198 lines
28 KiB
XML
198 lines
28 KiB
XML
<document id="02767FCC3685BDE7FF87FB293A0EBA46" ID-CLB-Dataset="28413" ID-DOI="10.11646/zootaxa.4564.2.9" ID-GBIF-Dataset="8ca25900-3a6f-4f10-9331-dff9eeccbb8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589400" ID-ZooBank="FDADF9CA-379C-4980-8AE4-D47F58537E3F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552236971884" checkinUser="plazi" docAuthor="Rocha, Rosana M. & Counts, Bailey Keegan" docDate="2019" docId="03DA3E07B05EFFFD85E1D9CB04D8F816" docLanguage="en" docName="zootaxa.4564.2.9.pdf" docOrigin="Zootaxa 4564 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pyura lignosa Michaelsen 1908" docType="treatment" docVersion="13" lastPageNumber="498" masterDocId="FFE3467FB058FFFA8576DD040707FB3D" masterDocTitle="Pyura (Tunicata: Ascidiacea: Pyuridae) on the coasts of Panama" masterLastPageNumber="513" masterPageNumber="491" pageNumber="497" updateTime="1698701146666" updateUser="ExternalLinkService">
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<mods:title id="E37C0505BEBF71714367E35AB2195461">Pyura (Tunicata: Ascidiacea: Pyuridae) on the coasts of Panama</mods:title>
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<mods:namePart id="847803546665297D424B71B8A74E947F">Rocha, Rosana M.</mods:namePart>
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<mods:namePart id="D39A807C2949233D4A186BCD74FC30AC">Counts, Bailey Keegan</mods:namePart>
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<mods:title id="26EFB270C4D2CB8A0542F73D675A0101">Zootaxa</mods:title>
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<mods:date id="8BF863DDFAFFC2C12A08C9011A26ADBC">2019</mods:date>
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<treatment id="03DA3E07B05EFFFD85E1D9CB04D8F816" ID-DOI="http://doi.org/10.5281/zenodo.5930328" ID-GBIF-Taxon="154366707" ID-Zenodo-Dep="5930328" LSID="urn:lsid:plazi:treatment:03DA3E07B05EFFFD85E1D9CB04D8F816" httpUri="http://treatment.plazi.org/id/03DA3E07B05EFFFD85E1D9CB04D8F816" lastPageId="7" lastPageNumber="498" pageId="6" pageNumber="497">
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<subSubSection id="C369DC9AB05EFFFC85E1D9CB0519FFD7" box="[151,542,1231,1258]" pageId="6" pageNumber="497" type="nomenclature">
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<paragraph id="8BCC8F11B05EFFFC85E1D9CB0519FFD7" blockId="6.[151,542,1231,1291]" box="[151,542,1231,1258]" pageId="6" pageNumber="497">
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<heading id="D084387DB05EFFFC85E1D9CB0519FFD7" bold="true" box="[151,542,1231,1258]" fontSize="11" level="1" pageId="6" pageNumber="497" reason="1">
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<taxonomicName id="4C73F492B05EFFFC85E1D9CB0519FFD7" ID-CoL="4QY9J" authority="Michaelsen, 1908" authorityName="Michaelsen" authorityYear="1908" box="[151,542,1231,1258]" class="Ascidiacea" family="Pyuridae" genus="Pyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stolidobranchia" pageId="6" pageNumber="497" phylum="Chordata" rank="species" species="lignosa">
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<emphasis id="B9075303B05EFFFC85E1D9CB0519FFD7" bold="true" box="[151,542,1231,1258]" pageId="6" pageNumber="497">
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<emphasis id="B9075303B05EFFFC85E1D9CB0638FFD4" bold="true" box="[151,319,1231,1257]" italics="true" pageId="6" pageNumber="497">Pyura lignosa</emphasis>
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<bibRefCitation id="EFE2F2E0B05EFFFC8430D9D40519FFD7" author="Michaelsen, W." box="[326,542,1232,1258]" pageId="6" pageNumber="497" pagination="227 - 287" refId="ref11681" refString="Michaelsen, W. (1908) Die Pyuriden [Halocynthiiden] des NaturhistorischenMuseums zu Hamburg. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 25, suppl. 2, 227 - 287." type="journal article" year="1908">Michaelsen, 1908</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DC9AB05EFFFC85E1D9F60628FE36" box="[151,303,1266,1291]" pageId="6" pageNumber="497" type="description">
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<paragraph id="8BCC8F11B05EFFFC85E1D9F60628FE36" blockId="6.[151,542,1231,1291]" box="[151,303,1266,1291]" pageId="6" pageNumber="497">
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(
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<figureCitation id="13489394B05EFFFC85E8D9F6060CFE36" box="[158,267,1266,1291]" captionStart="FIGURE 5" captionStartId="7.[151,250,1719,1741]" captionTargetBox="[176,1414,856,1697]" captionTargetId="figure@7.[173,1414,856,1697]" captionTargetPageId="7" captionText="FIGURE 5. Pyura lignosa Michaelsen, 1908 A. Animal in the field showing open siphons with red margins (arrowheads). B. External view of preserved animal. C. Right side, tunic removed. D. Left side, tunic removed. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2589410/files/figure.png" pageId="6" pageNumber="497">Figures 5</figureCitation>
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,
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<figureCitation id="13489394B05EFFFC8461D9F60621FE36" box="[279,294,1266,1291]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="6" pageNumber="497">6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DC9AB05EFFFC85E1D83F045DFE52" pageId="6" pageNumber="497" type="reference_group">
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<paragraph id="8BCC8F11B05EFFFC85E1D83F04C8FE6D" blockId="6.[151,975,1338,1391]" box="[151,975,1338,1360]" pageId="6" pageNumber="497">
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<treatmentCitationGroup id="AB63A83FB05EFFFC85E1D83F04C8FE6D" box="[151,975,1338,1360]" pageId="6" pageNumber="497">
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<taxonomicName id="4C73F492B05EFFFC85E1D83F05ADFE6D" ID-CoL="4QY9J" authority="Michaelsen, 1908: 256" authorityName="Michaelsen" authorityPageNumber="256" authorityYear="1908" box="[151,682,1338,1360]" class="Ascidiacea" family="Pyuridae" genus="Pyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stolidobranchia" pageId="6" pageNumber="497" phylum="Chordata" rank="species" species="lignosa" subGenus="Halocynthia">
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<emphasis id="B9075303B05EFFFC85E1D83F06BDFE6D" box="[151,442,1338,1360]" italics="true" pageId="6" pageNumber="497">Pyura (Halocynthia) lignosa</emphasis>
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<treatmentCitation id="0AD2A900B05EFFFC84B6D83E05ADFE6D" author="Michaelsen, W." box="[448,682,1338,1360]" page="256" pageId="6" pageNumber="497" year="1908">
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<bibRefCitation id="EFE2F2E0B05EFFFC84B6D83E057FFE6D" author="Michaelsen, W." box="[448,632,1338,1360]" pageId="6" pageNumber="497" pagination="227 - 287" refId="ref11681" refString="Michaelsen, W. (1908) Die Pyuriden [Halocynthiiden] des NaturhistorischenMuseums zu Hamburg. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 25, suppl. 2, 227 - 287." type="journal article" year="1908">Michaelsen, 1908</bibRefCitation>
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: 256
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</treatmentCitation>
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</taxonomicName>
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, pl.1, fig. 9; pl. 2, figs 21, 22
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCC8F11B05EFFFC85E1D85E045DFE52" blockId="6.[151,975,1338,1391]" box="[151,858,1369,1391]" pageId="6" pageNumber="497">
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<treatmentCitationGroup id="AB63A83FB05EFFFC85E1D85E045DFE52" box="[151,858,1369,1391]" pageId="6" pageNumber="497">
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<taxonomicName id="4C73F492B05EFFFC85E1D85E06F4FE52" ID-CoL="4QY9J" authority=": Monniot, 1994: 44" authorityName=": Monniot" authorityPageNumber="44" authorityYear="1994" box="[151,499,1369,1391]" class="Ascidiacea" family="Pyuridae" genus="Pyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stolidobranchia" pageId="6" pageNumber="497" phylum="Chordata" rank="species" species="lignosa">
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<emphasis id="B9075303B05EFFFC85E1D85E0621FE52" box="[151,294,1369,1391]" italics="true" pageId="6" pageNumber="497">Pyura lignosa</emphasis>
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:
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<treatmentCitation id="0AD2A900B05EFFFC8445D85D06F4FE52" author="Monniot, C." box="[307,499,1369,1391]" page="44" pageId="6" pageNumber="497" year="1994">
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||
<bibRefCitation id="EFE2F2E0B05EFFFC8445D85D06CAFE52" author="Monniot, C." box="[307,461,1369,1391]" pageId="6" pageNumber="497" pagination="41 - 51" refId="ref11817" refString="Monniot, C. (1994) Pyura lignosa (s. s.), a Pacific ascidian from Central American, with descriptions of species confused under this name. Zoological Journal of the Linnean Society, 110, 41 - 51. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 01470. x" type="journal article" year="1994">Monniot, 1994</bibRefCitation>
