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<document ID-DOI="http://doi.org/10.5281/zenodo.3792782" ID-GBIF-Dataset="142a25ed-47c3-4361-bc24-6f1b3e8dba5c" ID-GBIF-Taxon="163755057" ID-ISSN="13132970" ID-Zenodo-Dep="3792782" ID-ZooBank="664C49F1-5384-43C4-8BF1-CE76AC11D32E" approvalRequired="10" approvalRequired_for_treatments="10" checkinTime="1587998787238" checkinUser="plazi" docAuthor="Klimaszewski, Jan, Savard, Karine, Pelletier, Georges &amp; Webster, Reginald" docDate="2008" docId="D43E034BFF814172FF17FDB764E9FC06" docLanguage="en" docName="ZK_article_1943.pdf" docOrigin="ZooKeys 2 (2)" docStyle="DocumentStyle:BEF2356135E79670CA200BCAD40C4E3A:zookeys.2008.journal_article" docStyleId="BEF2356135E79670CA200BCAD40C4E3A" docStyleName="zookeys.2008.journal_article" docTitle="Gnypeta groenlandica Lohse 1989" docType="treatment" docVersion="14" lastPageId="30" lastPageNumber="41" masterDocId="28077B33FF9D416CFF9BFFC76718FFEC" masterDocTitle="Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution" masterLastPageNumber="84" masterPageNumber="11" pageId="28" pageNumber="39" updateTime="1643574247635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klimaszewski, Jan</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Savard, Karine</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pelletier, Georges</mods:namePart>
<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
<mods:affiliation>Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, Quebec, QC, Canada &amp; C 136 Eddc &amp; A- 86 C</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Webster, Reginald</mods:namePart>
<mods:affiliation>24 Millstream Drive, Fredericton, NB, Canada &amp; F-</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-09-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>2</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>11</mods:start>
<mods:end>84</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">142a25ed-47c3-4361-bc24-6f1b3e8dba5c</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576410</mods:identifier>
<mods:identifier type="ZooBank">664C49F1-5384-43C4-8BF1-CE76AC11D32E</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792782" ID-GBIF-Taxon="163755057" ID-Zenodo-Dep="3792782" LSID="urn:lsid:plazi:treatment:D43E034BFF814172FF17FDB764E9FC06" httpUri="http://treatment.plazi.org/id/D43E034BFF814172FF17FDB764E9FC06" lastPageId="30" lastPageNumber="41" pageId="28" pageNumber="39">
<subSubSection box="[140,510,623,650]" pageId="28" pageNumber="39" type="nomenclature">
<paragraph blockId="28.[140,510,623,686]" box="[140,510,623,650]" pageId="28" pageNumber="39">
<heading box="[140,510,623,650]" fontSize="11" level="2" pageId="28" pageNumber="39" reason="3">
<emphasis bold="true" box="[140,510,623,650]" pageId="28" pageNumber="39">
6.
<taxonomicName authority="Lohse" authorityName="Lohse" authorityYear="1989" box="[170,510,623,650]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="39" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis bold="true" box="[170,432,623,650]" italics="true" pageId="28" pageNumber="39">Gnypeta groenlandica</emphasis>
Lohse
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,459,659,686]" pageId="28" pageNumber="39" type="description">
<paragraph blockId="28.[140,510,623,686]" box="[140,459,659,686]" pageId="28" pageNumber="39">
(
<figureCitation box="[147,217,659,685]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Figs 8</figureCitation>
,
<figureCitation box="[231,311,659,686]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="28" pageNumber="39">26 a, b</figureCitation>
, 83-90, 195)
</paragraph>
</subSubSection>
<paragraph blockId="28.[140,1108,729,897]" pageId="28" pageNumber="39">
<subSubSection box="[140,577,729,756]" pageId="28" pageNumber="39" type="reference_group">
<taxonomicName authority="Lohse 1989: 58" authorityName="Lohse" authorityPageNumber="58" authorityYear="1989" box="[140,572,729,756]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="39" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis box="[140,381,730,756]" italics="true" pageId="28" pageNumber="39">Gnypeta groenlandica</emphasis>
<materialsCitation ID-GBIF-Occurrence="2608994651" box="[388,572,729,756]" collectingDate="1966-07-02" collectionCode="CNC" country="Greenland" location="Nedre Midsommer So" pageId="28" pageNumber="39" specimenCount="1" typeStatus="holotype">
<bibRefCitation author="Lohse" box="[388,572,729,756]" firstAuthor="Lohse" page="58" pageId="28" pageNumber="39" refId="ref23646" refString="Lohse GA (1989 a) Staphylinidae II (Hypocyphtinae und Aleocharinae). In: Lohse GA, Lucht WH (Ed). Die Kafer Mitteleuropas. Band 12. Supplementband mit Katalogteil. Goecke &amp; Evers, Krefeld." type="book" year="1989">Lohse 1989: 58</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
</subSubSection>
<subSubSection pageId="28" pageNumber="39" type="reference_group">
<emphasis bold="true" box="[585,743,729,755]" pageId="28" pageNumber="39">
<typeStatus box="[585,743,729,755]" pageId="28" pageNumber="39">HOLOTYPE</typeStatus>
</emphasis>
(male):
<collectingCountry box="[840,1024,730,756]" name="Greenland" pageId="28" pageNumber="39">GREENLAND</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:D43E034BFF814172FF17FDB764E9FC06:5948E486FF814170FB96FD1D6669FCFB" country="Greenland" name="Nedre Midsommer So" pageId="28" pageNumber="39">Nedre Midsommer Sö</location>
,
<date box="[382,515,765,791]" pageId="28" pageNumber="39" value="1966-07-02">
<collectingDate box="[382,515,765,791]" pageId="28" pageNumber="39" value="1966-07-02">2.VII.1966</collectingDate>
</date>
, Can. Peary Land Expd. [Expedition] (CNC). Examined.
</subSubSection>
</paragraph>
<subSubSection pageId="28" pageNumber="39" type="materials_examined">
<paragraph blockId="28.[140,1108,729,897]" box="[140,363,835,861]" pageId="28" pageNumber="39">
<emphasis bold="true" box="[140,363,835,861]" pageId="28" pageNumber="39">
<emphasis box="[140,241,835,861]" inLineHeading="true" pageId="28" pageNumber="39" reason="1">Material</emphasis>
examined
</emphasis>
</paragraph>
<paragraph blockId="28.[140,1108,729,897]" box="[185,592,870,897]" pageId="28" pageNumber="39">Specimens are listed in Appendix A.</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="39" type="diagnosis">
<paragraph blockId="28.[140,1108,941,1320]" box="[140,259,941,967]" pageId="28" pageNumber="39">
<emphasis bold="true" box="[140,259,941,967]" inLineHeading="true" pageId="28" pageNumber="39" reason="1">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="28.[140,1108,941,1320]" pageId="28" pageNumber="39">
This species can be recognized by the following combination of characters: body robust, length 3.2-3.7 mm (
<figureCitation box="[431,498,1011,1037]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); elytra (at suture) about as long as pronotum and 1/5 wider than maximum width of pronotum (
<figureCitation box="[628,695,1046,1073]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); abdomen at base as broad as elytra and slightly broadening posteriorly (
<figureCitation box="[552,620,1082,1108]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); antennal article 4 subquadrate or slightly elongate, 3-6 slightly to strongly transverse, 7-9 strongly transverse (
<figureCitation box="[907,1042,1117,1144]" captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="28" pageNumber="39">Figs 26 a, b</figureCitation>
); median lobe of aedeagus with apical part narrowly produced and sinuate laterally (
<figureCitation captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="28" pageNumber="39">Fig. 83</figureCitation>
), bulbus in dorsal view with broad lateral swells (
<figureCitation box="[719,801,1187,1213]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="28" pageNumber="39">Fig. 84</figureCitation>
); spermatheca with capsule longer than wide, funnel-shaped, lateral sides subparallel (
<figureCitation box="[794,878,1222,1249]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="28" pageNumber="39">Fig. 88</figureCitation>
); stem curved, sinuate apically and slightly swollen basally (
<figureCitation box="[588,671,1258,1284]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="28" pageNumber="39">Fig. 88</figureCitation>
); male tergite 8 strongly transverse and truncate apically (
<figureCitation box="[341,426,1293,1320]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="28" pageNumber="39">Fig. 86</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="41" pageId="28" pageNumber="39" type="description">
<paragraph blockId="28.[140,1107,1363,1671]" box="[140,282,1363,1389]" pageId="28" pageNumber="39">
<emphasis bold="true" box="[140,282,1363,1389]" inLineHeading="true" pageId="28" pageNumber="39" reason="1">Description</emphasis>
</paragraph>
<paragraph blockId="28.[140,1107,1363,1671]" pageId="28" pageNumber="39">
Body length 3.2-3.7 mm; uniformly dark brown to black (
<figureCitation box="[797,864,1398,1425]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); integument moderately glossy; pubescence yellowish grey and moderately long and dense; antennal article 4 subquadrate or slightly elongate, 5-6 slightly and 7-9 strongly transverse (
<figureCitation captionStart="Figs 21-38" captionStartId="10.[140,184,1462,1484]" captionTargetBox="[230,1007,164,1440]" captionTargetId="figure@10.[176,1072,134,1444]" captionTargetPageId="10" captionText="Figs 21-38. Antennae of Gnypeta species: G. ashei: 21a. male, 21b. female; G. brincki: 22a. male, 22b. female; G. atrolucens: 23a. male, 23b. female; G. sellmani: 24a. male, 24b. female; G. dentata: 25a. male, 25b. female; G. groenlandica: 26a. male, 26b. female: G. brevicornis: 27a. male, 27b. female; G. caerulea: 28a. male, 28b. female; G. lohsei: 29a. male, 29b. female; G. crebrepunctata: 30a. male, 30b. female; G. manitobae: 31b. female; G. uteana: 32a. male, 32b. female; G. carbonaria: 33a. male, 33b. female; G. helenae: 34a. male, 34b. female; G. canadensis: 35a. male, 35b. female; G. nigrella: 36a. male, 36b. female; G. saccharina: 37a. male, 37b. female; G. minuta: 38a. male, 38b. female. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="28" pageNumber="39">Figs 26 a, b</figureCitation>
); head and pronotum of about the same width (
<figureCitation box="[715,783,1504,1530]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); elytra and abdomen wider than either head or pronotum; elytra small and short (
<figureCitation box="[765,834,1539,1565]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
); head rounded basally; pronotum broadest in apical third, pubescence directed anterad along midline and laterad elsewhere; elytra (at suture) about as long as pronotum and 1/5 broader than maximum width of pronotum (
<figureCitation box="[497,564,1645,1671]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="28" pageNumber="39">Fig. 8</figureCitation>
), pubescence directed obliquely postero-laterad,
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774106" ID-Zenodo-Dep="3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="29" pageNumber="40" startId="29.[140,184,1590,1612]" targetBox="[148,1105,214,1510]" targetPageId="29">
<paragraph blockId="29.[140,1108,1590,1676]" pageId="29" pageNumber="40">
<emphasis bold="true" box="[140,259,1590,1612]" pageId="29" pageNumber="40">Figs 83-90.</emphasis>
Genital structures of
<taxonomicName authorityName="Lohse" authorityYear="1989" box="[462,655,1590,1612]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="40" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis box="[462,655,1590,1612]" italics="true" pageId="29" pageNumber="40">Gnypeta groenlandica</emphasis>
</taxonomicName>
: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8.
</paragraph>
</caption>
<paragraph blockId="30.[140,1108,166,509]" pageId="30" pageNumber="41">
in wavy pattern medially on each side; abdomen swollen medially, as broad as elytra at base (
<figureCitation box="[235,304,202,228]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="30" pageNumber="41">Fig. 8</figureCitation>
); metatarsus with two basal articles of about the same length and the third one slightly shorter.
<emphasis bold="true" box="[433,493,237,263]" pageId="30" pageNumber="41">Male</emphasis>
. Tergite 8 transverse and truncate apically and slightly concave medially (
<figureCitation box="[349,433,272,299]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 86</figureCitation>
). Sternite 8 elongate and slightly produced posteriorly (
<figureCitation captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 87</figureCitation>
). Median lobe of aedeagus with narrowly produced and sinuate apical part of tubus in lateral view (
<figureCitation box="[321,407,342,369]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 83</figureCitation>
), sides swollen in dorsal view; bulbus moderately large (
<figureCitation captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 84</figureCitation>
); internal sac with structures as illustrated (
<figureCitation box="[658,789,378,404]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Figs 83, 84</figureCitation>
).
