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<mods:title id="8D5F5D99FBA1D4CC952657D0F5171BC7">Catalogue of the Australian social wasps (Hymenoptera: Vespidae)</mods:title>
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<emphasis id="B9B1EAE4B128FF8BD3AF545464C8C103" box="[247,426,1339,1365]" italics="true" pageId="7" pageNumber="458">Vespa variabilis</emphasis>
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male London); 1787: 288. -
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<bibRefCitation id="EF544B07B128FF8BD619545464B8C12F" author="Gmelin" firstAuthor="Gmelin" page="2756" pageId="7" pageNumber="458" pagination="2225 - 3020" refId="ref35012" refString="Gmelin, J. E. (1790) Caroli a Linne, Systema Naturae. Vol. I. Par V. Impensis Georg. Emanuel. Beer, Lipsiae, 796 pp. [pp. 2225 - 3020]" type="book chapter" year="1790">Gmelin, 1790: 2756</bibRefCitation>
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. -
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. -
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(list).
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<emphasis id="B9B1EAE4B128FF8BD3AF54EC64A7C1CB" box="[247,453,1411,1437]" italics="true" pageId="7" pageNumber="458">Polistes variabilis</emphasis>
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;
<treatmentCitation id="0A6410E7B128FF8BD28054EC67ABC1CB" author="Fabricius, J. C." box="[472,713,1411,1437]" page="273" pageId="7" pageNumber="458" year="1804">
<bibRefCitation id="EF544B07B128FF8BD28054EC67ABC1CB" author="Fabricius" box="[472,713,1411,1437]" firstAuthor="Fabricius" page="273" pageId="7" pageNumber="458" pagination="15 - 440" refId="ref32093" refString="Fabricius, J. C. (1804) Systema Piezatorum. Carolum Reichard, Brunsvigae, xiv + 15 - 440 + 30 pp." type="book chapter" year="1804">Fabricius, 1804: 273</bibRefCitation>
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. - de Saussure, 1853: 66. -
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<bibRefCitation id="EF544B07B128FF8BD74954EC6194C1CB" author="Froggatt" box="[1041,1270,1411,1437]" firstAuthor="Froggatt" page="233" pageId="7" pageNumber="458" pagination="205 - 248" refId="ref32309" refString="Froggatt, W. W. (1892) Catalogue of the described Hymenoptera of Australia. Part II. Proceedings of the Linnaean Society of New South Wales, Series 2, 7, 205 - 248. https: // doi. org / 10.5962 / bhl. part. 26049" type="journal article" year="1892">Froggatt, 1892: 233</bibRefCitation>
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(cat.). -
<treatmentCitation id="0A6410E7B128FF8BD63854EC6778C197" author="Dalla Torre, K. W." page="135" pageId="7" pageNumber="458" year="1894">
<bibRefCitation id="EF544B07B128FF8BD63854EC6778C197" author="Dalla Torre" firstAuthor="Dalla Torre" page="135" pageId="7" pageNumber="458" refId="ref30910" refString="Dalla Torre, K. W. (1894) Catalogus Hymenopterorum. Vol. 9. Vespidae (Diploptera). G. Engelmann, Leipzig, 181 pp." type="book" year="1894">Dalla Torre, 1894: 135</bibRefCitation>
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(cat.); 1904: 71 (cat.). -
<treatmentCitation id="0A6410E7B128FF8BD01D54C8610BC197" author="Meade-Waldo, G." box="[837,1129,1447,1473]" page="104" pageId="7" pageNumber="458" year="1911">
<bibRefCitation id="EF544B07B128FF8BD01D54C8610BC197" author="Meade-Waldo" box="[837,1129,1447,1473]" firstAuthor="Meade-Waldo" page="104" pageId="7" pageNumber="458" pagination="98 - 113" refId="ref43190" refString="Meade-Waldo, G. (1911) X. - New species of Diploptera in the collection of the British Museum. Annals and Magazine of Natural History, Series 8, 7, 98 - 113. https: // doi. org / 10.1080 / 00222931108692910" type="journal article" year="1911">Meade-Waldo, 1911: 104</bibRefCitation>
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(key). -
<treatmentCitation id="0A6410E7B128FF8BD78E54C864E3C1B3" author="Cheesman, L. E." page="991" pageId="7" pageNumber="458" year="1951">
<bibRefCitation id="EF544B07B128FF8BD78E54C864E3C1B3" author="Cheesman" firstAuthor="Cheesman" page="991" pageId="7" pageNumber="458" pagination="982 - 993" refId="ref29949" refString="Cheesman, L. E. (1951) A collection of Polistes from Papuasia in the British Museum. Annals and Magazine of Natural History, Series 12, 4, 982 - 993. https: // doi. org / 10.1080 / 00222935108654229" type="journal article" year="1951">Cheesman, 1951: 991</bibRefCitation>