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: 44
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</treatmentCitation>
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||
</taxonomicName>
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, fig. 1;
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<treatmentCitation id="0AD2A900B05EFFFC8749D85E041BFE52" author="Van Name, W. G." box="[575,796,1369,1391]" page="336" pageId="6" pageNumber="497" year="1945">
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<bibRefCitation id="EFE2F2E0B05EFFFC8749D85E05EBFE52" author="Van Name, W. G." box="[575,748,1369,1391]" pageId="6" pageNumber="497" pagination="1 - 476" refId="ref12263" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name, 1945</bibRefCitation>
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||
: 336
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||
</treatmentCitation>
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||
(part)
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C369DC9AB05EFFFC85E1D898061BFEE5" pageId="6" pageNumber="497" type="materials_examined">
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<paragraph id="8BCC8F11B05EFFFC85E1D898061BFEE5" blockId="6.[151,1437,1436,2037]" pageId="6" pageNumber="497">
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||
<emphasis id="B9075303B05EFFFC85E1D8980689FE88" bold="true" box="[151,398,1436,1461]" pageId="6" pageNumber="497">Material Examined:</emphasis>
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<materialsCitation id="3B1B854CB05EFFFC84ECD8990290FE88" ID-GBIF-Occurrence="2012768558" box="[410,1431,1436,1461]" collectionCode="DZUP" collectorName="de Tierra & R. M. Rocha" country="Iceland" latitude="7.7443614" location="Canales de Tierra" longLatPrecision="1" longitude="-81.57922" pageId="6" pageNumber="497" specimenCode="PYU-144" specimenCount="1">
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<collectionCode id="ED6217D4B05EFFFC84ECD89906E4FE89" box="[410,483,1437,1460]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="497" type="University or college">DZUP</collectionCode>
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<specimenCode id="DBD5276AB05EFFFC8499D898055BFE89" box="[495,604,1436,1460]" collectionCode="PYU-" pageId="6" pageNumber="497">PYU-144</specimenCode>
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,
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<collectingCountry id="F364CF81B05EFFFC871BD8980592FE88" box="[621,661,1436,1461]" name="Iceland" pageId="6" pageNumber="497">Isla</collectingCountry>
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<location id="8EACD9CAB05EFFFC87D4D8980474FE88" LSID="urn:lsid:plazi:treatment:03DA3E07B05EFFFD85E1D9CB04D8F816:8EACD9CAB05EFFFC87D4D8980474FE88" box="[674,883,1436,1461]" country="Iceland" latitude="7.7443614" longLatPrecision="1" longitude="-81.57922" name="Canales de Tierra" pageId="6" pageNumber="497">
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Canales
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<collectorName id="2686EAC7B05EFFFC8670D8980474FE88" box="[774,883,1436,1461]" pageId="6" pageNumber="497">de Tierra</collectorName>
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</location>
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,
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<geoCoordinate id="EE47E9D6B05EFFFC86F5D898030BFE89" box="[899,1036,1436,1460]" direction="north" orientation="latitude" pageId="6" pageNumber="497" precision="1" value="7.7443614">7°44'39.7”N</geoCoordinate>
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<geoCoordinate id="EE47E9D6B05EFFFC816FD89803B0FE88" box="[1049,1207,1436,1461]" direction="west" orientation="longitude" pageId="6" pageNumber="497" precision="1" value="-81.57922">81°34'45.2”W</geoCoordinate>
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,
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<emphasis id="B9075303B05EFFFC81B0D89803F7FE89" box="[1222,1264,1436,1460]" italics="true" pageId="6" pageNumber="497">leg.</emphasis>
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<collectorName id="2686EAC7B05EFFFC818AD8990290FE88" box="[1276,1431,1436,1461]" pageId="6" pageNumber="497">R. M. Rocha</collectorName>
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</materialsCitation>
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, 0 8.01.2009.