<emphasis bold="true" box="[809,893,378,404]" pageId="30" pageNumber="41">Female</emphasis>
. Tergite 8 truncate apically (
<figureCitation box="[241,326,413,439]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 89</figureCitation>
). Sternite 8 broadly rounded posteriorly (
<figureCitation box="[798,883,413,439]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 90</figureCitation>
). Spermatheca with capsule elongate, funnel-shaped with sides subparallel (
<figureCitation box="[767,852,448,474]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 88</figureCitation>
); stem curved, sinuate apically and swollen basally (
<figureCitation box="[464,549,483,509]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="30" pageNumber="41">Fig. 88</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="41" type="distribution">
<paragraph blockId="30.[140,1107,553,651]" box="[140,412,553,580]" pageId="30" pageNumber="41">
<emphasis bold="true" box="[140,291,553,579]" inLineHeading="true" pageId="30" pageNumber="41" reason="1">Distribution</emphasis>
(Fig. 195)
</paragraph>
<paragraph blockId="30.[140,1107,553,651]" pageId="30" pageNumber="41">
<taxonomicName authorityName="Lohse" authorityYear="1989" box="[140,375,589,615]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="41" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis box="[140,375,589,615]" italics="true" pageId="30" pageNumber="41">Gnypeta groenlandica</emphasis>
</taxonomicName>
is a Nearctic species recorded here from
<collectingCountry box="[829,951,589,615]" name="Greenland" pageId="30" pageNumber="41">Greenland</collectingCountry>
, and newly in
<collectingCountry box="[140,226,624,650]" name="Canada" pageId="30" pageNumber="41">Canada</collectingCountry>
(
<collectingRegion box="[241,357,624,650]" country="Canada" name="Manitoba" pageId="30" pageNumber="41">Manitoba</collectingRegion>
,
<collectingRegion box="[368,472,625,651]" country="Canada" name="Nunavut" pageId="30" pageNumber="41">Nunavut</collectingRegion>
,
<collectingRegion box="[484,669,624,651]" country="Canada" name="Yukon" pageId="30" pageNumber="41">Yukon Territory</collectingRegion>
), and the
<collectingCountry box="[781,932,624,650]" name="United States of America" pageId="30" pageNumber="41">United States</collectingCountry>
(
<collectingRegion box="[947,1027,624,651]" country="United States of America" name="Alaska" pageId="30" pageNumber="41">Alaska</collectingRegion>
).
</paragraph>
<paragraph blockId="30.[140,1108,694,826]" box="[140,467,694,721]" pageId="30" pageNumber="41">
<emphasis bold="true" box="[140,467,694,721]" pageId="30" pageNumber="41">
<emphasis box="[140,265,694,720]" inLineHeading="true" pageId="30" pageNumber="41" reason="1">Collection</emphasis>
and habitat data
</emphasis>
</paragraph>
<paragraph blockId="30.[140,1108,694,826]" pageId="30" pageNumber="41">
Adults were collected in June, July and August, mainly in alpine habitats from
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" box="[1053,1107,730,756]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="30" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis box="[1053,1107,730,756]" italics="true" pageId="30" pageNumber="41">Salix</emphasis>
</taxonomicName>
litter, litter on shore of a lagoon, and in crown of Rosss Avens (
<taxonomicName box="[852,978,765,791]" class="Magnoliopsida" family="Rosaceae" genus="Geum" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="41" phylum="Tracheophyta" rank="species" species="rossii">
<emphasis box="[852,978,765,791]" italics="true" pageId="30" pageNumber="41">Geum rossii</emphasis>
</taxonomicName>
), rose family (
<taxonomicName authorityName="de Jussieu" authorityYear="1789" box="[182,291,800,826]" class="Magnoliopsida" family="Rosaceae" kingdom="Plantae" order="Rosales" pageId="30" pageNumber="41" phylum="Tracheophyta" rank="family">Rosaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="30" pageNumber="41" type="discussion">
<paragraph blockId="30.[140,1108,870,1003]" box="[140,271,870,896]" pageId="30" pageNumber="41">
<emphasis bold="true" box="[140,271,870,896]" inLineHeading="true" pageId="30" pageNumber="41" reason="1">Comments</emphasis>
</paragraph>
<paragraph blockId="30.[140,1108,870,1003]" pageId="30" pageNumber="41">
The late Steve Ashe (University of
<collectingRegion box="[533,613,906,932]" country="United States of America" name="Kansas" pageId="30" pageNumber="41">Kansas</collectingRegion>
) worked on a revision of Nearctic
<taxonomicName authorityName=": Casey" authorityYear="1906" box="[1010,1100,906,932]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis box="[1010,1100,906,932]" italics="true" pageId="30" pageNumber="41">Gnypeta</emphasis>
</taxonomicName>
, but was not able to complete it. Some CNC specimens of this species bear his identification labels as
<taxonomicName authority="Ashe, 1987" authorityName="Ashe" authorityYear="1987" box="[315,608,976,1002]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="41" phylum="Arthropoda" rank="species" species="praeminosa">
<emphasis box="[315,472,976,1002]" italics="true" pageId="30" pageNumber="41">G. praeminosa</emphasis>
Ashe, 1987
</taxonomicName>
[manuscript unpublished name].
</paragraph>
</subSubSection>
</treatment>
</document>