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<document id="6461C4E6E00D2B868FC86085885CD234" ID-DOI="10.5281/zenodo.188674" ID-GBIF-Dataset="d9b0d696-f8a6-4ea2-9f53-f62192913f01" ID-ISSN="1175-5326" ID-Zenodo-Dep="188674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461095943464" checkinUser="plazi" docAuthor="Shattuck, Steve" docDate="2009" docId="03F1FB3AE205FFAB0FDEBE63D661FA49" docLanguage="en" docName="zt02146p021.pdf" docOrigin="Zootaxa 2146" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Myrmecina alpina Shattuck, 2009, sp. n." docType="treatment" docUuid="622EE04D-1619-47EF-B9D9-4DCF97CC5EAE" docUuidSource="ZooBank" docVersion="10" lastPageNumber="6" masterDocId="FFC88342E207FFAE0F49BB0DD502FF81" masterDocTitle="A revision the Australian species of the ant genus Myrmecina (Hymenoptera: Formicidae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="3" updateTime="1698589194258" updateUser="plazi">
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<mods:title id="E0C51BFE4C15AE584E6FA3C767D014B2">A revision the Australian species of the ant genus Myrmecina (Hymenoptera: Formicidae)</mods:title>
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<mods:date id="4013F21A04A02532D3557DCF4F324529">2009</mods:date>
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<treatment id="03F1FB3AE205FFAB0FDEBE63D661FA49" ID-DOI="http://doi.org/10.5281/zenodo.6218936" ID-GBIF-Taxon="119620445" ID-Zenodo-Dep="6218936" LSID="urn:lsid:zoobank.org:act:622EE04D-1619-47EF-B9D9-4DCF97CC5EAE" httpUri="http://treatment.plazi.org/id/03F1FB3AE205FFAB0FDEBE63D661FA49" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="C34219A7E205FFAC0FDEBE63D43EFA2A" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8BE74A2CE205FFAC0FDEBE63D4BCFA09" blockId="2.[151,446,1390,1451]" box="[151,446,1390,1416]" pageId="2" pageNumber="3">
<heading id="D0AFFD40E205FFAC0FDEBE63D4BCFA09" bold="true" box="[151,446,1390,1416]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B92C963EE205FFAC0FDEBE63D4BCFA09" bold="true" box="[151,446,1390,1416]" pageId="2" pageNumber="3">
<taxonomicName id="4C5831AFE205FFAC0FDEBE63D477FA09" box="[151,373,1390,1416]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B92C963EE205FFAC0FDEBE63D477FA09" bold="true" box="[151,373,1390,1416]" italics="true" pageId="2" pageNumber="3">Myrmecina alpina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F2B45E205FFAC0E35BE63D4BCFA09" box="[380,446,1390,1416]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE74A2CE205FFAC0FDEBE9CD43EFA2A" blockId="2.[151,446,1390,1451]" box="[151,316,1425,1451]" pageId="2" pageNumber="3">
(
<figureCitation id="136356A9E205FFAC0FD6BE9CD40AFA2A" box="[159,264,1425,1451]" captionStart="FIGURES 1 4" captionStartId="3.[151,269,1615,1639]" captionTargetBox="[151,1436,197,1589]" captionTargetId="figure@3.[151,1436,197,1590]" captionTargetPageId="3" captionText="FIGURES 1 4. Myrmecina alpina sp. n. worker. Fig. 1, front of head; Fig. 2, side of head; Fig. 3, lateral view of body; Fig. 4, dorsal view of body." httpUri="https://zenodo.org/record/188675/files/figure.png" pageId="2" pageNumber="3">Figs 14</figureCitation>
,
<figureCitation id="136356A9E205FFAC0E5CBE9CD436FA2A" box="[277,308,1425,1451]" captionStart="FIGURES 47 52" captionStartId="19.[151,269,1945,1969]" captionTargetBox="[202,1388,194,1910]" captionTargetId="figure@19.[202,1389,194,1910]" captionTargetPageId="19" captionText="FIGURES 47 52. Distribution of material examined during this study. Fig. 47, M. alpina, M. pumila, M. silvalaeva, M. silvampla, M. silvangula, M. silvarugosa and M. silvatransversa; Fig. 48, M. australis; Fig. 49, M. difficulta; Fig. 50, M. eruga (circles) and M. rugosa (square); Fig. 51, M. inaequala; Fig. 52, M. wesselensis." httpUri="https://zenodo.org/record/188685/files/figure.png" pageId="2" pageNumber="3">47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34219A7E205FFAC0FDEBED3D178F9C7" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE74A2CE205FFAC0FDEBED3D178F9C7" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="3">
<emphasis id="B92C963EE205FFAC0FDEBED3D5E7FA79" bold="true" box="[151,229,1502,1528]" pageId="2" pageNumber="3">
<typeStatus id="54E3F48EE205FFAC0FDEBED3D5E2FA79" box="[151,224,1502,1528]" pageId="2" pageNumber="3">Types</typeStatus>
.
</emphasis>
<typeStatus id="54E3F48EE205FFAC0FA7BED3D459FA79" box="[238,347,1502,1528]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
worker from Mt. Demi,
<quantity id="4CA0E7C9E205FFAC0D36BED3D7B1FA79" box="[639,691,1502,1528]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="3" unit="km" value="7.0">7km</quantity>
SW Mossman,
<geoCoordinate id="EE6C2CEBE205FFAC0C26BED3D6C9FA79" box="[879,971,1502,1528]" direction="south" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="-16.5">16°30'S</geoCoordinate>
<geoCoordinate id="EE6C2CEBE205FFAC0C9ABED3D143FA79" box="[979,1089,1502,1528]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="145.31667">145°19'E</geoCoordinate>
,
<quantity id="4CA0E7C9E205FFAC0B04BED3D1A2FA79" box="[1101,1184,1502,1528]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="3" unit="m" value="1100.0">1100m</quantity>
, Queensland,
<date id="FFE66CECE205FFAC0A0CBED3D5DAF99E" pageId="2" pageNumber="3" value="1983-10-29">29 Oct. 1983</date>
, D. K. Yeates &amp; G. I. Thompson, rainforest, sieved litter berlesate (ANIC, ANIC32-047350); 13
<typeStatus id="54E3F48EE205FFAC0FDEBD21D5FEF9C7" box="[151,252,1580,1606]" pageId="2" pageNumber="3" type="paratype">paratype</typeStatus>
workers, same data as
<typeStatus id="54E3F48EE205FFAC0D42BD21D773F9C7" box="[523,625,1580,1606]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
(ANIC, BMNH, MCZC, ANIC32-047232).
</paragraph>
</subSubSection>
<subSubSection id="C34219A7E205FFAC0F8FBD5FD1A1F961" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="8BE74A2CE205FFAC0F8FBD5FD1A1F961" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="3">
<emphasis id="B92C963EE205FFAC0F8FBD5FD444F9ED" bold="true" box="[198,326,1618,1644]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
Sides of head behind compound eyes with 23 well defined longitudinal carinae running nearly the length of the head. The presence of these carinae will separate this species from all others in
<collectingCountry id="F34F0ABCE205FFAC0FDEBDADD406F93B" box="[151,260,1696,1722]" name="Australia" pageId="2" pageNumber="3">Australia</collectingCountry>
with the exception of some higher-elevation populations of
<taxonomicName id="4C5831AFE205FFAC0C9CBDADD178F938" box="[981,1146,1696,1721]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="silvangula">
<emphasis id="B92C963EE205FFAC0C9CBDADD178F938" box="[981,1146,1696,1721]" italics="true" pageId="2" pageNumber="3">M. silvangula</emphasis>
</taxonomicName>
. The longitudinal rather than transverse sculpturing on the pronotum will separate this taxon from these workers.
</paragraph>
</subSubSection>
<subSubSection id="C34219A7E205FFAD0F8FBDE0D4CFF979" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="description">
<paragraph id="8BE74A2CE205FFAC0F8FBDE0D41CF86E" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="3">
<emphasis id="B92C963EE205FFAC0F8FBDE0D4C1F886" bold="true" box="[198,451,1773,1799]" pageId="2" pageNumber="3">Worker description.</emphasis>
Antennal scapes smooth to weakly ridged. First segment of funiculus cone-shaped. Sides of head behind compound eyes with 23 longitudinal carinae running the length of the head. Sculpturing on dorsal surface of mesosoma running longitudinally and generally with the central carina (or carinae) running nearly the length (the pronotum not differentiated from the mesonotum). Carinae extending continuously from the dorsal surface onto the lateral surfaces of the mesosoma. Metanotal spines short. Propodeal spines long. Erect hairs abundant, straight. Colour dark brown-black, antennae, mandibles and legs yellow-red.
</paragraph>
<caption id="DF271AA4E204FFAD0FDEBD42D4BFF908" httpUri="https://zenodo.org/record/188675/files/figure.png" pageId="3" pageNumber="4" targetBox="[151,1436,197,1589]" targetPageId="3">
<paragraph id="8BE74A2CE204FFAD0FDEBD42D4BFF908" blockId="3.[151,1436,1615,1673]" pageId="3" pageNumber="4">
<emphasis id="B92C963EE204FFAD0FDEBD42D440F9E6" bold="true" box="[151,322,1615,1639]" pageId="3" pageNumber="4">FIGURES 14.</emphasis>
<taxonomicName id="4C5831AFE204FFAD0E03BD42D70CF9E7" box="[330,526,1615,1638]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alpina" status="sp. nov.">
<emphasis id="B92C963EE204FFAD0E03BD42D70CF9E7" box="[330,526,1615,1638]" italics="true" pageId="3" pageNumber="4">Myrmecina alpina</emphasis>
</taxonomicName>
<emphasis id="B92C963EE204FFAD0D5CBD42D753F9E6" bold="true" box="[533,593,1615,1639]" pageId="3" pageNumber="4">
<taxonomicNameLabel id="A21F2B45E204FFAD0D5CBD42D753F9E6" box="[533,593,1615,1639]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
worker. Fig. 1, front of head; Fig. 2, side of head; Fig. 3, lateral view of body; Fig. 4, dorsal view of body.
</paragraph>
</caption>
<paragraph id="8BE74A2CE204FFAD0F8FBDBAD4CFF979" blockId="3.[151,1437,1719,2016]" pageId="3" pageNumber="4">
<emphasis id="B92C963EE204FFAD0F8FBDBAD487F950" bold="true" box="[198,389,1719,1745]" pageId="3" pageNumber="4">Measurements.</emphasis>
<emphasis id="B92C963EE204FFAD0EC5BDBAD4DCF951" box="[396,478,1719,1744]" italics="true" pageId="3" pageNumber="4">Worker</emphasis>
(
<emphasis id="B92C963EE204FFAD0EB8BDBAD702F951" box="[497,512,1719,1744]" italics="true" pageId="3" pageNumber="4">n</emphasis>
= 5) - CI 9599; HL 0.720.84; HW 0.690.80; MTL 0.380.45; SI 8594; SL 0.650.72; WL 0.871.06.
</paragraph>
</subSubSection>
<subSubSection id="C34219A7E204FFAD0F8FBC09D009F861" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="8BE74A2CE204FFAD0F8FBC09D009F861" blockId="3.[151,1437,1719,2016]" pageId="3" pageNumber="4">
<emphasis id="B92C963EE204FFAD0F8FBC09D13CF89F" bold="true" box="[198,1086,1796,1822]" pageId="3" pageNumber="4">
Additional material examined (
<collectionCode id="ED49D2E9E204FFAD0D05BC09D79BF89F" LSID="urn:lsid:biocol.org:col:32981" box="[588,665,1796,1822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="3" pageNumber="4">ANIC</collectionCode>
except where noted). Queensland
</emphasis>
:
<quantity id="4CA0E7C9E204FFAD0B07BC09D180F89F" box="[1102,1154,1796,1822]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="4" unit="km" value="1.0">1km</quantity>
S Mt. Lewis (Calder,A. &amp; Weir,T.);
<quantity id="4CA0E7C9E204FFAD0E61BC26D476F8C4" box="[296,372,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="4" unit="km" value="2.5">2.5km</quantity>
N Mt. Lewis via Julatten (Yeates,D.K. &amp; Thompson,G.I.);
<quantity id="4CA0E7C9E204FFAD0B77BC26D171F8C4" box="[1086,1139,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="4" unit="km" value="4.0">4km</quantity>
WNW Round Mt. site 3 (Burwell,C.) (
<collectionCode id="ED49D2E9E204FFAD0E0CBC5FD4A5F8ED" box="[325,423,1874,1900]" pageId="3" pageNumber="4">QMBA</collectionCode>
);
<quantity id="4CA0E7C9E204FFAD0EF4BC5FD4F6F8ED" box="[445,500,1874,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="4" unit="km" value="4.0">4km</quantity>
WNW Round Mt. site 5 (Yek,S) (
<collectionCode id="ED49D2E9E204FFAD0CD7BC5FD102F8ED" box="[926,1024,1874,1900]" pageId="3" pageNumber="4">QMBA</collectionCode>
); Mary Creek site 4 (Burwell,C.) (
<collectionCode id="ED49D2E9E204FFAD0FE9BC75D404F813" box="[160,262,1912,1938]" pageId="3" pageNumber="4">QMBA</collectionCode>
); Mary Creek site 5 (Burwell,C.) (
<collectionCode id="ED49D2E9E204FFAD0D80BC75D62DF813" box="[713,815,1912,1938]" pageId="3" pageNumber="4">QMBA</collectionCode>
); Mt. Demi,
<quantity id="4CA0E7C9E204FFAD0C9FBC75D10CF813" box="[982,1038,1912,1938]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="4" unit="km" value="7.0">7km</quantity>
SW Mossman (Yeates,D.K. &amp; Thompson,G.I.); Mt. Lewis (Taylor &amp; Feehan); Mt. Lewis (Van Ingen,L.) (
<collectionCode id="ED49D2E9E204FFAD0B69BC92D171F838" box="[1056,1139,1951,1977]" pageId="3" pageNumber="4">TERC</collectionCode>
); Mt. Lewis summit, via Julatten (Monteith,G. &amp; Cook,D.); The Bluff,
<quantity id="4CA0E7C9E204FFAD0DF8BCCBD7F1F861" box="[689,755,1990,2016]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="4" unit="km" value="11.0">11km</quantity>
W Mossman (Monteith,G.B. &amp; Yeates,D.K.).
</paragraph>
</subSubSection>
<caption id="DF271AA4E203FFAA0FDEBCB9D13CF86F" httpUri="https://zenodo.org/record/188676/files/figure.png" pageId="4" pageNumber="5" targetBox="[214,1372,187,1947]" targetPageId="4">
<paragraph id="8BE74A2CE203FFAA0FDEBCB9D13CF86F" blockId="4.[151,1436,1972,2031]" pageId="4" pageNumber="5">
<emphasis id="B92C963EE203FFAA0FDEBCB9D446F84D" bold="true" box="[151,324,1972,1996]" pageId="4" pageNumber="5">FIGURES 56.</emphasis>
Fig. 5, graph of head length versus head width for
<emphasis id="B92C963EE203FFAA0C23BCB8D11FF84D" box="[874,1053,1973,1996]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="4C5831AFE203FFAA0C23BCB8D6D1F84D" box="[874,979,1973,1996]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ampla">M. ampla</taxonomicName>
,
<taxonomicName id="4C5831AFE203FFAA0C96BCB8D11FF84D" box="[991,1053,1973,1996]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="eruga">eruga</taxonomicName>
</emphasis>
,
<emphasis id="B92C963EE203FFAA0B62BCB8D187F84D" box="[1067,1157,1973,1996]" italics="true" pageId="4" pageNumber="5">orbicula</emphasis>
and
<taxonomicName id="4C5831AFE203FFAA0BF3BCB8D006F84D" box="[1210,1284,1973,1996]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pumila">
<emphasis id="B92C963EE203FFAA0BF3BCB8D006F84D" box="[1210,1284,1973,1996]" italics="true" pageId="4" pageNumber="5">pumila</emphasis>
</taxonomicName>
; Fig. 6, graph of head length versus head width for
<taxonomicName id="4C5831AFE203FFAA0D6ABCD5D7A4F86E" box="[547,678,2008,2031]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="difficulta">
<emphasis id="B92C963EE203FFAA0D6ABCD5D7A4F86E" box="[547,678,2008,2031]" italics="true" pageId="4" pageNumber="5">M. difficulta</emphasis>
</taxonomicName>
and the remaining Australian species.
</paragraph>
</caption>
<caption id="DF271AA4E202FFAB0FDEBFA6D443FB64" httpUri="https://zenodo.org/record/188677/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1435,195,1170]" targetPageId="5">
<paragraph id="8BE74A2CE202FFAB0FDEBFA6D443FB64" blockId="5.[151,1436,1195,1253]" pageId="5" pageNumber="6">
<emphasis id="B92C963EE202FFAB0FDEBFA6D41FFB42" bold="true" box="[151,285,1195,1219]" pageId="5" pageNumber="6">FIGURE 7.</emphasis>
Fig. 7, graph of scape length versus head width for
<taxonomicName id="4C5831AFE202FFAB0C14BFA1D6EFFB42" box="[861,1005,1196,1219]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silvampla">
<emphasis id="B92C963EE202FFAB0C14BFA1D6EFFB42" box="[861,1005,1196,1219]" italics="true" pageId="5" pageNumber="6">M. silvampla</emphasis>
</taxonomicName>
,
<emphasis id="B92C963EE202FFAB0CB5BFA1D06FFB42" box="[1020,1389,1196,1219]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C5831AFE202FFAB0CB5BFA1D173FB42" box="[1020,1137,1196,1219]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silvangula">silvangula</taxonomicName>
,
<taxonomicName id="4C5831AFE202FFAB0B35BFA1D1FCFB42" box="[1148,1278,1196,1219]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silvarugosa">silvarugosa</taxonomicName>
, livelaeva
</emphasis>
and
<taxonomicName id="4C5831AFE202FFAB0FDEBFC3D438FB64" box="[151,314,1230,1253]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silvatransversa">
<emphasis id="B92C963EE202FFAB0FDEBFC3D438FB64" box="[151,314,1230,1253]" italics="true" pageId="5" pageNumber="6">silvatransversa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C34219A7E202FFAB0F8FBE1ED661FA49" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BE74A2CE202FFAB0F8FBE1ED661FA49" blockId="5.[151,1437,1299,1480]" pageId="5" pageNumber="6">
<emphasis id="B92C963EE202FFAB0F8FBE1ED457FAAC" bold="true" box="[198,341,1299,1325]" pageId="5" pageNumber="6">Comments.</emphasis>
This is a high-elevation specialist known from a limited number of mountain-tops, the lowest collection site being just less than
<quantity id="4CA0E7C9E202FFAB0D71BE37D78CFAD5" box="[568,654,1338,1364]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="m" value="1000.0">1000m</quantity>
, and is confined to an area less than 20 square kilometres within Queenslands wet tropics. It is found in rainforest where it has been collected in leaf litter samples. It is sympatric with
<taxonomicName id="4C5831AFE202FFAB0E1CBE85D4E0FA20" box="[341,482,1416,1441]" class="Insecta" family="Formicidae" genus="Myrmecina" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="silvarugosa">
<emphasis id="B92C963EE202FFAB0E1CBE85D4E0FA20" box="[341,482,1416,1441]" italics="true" pageId="5" pageNumber="6">silvarugosa</emphasis>
</taxonomicName>
, another high-elevation species, and these two species are the only Australian species with carinae on the sides of the head behind the eyes.
</paragraph>
</subSubSection>
</treatment>
</document>