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</paragraph>
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</subSubSection>
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<subSubSection id="C369DC9AB05EFFFD85B1D8E0020CF9A6" lastPageId="7" lastPageNumber="498" pageId="6" pageNumber="497" type="description">
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<paragraph id="8BCC8F11B05EFFFC85B1D8E007D5FDE8" blockId="6.[151,1437,1436,2037]" pageId="6" pageNumber="497">
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<emphasis id="B9075303B05EFFFC85B1D8E0065EFEC0" bold="true" box="[199,345,1508,1533]" pageId="6" pageNumber="497">Description.</emphasis>
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This Pacific species is common in the region in shallow waters
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<quantity id="4C8B22F4B05EFFFC8132D8E00389FEC1" box="[1092,1166,1508,1533]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="6" pageNumber="497" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">2–5 m</quantity>
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deep (low tide). It was found on rocky substrates in mainland,
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<collectingCountry id="F364CF81B05EFFFC8727DB03057EFD1D" box="[593,633,1543,1568]" name="Iceland" pageId="6" pageNumber="497">Isla</collectingCountry>
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Canales de Tierra and
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<collectingCountry id="F364CF81B05EFFFC860DDB0304A4FD1D" box="[891,931,1543,1568]" name="Iceland" pageId="6" pageNumber="497">Isla</collectingCountry>
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Canales de Afuera, but only one sample was collected due to the difficulty of detaching the animal from the substrate. It is extremely camouflaged by encrustations on the dark tunic, which is thick and hard. The siphons are red with four small lobes, but they are situated in depressions in the tunic (
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<figureCitation id="13489394B05EFFFC8737DB7005CFFDB1" box="[577,712,1652,1677]" captionStart="FIGURE 5" captionStartId="7.[151,250,1719,1741]" captionTargetBox="[176,1414,856,1697]" captionTargetId="figure@7.[173,1414,856,1697]" captionTargetPageId="7" captionText="FIGURE 5. Pyura lignosa Michaelsen, 1908 A. Animal in the field showing open siphons with red margins (arrowheads). B. External view of preserved animal. C. Right side, tunic removed. D. Left side, tunic removed. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2589410/files/figure.png" pageId="6" pageNumber="497">Figs 5A, B</figureCitation>
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), and when closed, it is impossible to spot the animal on the substrate. The size of our specimen with tunic was 7.5 x 6.5
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<date id="FFCDA9D1B05EFFFC8641DB9C045DFD92" box="[823,858,1688,1711]" pageId="6" pageNumber="497">
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||
x
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||
<quantity id="4C8B22F4B05EFFFC863ADB9C045DFD92" box="[844,858,1688,1711]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="497" unit="cm" value="7.0">7</quantity>
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</date>
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||
cm, but larger and smaller animals were seen in the field.
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</paragraph>
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<paragraph id="8BCC8F11B05EFFFC85B1DBDB059EFCB5" blockId="6.[151,1437,1436,2037]" pageId="6" pageNumber="497">
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The body wall is thick and opaque because the musculature forms a dense net of fibers in all directions (
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<figureCitation id="13489394B05EFFFC801CDBDB07E7FC21" captionStart="FIGURE 5" captionStartId="7.[151,250,1719,1741]" captionTargetBox="[176,1414,856,1697]" captionTargetId="figure@7.[173,1414,856,1697]" captionTargetPageId="7" captionText="FIGURE 5. Pyura lignosa Michaelsen, 1908 A. Animal in the field showing open siphons with red margins (arrowheads). B. External view of preserved animal. C. Right side, tunic removed. D. Left side, tunic removed. Scale bar = 1 cm." httpUri="https://zenodo.org/record/2589410/files/figure.png" pageId="6" pageNumber="497">Figs 5C, D</figureCitation>
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). Longitudinal muscles run from the siphons until the ventral margin. The atrial siphon is at half distance from the posterior region. The internal velum of the oral siphon has undulations covering the whole siphon, and there is a layer of tunic between the velum and the siphon musculature. The anterior third of the siphon lining is covered by spinules 50 µm long (
|
||
<figureCitation id="13489394B05EFFFC877BDA6B058BFCB5" box="[525,652,1903,1928]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="6" pageNumber="497">Figs 6C, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC8F11B05EFFFD85B1DA9003FFFBE8" blockId="6.[151,1437,1436,2037]" lastBlockId="7.[151,1436,151,320]" lastPageId="7" lastPageNumber="498" pageId="6" pageNumber="497">
|
||
There are 16 oral tentacles, flat and triangular. The first order ramifications appear along the posterior margin of the tentacle and are longer, while the second order ramifications are short and third order are minute (
|
||
<figureCitation id="13489394B05EFFFC804FDABC0288FCEC" box="[1337,1423,1976,2001]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="6" pageNumber="497">Fig. 6F</figureCitation>
|
||
). The dorsal tubercle is rounded, not protruding, with meandering aperture, while the peritubercle area forms a deep V (
|
||
<figureCitation id="13489394B05FFFFD85CDDD930611FB8D" box="[187,278,151,176]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="7" pageNumber="498">Fig. 6D</figureCitation>
|
||
). The dorsal lamina is divided in short triangular languets,
|
||
<quantity id="4C8B22F4B05FFFFD86C5DD9C04FCFB8D" box="[947,1019,151,176]" metricMagnitude="0" metricUnit="m" metricValue="2.5654" pageId="7" pageNumber="498" unit="in" value="101.0">101 in</quantity>
|
||
total, many of which are not aligned (two or three were seen side by side). The languets are very delicate and break easily when touched.
|
||
</paragraph>
|
||
<paragraph id="8BCC8F11B05FFFFD85B1DDDB05A5FA7D" blockId="7.[151,1436,151,320]" pageId="7" pageNumber="498">
|
||
The pharynx has six folds in each side and they are high but almost do not overlap. The folds on the left side are more curved than the ones on the right side (
|
||
<figureCitation id="13489394B05FFFFD87A8DC00043AFA20" box="[734,829,260,285]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="7" pageNumber="498">Fig. 6A</figureCitation>
|
||
). A total of 445 longitudinal vessels are distributed according the following formula (right to left):
|
||
</paragraph>
|
||
<paragraph id="8BCC8F11B05FFFFD85E1DC6E03CFFABC" blockId="7.[151,1224,361,386]" box="[151,1224,361,386]" pageId="7" pageNumber="498">E 4 (18) 11 (25) 12 (30) 15 (34) 8 (30) 10 DL 20 (27) 10 (33) 9 (35) 10 (32) 13 (28) 10 (16) 5 E</paragraph>
|
||
<paragraph id="8BCC8F11B05FFFFD85B1DCAE0435F936" blockId="7.[151,1436,426,811]" pageId="7" pageNumber="498">
|
||
There is a large amount of short vessels linking the pharynx with the body wall or with other organs, meaning that the atrial cavity is small. Also we confirm the thick tissue and projections described by
|
||
<bibRefCitation id="EFE2F2E0B05FFFFD81EBDCCA0257FADA" author="Monniot, C." box="[1181,1360,462,487]" pageId="7" pageNumber="498" pagination="41 - 51" refId="ref11817" refString="Monniot, C. (1994) Pyura lignosa (s. s.), a Pacific ascidian from Central American, with descriptions of species confused under this name. Zoological Journal of the Linnean Society, 110, 41 - 51. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 01470. x" type="journal article" year="1994">Monniot (1994)</bibRefCitation>
|
||
on the atrial side of the pharynx, along the dorsal region (
|
||
<figureCitation id="13489394B05FFFFD87B8DCF60422F936" box="[718,805,498,523]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="7" pageNumber="498">Fig. 6E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCC8F11B05FFFFD85B1DF12020CF9A6" blockId="7.[151,1436,426,811]" pageId="7" pageNumber="498">
|
||
The alimentary canal forms a wide first loop occupying the whole left side, and a short and close secondary loop (
|
||
<figureCitation id="13489394B05FFFFD85AFDF3E0636F96E" box="[217,305,570,595]" captionStart="FIGURE 6" captionStartId="8.[151,250,1713,1735]" captionTargetBox="[161,1425,517,1691]" captionTargetId="figure@8.[161,1425,517,1691]" captionTargetPageId="8" captionText="FIGURE 6. Pyura lignosa Michaelsen, 1908. A. Dissected animal showing the pharynx (stained). B. Dissected animal without the pharynx. C. Lining of the oral siphon. D. Stained anterior region showing oral tentacles and dorsal tubercle with meandering aperture (arrowhead). E. Detail of the atrial side of the dorsal region of the pharynx (stained). F. Detail of the oral tentacle (stained). G. Spinules in the oral siphon. Scale bar = 20 µm." httpUri="https://zenodo.org/record/2589412/files/figure.png" pageId="7" pageNumber="498">Fig. 6B</figureCitation>
|
||
). There are a few large endocarps on the intestinal wall and also on the gonads. The green digestive gland forms one large portion. The anus contains a typhlosole, but the margin is smooth. The gonads are large and occupy most of the space of the atrial cavity in each side. They do not form well individualized lobes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C369DC9AB05FFFFD85B1DFA204D8F816" pageId="7" pageNumber="498" type="discussion">
|
||
<paragraph id="8BCC8F11B05FFFFD85B1DFA204D8F816" blockId="7.[151,1436,426,811]" pageId="7" pageNumber="498">
|
||
<emphasis id="B9075303B05FFFFD85B1DFA2063CF982" bold="true" box="[199,315,678,703]" pageId="7" pageNumber="498">Remarks.</emphasis>
|
||
Except by small differences in the number of tentacles and longitudinal vessels, this specimen agrees well with the description of
|
||
<taxonomicName id="4C73F492B05FFFFD8742DFCF05A2F9DE" authorityName=": Monniot" authorityYear="1994" box="[564,677,715,739]" class="Ascidiacea" family="Pyuridae" genus="Pyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stolidobranchia" pageId="7" pageNumber="498" phylum="Chordata" rank="species" species="lignosa">
|
||
<emphasis id="B9075303B05FFFFD8742DFCF05A2F9DE" box="[564,677,715,739]" italics="true" pageId="7" pageNumber="498">P. lignosa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFE2F2E0B05FFFFD87A3DFCE048AF9DE" author="Monniot, C." box="[725,909,714,739]" pageId="7" pageNumber="498" pagination="41 - 51" refId="ref11817" refString="Monniot, C. (1994) Pyura lignosa (s. s.), a Pacific ascidian from Central American, with descriptions of species confused under this name. Zoological Journal of the Linnean Society, 110, 41 - 51. https: // doi. org / 10.1111 / j. 1096 - 3642.1994. tb 01470. x" type="journal article" year="1994">Monniot (1994)</bibRefCitation>
|
||
. The individuals from Gulf of California and
|
||
<collectingCountry id="F364CF81B05FFFFD85E1DFEA060DF83A" box="[151,266,750,775]" name="Nicaragua" pageId="7" pageNumber="498">Nicaragua</collectingCountry>
|
||
studied by
|
||
<bibRefCitation id="EFE2F2E0B05FFFFD84E0DFEB0563F83A" author="Van Name, W. G." box="[406,612,750,775]" pageId="7" pageNumber="498" pagination="1 - 476" refId="ref12263" refString="Van Name, W. G. (1945) The North and South American ascidians. Bulletin of the American Museum of Natural History, 84, 1 - 476." type="journal article" year="1945">Van Name (1945)</bibRefCitation>
|
||
and reported under the name
|
||
<taxonomicName id="4C73F492B05FFFFD86B1DFEB033EF83B" authorityName=": Monniot" authorityYear="1994" box="[967,1081,750,774]" class="Ascidiacea" family="Pyuridae" genus="Pyura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stolidobranchia" pageId="7" pageNumber="498" phylum="Chordata" rank="species" species="lignosa">
|
||
<emphasis id="B9075303B05FFFFD86B1DFEB033EF83B" box="[967,1081,750,774]" italics="true" pageId="7" pageNumber="498">P. lignosa</emphasis>
|
||
</taxonomicName>
|
||
do not seem to belong to this species and should be reviewed to have their identity correctly determined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |