346 lines
49 KiB
XML
346 lines
49 KiB
XML
<document id="DACA9BCA45BB0509C5C258DBBB59996F" ID-CLB-Dataset="54646" ID-DOI="10.11646/zootaxa.5278.1.1" ID-GBIF-Dataset="1528757a-5499-40b9-a728-c5bf0c770b27" ID-ISSN="1175-5326" ID-Zenodo-Dep="7894826" ID-ZooBank="BE0A07DB-170A-4B72-8178-9B8144C1FBF6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1683182638917" checkinUser="plazi" docAuthor="Archibald, S. Bruce & Rasnitsyn, Alexandr P." docDate="2023" docId="A148D509FFACFFE1FF6D6E60CDE5FED4" docLanguage="en" docName="zootaxa.5278.1.1.pdf" docOrigin="Zootaxa 5278 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5278.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptostigma brevilatum Archibald & Rasnitsyn 2023, n. sp." docType="treatment" docUuid="B29C5890-4E50-4408-8D67-0E073DB25546" docUuidSource="ZooBank" docVersion="9" lastPageNumber="13" masterDocId="5D71AD71FFABFFEDFFFA6B55C878FFC9" masterDocTitle="Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands" masterLastPageNumber="38" masterPageNumber="1" pageNumber="8" updateTime="1728574925831" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="EE20EEB91207DDC9C196854F4535DE11">Cimbicidae (Hymenoptera, ‘ Symphyta’) in the Paleogene: revision, the new subfamily Cenocimbicinae, and new taxa from the Eocene Okanagan Highlands</mods:title>
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<mods:name id="A4450C32776D256C50E1E14787F8AE03" type="personal">
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<mods:roleTerm id="F9D976CE8863497BACE69F40E65AE97E">Author</mods:roleTerm>
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<mods:namePart id="2AD3D14CBA79F6D594A575E1587A6152">Archibald, S. Bruce</mods:namePart>
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<mods:affiliation id="C25B97432EA75FA48052C7E9234D86DB">Department of Earth, Ocean and Atmospheric Sciences, University of British Columbia, 2020 - 2207 Main Mall, Vancouver, British Columbia, Canada, V 6 T 1 Z 4, Canada. bruce. archibald @ ubc. ca; https: // orcid. org / 0000 - 0002 - 4397 - 2497 & Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V 5 A 1 S 6, Canada Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America & Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V 8 W 9 W 2, Canada</mods:affiliation>
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<mods:nameIdentifier id="5497A150E82DFE4D40CE89BCD5895A24" type="email">bruce.archibald@ubc.ca</mods:nameIdentifier>
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<mods:namePart id="B4EC21C1750E77396E14950D3AF0B1D0">Rasnitsyn, Alexandr P.</mods:namePart>
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<mods:affiliation id="288D037BCFC7BFFD0FEF370C01461E56">A. A. Borissiak Paleontological Institute, Russian Academy of Sciences, Moscow 117647, Russia & Invertebrate Paleontology Department, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom Corresponding author. bruce. archibald @ ubc. ca</mods:affiliation>
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<mods:title id="9E82D186C6CE1DAFE77B1A80F61A6B2C">Zootaxa</mods:title>
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<mods:part id="C7EEB88393C3D6AE345B6A838D3D1268">
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<mods:date id="3587E84FE56ADE8EAB313F070666854D">2023</mods:date>
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<mods:number id="9522AE1CE4AA6462CB67B108454CFB08">2023-05-04</mods:number>
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<mods:number id="471656104B2DBBE93E02523806BD4A91">5278</mods:number>
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<treatment id="A148D509FFACFFE1FF6D6E60CDE5FED4" ID-DOI="http://doi.org/10.5281/zenodo.7894856" ID-GBIF-Taxon="209583692" ID-Zenodo-Dep="7894856" LSID="urn:lsid:zoobank.org:act:B29C5890-4E50-4408-8D67-0E073DB25546" httpUri="http://treatment.plazi.org/id/A148D509FFACFFE1FF6D6E60CDE5FED4" lastPageId="12" lastPageNumber="13" pageId="7" pageNumber="8">
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<subSubSection id="61FB3794FFACFFEAFF6D6E60C987FA86" box="[151,511,1333,1360]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="295E641FFFACFFEAFF6D6E60C987FA86" blockId="7.[151,1163,1333,1432]" box="[151,511,1333,1360]" pageId="7" pageNumber="8">
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<heading id="7216D373FFACFFEAFF6D6E60C987FA86" bold="true" box="[151,511,1333,1360]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="1B95B80DFFACFFEAFF6D6E60C987FA86" bold="true" box="[151,511,1333,1360]" pageId="7" pageNumber="8">
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<taxonomicName id="EEE11F9CFFACFFEAFF6D6E60C9CDFA99" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,437,1333,1360]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
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<emphasis id="1B95B80DFFACFFEAFF6D6E60C9CDFA99" bold="true" box="[151,437,1333,1360]" italics="true" pageId="7" pageNumber="8">Leptostigma brevilatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00A60576FFACFFEAFE476E60C987FA86" box="[445,511,1333,1359]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="61FB3794FFACFFEAFF6D6E0CC91EFA51" pageId="7" pageNumber="8" type="description">
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<uuid id="5D475ECAFFACFFEAFD796E0CCCF3FABD" box="[643,1163,1369,1396]" pageId="7" pageNumber="8">B29C5890-4E50-4408-8D67-0E073DB25546</uuid>
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</paragraph>
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<paragraph id="295E641FFFACFFEAFF6D6E28C91EFA51" blockId="7.[151,1163,1333,1432]" box="[151,358,1405,1432]" pageId="7" pageNumber="8">
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<figureCitation id="B1DA789AFFACFFEAFF6D6E28C963FA5E" box="[151,283,1405,1431]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="7" pageNumber="8">Figs. 3A–B</figureCitation>
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,
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<figureCitation id="B1DA789AFFACFFEAFEDC6E28C91AFA51" box="[294,354,1405,1432]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStart-5="FIGURE 10" captionStartId-0="8.[151,250,1970,1994]" captionStartId-1="9.[151,250,1927,1951]" captionStartId-2="10.[151,250,838,862]" captionStartId-3="10.[151,250,1977,2001]" captionStartId-4="11.[151,250,1136,1160]" captionStartId-5="11.[151,250,1987,2011]" captionTargetBox-0="[193,1394,741,1940]" captionTargetBox-1="[163,1426,705,1888]" captionTargetBox-2="[287,1329,181,796]" captionTargetBox-3="[253,1334,922,1945]" captionTargetBox-4="[310,1276,181,1104]" captionTargetBox-5="[303,1284,1221,1950]" captionTargetId-0="figure-305@8.[193,1394,741,1946]" captionTargetId-1="figure-293@9.[151,1436,705,1903]" captionTargetId-2="figure-20@10.[258,1329,181,814]" captionTargetId-3="figure-42@10.[253,1334,922,1953]" captionTargetId-4="figure-18@11.[310,1276,181,1112]" captionTargetId-5="figure-64@11.[303,1284,1221,1963]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="11" captionText-0="FIGURE 5. Leptostigma brevilatum, holotype GSC 142839. A, photograph, and B–D, drawings: B, right forewing; C, left forewing; D, right hind wing. All to scale, 5 mm." captionText-1="FIGURE 6. F-1182, tentative Leptostigma brevilatum, A, photograph, and B–E, drawings of: B, left forewing; C, right forewing; D, left hindwing, and; E, right hind wing. All to scale, 5 mm." captionText-2="FIGURE 7. UWBM-54840, tentative Leptostigma brevilatum: A, photograph; B, drawing from part and counterpart. Scale bar 5 mm." captionText-3="FIGURE 8. RBCM-EH2017-050-4628-001, tentative Leptostigma brevilatum, A, photograph, and B–C, drawings of the wings: B, right forewing; C, right hindwing.All to scale, 5 mm." captionText-4="FIGURE 9. F-774, tentative Leptostigma brevilatum:A–B photographs and C–D, wing drawings:A, the whole fossil; B, closeup with arrow indicating the hind apical spur; C, right forewing; D, left forewing.A, C–D to 5 mm scale; B scale is 2 mm." captionText-5="FIGURE 10. GSC 142840, tentative Leptostigma brevilatum: A, photograph, B, drawing of the forewing; scale is 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7894846" figureDoi-1="http://doi.org/10.5281/zenodo.7894854" figureDoi-2="http://doi.org/10.5281/zenodo.7894858" figureDoi-3="http://doi.org/10.5281/zenodo.7894860" figureDoi-4="http://doi.org/10.5281/zenodo.7894866" figureDoi-5="http://doi.org/10.5281/zenodo.7894868" httpUri-0="https://zenodo.org/record/7894846/files/figure.png" httpUri-1="https://zenodo.org/record/7894854/files/figure.png" httpUri-2="https://zenodo.org/record/7894858/files/figure.png" httpUri-3="https://zenodo.org/record/7894860/files/figure.png" httpUri-4="https://zenodo.org/record/7894866/files/figure.png" httpUri-5="https://zenodo.org/record/7894868/files/figure.png" pageId="7" pageNumber="8">5–10</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFACFFEAFF6D6E95CBBCF9EB" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="295E641FFFACFFEAFF6D6E95CBBCF9EB" blockId="7.[151,1437,1472,2039]" pageId="7" pageNumber="8">
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<materialsCitation id="99896E42FFACFFEAFF6D6E95CBBCF9EB" ID-GBIF-Occurrence="4100387312" collectionCode="GSC" pageId="7" pageNumber="8" specimenCode="GSC 142839" specimenCount="1" typeStatus="holotype">
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<emphasis id="1B95B80DFFACFFEAFF6D6E95C97CFA13" bold="true" box="[151,260,1472,1498]" pageId="7" pageNumber="8">
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<typeStatus id="F65ADABDFFACFFEAFF6D6E95C97CFA13" box="[151,260,1472,1498]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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</emphasis>
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.
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<specimenCode id="7947CC64FFACFFEAFEEE6E95C9DFFA13" box="[276,423,1472,1498]" collectionCode="GSC" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="7" pageNumber="8">GSC 142839</specimenCode>
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(
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<figureCitation id="B1DA789AFFACFFEAFE426E95CA4CFA13" box="[440,564,1472,1498]" captionStart="FIGURE 3" captionStartId="5.[151,250,1833,1857]" captionTargetBox="[192,1395,191,1809]" captionTargetId="figure-18@5.[192,1395,181,1809]" captionTargetPageId="5" captionText="FIGURE 3. Heads of Cenocimbicinae: A, Leptostigma brevilatum GSC 142839, drawing; B, L. brevilatum GSC 142839, photograph; C, Leptostigma longipallidum GSC 142841, drawing; D, L. longipallidum GSC 142841, photograph; E, Cenocimbex menatensis MNHN-LP-R.11195, drawing; F, C. menatensis MNHN-LP-R.11195, photograph; G, Leptostigma longipallidum F-771, drawing; H, L. longipallidum F-771, photograph; I, Leptostigma longitenebricum F-1565, drawing; J, L. longitenebricum F-1565, photograph; K, Leptostigma longiclava RBCM.EH2017.050.4085.0001, drawing; L, L. longiclava RBCM.EH2017.050.4085.0001, photograph. Scale bars 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894836" httpUri="https://zenodo.org/record/7894836/files/figure.png" pageId="7" pageNumber="8">Fig. 3A–B</figureCitation>
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,
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<figureCitation id="B1DA789AFFACFFEAFDBB6E95CA29FA12" box="[577,593,1472,1499]" captionStart="FIGURE 5" captionStartId="8.[151,250,1970,1994]" captionTargetBox="[193,1394,741,1940]" captionTargetId="figure-305@8.[193,1394,741,1946]" captionTargetPageId="8" captionText="FIGURE 5. Leptostigma brevilatum, holotype GSC 142839. A, photograph, and B–D, drawings: B, right forewing; C, left forewing; D, right hind wing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894846" httpUri="https://zenodo.org/record/7894846/files/figure.png" pageId="7" pageNumber="8">5</figureCitation>
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). Head with antennae (one incomplete), thorax, perhaps part of abdomen, parts of several legs, both forewings, both hind wings poorly preserved, collector SBA at McAbee (
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<collectionCode id="4FF0FCDAFFACFFEAFAF26EB1CD38FA37" box="[1288,1344,1508,1534]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="7" pageNumber="8">GSC</collectionCode>
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locality V-016800), pre-1999, housed in the collections of the
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<collectionCode id="4FF0FCDAFFACFFEAFD0F6D5DCB53F9EB" box="[757,811,1544,1570]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="7" pageNumber="8">GSC</collectionCode>
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(Vancouver).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFACFFEAFF3D6D79C9C7F91F" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="295E641FFFACFFEAFF3D6D79C9C7F91F" blockId="7.[151,1437,1472,2039]" pageId="7" pageNumber="8">
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<emphasis id="1B95B80DFFACFFEAFF3D6D79C93AF98F" bold="true" box="[199,322,1580,1606]" pageId="7" pageNumber="8">Diagnosis.</emphasis>
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Distinguished from other species of
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<taxonomicName id="EEE11F9CFFACFFEAFD246D78CB10F98F" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[734,872,1581,1606]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="1B95B80DFFACFFEAFD246D78CB10F98F" box="[734,872,1581,1606]" italics="true" pageId="7" pageNumber="8">Leptostigma</emphasis>
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</taxonomicName>
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by a combination of antenna dark, with club short and wide, with two basal flagellar segments comparatively short (combined shorter than half head width). Forewing lightly, evenly infuscate throughout. Tentatively diagnostic: R+M shorter than 1-M, 2r-rs enter cell 3rm at or before its middle, 2r-m enters M far behind 2m-cu, anal cells meet at crossvein or at most minimally separated by short fusion of 1A with 2A+3A.
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</paragraph>
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</subSubSection>
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<subSubSection id="61FB3794FFACFFE5FF3D6DB5C891FD04" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
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<paragraph id="295E641FFFACFFE5FF3D6DB5C891FD04" blockId="7.[151,1437,1472,2039]" lastBlockId="8.[151,1437,151,717]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="1B95B80DFFACFFEAFF3D6DB5C921F933" bold="true" box="[199,345,1760,1786]" pageId="7" pageNumber="8">Description.</emphasis>
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<typeStatus id="F65ADABDFFACFFEAFE9E6DB5C9A8F933" box="[356,464,1760,1786]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
|
||
. Body length not known by preservation. Body as preserved (including head with antenna, thorax, femora, tibiae, abdominal base) and pterostigma dark, membrane lightly infuscate throughout, tarsal fragments (at least hind tarsomeres 1–3) darkened. Head (interpretation tentative by imperfect preservation) wider than long, with compound eyes oval, rather short, converging below, rather close to mandibles; ocelli not known, toruli close to each other, touching clypeus; clypeus small, excised apically; labrum oval, narrower and slightly shorter than clypeus; temples not inflated, scarcely exceeding eye contour.Antenna apparently with 9 antennomeres; scape, pedicel small, poorly preserved; flagellomeres possibly gradually becoming shorter, wider (but precise length sometimes obscure) except apical two of similar length; club somewhat unclear, apparently 3-segmented, short, rounded apically. Thorax poorly known by preservation, with notauli and median longitudinal mesonotal sulcus distinct, meeting well before transverse scutellum. Forewing length 13 (
|
||
<typeStatus id="F65ADABDFFA3FFE5FC286BEECC41FF1C" box="[978,1081,187,213]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
)–
|
||
<quantity id="EE19C9FAFFA3FFE5FBB56BEECCD8FF1C" box="[1103,1184,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="8" pageNumber="9" unit="mm" value="17.0">17 mm</quantity>
|
||
, maximum width
|
||
<quantity id="EE19C9FAFFA3FFE5FA886BEFC8BDFF30" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="8" pageNumber="9" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
, length/width: 3.1–3.3. Pterostigma narrow, similar to width of C-R; R+M shorter than 1-M; 2r-rs before middle of cell 3rm (near middle in UWBM 54840); 2r-rm distinctly curved (F-1182 with aberrant short stub near middle in both wings), joins Rs about same level as 2m-cu joining M, joining M behind 2m-cu for more than half of 2r-m length (in F-1182, slightly less than that in right wing); 1-M straight or almost so; 3r-m almost straight; cell 1mcu with M and 1m-cu distinctly to slightly (in GSC 142840) diverging toward anterior margin, aberrant (split) in left wing of F-1182; cell 2mcu with M usually shorter than 1m-cu (as long as that in GSC 142840, slightly longer than that in F-1182); 2m-cu curved distinctly toward base (weakly so in GSC 142840); cu-a distinctly postfurcal; anal cells meet at crossvein. Hind wing with 3r-m, m-cu, cu-a present, 3r-m slightly more than its own length from m-cu; cu-a before middle of cell mcu, more than for its length before m-cu (aberrantly two cu-a present in
|
||
<typeStatus id="F65ADABDFFA3FFE5FB116AAACD35FDD0" box="[1259,1357,511,537]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
placed before and behind middle of cell mcu: which is true crossvein unclear). Legs long, narrow, hind femur widest subbasally, 2.9 times as long as wide, with lower margin smooth, hind tibia
|
||
<emphasis id="1B95B80DFFA3FFE5FC366912CB9EFDA9" box="[972,998,583,608]" italics="true" pageId="8" pageNumber="9">ca</emphasis>
|
||
. 6 times as long as wide apically, hind apical spur long (as long as basitarsus, almost 0.4 times as long as tibia,
|
||
<figureCitation id="B1DA789AFFA3FFE5FC3F693ECC6CFD4C" box="[965,1044,619,645]" captionStart="FIGURE 9" captionStartId="11.[151,250,1136,1160]" captionTargetBox="[310,1276,181,1104]" captionTargetId="figure-18@11.[310,1276,181,1112]" captionTargetPageId="11" captionText="FIGURE 9. F-774, tentative Leptostigma brevilatum:A–B photographs and C–D, wing drawings:A, the whole fossil; B, closeup with arrow indicating the hind apical spur; C, right forewing; D, left forewing.A, C–D to 5 mm scale; B scale is 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894866" httpUri="https://zenodo.org/record/7894866/files/figure.png" pageId="8" pageNumber="9">Fig 9B</figureCitation>
|
||
), sharp, tarsomere 1–3 length ratio 1: 0.7: 0.45 (in GSC 142840, femora, tibiae apparently narrower, but difficult to measure), with claw apparently simple.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFA3FFE5FF6D6CE7CAF0F827" ID-DOI="http://doi.org/10.5281/zenodo.7894846" ID-Zenodo-Dep="7894846" httpUri="https://zenodo.org/record/7894846/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1970,1994]" targetBox="[193,1394,741,1940]" targetPageId="8" targetType="figure">
|
||
<paragraph id="295E641FFFA3FFE5FF6D6CE7CAF0F827" blockId="8.[151,1436,1970,2030]" pageId="8" pageNumber="9">
|
||
<emphasis id="1B95B80DFFA3FFE5FF6D6CE7C96EF803" bold="true" box="[151,278,1970,1994]" pageId="8" pageNumber="9">FIGURE 5.</emphasis>
|
||
<taxonomicName id="EEE11F9CFFA3FFE5FEE56CE6CA6AF803" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[287,530,1970,1994]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA3FFE5FEE56CE6CA6AF803" box="[287,530,1970,1994]" italics="true" pageId="8" pageNumber="9">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
, holotype GSC 142839. A, photograph, and B–D, drawings: B, right forewing; C, left forewing; D, right hind wing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFA2FFE4FF3D6BC2CC61FE40" pageId="9" pageNumber="10" type="discussion">
|
||
<paragraph id="295E641FFFA2FFE4FF3D6BC2CC61FE40" blockId="9.[151,1437,151,681]" pageId="9" pageNumber="10">
|
||
<emphasis id="1B95B80DFFA2FFE4FF3D6BC2C943FF78" bold="true" box="[199,315,151,177]" pageId="9" pageNumber="10">Remarks.</emphasis>
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FEB26BC2C99BFF78" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[328,483,151,177]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FEB26BC2C99BFF78" box="[328,483,151,177]" italics="true" pageId="9" pageNumber="10">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FE0A6BC2CA4DFF78" bold="true" box="[496,565,151,177]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FE0A6BC2CA4DFF78" box="[496,565,151,177]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from all congeners with known antenna (
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FB7D6BC2CD58FF78" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1159,1312,151,177]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FB7D6BC2CD58FF78" box="[1159,1312,151,177]" italics="true" pageId="9" pageNumber="10">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FAD76BC2CD09FF78" bold="true" box="[1325,1393,151,177]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FAD76BC2CD09FF78" box="[1325,1393,151,177]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FA7C6BC2C94CFF1C" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FA7C6BC2C94CFF1C" italics="true" pageId="9" pageNumber="10">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FEC16BEEC902FF1C" bold="true" box="[315,378,187,213]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FEC16BEEC902FF1C" box="[315,378,187,213]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FE726BEECA25FF1C" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[392,605,187,213]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FE726BEECA25FF1C" box="[392,605,187,213]" italics="true" pageId="9" pageNumber="10">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FD9E6BEECADAFF1C" bold="true" box="[612,674,187,213]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FD9E6BEECADAFF1C" box="[612,674,187,213]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) by its short and wide club. It additionally differs venationally from
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FF6D6B8AC954FF30" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,300,223,249]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FF6D6B8AC954FF30" box="[151,300,223,249]" italics="true" pageId="9" pageNumber="10">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FECF6B8AC90DFF30" bold="true" box="[309,373,223,249]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FECF6B8AC90DFF30" box="[309,373,223,249]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having 2r-rs at or before middle of cell 3rm, from
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FC206B8ACCEEFF30" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[986,1174,223,249]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longipallidum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FC206B8ACCEEFF30" box="[986,1174,223,249]" italics="true" pageId="9" pageNumber="10">L. longipallidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FB646B8ACCA6FF30" bold="true" box="[1182,1246,223,249]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FB646B8ACCA6FF30" box="[1182,1246,223,249]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, as well as from
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FF6D6A56C916FED4" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[151,366,259,285]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longitenebricum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FF6D6A56C916FED4" box="[151,366,259,285]" italics="true" pageId="9" pageNumber="10">L. longitenebricum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FE8C6A56C9CFFED4" bold="true" box="[374,439,259,285]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FE8C6A56C9CFFED4" box="[374,439,259,285]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in having R+M shorter than 1-M and by its anal cells being either separated by a short crossvein or at most by 1A and 2A+3A fused for a very short distance. Differs from
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FBCE6A72CCBAFE89" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1076,1218,295,320]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="proxrvena" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FBCE6A72CCBAFE89" box="[1076,1218,295,320]" italics="true" pageId="9" pageNumber="10">L. proxrvena</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FB326A72CD7EFE88" bold="true" box="[1224,1286,295,321]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FB326A72CD7EFE88" box="[1224,1286,295,321]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FAE96A72CDE4FE88" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[1299,1436,295,321]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fasciatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FAE96A72CDE4FE88" box="[1299,1436,295,321]" italics="true" pageId="9" pageNumber="10">L. fasciatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FF6D6A1EC8AFFEAC" bold="true" box="[151,215,331,357]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FF6D6A1EC8AFFEAC" box="[151,215,331,357]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FF1C6A1EC9FFFEAC" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[230,391,331,357]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="alaemacula" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FF1C6A1EC9FFFEAC" box="[230,391,331,357]" italics="true" pageId="9" pageNumber="10">L. alaemacula</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FE746A1EC9B5FEAC" bold="true" box="[398,461,331,357]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FE746A1EC9B5FEAC" box="[398,461,331,357]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with unknown antenna in having wing membrane lightly, evenly infuscate throughout (neither fascia nor macula) and 2r-m entering M far behind, not close to, 2m-cu.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFA2FFE4FF3D6AC7CB42FD60" pageId="9" pageNumber="10" type="materials_examined">
|
||
<paragraph id="295E641FFFA2FFE4FF3D6AC7CB42FD60" blockId="9.[151,1437,151,681]" pageId="9" pageNumber="10">
|
||
<emphasis id="1B95B80DFFA2FFE4FF3D6AC7C957FE65" bold="true" box="[199,303,402,428]" pageId="9" pageNumber="10">Material</emphasis>
|
||
.
|
||
<typeStatus id="F65ADABDFFA2FFE4FEC06AC6C9DAFE64" box="[314,418,403,429]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
|
||
and tentatively (by venational characters) specimens F-1182/1183 (
|
||
<figureCitation id="B1DA789AFFA2FFE4FB866AC6CCC5FE64" box="[1148,1213,403,429]" captionStart="FIGURE 6" captionStartId="9.[151,250,1927,1951]" captionTargetBox="[163,1426,705,1888]" captionTargetId="figure-293@9.[151,1436,705,1903]" captionTargetPageId="9" captionText="FIGURE 6. F-1182, tentative Leptostigma brevilatum, A, photograph, and B–E, drawings of: B, left forewing; C, right forewing; D, left hindwing, and; E, right hind wing. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894854" httpUri="https://zenodo.org/record/7894854/files/figure.png" pageId="9" pageNumber="10">Fig.6</figureCitation>
|
||
), unknown collector at McAbee, in the collections of
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FDF76AE2CA31FE18" box="[525,585,439,465]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="9" pageNumber="10" type="Herbarium">TRU</collectionCode>
|
||
;
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FDAC6AE3CAC1FE18" box="[598,697,438,465]" pageId="9" pageNumber="10">UWBM-</collectionCode>
|
||
54840 (
|
||
<figureCitation id="B1DA789AFFA2FFE4FCEA6AE2CB2DFE18" box="[784,853,439,465]" captionStart="FIGURE 7" captionStartId="10.[151,250,838,862]" captionTargetBox="[287,1329,181,796]" captionTargetId="figure-20@10.[258,1329,181,814]" captionTargetPageId="10" captionText="FIGURE 7. UWBM-54840, tentative Leptostigma brevilatum: A, photograph; B, drawing from part and counterpart. Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894858" httpUri="https://zenodo.org/record/7894858/files/figure.png" pageId="9" pageNumber="10">Fig. 7</figureCitation>
|
||
) (
|
||
<bibRefCitation id="4D7019EEFFA2FFE4FC976AE2CB8FFE18" author="Lewis, S. E." box="[877,1015,439,465]" pageId="9" pageNumber="10" pagination="15 - 19" refId="ref18244" refString="Lewis, S. E. (1992) Insects of the Klondike Mountain Formation, Republic, Washington. Washington Geology, 20, 15 - 19." type="journal article" year="1992">Lewis 1992</bibRefCitation>
|
||
:
|
||
<figureCitation id="B1DA789AFFA2FFE4FBFE6AE2CC23FE18" box="[1028,1115,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1147,1171]" captionTargetBox="[151,1391,564,1120]" captionTargetId="figure-185@4.[151,1436,561,1124]" captionTargetPageId="4" captionText="FIGURE 2. Cenocimbex menatensis Nel, holotype MNHN-LP-R.11195: A, extract from a reflectance transformation imaging (RTI) file (Cultural Heritage Imaging 2012); B, drawing of left forewing; C, drawing of right forewing preserved portion; D, left hind wing. Scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894834" httpUri="https://zenodo.org/record/7894834/files/figure.png" pageId="9" pageNumber="10">Fig. 2N</figureCitation>
|
||
as unknown
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FB096AE2CDE8FE18" authorityName="Linnaeus" authorityYear="1758" box="[1267,1424,439,465]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
|
||
), unknown collector at Republic, in the collections of the
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FCF36A8ECB1FFE3C" box="[777,871,475,501]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="9" pageNumber="10" type="Museum">UWBM</collectionCode>
|
||
;
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FC8B6A8FCBBBFE3C" box="[881,963,474,501]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="9" pageNumber="10" type="Museum">RBCM</collectionCode>
|
||
.EH2017.050.4628 (
|
||
<figureCitation id="B1DA789AFFA2FFE4FB5C6A8ECC91FE3C" box="[1190,1257,475,501]" captionStart="FIGURE 8" captionStartId="10.[151,250,1977,2001]" captionTargetBox="[253,1334,922,1945]" captionTargetId="figure-42@10.[253,1334,922,1953]" captionTargetPageId="10" captionText="FIGURE 8. RBCM-EH2017-050-4628-001, tentative Leptostigma brevilatum, A, photograph, and B–C, drawings of the wings: B, right forewing; C, right hindwing.All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894860" httpUri="https://zenodo.org/record/7894860/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
|
||
), collector John Leahy at McAbee, in the collections of the
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FD8C6AAACAB3FDD0" box="[630,715,511,537]" country="Canada" httpUri="http://grbio.org/cool/gayt-20cg" name="Royal British Columbia Museum" pageId="9" pageNumber="10" type="Museum">RBCM</collectionCode>
|
||
; F-774 (
|
||
<figureCitation id="B1DA789AFFA2FFE4FCDD6AAACB12FDD0" box="[807,874,511,537]" captionStart="FIGURE 9" captionStartId="11.[151,250,1136,1160]" captionTargetBox="[310,1276,181,1104]" captionTargetId="figure-18@11.[310,1276,181,1112]" captionTargetPageId="11" captionText="FIGURE 9. F-774, tentative Leptostigma brevilatum:A–B photographs and C–D, wing drawings:A, the whole fossil; B, closeup with arrow indicating the hind apical spur; C, right forewing; D, left forewing.A, C–D to 5 mm scale; B scale is 2 mm." figureDoi="http://doi.org/10.5281/zenodo.7894866" httpUri="https://zenodo.org/record/7894866/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
|
||
), unknown collector at McAbee, in the collections of
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FF4E6976C888FDF4" box="[180,240,547,573]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13947" lsid="urn:lsid:biocol.org:col:13947" name="Royal Cornwall Museum" pageId="9" pageNumber="10" type="Herbarium">TRU</collectionCode>
|
||
.
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FF006976C948FDF4" box="[250,304,547,573]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="9" pageNumber="10">GSC</collectionCode>
|
||
142840 (
|
||
<figureCitation id="B1DA789AFFA2FFE4FE606976C993FDF4" box="[410,491,547,573]" captionStart="FIGURE 10" captionStartId="11.[151,250,1987,2011]" captionTargetBox="[303,1284,1221,1950]" captionTargetId="figure-64@11.[303,1284,1221,1963]" captionTargetPageId="11" captionText="FIGURE 10. GSC 142840, tentative Leptostigma brevilatum: A, photograph, B, drawing of the forewing; scale is 5 mm." figureDoi="http://doi.org/10.5281/zenodo.7894868" httpUri="https://zenodo.org/record/7894868/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
|
||
), collector John Leahy at McAbee (
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FC836976CBC9FDF4" box="[889,945,547,573]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="9" pageNumber="10">GSC</collectionCode>
|
||
locality V-016800), in the collections of the
|
||
<collectionCode id="4FF0FCDAFFA2FFE4FF6D6912C8A9FDA8" box="[151,209,583,609]" country="Canada" httpUri="http://grbio.org/cool/52gs-z31g" name="Geological Survey of Canada" pageId="9" pageNumber="10">GSC</collectionCode>
|
||
; this specimen is very tentatively attributed to the species, as it is missing 3r-m, which prevents identification of the precise position of 2r-rs with respect to cell 3rm: 2r-rs is distant from 2r-m for less that its own length, more consistent with
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FEBD69DAC9A5FD60" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[327,477,655,681]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevilatum" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FEBD69DAC9A5FD60" box="[327,477,655,681]" italics="true" pageId="9" pageNumber="10">L. brevilatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FE1E69DACA5AFD60" bold="true" box="[484,546,655,681]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FE1E69DACA5AFD60" box="[484,546,655,681]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FD9B69DACA8DFD60" authority="Archibald & Rasnitsyn, 2023" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[609,757,655,681]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="longiclava" status="sp. nov.">
|
||
<emphasis id="1B95B80DFFA2FFE4FD9B69DACA8DFD60" box="[609,757,655,681]" italics="true" pageId="9" pageNumber="10">L. longiclava</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B95B80DFFA2FFE4FD0669DACB42FD60" bold="true" box="[764,826,655,681]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="00A60576FFA2FFE4FD0669DACB42FD60" box="[764,826,655,681]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="7D9E3497FFA2FFE4FF6D6CD2CB7AF80B" ID-DOI="http://doi.org/10.5281/zenodo.7894854" ID-Zenodo-Dep="7894854" httpUri="https://zenodo.org/record/7894854/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,250,1927,1951]" targetBox="[163,1426,705,1888]" targetPageId="9" targetType="figure">
|
||
<paragraph id="295E641FFFA2FFE4FF6D6CD2CB7AF80B" blockId="9.[151,1437,1927,1987]" pageId="9" pageNumber="10">
|
||
<emphasis id="1B95B80DFFA2FFE4FF6D6CD2C969F856" bold="true" box="[151,273,1927,1951]" pageId="9" pageNumber="10">FIGURE 6.</emphasis>
|
||
F-1182, tentative
|
||
<taxonomicName id="EEE11F9CFFA2FFE4FE3C6CDDCACDF856" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[454,693,1927,1951]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA2FFE4FE3C6CDDCACDF856" box="[454,693,1927,1951]" italics="true" pageId="9" pageNumber="10">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
, A, photograph, and B–E, drawings of: B, left forewing; C, right forewing; D, left hindwing, and; E, right hind wing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7D9E3497FFA1FFE7FF6D6813C8A1FC4B" ID-DOI="http://doi.org/10.5281/zenodo.7894858" ID-Zenodo-Dep="7894858" httpUri="https://zenodo.org/record/7894858/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,838,862]" targetBox="[287,1329,181,796]" targetPageId="10" targetType="figure">
|
||
<paragraph id="295E641FFFA1FFE7FF6D6813C8A1FC4B" blockId="10.[151,1436,838,898]" pageId="10" pageNumber="11">
|
||
<emphasis id="1B95B80DFFA1FFE7FF6D6813C96BFC97" bold="true" box="[151,275,838,862]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
|
||
UWBM-54840, tentative
|
||
<taxonomicName id="EEE11F9CFFA1FFE7FDE76812CB75FC97" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[541,781,838,862]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA1FFE7FDE76812CB75FC97" box="[541,781,838,862]" italics="true" pageId="10" pageNumber="11">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
: A, photograph; B, drawing from part and counterpart. Scale bar 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7D9E3497FFA1FFE7FF6D6CECCAACF83C" ID-DOI="http://doi.org/10.5281/zenodo.7894860" ID-Zenodo-Dep="7894860" httpUri="https://zenodo.org/record/7894860/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1977,2001]" targetBox="[253,1334,922,1945]" targetPageId="10" targetType="figure">
|
||
<paragraph id="295E641FFFA1FFE7FF6D6CECCAACF83C" blockId="10.[151,1437,1977,2038]" pageId="10" pageNumber="11">
|
||
<emphasis id="1B95B80DFFA1FFE7FF6D6CECC96AF818" bold="true" box="[151,274,1977,2001]" pageId="10" pageNumber="11">FIGURE 8.</emphasis>
|
||
RBCM-EH2017-050-4628-001, tentative
|
||
<taxonomicName id="EEE11F9CFFA1FFE7FD476CEFCBD5F818" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[701,941,1977,2001]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA1FFE7FD476CEFCBD5F818" box="[701,941,1977,2001]" italics="true" pageId="10" pageNumber="11">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
, A, photograph, and B–C, drawings of the wings: B, right forewing; C, right hindwing. All to scale, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7D9E3497FFA0FFE6FF6D6F25CD37FB65" ID-DOI="http://doi.org/10.5281/zenodo.7894866" ID-Zenodo-Dep="7894866" httpUri="https://zenodo.org/record/7894866/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1136,1160]" targetBox="[310,1276,181,1104]" targetPageId="11" targetType="figure">
|
||
<paragraph id="295E641FFFA0FFE6FF6D6F25CD37FB65" blockId="11.[151,1436,1136,1197]" pageId="11" pageNumber="12">
|
||
<emphasis id="1B95B80DFFA0FFE6FF6D6F25C969FB41" bold="true" box="[151,273,1136,1160]" pageId="11" pageNumber="12">FIGURE 9.</emphasis>
|
||
F-774, tentative
|
||
<taxonomicName id="EEE11F9CFFA0FFE6FE406F24CAD1FB41" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[442,681,1136,1160]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA0FFE6FE406F24CAD1FB41" box="[442,681,1136,1160]" italics="true" pageId="11" pageNumber="12">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
: A–B photographs and C–D, wing drawings:A, the whole fossil; B, closeup with arrow indicating the hind apical spur; C, right forewing; D, left forewing. A, C–D to 5 mm scale; B scale is 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7D9E3497FFA0FFE6FF6D6C96CD1AF812" ID-DOI="http://doi.org/10.5281/zenodo.7894868" ID-Zenodo-Dep="7894868" box="[151,1378,1987,2011]" httpUri="https://zenodo.org/record/7894868/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,250,1987,2011]" targetBox="[303,1284,1221,1950]" targetPageId="11" targetType="figure">
|
||
<paragraph id="295E641FFFA0FFE6FF6D6C96CD1AF812" blockId="11.[151,1378,1987,2011]" box="[151,1378,1987,2011]" pageId="11" pageNumber="12">
|
||
<emphasis id="1B95B80DFFA0FFE6FF6D6C96C958F812" bold="true" box="[151,288,1987,2011]" pageId="11" pageNumber="12">FIGURE 10.</emphasis>
|
||
GSC 142840, tentative
|
||
<taxonomicName id="EEE11F9CFFA0FFE6FDEE6C91CB7CF812" authorityName="Archibald & Rasnitsyn" authorityYear="2023" box="[532,772,1987,2011]" class="Insecta" family="Cimbicidae" genus="Leptostigma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="brevilatum">
|
||
<emphasis id="1B95B80DFFA0FFE6FDEE6C91CB7CF812" box="[532,772,1987,2011]" italics="true" pageId="11" pageNumber="12">Leptostigma brevilatum</emphasis>
|
||
</taxonomicName>
|
||
: A, photograph, B, drawing of the forewing; scale is 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="61FB3794FFA7FFE1FF3D6BC2C95DFF1C" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="295E641FFFA7FFE1FF3D6BC2C95DFF1C" blockId="12.[151,1437,151,285]" pageId="12" pageNumber="13">
|
||
<emphasis id="1B95B80DFFA7FFE1FF3D6BC2C936FF78" bold="true" box="[199,334,151,177]" pageId="12" pageNumber="13">Etymology.</emphasis>
|
||
From the Latin
|
||
<emphasis id="1B95B80DFFA7FFE1FDF96BC2CA3EFF78" box="[515,582,151,177]" italics="true" pageId="12" pageNumber="13">brevis</emphasis>
|
||
, meaning short and
|
||
<emphasis id="1B95B80DFFA7FFE1FCD06BC2CB18FF78" box="[810,864,151,177]" italics="true" pageId="12" pageNumber="13">latus</emphasis>
|
||
, wide, referring to the antennal club. The epithet is an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61FB3794FFA7FFE1FF3D6B8BCDE5FED4" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="295E641FFFA7FFE1FF3D6B8BCDE5FED4" blockId="12.[151,1437,151,285]" pageId="12" pageNumber="13">
|
||
<emphasis id="1B95B80DFFA7FFE1FF3D6B8BC9EAFF30" bold="true" box="[199,402,222,249]" pageId="12" pageNumber="13">Locality and age.</emphasis>
|
||
Republic,
|
||
<collectingRegion id="EB25AAFDFFA7FFE1FDF16B8ACAEBFF30" box="[523,659,223,249]" country="United States of America" name="Washington" pageId="12" pageNumber="13">Washington</collectingRegion>
|
||
,
|
||
<collectingCountry id="51F6248FFFA7FFE1FD676B8ACBC0FF30" box="[669,952,223,249]" name="United States of America" pageId="12" pageNumber="13">United States of America</collectingCountry>
|
||
, Tom Thumb Tuff Member of the Klondike Mountain Formation exposureA0307;and McAbee,
|
||
<collectingRegion id="EB25AAFDFFA7FFE1FD2B6A56CBE9FED4" box="[721,913,259,285]" country="Canada" name="British Columbia" pageId="12" pageNumber="13">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="51F6248FFFA7FFE1FC636A56CB88FED4" box="[921,1008,259,285]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
|
||
,Hoodoo Face beds;both mid-Ypresian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |