby fixed as the
<typeStatus id="54C0886DFF8FFFAE32BCFFB139E524B7" box="[417,465,689,713]" pageId="2">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8FFFAE3176FFB23B3424B7" authorityName="Schellenberg" authorityYear="1937" box="[619,768,690,713]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3176FFB23B3424B7" box="[619,768,690,713]" italics="true" pageId="2">Eocrangonyx</emphasis>
</taxonomicName>
, by monotypy.
<bibRefCitation id="EFEA4B3EFF8FFFAE30D0FFB03CF224B7" author="Schellenberg, A." box="[973,1222,688,713]" pageId="2" pagination="31 - 44" refId="ref6294" refString="Schellenberg, A. (1937 a) Die Amphipodengattungen um Crangonyx, ihre Verbreitung und ihre Arten. Mitteilungen aus dem Zoologischen Museum in Berlin, 22, 31 - 44. https: // doi. org / 10.1002 / mmnz. 19370220103" type="journal article" year="1937">Schellenberg (1937a)</bibRefCitation>
never mentioned
<taxonomicName id="4C7B4D4CFF8FFFAE338AFFD639022493" authorityName=", Schellenberg" authorityYear="1934" box="[151,310,726,749]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AFFD639022493" box="[151,310,726,749]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC436CFFF8FFFAE33DAFFF83CB82209" blockId="2.[151,1437,151,2038]" pageId="2">
When erecting the new family
<taxonomicName id="4C7B4D4CFF8FFFAE3122FFF83B0F256F" authorityName="Holsinger" authorityYear="1989" box="[575,827,760,785]" class="Malacostraca" family="Pseudocrangonyctidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="family">Pseudocrangonyctidae</taxonomicName>
consisting of
<taxonomicName id="4C7B4D4CFF8FFFAE30F6FFF83C80256F" authorityName="Akatsuka &amp; Komai" authorityYear="1922" box="[1003,1204,760,785]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Pseudocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE30F6FFF83C80256F" box="[1003,1204,760,785]" italics="true" pageId="2">Pseudocrangonyx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D4CFF8FFFAE37EBFFF9396D254A" authority=", Holsinger (1989)" authorityName=", Holsinger" authorityYear="1989" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE37EBFFF93DA1256E" box="[1270,1429,761,784]" italics="true" pageId="2">Procrangonyx</emphasis>
,
<bibRefCitation id="EFEA4B3EFF8FFFAE338AFE1C396D254A" author="Holsinger, J. R." box="[151,345,796,821]" pageId="2" pagination="947 - 959" refId="ref4648" refString="Holsinger, J. R. (1989) Allocrangonyctidae and Pseudocrangonyctidae, two new families of Holarctic subterranean amphipod crustaceans (Gammaridea), with comments on their phylogenetic and zoogeographic relationships. Proceedings of the Biological Society of Washington, 102, 947 - 959." type="journal article" year="1989">Holsinger (1989)</bibRefCitation>
</taxonomicName>
treated
<taxonomicName id="4C7B4D4CFF8FFFAE32A5FE1D3A79254A" authorityName="Schellenberg" authorityYear="1937" box="[440,589,797,820]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE32A5FE1D3A79254A" box="[440,589,797,820]" italics="true" pageId="2">Eocrangonyx</emphasis>
</taxonomicName>
as an objective junior synonym of
<taxonomicName id="4C7B4D4CFF8FFFAE30F5FE1D3CB3254A" authorityName=", Holsinger" authorityYear="1989" box="[1000,1159,797,820]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE30F5FE1D3CB3254A" box="[1000,1159,797,820]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
and stated that the
<typeStatus id="54C0886DFF8FFFAE3671FE1D3DA8254B" box="[1388,1436,797,821]" pageId="2">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8FFFAE3213FE4139992526" authorityName=", Holsinger" authorityYear="1989" box="[270,429,833,856]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3213FE4139992526" box="[270,429,833,856]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
is
<taxonomicName id="4C7B4D4CFF8FFFAE32D3FE413B572526" authority="Ueno, 1930" authorityName="Ueno" authorityYear="1930" box="[462,867,832,856]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE32D3FE413AEC2526" box="[462,728,833,856]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE31C2FE403B572526" author="Ueno, M." box="[735,867,832,856]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
</taxonomicName>
fixed by monotypy. Holsingers contentions were, however, not supported by explicit citations of relevant provisions of the Code. In contrast,
<bibRefCitation id="EFEA4B3EFF8FFFAE378EFE6338FF25DE" author="Stock, J. H. &amp; Jo, Y. W." pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990: 120 121)</bibRefCitation>
pointed out that the expression “z. B.” (i.e., “zum Beispiel”, meaning “for example”) in the original description of
<taxonomicName id="4C7B4D4CFF8FFFAE3223FEAC39E925BD" authorityName=", Holsinger" authorityYear="1989" box="[318,477,940,963]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3223FEAC39E925BD" box="[318,477,940,963]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
cannot be regarded as a
<typeStatus id="54C0886DFF8FFFAE3019FEAC3B0025BA" box="[772,820,940,964]" pageId="2">type</typeStatus>
species designation according to Article 67(c) of the International Code of Zoological Nomenclature, Third Edition (International Commission on
<bibRefCitation id="EFEA4B3EFF8FFFAE363EFECF394E2275" author="International Commission on Zoological Nomenclature" pageId="2" refId="ref5902" refString="International Commission on Zoological Nomenclature. (1985) International Code of Zoological Nomenclature. 3 rd Edition. International Trust for Zoological Nomenclature, London, xx + 338 pp." type="book" year="1985">Zoological Nomenclature 1985</bibRefCitation>
). They judged the generic name
<taxonomicName id="4C7B4D4CFF8FFFAE31EFFEF43BA52275" authorityName=", Holsinger" authorityYear="1989" box="[754,913,1012,1035]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE31EFFEF43BA52275" box="[754,913,1012,1035]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
to be unavailable because, as they saw it, the genus was erected without a
<typeStatus id="54C0886DFF8FFFAE32F4F9173A2D2251" box="[489,537,1047,1071]" pageId="2">type</typeStatus>
species after 1930, contrary to the Third Codes Article 67(c)(1). In place of
<taxonomicName id="4C7B4D4CFF8FFFAE338AF93B3A22222D" authority="Stock &amp; Jo (1990)" authorityName="Stock &amp; Jo" authorityYear="1990" box="[151,534,1082,1107]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AF93B3902222C" box="[151,310,1083,1106]" italics="true" pageId="2">Procrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE325DF93A3A22222D" author="Stock, J. H. &amp; Jo, Y. W." box="[320,534,1082,1107]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990)</bibRefCitation>
</taxonomicName>
used the available name
<taxonomicName id="4C7B4D4CFF8FFFAE3023F93B3BE7222C" authorityName="Schellenberg" authorityYear="1937" box="[830,979,1083,1106]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3023F93B3BE7222C" box="[830,979,1083,1106]" italics="true" pageId="2">Eocrangonyx</emphasis>
</taxonomicName>
for the Japanese species, and they also described a new Far-East Russian species as
<taxonomicName id="4C7B4D4CFF8FFFAE3190F95F3CBD2209" authority="Stock &amp; Jo, 1990" authorityName="Stock &amp; Jo" authorityYear="1990" box="[653,1161,1118,1143]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="primoryensis">
<emphasis id="B90FEADDFF8FFFAE3190F95F3B882208" box="[653,956,1119,1142]" italics="true" pageId="2">Eocrangonyx primoryensis</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE30DEF95E3CBD2209" author="Stock, J. H. &amp; Jo, Y. W." box="[963,1161,1118,1143]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo, 1990</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC436CFFF8FFFAE33DAF9823B492333" blockId="2.[151,1437,151,2038]" pageId="2">
After 2000,
<bibRefCitation id="EFEA4B3EFF8FFFAE324FF9833A2122E4" author="Hou, Z. - E. &amp; Li, S." box="[338,533,1154,1179]" pageId="2" pagination="1179 - 1188" refId="ref5802" refString="Hou, Z. - E. &amp; Li, S. (2003) A new subterranean amphipod of the genus Procrangonyx from Beijing, China. Crustaceana, 76, 1179 - 1188. https: // doi. org / 10.1163 / 156854003773123410" type="journal article" year="2003">Hou &amp; Li (2003)</bibRefCitation>
and
<bibRefCitation id="EFEA4B3EFF8FFFAE3152F9823BA422E4" author="Sidorov, D. &amp; Holsinger, J. R." box="[591,912,1154,1179]" pageId="2" pagination="417 - 430" refId="ref6467" refString="Sidorov, D. [A.] &amp; Holsinger, J. R. (2007) Procrangonyx stygoedincus, a new species of subterranean amphipod (Pseudocrangonyctidae) from the Far East of Russia, with remarks on biogeographic relationships. Crustaceana, 80, 417 - 430. https: // doi. org / 10.1163 / 156854007780440984" type="journal article" year="2007">Sidorov &amp; Holsinger (2007)</bibRefCitation>
each described a new subterranean amphipod species with a uniramous and uniarticulate uropod 3, the two species being placed in
<taxonomicName id="4C7B4D4CFF8FFFAE3744F9A73CCC22C0" authorityName="Stock &amp; Jo" authorityYear="1990" box="[1113,1272,1191,1214]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3744F9A73CCC22C0" box="[1113,1272,1191,1214]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
as
<taxonomicName id="4C7B4D4CFF8FFFAE3602F9A7397C229C" authority="Hou &amp; Li, 2003" authorityName="Hou &amp; Li" authorityYear="2003" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="limpidus">
<emphasis id="B90FEADDFF8FFFAE3602F9A73D1922C0" box="[1311,1325,1191,1214]" italics="true" pageId="2">P</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE3621F9A53DA822C0" box="[1340,1436,1189,1214]" italics="true" pageId="2">limpidus</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE338AF9CA397C229C" author="Hou, Z. - E. &amp; Li, S." box="[151,328,1225,1250]" pageId="2" pagination="1179 - 1188" refId="ref5802" refString="Hou, Z. - E. &amp; Li, S. (2003) A new subterranean amphipod of the genus Procrangonyx from Beijing, China. Crustaceana, 76, 1179 - 1188. https: // doi. org / 10.1163 / 156854003773123410" type="journal article" year="2003">Hou &amp; Li, 2003</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7B4D4CFF8FFFAE3260F9CA3B56229C" authority="Sidorov &amp; Holsinger, 2007" authorityName="Sidorov &amp; Holsinger" authorityYear="2007" box="[381,866,1225,1250]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="stygoedincus">
<emphasis id="B90FEADDFF8FFFAE3260F9CA39BF229F" box="[381,395,1226,1249]" italics="true" pageId="2">P</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE3284F9C93A1E229C" box="[409,554,1225,1250]" italics="true" pageId="2">stygoedincus</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE312DF9C93B56229C" author="Sidorov, D. &amp; Holsinger, J. R." box="[560,866,1225,1250]" pageId="2" pagination="417 - 430" refId="ref6467" refString="Sidorov, D. [A.] &amp; Holsinger, J. R. (2007) Procrangonyx stygoedincus, a new species of subterranean amphipod (Pseudocrangonyctidae) from the Far East of Russia, with remarks on biogeographic relationships. Crustaceana, 80, 417 - 430. https: // doi. org / 10.1163 / 156854007780440984" type="journal article" year="2007">Sidorov &amp; Holsinger, 2007</bibRefCitation>
</taxonomicName>
. Neither work referred to
<bibRefCitation id="EFEA4B3EFF8FFFAE379FF9C93D65229C" author="Stock, J. H. &amp; Jo, Y. W." box="[1154,1361,1225,1250]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990)</bibRefCitation>
. Later,
<bibRefCitation id="EFEA4B3EFF8FFFAE338AF9ED3A442378" author="Sidorov, D. A. &amp; Barabanschikov, E. I." box="[151,624,1261,1286]" pageId="2" pagination="70 - 75" refId="ref6394" refString="Sidorov, D. A. &amp; Barabanschikov, E. I. (2010) Findings of stygobiotic and amphibiotic Peracarida (Amphipoda) in subsurface waters of the Samarga R. Basin (Northern Primorye) and some considerations about the taxonomic position of &quot; Orchestia &quot; solifuga Iwasa. Bulletin of the North-East Scientific Center, Russian Academy of Sciences Far East Branch, 2010, 70 - 75." type="journal article" year="2010">Sidorov &amp; Barabanschikov (2010: 7071)</bibRefCitation>
cited
<bibRefCitation id="EFEA4B3EFF8FFFAE31A8F9ED3BB02378" author="Stock, J. H. &amp; Jo, Y. W." box="[693,900,1261,1286]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990)</bibRefCitation>
and argued that the earlier name
<taxonomicName id="4C7B4D4CFF8FFFAE37E0F9EE3DA8237B" authorityName="Stock &amp; Jo" authorityYear="1990" box="[1277,1436,1262,1285]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE37E0F9EE3DA8237B" box="[1277,1436,1262,1285]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
was valid. Additionally, they treated
<taxonomicName id="4C7B4D4CFF8FFFAE3121F8123ADE2354" authorityName="Sidorov &amp; Holsinger" authorityYear="2007" box="[572,746,1297,1322]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="stygoedincus">
<emphasis id="B90FEADDFF8FFFAE3121F8123A7E2357" box="[572,586,1298,1321]" italics="true" pageId="2">P</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE3144F8113ADE2354" box="[601,746,1297,1322]" italics="true" pageId="2">stygoedincus</emphasis>
</taxonomicName>
as a subjective junior synonym of
<taxonomicName id="4C7B4D4CFF8FFFAE3761F81239352333" authority="(Stock &amp; Jo, 1990)" baseAuthorityName="Stock &amp; Jo" baseAuthorityYear="1990" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="primoryensis">
<emphasis id="B90FEADDFF8FFFAE3761F8123CBE2357" box="[1148,1162,1298,1321]" italics="true" pageId="2">P</emphasis>
.
<emphasis id="B90FEADDFF8FFFAE3787F8123D182357" box="[1178,1324,1298,1321]" italics="true" pageId="2">primoryensis</emphasis>
(
<bibRefCitation id="EFEA4B3EFF8FFFAE3621F81138CE2333" author="Stock, J. H. &amp; Jo, Y. W." pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo, 1990</bibRefCitation>
)
</taxonomicName>
, the latter being shifted to this genus from
<taxonomicName id="4C7B4D4CFF8FFFAE31FCF8363B422333" authorityName="Schellenberg" authorityYear="1937" box="[737,886,1334,1357]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE31FCF8363B422333" box="[737,886,1334,1357]" italics="true" pageId="2">Eocrangonyx</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC436CFFF8FFFAE33DAF8583A3C20A9" blockId="2.[151,1437,151,2038]" pageId="2">
<bibRefCitation id="EFEA4B3EFF8FFFAE33DAF8583A21230F" author="Tomikawa, K. &amp; Shinoda, S." box="[199,533,1368,1393]" pageId="2" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda (2016)</bibRefCitation>
redescribed
<taxonomicName id="4C7B4D4CFF8FFFAE31B4F85A3C73230F" authority="Ueno, 1930" authorityName="Ueno" authorityYear="1930" box="[681,1095,1369,1393]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE31B4F85A3B81230F" box="[681,949,1370,1393]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
<bibRefCitation id="EFEA4B3EFF8FFFAE30A3F8593C73230F" author="Ueno, M." box="[958,1095,1369,1393]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
</taxonomicName>
. Citing relevant provisions of the current Fourth Edition of the Code (International Commission on
<bibRefCitation id="EFEA4B3EFF8FFFAE30FEF87C3D6623EB" author="International Commission on Zoological Nomenclature" box="[995,1362,1404,1429]" pageId="2" refId="ref5934" refString="International Commission on Zoological Nomenclature. (1999) International Code of Zoological Nomenclature. 4 th Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">Zoological Nomenclature 1999</bibRefCitation>
), they reaffirmed
<bibRefCitation id="EFEA4B3EFF8FFFAE3205F8A03A3323C6" author="Stock, J. H. &amp; Jo, Y. W." box="[280,519,1440,1465]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jos (1990)</bibRefCitation>
nomenclatural conclusion on the invalidity [sic; unavailability was meant] of
<taxonomicName id="4C7B4D4CFF8FFFAE338AF8C5390223A2" authorityName="Stock &amp; Jo" authorityYear="1990" box="[151,310,1477,1500]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE338AF8C5390223A2" box="[151,310,1477,1500]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
and used the binomen
<taxonomicName id="4C7B4D4CFF8FFFAE314DF8C53B6923A2" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[592,861,1477,1500]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE314DF8C53B6923A2" box="[592,861,1477,1500]" italics="true" pageId="2">Eocrangonyx japonicus</emphasis>
</taxonomicName>
(pp. 584585). Following
<bibRefCitation id="EFEA4B3EFF8FFFAE3780F8C438D0207E" author="Tomikawa, K. &amp; Shinoda, S." pageId="2" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda (2016)</bibRefCitation>
, all species assigned to
<taxonomicName id="4C7B4D4CFF8FFFAE311BF8E93A91207E" authorityName="Stock &amp; Jo" authorityYear="1990" box="[518,677,1513,1536]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE311BF8E93A91207E" box="[518,677,1513,1536]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
were placed under
<taxonomicName id="4C7B4D4CFF8FFFAE308EF8E93C1C207E" authorityName="Schellenberg" authorityYear="1937" box="[915,1064,1513,1536]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE308EF8E93C1C207E" box="[915,1064,1513,1536]" italics="true" pageId="2">Eocrangonyx</emphasis>
</taxonomicName>
in the influential web resource WoRMS—in which amphipod records are sourced from World
<taxonomicName id="4C7B4D4CFF8FFFAE309CFB0B3C43205A" authority="Database" authorityName="Database" box="[897,1143,1547,1572]" class="Malacostraca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="order">Amphipoda Database</taxonomicName>
(= WAD) (
<bibRefCitation id="EFEA4B3EFF8FFFAE361EFB0C38E92036" author="Horton, T. &amp; Lowry, J. &amp; De Broyer, C. &amp; Bellan-Santini, D. &amp; Coleman, C. O. &amp; Corbari, L. &amp; Costello, M. J. &amp; Daneliya, M. &amp; Dauvin, J. - C. &amp; Fiser, C. &amp; Gasca, R. &amp; Grabowski, M. &amp; Guerra-Garcia, J. M. &amp; Hendrycks, E. &amp; Hughes, L. &amp; Jaume, D. &amp; Jazdzewski, K. &amp; Kim, Y. - H. &amp; King, R. &amp; Krapp-Schickel, T. &amp; LeCroy, S. &amp; Lorz, A. - N. &amp; Mamos, T. &amp; Senna, A. R. &amp; Serejo, C. &amp; Sket, B. &amp; Souza-Filho, J. F. &amp; Tandberg, A. H. &amp; Thomas, J. &amp; Thurston, M. &amp; Vader, W. &amp; Vainola, R. &amp; Vonk, R. &amp; White, K. &amp; Zeidler, W." pageId="2" refId="ref4840" refString="Horton, T., Lowry, J., De Broyer, C., Bellan-Santini, D., Coleman, C. O., Corbari, L., Costello, M. J., Daneliya, M., Dauvin, J. - C., Fiser, C., Gasca, R., Grabowski, M., Guerra-Garcia, J. M., Hendrycks, E., Hughes, L., Jaume, D., Jazdzewski, K., Kim, Y. - H., King, R., Krapp-Schickel, T., LeCroy, S., Lorz, A. - N., Mamos, T., Senna, A. R., Serejo, C., Sket, B., Souza-Filho, J. F., Tandberg, A. H., Thomas, J., Thurston, M., Vader, W., Vainola, R., Vonk, R., White, K. &amp; Zeidler, W. (2018 a) World Amphipoda Database. Procrangonyx Schellenberg, 1934. Available from: http: // www. marinespecies. org / amphipoda / aphia. php? p = taxdetails &amp; id = 550654 (accessed 2 October 2018)" type="url" year="2018">
Horton
<emphasis id="B90FEADDFF8FFFAE3642FB0D3DA8205A" box="[1375,1436,1547,1572]" italics="true" pageId="2">et al.</emphasis>
2018a
</bibRefCitation>
)—, the former genus being treated there as an invalid name without
<typeStatus id="54C0886DFF8FFFAE30FCFB303C252036" box="[993,1041,1584,1608]" pageId="2">type</typeStatus>
species designation. In accordance with the manual for editors of WoRMS (Available from: http://www.marinespecies.org/aphia.php?p=manual#top; see also
<bibRefCitation id="EFEA4B3EFF8FFFAE33EAFB7839FE20F1" author="Horton, T. &amp; Gofas, S. &amp; Kroh, A. &amp; Poore, G. C. B. &amp; Read, G. &amp; Rosenberg, G. &amp; Stohr, S. &amp; Bailly, N. &amp; Boury-Esnault, N. &amp; Brandao, S. N. &amp; Costello, M. J. &amp; Decock, W. &amp; Dekeyzer, S. &amp; Hernandez, F. &amp; Mees, J. &amp; Paulay, G. &amp; Vandepitte, L. &amp; Vanhoorne, B. &amp; Vranken, S." box="[247,458,1654,1679]" pageId="2" pagination="1 - 24" refId="ref4696" refString="Horton, T., Gofas, S., Kroh, A., Poore, G. C. B., Read, G., Rosenberg, G., Stohr, S., Bailly, N., Boury-Esnault, N., Brandao, S. N., Costello, M. J., Decock, W., Dekeyzer, S., Hernandez, F., Mees, J., Paulay, G., Vandepitte, L., Vanhoorne, B. &amp; Vranken, S. (2017) Improving nomenclatural consistency: a decade of experience in the World Register of Marine Species. European Journal of Taxonomy, 389, 1 - 24. https: // doi. org / 10.5852 / ejt. 2017.389" type="journal article" year="2017">
Horton
<emphasis id="B90FEADDFF8FFFAE3250FB7839B220F1" box="[333,390,1654,1679]" italics="true" pageId="2">et al.</emphasis>
2017
</bibRefCitation>
),
<taxonomicName id="4C7B4D4CFF8FFFAE32C0FB783A4820F1" authorityName="Stock &amp; Jo" authorityYear="1990" box="[477,636,1656,1679]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE32C0FB783A4820F1" box="[477,636,1656,1679]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
was treated in the same way as the “present literature” does, i.e., as an “unaccepted” (= “invalid”) name that lacks a
<typeStatus id="54C0886DFF8FFFAE318BFB9B3AF220CD" box="[662,710,1691,1715]" pageId="2">type</typeStatus>
species designation; however, a note acknowledged the ongoing disagreement on this latter point.
</paragraph>
<paragraph id="8BC436CFFF8FFFAF33DAFBE23932273E" blockId="2.[151,1437,151,2038]" lastBlockId="3.[151,1437,151,716]" lastPageId="3" pageId="2">
<emphasis id="B90FEADDFF8FFFAE33DAFBE23A382085" bold="true" box="[199,524,1762,1787]" pageId="2">Nomenclatural conclusion.</emphasis>
In considering the potential availability and validity of
<taxonomicName id="4C7B4D4CFF8FFFAE37A5FBE33D632084" authorityName="Stock &amp; Jo" authorityYear="1990" box="[1208,1367,1763,1786]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE37A5FBE33D632084" box="[1208,1367,1763,1786]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
, it is necessary to clearly distinguish the two concepts of
<typeStatus id="54C0886DFF8FFFAE31E9FA073B102161" box="[756,804,1799,1823]" pageId="2">type</typeStatus>
species “fixation” (Article 67.4 of the Code) and
<typeStatus id="54C0886DFF8FFFAE3671FA073DA82161" box="[1388,1436,1799,1823]" pageId="2">type</typeStatus>
species “designation” (Article 67.5). The nomenclatural arguments of both
<bibRefCitation id="EFEA4B3EFF8FFFAE30EAFA2A3CF8213C" author="Stock, J. H. &amp; Jo, Y. W." box="[1015,1228,1834,1859]" pageId="2" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990)</bibRefCitation>
and
<bibRefCitation id="EFEA4B3EFF8FFFAE361AFA2A39722118" author="Tomikawa, K. &amp; Shinoda, S." pageId="2" pagination="583 - 594" refId="ref7043" refString="Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, Eocrangonyx japonicus (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. Crustaceana, 89, 583 - 594. https: // doi. org / 10.1163 / 15685403 - 00003544" type="journal article" year="2016">Tomikawa &amp; Shinoda (2016)</bibRefCitation>
were partially correct in that
<bibRefCitation id="EFEA4B3EFF8FFFAE3180FA4D3BB72118" author="Schellenberg, A." box="[669,899,1869,1894]" pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg (1934)</bibRefCitation>
did not “designate”
<taxonomicName id="4C7B4D4CFF8FFFAE376DFA4F3D48211B" authorityName="Ueno" authorityYear="1930" box="[1136,1404,1870,1894]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE376DFA4F3D48211B" box="[1136,1404,1870,1894]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
</taxonomicName>
as the
<typeStatus id="54C0886DFF8FFFAE33DFFA7238C621F4" box="[194,242,1906,1930]" pageId="2">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8FFFAE3268FA723A2021F7" authorityName="Stock &amp; Jo" authorityYear="1990" box="[373,532,1906,1929]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3268FA723A2021F7" box="[373,532,1906,1929]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
in a manner recognized by Article 67.5 of the Code. Under Article 67.2.1 and Article 68.3, however, because
<taxonomicName id="4C7B4D4CFF8FFFAE311BFA963BF521D0" authority="(Ueno, 1930)" baseAuthorityName="Ueno" baseAuthorityYear="1930" box="[518,961,1942,1966]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE311BFA963B2921D3" box="[518,797,1942,1965]" italics="true" pageId="2">Procrangonyx japonicus</emphasis>
(
<bibRefCitation id="EFEA4B3EFF8FFFAE302DFA963B8D21D0" author="Ueno, M." box="[816,953,1942,1966]" pageId="2" pagination="21 - 23" refId="ref7093" refString="Ueno, M. (1930) A new subterranean amphipod from Japan. Annotationes Zoologicae Japonenses, 13, 21 - 23." type="journal article" year="1930">Uéno, 1930</bibRefCitation>
)
</taxonomicName>
was the only available nominal speciesgroup taxon originally included in
<taxonomicName id="4C7B4D4CFF8FFFAE3101FABA3A8F21AF" authorityName="Stock &amp; Jo" authorityYear="1990" box="[540,699,1978,2001]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE3101FABA3A8F21AF" box="[540,699,1978,2001]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
, this species (
<taxonomicName id="4C7B4D4CFF8FFFAE3047FABA3C5721AF" authorityName="Ueno" authorityYear="1930" box="[858,1123,1978,2001]" class="Malacostraca" family="Crangonyctidae" genus="Eucrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="species" species="japonicus">
<emphasis id="B90FEADDFF8FFFAE3047FABA3C5721AF" box="[858,1123,1978,2001]" italics="true" pageId="2">Eucrangonyx japonicus</emphasis>
</taxonomicName>
in its original combination) was fixed by
<bibRefCitation id="EFEA4B3EFF8FFFAE3233FADD3A26218B" author="Schellenberg, A." box="[302,530,2013,2038]" pageId="2" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg (1934)</bibRefCitation>
as the
<typeStatus id="54C0886DFF8FFFAE317CFADE3AA52188" box="[609,657,2014,2038]" pageId="2">type</typeStatus>
species of
<taxonomicName id="4C7B4D4CFF8FFFAE300DFADE3B9B218B" authorityName="Stock &amp; Jo" authorityYear="1990" box="[784,943,2014,2037]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="2" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8FFFAE300DFADE3B9B218B" box="[784,943,2014,2037]" italics="true" pageId="2">Procrangonyx</emphasis>
</taxonomicName>
. [N.B.: Under Recommendation 67B of the Code, the name of a
<typeStatus id="54C0886DFF8EFFAF329FFD98398626CE" box="[386,434,152,176]" pageId="3">type</typeStatus>
species should be cited by its original binomen.] Since a
<typeStatus id="54C0886DFF8EFFAF3722FD983C5B26CE" box="[1087,1135,152,176]" pageId="3">type</typeStatus>
species was “fixed” in the original publication, even if not explicitly “designated”, and a diagnostic character for the genus was proposed in the same work,
<taxonomicName id="4C7B4D4CFF8EFFAF324DFDE03AE72686" authority="Schellenberg, 1934" authorityName=", Schellenberg" authorityYear="1934" box="[336,723,223,248]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF324DFDE039DB2689" box="[336,495,224,247]" italics="true" pageId="3">Procrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8EFFAF32E5FDDF3AE72686" author="Schellenberg, A." box="[504,723,223,248]" pageId="3" pagination="215 - 218" refId="ref6263" refString="Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen Hohle, nebst Bemerkungen uber die Gattung Crangonyx. Zoologischer Anzeiger, 106, 215 - 218." type="journal article" year="1934">Schellenberg, 1934</bibRefCitation>
</taxonomicName>
is available and is the valid name of the genus.
<taxonomicName id="4C7B4D4CFF8EFFAF361BFDE0394A2762" authority="Schellenberg, 1937 a" authorityName="Schellenberg" authorityYear="1937" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF361BFDE03DAF2689" box="[1286,1435,224,247]" italics="true" pageId="3">Eocrangonyx</emphasis>
<bibRefCitation id="EFEA4B3EFF8EFFAF338AFC03394A2762" author="Schellenberg, A." box="[151,382,259,284]" pageId="3" pagination="31 - 44" refId="ref6294" refString="Schellenberg, A. (1937 a) Die Amphipodengattungen um Crangonyx, ihre Verbreitung und ihre Arten. Mitteilungen aus dem Zoologischen Museum in Berlin, 22, 31 - 44. https: // doi. org / 10.1002 / mmnz. 19370220103" type="journal article" year="1937">Schellenberg, 1937a</bibRefCitation>
</taxonomicName>
, with the same
<typeStatus id="54C0886DFF8EFFAF3130FC043A692762" box="[557,605,260,284]" pageId="3">type</typeStatus>
species, is then a junior objective synonym of
<taxonomicName id="4C7B4D4CFF8EFFAF3770FC043D382765" authorityName=", Schellenberg" authorityYear="1934" box="[1133,1292,260,283]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF3770FC043D382765" box="[1133,1292,260,283]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
, and as such is invalid.
</paragraph>
<paragraph id="8BC436CFFF8EFFAF33DAFC4A39F524B2" blockId="3.[151,1437,151,716]" pageId="3">
<bibRefCitation id="EFEA4B3EFF8EFFAF33DAFC4A39A5271D" author="Holsinger, J. R." box="[199,401,330,355]" pageId="3" pagination="947 - 959" refId="ref4648" refString="Holsinger, J. R. (1989) Allocrangonyctidae and Pseudocrangonyctidae, two new families of Holarctic subterranean amphipod crustaceans (Gammaridea), with comments on their phylogenetic and zoogeographic relationships. Proceedings of the Biological Society of Washington, 102, 947 - 959." type="journal article" year="1989">Holsinger (1989)</bibRefCitation>
was unquestionably correct about the availability and the
<typeStatus id="54C0886DFF8EFFAF3764FC4B3C9D271D" box="[1145,1193,331,355]" pageId="3">type</typeStatus>
species fixation of
<taxonomicName id="4C7B4D4CFF8EFFAF338AFC70390227F9" authorityName=", Schellenberg" authorityYear="1934" box="[151,310,368,391]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF338AFC70390227F9" box="[151,310,368,391]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
, despite his lack of rigorous reference to the Code. The erroneous status of
<taxonomicName id="4C7B4D4CFF8EFFAF37D5FC703D5327F9" authorityName=", Schellenberg" authorityYear="1934" box="[1224,1383,368,391]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF37D5FC703D5327F9" box="[1224,1383,368,391]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D4CFF8EFFAF338AFC94391827D5" authorityName="Schellenberg" authorityYear="1937" box="[151,300,404,427]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Eocrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF338AFC94391827D5" box="[151,300,404,427]" italics="true" pageId="3">Eocrangonyx</emphasis>
</taxonomicName>
in WAD should be emended promptly to avoid additional confusion about which generic name to use for the species now assigned to these genera. The nomenclatural disorder concerning the availability, and thus the validity, of
<taxonomicName id="4C7B4D4CFF8EFFAF323CFCDC39F4278D" authorityName=", Schellenberg" authorityYear="1934" box="[289,448,476,499]" class="Malacostraca" family="Pseudocrangonyctidae" genus="Procrangonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="3" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEADDFF8EFFAF323CFCDC39F4278D" box="[289,448,476,499]" italics="true" pageId="3">Procrangonyx</emphasis>
</taxonomicName>
highlights the common error of confusing the two terms “
<typeStatus id="54C0886DFF8EFFAF3760FCDB3C84278D" box="[1149,1200,475,499]" pageId="3">type</typeStatus>
fixation” and “
<typeStatus id="54C0886DFF8EFFAF3677FCDB3DA8278D" box="[1386,1436,475,499]" pageId="3">type</typeStatus>
designation”. Zoologists are generally aware that to be available, every new genus-group name proposed after 1930 must be accompanied by
<typeStatus id="54C0886DFF8EFFAF32A0FF2339D92445" box="[445,493,547,571]" pageId="3">type</typeStatus>
species “fixation”, but, like
<bibRefCitation id="EFEA4B3EFF8EFFAF3025FF223C392445" author="Stock, J. H. &amp; Jo, Y. W." box="[824,1037,546,571]" pageId="3" pagination="119 - 127" refId="ref6723" refString="Stock, J. H. &amp; Jo, Y. W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. Stygologia, 5, 119 - 127." type="journal article" year="1990">Stock &amp; Jo (1990)</bibRefCitation>
they tend to misunderstand this to mean “original designation” by the author of the genus. Indeed, explicit designation is probably best practice, but several other permissible means of
<typeStatus id="54C0886DFF8EFFAF3127FF6C3A5E24FA" box="[570,618,620,644]" pageId="3">type</typeStatus>
species fixation are also listed in Article 68 of the Code: monotypy, absolute tautotomy and “Linnean tautotomy”. In the present instance,
<typeStatus id="54C0886DFF8EFFAF30B3FF903BEA24D6" box="[942,990,656,680]" pageId="3">type</typeStatus>
species fixation occurred by monotypy in the original publication.
</paragraph>
</subSubSection>
</treatment>
</document>