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(
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compared to
<taxonomicName id="4CC54D75B128FF8BD13854A467DCC1B2" ID-CoL="78522" authority="Carpenter &amp; Brown, 2022" authorityName="Carpenter &amp; Brown" authorityYear="2022" box="[608,702,1483,1508]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="comis" status="sp. nov.">
<emphasis id="B9B1EAE4B128FF8BD13854A467DCC1B2" box="[608,702,1483,1508]" italics="true" pageId="7" pageNumber="458">P. comis</emphasis>
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<taxonomicNameLabel id="A282579FB128FF8BD19F54A46665C1B3" box="[711,775,1483,1509]" pageId="7" pageNumber="458" rank="species">n. sp.</taxonomicNameLabel>
). -
<treatmentCitation id="0A6410E7B128FF8BD07654A4617EC1B3" author="Yoshikawa, K." box="[814,1052,1483,1509]" page="40" pageId="7" pageNumber="458" year="1962">
<bibRefCitation id="EF544B07B128FF8BD07654A4617EC1B3" author="Yoshikawa" box="[814,1052,1483,1509]" firstAuthor="Yoshikawa" page="40" pageId="7" pageNumber="458" pagination="19 - 44" refId="ref53894" refString="Yoshikawa, K. (1962) Introductory studies on the life economy of polistine wasps VI. Geographical Distribution and its ecological significances. Journal of Biology Osaka City University, 13, 19 - 44." type="journal article" year="1962">Yoshikawa, 1962: 40</bibRefCitation>
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(list). -
<treatmentCitation id="0A6410E7B128FF8BD72354A46030C1B3" author="Richards, O. W." box="[1147,1362,1483,1509]" page="10" pageId="7" pageNumber="458" year="1978">
<bibRefCitation id="EF544B07B128FF8BD72354A46030C1B3" author="Richards" box="[1147,1362,1483,1509]" firstAuthor="Richards" page="10" pageId="7" pageNumber="458" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 10</bibRefCitation>
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(key), 40 (in subgenus
<taxonomicName id="4CC54D75B128FF8BD144548067E5C25F" authorityName="Ashmead, 1904" box="[540,647,1519,1545]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="subGenus" subGenus="Polistella">
<emphasis id="B9B1EAE4B128FF8BD144548067E5C25F" box="[540,647,1519,1545]" italics="true" pageId="7" pageNumber="458">Polistella</emphasis>
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; redescription; Qld.). -
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<bibRefCitation id="EF544B07B128FF8BD0F9548061E2C25F" author="Cardale" box="[929,1152,1518,1545]" firstAuthor="Cardale" page="213" pageId="7" pageNumber="458" pagination="150 - 303" refId="ref28558" refString="Cardale, J. C. (1985) Vespoidea and Sphecoidea. Zoological Catalogue of Australia, 2, 150 - 303." type="journal article" year="1985">Cardale, 1985: 213</bibRefCitation>
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(in subgenus
<taxonomicName id="4CC54D75B128FF8BD671548060F6C25F" authorityName="Ashmead, 1904" box="[1321,1428,1519,1545]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="subGenus" subGenus="Polistella">
<emphasis id="B9B1EAE4B128FF8BD671548060F6C25F" box="[1321,1428,1519,1545]" italics="true" pageId="7" pageNumber="458">Polistella</emphasis>
</taxonomicName>
; cat.). -
<treatmentCitation id="0A6410E7B128FF8BD2F3577C67F8C27B" author="Carpenter, J. M." box="[427,666,1554,1581]" page="54" pageId="7" pageNumber="458" year="1996">
<bibRefCitation id="EF544B07B128FF8BD2F3577C67F8C27B" author="Carpenter" box="[427,666,1554,1581]" firstAuthor="Carpenter" page="54" pageId="7" pageNumber="458" pagination="18 - 57" refId="ref28663" refString="Carpenter, J. M. (1996 a) Phylogeny and biogeography of Polistes. In: Turillazzi, S. &amp; West-Eberhard, M. J. (Eds.), Natural History and Evolution of Paper-Wasps. Oxford University Press, Oxford, pp. 18 - 57." type="book chapter" year="1996" yearSuffix="a">Carpenter, 1996a: 54</bibRefCitation>
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(in subgenus
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<emphasis id="B9B1EAE4B128FF8BD066577C66CBC27B" box="[830,937,1555,1581]" italics="true" pageId="7" pageNumber="458">Polistella</emphasis>
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); 1996b: 17 (in subgenus
<taxonomicName id="4CC54D75B128FF8BD782577C6027C27B" authorityName="Ashmead, 1904" box="[1242,1349,1555,1581]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="subGenus" subGenus="Polistella">
<emphasis id="B9B1EAE4B128FF8BD782577C6027C27B" box="[1242,1349,1555,1581]" italics="true" pageId="7" pageNumber="458">Polistella</emphasis>
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; list). -
<treatmentCitation id="0A6410E7B128FF8BD20F5758677CC207" author="Kojima, J." box="[343,542,1591,1617]" page="27" pageId="7" pageNumber="458" year="1998">
<bibRefCitation id="EF544B07B128FF8BD20F5758677CC207" author="Kojima" box="[343,542,1591,1617]" firstAuthor="Kojima" page="27" pageId="7" pageNumber="458" pagination="7 - 227" refId="ref39791" refString="Kojima, J. (1998) Larvae of social wasps (Insecta: Hymenoptera; Vespidae). Natural History Bulletin of Ibaraki University, 2, 7 - 227." type="journal article" year="1998">Kojima, 1998: 27</bibRefCitation>
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, 213, Figs 126141 (larva). -
<treatmentCitation id="0A6410E7B128FF8BD03257586076C207" author="Kojima, J. &amp; Carpenter, J. M." box="[874,1300,1591,1617]" page="117" pageId="7" pageNumber="458" year="1997">
<bibRefCitation id="EF544B07B128FF8BD03257586076C207" author="Kojima &amp; Carpenter" box="[874,1300,1591,1617]" firstAuthor="Kojima" page="117 - 122" pageId="7" pageNumber="458" pagination="117 - 122" refId="ref40349" refString="Kojima, J. &amp; Carpenter, J. M. (1997 b) A taxonomic note and nest description of an Australian paper wasp, Polistes variabilis (Fabricius) (Hymenoptera, Vespidae, Polistinae). Japanese Journal of Systematic Entomology, 3 (2), 117 - 122." type="journal article" year="1997" yearSuffix="b">Kojima &amp; Carpenter, 1997b: 117122</bibRefCitation>
</treatmentCitation>
, Figs 1, 3, 6 (diagnostic characters; nest; Qld.). -
<bibRefCitation id="EF544B07B128FF8BD1AB573466ABC223" author="Santos" box="[755,969,1626,1653]" etAl="et al." firstAuthor="Santos" pageId="7" pageNumber="458" pagination="535 - 549" refId="ref47729" refString="Santos, B. F., Payne, A., Pickett, K. M. &amp; Carpenter, J. M. (2015) Phylogeny and historical biogeography of the paper wasp genus Polistes (Hymenoptera: Vespidae): implications for the overwintering hypothesis of social evolution. Cladistics, 31, 535 - 549. https: // doi. org / 10.1111 / cla. 12103" type="journal article" year="2015">
Santos
<emphasis id="B9B1EAE4B128FF8BD01E5734661DC223" box="[838,895,1627,1653]" italics="true" pageId="7" pageNumber="458">et al.</emphasis>
, 2015
</bibRefCitation>
: Figs 18, 9 (phylogenetic relationships). -
<bibRefCitation id="EF544B07B128FF8BD2305710672BC2CF" author="Nguyen" box="[360,585,1663,1689]" etAl="et al." firstAuthor="Nguyen" pageId="7" pageNumber="458" pagination="638 - 644" refId="ref45147" refString="Nguyen, L. T. P., Nguyen, A. D., Nguyen, T. T. P., Perrard, A. &amp; Carpenter, J. M. (2018) Molecular phylogeny of the paper wasp subgenus Polistes (Polistella) Ashmead, 1904 (Hymenoptera: Vespidae: Polistinae) from Vietnam. Journal of Asia-Pacific Entomology, 21, 638 - 644. https: // doi. org / 10.1016 / j. aspen. 2018.04.006" type="journal article" year="2018">
Nguyen
<emphasis id="B9B1EAE4B128FF8BD29F5710649DC2CF" box="[455,511,1663,1689]" italics="true" pageId="7" pageNumber="458">et al.</emphasis>
, 2018
</bibRefCitation>
: Figs 13 (phylogenetic relationships).
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</paragraph>
<paragraph id="8B7A36F6B128FF8BD3AF57CC665AC2EB" blockId="7.[151,1437,1303,1833]" box="[247,824,1698,1725]" pageId="7" pageNumber="458">
<treatmentCitationGroup id="ABD511D8B128FF8BD3AF57CC665AC2EB" box="[247,824,1698,1725]" pageId="7" pageNumber="458">
<taxonomicName id="4CC54D75B128FF8BD3AF57CC672EC2EB" ID-CoL="4KYKB" box="[247,588,1699,1725]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="variety" species="humilis" variety="variabilis">
<emphasis id="B9B1EAE4B128FF8BD3AF57CC64CBC2EB" box="[247,425,1699,1725]" italics="true" pageId="7" pageNumber="458">Polistes humilis</emphasis>
var.
<emphasis id="B9B1EAE4B128FF8BD2B957CC672EC2EB" box="[481,588,1699,1725]" italics="true" pageId="7" pageNumber="458">variabilis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B128FF8BD10357CC6651C2EB" author="Bequaert, J." box="[603,819,1698,1725]" page="58" pageId="7" pageNumber="458" year="1928">
<bibRefCitation id="EF544B07B128FF8BD10357CC6651C2EB" author="Bequaert" box="[603,819,1698,1725]" firstAuthor="Bequaert" page="58" pageId="7" pageNumber="458" pagination="53 - 63" refId="ref26065" refString="Bequaert, J. (1928) The diplopterous wasps of Fabricius, in the Banksian collection of the British Museum. Bulletin of the Brooklyn Entomological Society, 23, 53 - 63." type="journal article" year="1928">Bequaert, 1928: 58</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B7A36F6B128FF8BD3AF57A86603C353" blockId="7.[151,1437,1303,1833]" pageId="7" pageNumber="458">
<treatmentCitationGroup id="ABD511D8B128FF8BD3AF57A86603C353" pageId="7" pageNumber="458">
<taxonomicName id="4CC54D75B128FF8BD3AF57A8677EC2B7" ID-CoL="4KYKB" box="[247,540,1735,1761]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="subSpecies" species="humilis" subSpecies="variabilis">
<emphasis id="B9B1EAE4B128FF8BD3AF57A8677EC2B7" box="[247,540,1735,1761]" italics="true" pageId="7" pageNumber="458">Polistes humilis variabilis</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A6410E7B128FF8BD17457A8662EC2B7" author="Giordani Soika, A." box="[556,844,1734,1761]" page="21" pageId="7" pageNumber="458" year="1975">
<bibRefCitation id="EF544B07B128FF8BD17457A8662EC2B7" author="Giordani Soika" box="[556,844,1734,1761]" firstAuthor="Giordani Soika" page="21" pageId="7" pageNumber="458" pagination="20 - 25" refId="ref34922" refString="Giordani Soika, A. (1975) Notulae vespidologicae XXXVII. Nuovi Polistes del continente australiano (Hymenoptera Vespidae). Bollettino della Societa Entomologica Italiana, 107 (1 - 2), 20 - 25." type="journal article" year="1975">Giordani Soika, 1975: 21</bibRefCitation>
</treatmentCitation>
. -
<treatmentCitation id="0A6410E7B128FF8BD03057A8615FC2B7" author="Richards, O. W." box="[872,1085,1735,1761]" page="42" pageId="7" pageNumber="458" year="1978">
<bibRefCitation id="EF544B07B128FF8BD03057A8615FC2B7" author="Richards" box="[872,1085,1735,1761]" firstAuthor="Richards" page="42" pageId="7" pageNumber="458" pagination="1 - 132" refId="ref46880" refString="Richards, O. W. (1978) The Australian social wasps (Hymenoptera: Vespidae). Australian Journal of Zoology, Supplementary Series, 61, 1 - 132. https: // doi. org / 10.1071 / AJZS 061" type="journal article" year="1978">Richards, 1978: 42</bibRefCitation>
</treatmentCitation>
(compared with
<taxonomicName id="4CC54D75B128FF8BD7A657A860F7C2B7" ID-CoL="4KYHK" authorityName="Giordani Soika" authorityYear="1975" box="[1278,1429,1735,1761]" class="Insecta" family="Eumenidae" genus="Polistes" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="sgarambus">
<emphasis id="B9B1EAE4B128FF8BD7A657A860F7C2B7" box="[1278,1429,1735,1761]" italics="true" pageId="7" pageNumber="458">P. sgarambus</emphasis>
</taxonomicName>
; “seems to be very rare in
<collectingRegion id="4901F814B128FF8BD12D5784662EC353" box="[629,844,1771,1797]" country="Australia" name="Northern Territory" pageId="7" pageNumber="458">Northern Territory</collectingRegion>
”).
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</paragraph>
</subSubSection>
<subSubSection id="C3DF657DB128FF8BD3AF5660672AC37F" box="[247,584,1807,1833]" pageId="7" pageNumber="458" type="distribution">
<paragraph id="8B7A36F6B128FF8BD3AF5660672AC37F" blockId="7.[151,1437,1303,1833]" box="[247,584,1807,1833]" pageId="7" pageNumber="458">Distribution: NE coastal; Qld.</paragraph>
</subSubSection>
</treatment>
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