533 lines
67 KiB
XML
533 lines
67 KiB
XML
<document id="A0940BF854069690A668ED4000A9E6DC" ID-CLB-Dataset="57492" ID-DOI="10.5852/ejt.2023.885.2183" ID-GBIF-Dataset="f1935cd8-83c6-45fd-aac5-8c6659a934aa" ID-ISSN="2118-9773" ID-Zenodo-Dep="8177540" ID-ZooBank="A928AB6A-0029-4CEA-B941-C2238643835A" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1690201302250" checkinUser="felipe" docAuthor="Arriaga-Jiménez, Alfonsina, Zunino, Mario, Halffter, Gonzalo, Escobar-Hernández, Fernando & Rossini, Michele" docDate="2023" docId="03CF324D047D1F40FDF3F7EAFB67F9A0" docLanguage="en" docName="ejt-885_1-20.pdf" docOrigin="European Journal of Taxonomy 885" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ceratotrupes gonzaloi Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini 2023, sp. nov." docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="FFF64A3504761F4FFF97F328FF93FFF7" masterDocTitle="A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur" masterLastPageNumber="20" masterPageNumber="1" pageNumber="12" updateTime="1699544954118" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
|
||
<mods:mods id="30398B2F976080901CAE7151F92B7DC7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="A3B773214A961A4E8E47355556E0050F">
|
||
<mods:title id="A8CAFC5AC8714E411308DD1631349505">A taxonomic review of the endemic Mexican genus Ceratotrupes Jekel, 1865 (Coleoptera: Geotrupidae: Ceratotrupini), with the description of an unexpected new species from the Sierra Madre del Sur</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DDB8BDBF7A485D0E20A66C9DCFFB4FE8" type="personal">
|
||
<mods:role id="13438A5EBCA978B48E448AEBE7C093A9">
|
||
<mods:roleTerm id="F64EA36ADDC35824C7A7F3E8AEFB2B35">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="97A57FACE4E515344CA8D669DEA0C5B8">Arriaga-Jiménez, Alfonsina</mods:namePart>
|
||
<mods:nameIdentifier id="D6B27B14AAA63A0CCBCBC09ACCB381B7" type="LSID">C63A2853-E857-4683-A5AE-626D2387A352</mods:nameIdentifier>
|
||
<mods:affiliation id="CA25BA3E69829E8D61270A4433D988FF">Insect Ecology Lab, School of Environmental and Rural Sciences, University of New England, Armidale NSW 2351, Australia.</mods:affiliation>
|
||
<mods:nameIdentifier id="DFC96F00570A993CC4EA619EE70F91B9" type="email">aarriaga@une.edu.au</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="930C746105E412BB7F343CAFA5059825" type="personal">
|
||
<mods:role id="47FAE99549493DE7891876777ACB4BD8">
|
||
<mods:roleTerm id="0EB70CCE12BC12051CF44C54A9B67A6C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D1A318064B1027B8DA714563D0C23907">Zunino, Mario</mods:namePart>
|
||
<mods:nameIdentifier id="B6C64B19F81D4492CCDC4ADB2DDBAD2D" type="LSID">A1EE6CAE-8FC5-413B-9984-625DF6C4C486</mods:nameIdentifier>
|
||
<mods:affiliation id="4B319FC618A7B48054BA5D8E27B31CBA">Instituto de Ecología A. C., Invited researcher, Carretera antigua a Coatepec 351, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
|
||
<mods:nameIdentifier id="6562DD74804B8B706FA8E213F5305BFF" type="email">mariozunino@hotmail.it</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="89E3E3D5A58E0CD75BFC725C238DC852" type="personal">
|
||
<mods:role id="03B10CC3C452C2101FC0809CF0ACBC9D">
|
||
<mods:roleTerm id="D02576E6EF503BAE0CECDCCAEB45D4B4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F466E40EAB601536F6FC5B350EF16379">Halffter, Gonzalo</mods:namePart>
|
||
<mods:nameIdentifier id="D09D974F77A395DF704A6FDD9E28371B" type="LSID">CF2EF74E-878A-4BC8-82A2-01F669049650</mods:nameIdentifier>
|
||
<mods:affiliation id="8B7E483835E634590FCD8F9650C92600">Deceased 11 March 2022. Former address: Aldama Street 17, Centro, 91500 Coatepec, Veracruz, Mexico.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="BED1D1D1BEC656A472188D2709B6E614" type="personal">
|
||
<mods:role id="2418BD170D77E27F62CAF5E02A9E5F9B">
|
||
<mods:roleTerm id="1E76283A5727960B5DBCA665C90103BC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A577D4D61271AB8561A36DA27BFDAE60">Escobar-Hernández, Fernando</mods:namePart>
|
||
<mods:nameIdentifier id="426258D16B4BFE5ADFFE992836894224" type="LSID">0F5EFB92-6896-4E9D-A3E3-6DB3F33B762A</mods:nameIdentifier>
|
||
<mods:affiliation id="882786B5E067C2DF7072B36CA6137C70">Red de Ecoetología, Instituto de Ecología, INECOL A. C. Xalapa, México, Carretera antigua a Coatepec 351, El Haya, Xalapa 91070, Veracruz, Mexico.</mods:affiliation>
|
||
<mods:nameIdentifier id="AC23239ECEFBE56DCFF69199D9AB76F2" type="email">fernando.escobar@inecol.mx</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="AC42EA16825150CA9E780C018A3DEB17" type="personal">
|
||
<mods:role id="BD37AA80F4B3AEFC3FD1615138809335">
|
||
<mods:roleTerm id="91DD12DECF7AB2F83F375215798CF492">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6AED491D605A9CE19683773D37527468">Rossini, Michele</mods:namePart>
|
||
<mods:nameIdentifier id="C49C0C2E2A3D75A644F6339D5E5BD995" type="LSID">2698881E-C265-42DB-8A8E-0BE3A5BDDB97</mods:nameIdentifier>
|
||
<mods:affiliation id="E5E3819E7038724DB35AE5A26E8C43AD">Finnish Museum of Natural History (LUOMUS), University of Helsinki, Pohjoinen Rautatiekatu 13, Helsinki, 00014, Finland. & Department of Agronomy, Food, Natural resources, Animals and Environment (DAFNAE), University of Padova, Via dell'Università 16, 35020, Legnaro (PD), Italy.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="8B612E6B99790A2E7DA3ABBCA8DD2F7B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="6631DB4A95318EF05C74232B202E583E" type="host">
|
||
<mods:titleInfo id="2D264DA275D6800A1320063BC7B8242F">
|
||
<mods:title id="8D30CFEA534B6717791B7412B5C5D20C">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="5BF87D3423C2C549EFEBF99D35A0CCC9">
|
||
<mods:date id="0DE62508A36B7DBDD9042854EFF06197">2023</mods:date>
|
||
<mods:detail id="971B65E3933A3381E933A4BE64F593E0" type="pubDate">
|
||
<mods:number id="8D1F4DFF8F65459289721C063056F1F7">2023-07-24</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="95486B2A7E576E24CF4D7D0D126A1C8B" type="volume">
|
||
<mods:number id="EA1AB3E021448DF0F93B26B287392A1E">885</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="081B1CEA9F97D75EC5897A58C7A72F1C" unit="page">
|
||
<mods:start id="60EC002BFEDC60CF6CB766CF6D72FA45">1</mods:start>
|
||
<mods:end id="499BB27158F3205582740D2D96FB848B">20</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="3778C75D4A9E769E828D9648540AAA96">journal article</mods:classification>
|
||
<mods:identifier id="C4EB8BCE84B8B9E65978ACCB59068596" type="CLB-Dataset">57492</mods:identifier>
|
||
<mods:identifier id="E612304EA2B1A521CA64D51F72452006" type="DOI">10.5852/ejt.2023.885.2183</mods:identifier>
|
||
<mods:identifier id="2B1FF67B691B2AD52EDD09ACF5725297" type="GBIF-Dataset">f1935cd8-83c6-45fd-aac5-8c6659a934aa</mods:identifier>
|
||
<mods:identifier id="ECC775675750EE34AE7E97197B384C61" type="ISSN">2118-9773</mods:identifier>
|
||
<mods:identifier id="C533B260079D9073B738319F7FA3A90A" type="Zenodo-Dep">8177540</mods:identifier>
|
||
<mods:identifier id="C6465CAD316613893130A18923E5FE8E" type="ZooBank">A928AB6A-0029-4CEA-B941-C2238643835A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03CF324D047D1F40FDF3F7EAFB67F9A0" ID-DOI="http://doi.org/10.5281/zenodo.8182243" ID-GBIF-Taxon="212419182" ID-Zenodo-Dep="8182243" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0" httpUri="http://treatment.plazi.org/id/03CF324D047D1F40FDF3F7EAFB67F9A0" lastPageId="15" lastPageNumber="16" pageId="11" pageNumber="12">
|
||
<subSubSection id="C37CD0D0047D1F44FDF3F7EAFC43FB2B" box="[612,976,1217,1244]" pageId="11" pageNumber="12" type="nomenclature">
|
||
<paragraph id="8BD9835B047D1F44FDF3F7EAFC43FB2B" blockId="11.[364,1223,1217,1314]" box="[612,976,1217,1244]" pageId="11" pageNumber="12">
|
||
<heading id="D0913437047D1F44FDF3F7EAFC43FB2B" box="[612,976,1217,1244]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="12" reason="2">
|
||
<taxonomicName id="4C66F8D8047D1F44FDF3F7EAFCFDFB2B" ID-CoL="BLZQ3" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[612,878,1217,1244]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047D1F44FDF3F7EAFCFDFB2B" bold="true" box="[612,878,1217,1244]" italics="true" pageId="11" pageNumber="12">Ceratotrupes gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047D1F44FCE1F7EAFC43FB2B" box="[886,976,1218,1244]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047D1F44FEFBF7CDFCAFFAD5" pageId="11" pageNumber="12" type="description">
|
||
<paragraph id="8BD9835B047D1F44FEFBF7CDFB54FAF7" blockId="11.[364,1223,1217,1314]" box="[364,1223,1253,1280]" pageId="11" pageNumber="12">
|
||
<uri id="FFF78F59047D1F44FEFBF7CDFB54FAF7" box="[364,1223,1253,1280]" pageId="11" pageNumber="12">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFC0B98E047D1F44FD18F7CDFB54FAF7" box="[655,1223,1253,1280]" pageId="11" pageNumber="12">49D5CB15-8785-47DB-A4A4-865994D4C1C5</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FD60F620FCAFFAD5" blockId="11.[364,1223,1217,1314]" box="[759,828,1288,1314]" pageId="11" pageNumber="12">
|
||
<figureCitation id="135D9FDE047D1F44FD60F620FCAFFAD5" box="[759,828,1288,1314]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047D1F44FF2AF661FE2BF97E" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BD9835B047D1F44FF2AF661FEA5FA94" blockId="11.[189,310,1353,1379]" box="[189,310,1353,1379]" pageId="11" pageNumber="12">
|
||
<heading id="D0913437047D1F44FF2AF661FEA5FA94" bold="true" box="[189,310,1353,1379]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B9125F49047D1F44FF2AF661FEA5FA94" bold="true" box="[189,310,1353,1379]" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF650FE2BF97E" blockId="11.[188,1399,1400,1673]" pageId="11" pageNumber="12">
|
||
The following characters separate
|
||
<taxonomicName id="4C66F8D8047D1F44FDC5F650FD49FA65" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[594,730,1400,1426]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047D1F44FDC5F650FD49FA65" box="[594,730,1400,1426]" italics="true" pageId="11" pageNumber="12">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047D1F44FD76F651FCAFFA64" box="[737,828,1401,1427]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from its close relative,
|
||
<taxonomicName id="4C66F8D8047D1F44FBC5F650FB53FA65" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1106,1216,1400,1426]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FBC5F650FB53FA65" box="[1106,1216,1400,1426]" italics="true" pageId="11" pageNumber="12">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
: posterior edge of pronotum without margin at middle (
|
||
<figureCitation id="135D9FDE047D1F44FD1DF6B3FD75FA42" box="[650,742,1435,1461]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4C</figureCitation>
|
||
); clypeal horn of male stronger, laterally wider (
|
||
<figureCitation id="135D9FDE047D1F44FA81F6B3FF47FA2E" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4A, D</figureCitation>
|
||
), tip depressed at middle; pronotal projection larger, with parallel sides (
|
||
<figureCitation id="135D9FDE047D1F44FB8CF696FB08FA2E" box="[1051,1179,1470,1497]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4A, D</figureCitation>
|
||
); elytral interstriae very finely and densely punctate. Male genitalia differ as follows: distal tip of right ventral lobe of phallobase straight and directed obliquely upward (
|
||
<figureCitation id="135D9FDE047D1F44FCB5F52DFCEAF9E8" box="[802,889,1541,1567]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4J</figureCitation>
|
||
); shorter branch of the bifurcation of right lobe of phallobase distinctly shorter than in
|
||
<taxonomicName id="4C66F8D8047D1F44FD53F500FCA2F9B5" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[708,817,1576,1602]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F49047D1F44FD53F500FCA2F9B5" box="[708,817,1576,1602]" italics="true" pageId="11" pageNumber="12">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="135D9FDE047D1F44FCD5F500FC0DF9B5" box="[834,926,1576,1602]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4L</figureCitation>
|
||
); parameres differently shaped (
|
||
<figureCitation id="135D9FDE047D1F44FA8FF500FAE1F9B5" box="[1304,1394,1576,1602]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4F</figureCitation>
|
||
; arrows indicate the main differences); in lateral view, left paramere simple, without distinct lobe beneath apical spur (
|
||
<figureCitation id="135D9FDE047D1F44FEDBF546FE38F97E" box="[332,427,1646,1673]" captionStart="Fig" captionStartId="12.[189,232,1752,1778]" captionTargetBox="[289,1301,279,1699]" captionTargetId="figure-19@12.[283,1305,265,1711]" captionTargetPageId="12" captionText="Fig. 4. Ceratotrupes gonzaloi sp. nov. (CMNC). A–B. Dorsal habitus of male (A) and female (B). C. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin. D. Details of cephalic horn and pronotal projection of male. E. Aedeagus in dorsal view. F. Parameres in dorsal view; arrows indicate main differences between C. gonzaloi sp. nov. and C. sturmi. G. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.H. Tergite.I. Pleurotergites. J. Phallobase in ventral view; arrow indicates straight margin of ventral lobe. K. Phallobase in dorsal view. L. Phallobase in lateral view; arrow indicates the short bifurcation. M. Internal sac (is) and median struts (ms) in dorsal view. N. Internal sac and median struts in lateral view." figureDoi="http://doi.org/10.5281/zenodo.8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="11" pageNumber="12">Fig. 4G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047D1F44FF2AF598FC39F872" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BD9835B047D1F44FF2AF598FED7F93D" blockId="11.[189,324,1712,1738]" box="[189,324,1712,1738]" pageId="11" pageNumber="12">
|
||
<heading id="D0913437047D1F44FF2AF598FED7F93D" bold="true" box="[189,324,1712,1738]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
|
||
<emphasis id="B9125F49047D1F44FF2AF598FED7F93D" bold="true" box="[189,324,1712,1738]" pageId="11" pageNumber="12">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047D1F44FF2AF5F6FC39F872" blockId="11.[188,1400,1758,1926]" pageId="11" pageNumber="12">
|
||
This species is dedicated to Gonzalo Halffter (1932–2022), a renowned academic, researcher and entomologist, who passed away during the preparation of the current paper. Founder of INECOL and Biosphere Reserves, UNESCO, his research in the biogeography and ecology of beetles is a corner stone for research in
|
||
<collectingCountry id="F371C3CB047D1F44FEE2F460FE5CF895" box="[373,463,1864,1890]" name="Mexico" pageId="11" pageNumber="12">Mexico</collectingCountry>
|
||
and worldwide. But above all, Gonzalo, Dr Halffter, our “Maestro”, was our mentor, teacher, colleague, and our endearing and lovely friend.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF19D3D3047A1F43FF2AF5F0FBB8F81F" ID-DOI="http://doi.org/10.5281/zenodo.8177550" ID-Zenodo-Dep="8177550" httpUri="https://zenodo.org/record/8177550/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1752,1778]" targetBox="[289,1301,279,1699]" targetPageId="12">
|
||
<paragraph id="8BD9835B047A1F43FF2AF5F0FBB8F81F" blockId="12.[187,1399,1752,2025]" pageId="12" pageNumber="13">
|
||
<emphasis id="B9125F49047A1F43FF2AF5F0FE9CF905" bold="true" box="[189,271,1752,1778]" pageId="12" pageNumber="13">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C66F8D8047A1F43FE8DF5F0FDB5F905" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[282,550,1752,1778]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047A1F43FE8DF5F0FDB5F905" box="[282,550,1752,1778]" italics="true" pageId="12" pageNumber="13">Ceratotrupes gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047A1F43FDA6F5F0FD03F905" box="[561,656,1752,1778]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(CMNC).
|
||
<emphasis id="B9125F49047A1F43FC8FF5F0FCC3F905" bold="true" box="[792,848,1752,1778]" pageId="12" pageNumber="13">A–B</emphasis>
|
||
. Dorsal habitus of male (A) and female (B).
|
||
<emphasis id="B9125F49047A1F43FF2AF5D3FF41F8E2" bold="true" box="[189,210,1787,1813]" pageId="12" pageNumber="13">C</emphasis>
|
||
. Details of head and prothorax; arrows indicate posteromedial edge of pronotum without margin.
|
||
<emphasis id="B9125F49047A1F43FF2AF436FF41F8CF" bold="true" box="[189,210,1822,1848]" pageId="12" pageNumber="13">D</emphasis>
|
||
. Details of cephalic horn and pronotal projection of male.
|
||
<emphasis id="B9125F49047A1F43FC1DF436FC0EF8CF" bold="true" box="[906,925,1822,1848]" pageId="12" pageNumber="13">E</emphasis>
|
||
. Aedeagus in dorsal view.
|
||
<emphasis id="B9125F49047A1F43FB4BF436FB7EF8CF" bold="true" box="[1244,1261,1822,1848]" pageId="12" pageNumber="13">F</emphasis>
|
||
. Parameres in dorsal view; arrows indicate main differences between
|
||
<taxonomicName id="4C66F8D8047A1F43FCFDF46AFC61F8AC" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[874,1010,1857,1884]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F49047A1F43FCFDF46AFC61F8AC" box="[874,1010,1857,1884]" italics="true" pageId="12" pageNumber="13">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E232047A1F43FC6DF46AFBC5F8AC" box="[1018,1110,1858,1884]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<emphasis id="B9125F49047A1F43FB06F46AFA94F8AC" box="[1169,1287,1858,1884]" italics="true" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C66F8D8047A1F43FB06F46AFA90F8AC" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[1169,1283,1858,1884]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sturmi">C. sturmi</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B9125F49047A1F43FA87F469FAB5F8AC" bold="true" box="[1296,1318,1857,1883]" pageId="12" pageNumber="13">G</emphasis>
|
||
. Right paramere in lateral view; arrow indicates absence of lobe beneath apical spur.
|
||
<emphasis id="B9125F49047A1F43FBA5F44DFBDAF888" bold="true" box="[1074,1097,1893,1919]" pageId="12" pageNumber="13">H</emphasis>
|
||
. Tergite.
|
||
<emphasis id="B9125F49047A1F43FB25F44DFB2EF888" bold="true" box="[1202,1213,1893,1919]" pageId="12" pageNumber="13">I</emphasis>
|
||
. Pleurotergites.
|
||
<emphasis id="B9125F49047A1F43FF2CF4A0FF59F855" bold="true" box="[187,202,1928,1954]" pageId="12" pageNumber="13">J</emphasis>
|
||
. Phallobase in ventral view; arrow indicates straight margin of ventral lobe.
|
||
<emphasis id="B9125F49047A1F43FBF5F4A0FBE4F855" bold="true" box="[1122,1143,1928,1954]" pageId="12" pageNumber="13">K</emphasis>
|
||
. Phallobase in dorsal view.
|
||
<emphasis id="B9125F49047A1F43FE96F483FE86F832" bold="true" box="[257,277,1963,1989]" pageId="12" pageNumber="13">L</emphasis>
|
||
. Phallobase in lateral view; arrow indicates the short bifurcation.
|
||
<emphasis id="B9125F49047A1F43FB9CF483FBB4F832" bold="true" box="[1035,1063,1963,1989]" pageId="12" pageNumber="13">M</emphasis>
|
||
. Internal sac (is) and median struts (ms) in dorsal view.
|
||
<emphasis id="B9125F49047A1F43FE63F4E6FD9AF81F" bold="true" box="[500,521,1998,2024]" pageId="12" pageNumber="13">N</emphasis>
|
||
. Internal sac and median struts in lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37CD0D0047B1F42FF2AF221FE4EFBA4" pageId="13" pageNumber="14" type="materials_examined">
|
||
<paragraph id="8BD9835B047B1F42FF2AF221FE7FFED5" blockId="13.[189,492,264,291]" box="[189,492,264,291]" pageId="13" pageNumber="14">
|
||
<heading id="D0913437047B1F42FF2AF221FE7FFED5" bold="true" box="[189,492,264,291]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B9125F49047B1F42FF2AF221FE7FFED5" bold="true" box="[189,492,264,291]" pageId="13" pageNumber="14">Type material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF7DF21FFEA8FE29" blockId="13.[187,1398,311,478]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B0E8906047B1F42FF7DF21FFEA8FE29" ID-GBIF-Occurrence="4164095317" collectingDate="1965-08-25" collectionCode="CMNC" collectorName="G. V. V. Halffter" country="Mexico" location="Canadian Museum of Nature" municipality="Omiltemi" pageId="13" pageNumber="14" specimenCode="CMNC 00031076" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="holotype">
|
||
<emphasis id="B9125F49047B1F42FF7DF21FFECFFEA6" bold="true" box="[234,348,311,337]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54DD3DF9047B1F42FF7DF21FFECFFEA6" box="[234,348,311,337]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
MEXICO •
|
||
<specimenCount id="9D6048D2047B1F42FED8F272FE13FE82" box="[335,384,346,373]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FE0EF272FD9BFE83" box="[409,520,346,372]" country="Mexico" name="Guerrero" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBD1921047B1F42FD80F272FD14FE83" box="[535,647,346,372]" pageId="13" pageNumber="14">Omiltemi</collectingMunicipality>
|
||
/
|
||
<date id="FFD8A59B047B1F42FD34F272FCDBFE82" box="[675,840,346,373]" pageId="13" pageNumber="14" value="1965-08-25">
|
||
<collectingDate id="EF9C5C73047B1F42FD34F272FCDBFE82" box="[675,840,346,373]" pageId="13" pageNumber="14" value="1965-08-25">25 Aug. 1965</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FCF4F272FB84FE83" box="[867,1047,346,372]" pageId="13" pageNumber="14">G.V.V. Halffter</collectorName>
|
||
leg. / Cebo, exterior bosque, I
|
||
<emphasis id="B9125F49047B1F42FF50F255FF40FE60" bold="true" box="[199,211,381,407]" pageId="13" pageNumber="14">°</emphasis>
|
||
Estación dia, bosque nebular, pino (modificado) / Halffter 73-18 / HOLOTYPE
|
||
<taxonomicName id="4C66F8D8047B1F42FBE6F256FD98FE4C" authority="Arriaga-Jimenez et al., 2022" authorityName="Arriaga-Jimenez" authorityYear="2022" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gonzaloi">
|
||
<emphasis id="B9125F49047B1F42FBE6F256FAE5FE60" box="[1137,1398,381,408]" italics="true" pageId="13" pageNumber="14">Ceratotrupes gonzaloi</emphasis>
|
||
Arriaga-Jiménez et al., 2022
|
||
</taxonomicName>
|
||
/
|
||
<location id="8EB9D580047B1F42FDB5F289FCE8FE4C" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FDB5F289FCE8FE4C" box="[546,891,417,443]" country="Mexico" municipality="Omiltemi" name="Canadian Museum of Nature" pageId="13" pageNumber="14" stateProvince="Guerrero">Canadian Museum of Nature</location>
|
||
, Musée canadien de la NATURE”;
|
||
<specimenCode id="DBC02B20047B1F42FA89F289FEA4FE29" collectionCode="CMNC" pageId="13" pageNumber="14">
|
||
<collectionCode id="ED771B9E047B1F42FA89F289FAE7FE4C" box="[1310,1396,417,443]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="13" pageNumber="14">CMNC</collectionCode>
|
||
00031076
|
||
</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF7DF12DFE4EFBA4" blockId="13.[189,1399,517,1107]" pageId="13" pageNumber="14">
|
||
<materialsCitation id="3B0E8906047B1F42FF7DF12DFBB8FD7E" ID-GBIF-Occurrence="4164095329" collectingDate="1966-08-14" collectingDateMax="1966-08-15" collectingDateMin="1966-08-14" collectionCode="CMNC" collectorName="Ball-Whitehead" country="Mexico" determinerName="H. F. Howden" location="Omiltemi" municipality="Omiltemi" pageId="13" pageNumber="14" specimenCode="CMNC 00031074" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<emphasis id="B9125F49047B1F42FF7DF12DFEF4FDE8" bold="true" box="[234,359,517,543]" pageId="13" pageNumber="14">
|
||
<typeStatus id="54DD3DF9047B1F42FF7DF12DFEF4FDE8" box="[234,359,517,543]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F371C3CB047B1F42FF2AF101FEA2FDB4" box="[189,305,553,579]" name="Mexico" pageId="13" pageNumber="14">MEXICO</collectingCountry>
|
||
–
|
||
<emphasis id="B9125F49047B1F42FEC7F100FE55FDB5" bold="true" box="[336,454,552,578]" pageId="13" pageNumber="14">
|
||
<collectingRegion id="49A24DB9047B1F42FEC7F100FE55FDB5" box="[336,454,552,578]" country="Mexico" name="Guerrero" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D6048D2047B1F42FE48F101FD9CFDB4" box="[479,527,553,579]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “Guerrero,
|
||
<collectingMunicipality id="6BBD1921047B1F42FD35F100FC85FDB5" box="[674,790,552,578]" pageId="13" pageNumber="14">Omiltemi</collectingMunicipality>
|
||
; 7300’ alt. /
|
||
<date id="FFD8A59B047B1F42FC3FF101FBE1FDB5" box="[936,1138,552,579]" pageId="13" pageNumber="14" value="1966-08-14" valueMax="1966-08-15" valueMin="1966-08-14">
|
||
<collectingDate id="EF9C5C73047B1F42FC3FF101FBE1FDB5" box="[936,1138,552,579]" pageId="13" pageNumber="14" value="1966-08-14" valueMax="1966-08-15" valueMin="1966-08-14">Aug. 14-15 1966</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FB1DF100FAD7FDB5" box="[1162,1348,552,578]" pageId="13" pageNumber="14">Ball-Whitehead</collectorName>
|
||
leg. / Specimén photographié par François Génier /
|
||
<taxonomicName id="4C66F8D8047B1F42FC93F164FBDEFD91" authority="(Jekel)" authorityName="CMNC" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[772,1101,588,614]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<originalDetermination id="A8783BA7047B1F42FC93F164FBDEFD91" box="[772,1101,588,614]" pageId="13" pageNumber="14">Ceratotrupes sturmi (Jekel)</originalDetermination>
|
||
</taxonomicName>
|
||
, Det.
|
||
<determinerName id="7E056F9F047B1F42FB0DF164FAA9FD91" box="[1178,1338,588,614]" pageId="13" pageNumber="14">H.F. Howden</determinerName>
|
||
69 / Canadian Museum of Musée canadien de la NATURE”;
|
||
<specimenCode id="DBC02B20047B1F42FCCFF147FBB8FD7E" box="[856,1067,623,649]" collectionCode="CMNC" pageId="13" pageNumber="14">CMNC 00031074</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FBD3F147FE1CFD27" ID-GBIF-Occurrence="4164095304" collectingDate="1965-08" collectionCode="CMNC" collectorName="P. Reyes C." location="Omiltemi" municipality="Omiltemi" pageId="13" pageNumber="14" specimenCode="CMNC 00031075" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FBD3F147FBE0FD7E" box="[1092,1139,623,649]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FB1CF147FB69FD7E" box="[1163,1274,623,649]" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBD1921047B1F42FA91F147FAE5FD7E" box="[1286,1398,623,649]" pageId="13" pageNumber="14">Omiltemi</collectingMunicipality>
|
||
/
|
||
<date id="FFD8A59B047B1F42FF5CF1BAFED5FD5A" box="[203,326,658,685]" pageId="13" pageNumber="14" value="1965-08">
|
||
<collectingDate id="EF9C5C73047B1F42FF5CF1BAFED5FD5A" box="[203,326,658,685]" pageId="13" pageNumber="14" value="1965-08">Aug. 1965</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FECBF1BBFE72FD5B" box="[348,481,658,685]" pageId="13" pageNumber="14">P. Reyes C.</collectorName>
|
||
leg. / Halffter 73-29 / Canadian Museum of Musée canadien de la NATURE”;
|
||
<specimenCode id="DBC02B20047B1F42FF2AF19DFE1CFD27" box="[189,399,693,720]" collectionCode="CMNC" pageId="13" pageNumber="14">CMNC 00031075</specimenCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FE3FF19DFAB1FD04" ID-GBIF-Occurrence="4164095328" collectingDate="2018-08-09" collectionCode="FMNH" collectorName="Arriaga J. A." elevation="2244" latitude="17.560528" location="Omiltemi" longLatPrecision="1" longitude="-99.691086" municipality="Omiltemi" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FE3FF19DFE44FD27" box="[424,471,693,720]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FE79F19DFDCEFD38" box="[494,605,693,719]" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BBD1921047B1F42FDFEF19DFD4AFD38" box="[617,729,693,719]" pageId="13" pageNumber="14">Omiltemi</collectingMunicipality>
|
||
/
|
||
<geoCoordinate id="EE52E59C047B1F42FD67F19DFC33FD38" box="[752,928,693,719]" degrees="99" direction="west" minutes="41" orientation="longitude" pageId="13" pageNumber="14" precision="1" seconds="27.9" value="-99.691086">99°41’27.9’’W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE52E59C047B1F42FC3BF19DFBC5FD38" box="[940,1110,693,719]" degrees="17" direction="north" minutes="33" orientation="latitude" pageId="13" pageNumber="14" precision="1" seconds="37.9" value="17.560528">17°33’37.9’’N</geoCoordinate>
|
||
/
|
||
<elevation id="004B6468047B1F42FBFBF19DFA90FD38" box="[1132,1283,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.244" pageId="13" pageNumber="14" unit="m" value="2244.0">
|
||
<quantity id="4C9E2EBE047B1F42FBFBF19DFB56FD27" box="[1132,1221,693,720]" metricMagnitude="3" metricUnit="m" metricValue="2.244" pageId="13" pageNumber="14" unit="m" value="2244.0">2244 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
/
|
||
<date id="FFD8A59B047B1F42FA8EF19DFF6BFD04" pageId="13" pageNumber="14" value="2018-08-09">
|
||
<collectingDate id="EF9C5C73047B1F42FA8EF19DFF6BFD04" pageId="13" pageNumber="14" value="2018-08-09">09 Aug. 2018</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FE85F1F0FE36FD04" box="[274,421,728,755]" pageId="13" pageNumber="14">Arriaga J.A.</collectorName>
|
||
leg. / bosque de galleria, coprotrampa / http://id.luomus.fi/NC.07;
|
||
<collectionCode id="ED771B9E047B1F42FB5AF1F1FAB1FD04" box="[1229,1314,729,755]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14" type="Museum">FMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FAD7F1F1FB81FCAA" ID-GBIF-Occurrence="4164095307" collectingDate="1965-07-25" collectionCode="FMNH" collectorName="G. y V. Halffter" determinerName="H. F. Howden, 1973" location="Omiltemi" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FAD7F1F1FAE2FD04" box="[1344,1393,729,755]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
;
|
||
<collectingRegion id="49A24DB9047B1F42FF2AF1D4FEB9FCE1" box="[189,298,764,790]" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<location id="8EB9D580047B1F42FEA0F1D4FE34FCE1" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FEA0F1D4FE34FCE1" box="[311,423,764,790]" country="Mexico" county="Guerrero" latitude="17.560528" longLatPrecision="1" longitude="-99.691086" municipality="Omiltemi" name="Omiltemi" pageId="13" pageNumber="14" stateProvince="Guerrero">Omiltemi</location>
|
||
/
|
||
<date id="FFD8A59B047B1F42FE55F1D3FDC6FCE1" box="[450,597,763,790]" pageId="13" pageNumber="14" value="1965-07-25">
|
||
<collectingDate id="EF9C5C73047B1F42FE55F1D3FDC6FCE1" box="[450,597,763,790]" pageId="13" pageNumber="14" value="1965-07-25">25 Jul. 1965</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FDF8F1D4FCBAFCE1" box="[623,809,764,790]" pageId="13" pageNumber="14">G. y V. Halffter</collectorName>
|
||
leg. / camino bosque nebular, pino (modificado), 1° estación, cebo excremento, excremento humano, noche-dia, Bosque nebular /
|
||
<taxonomicName id="4C66F8D8047B1F42FB1BF037FECDFCAB" authority="(Jekel, 1866)" baseAuthorityName="Jekel" baseAuthorityYear="1866" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<originalDetermination id="A8783BA7047B1F42FB1BF037FECDFCAB" pageId="13" pageNumber="14">Ceratotrupes sturmi (Jekel, 1866)</originalDetermination>
|
||
</taxonomicName>
|
||
,
|
||
<determinerName id="7E056F9F047B1F42FEF8F06BFDF7FCAB" box="[367,612,834,861]" pageId="13" pageNumber="14">H.F. Howden, 1973</determinerName>
|
||
”; http://id.luomus.fi/NC.08;
|
||
<collectionCode id="ED771B9E047B1F42FC2AF06BFB81FCAA" box="[957,1042,835,861]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14" type="Museum">FMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FBA2F06AFBB2FC31" ID-GBIF-Occurrence="4164095353" collectingDate="1965-07-25" collectionCode="FMNH" collectorName="G. y V. Halffter" determinerName="H. F. Howden, 1973" location="Omiltemi" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FBA2F06AFBF9FCAA" box="[1077,1130,834,861]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FB11F06AFB66FCAB" box="[1158,1269,834,860]" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<location id="8EB9D580047B1F42FA91F06AFAE5FCAB" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FA91F06AFAE5FCAB" box="[1286,1398,834,860]" country="Mexico" municipality="Guerrero" name="Omiltemi" pageId="13" pageNumber="14" stateProvince="Guerrero">Omiltemi</location>
|
||
/
|
||
<date id="FFD8A59B047B1F42FF47F04DFEF4FC77" box="[208,359,869,896]" pageId="13" pageNumber="14" value="1965-07-25">
|
||
<collectingDate id="EF9C5C73047B1F42FF47F04DFEF4FC77" box="[208,359,869,896]" pageId="13" pageNumber="14" value="1965-07-25">25 Jul. 1965</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="2693E68D047B1F42FE13F04DFDD7FC88" box="[388,580,869,896]" pageId="13" pageNumber="14">G. y V. Halffter</collectorName>
|
||
leg. / camino bosque nebular, pino (modificado), 1° estación, cebo excremento, excremento humano, noche-dia, Bosque nebular /
|
||
<taxonomicName id="4C66F8D8047B1F42FC45F0A1FAE1FC55" authority="(Jekel, 1866)" baseAuthorityName="Jekel" baseAuthorityYear="1866" box="[978,1394,904,931]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<originalDetermination id="A8783BA7047B1F42FC45F0A1FAE1FC55" box="[978,1394,904,931]" pageId="13" pageNumber="14">Ceratotrupes sturmi (Jekel, 1866)</originalDetermination>
|
||
</taxonomicName>
|
||
,
|
||
<determinerName id="7E056F9F047B1F42FF2AF084FE3EFC31" box="[189,429,940,966]" pageId="13" pageNumber="14">H.F. Howden, 1973</determinerName>
|
||
, Howden 73-26”; http://id.luomus.fi/NC.09;
|
||
<collectionCode id="ED771B9E047B1F42FC5BF084FBB2FC31" box="[972,1057,940,966]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14" type="Museum">FMNH</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FBD6F084FCFBFC1E" ID-GBIF-Occurrence="4164095308" collectingDate="1990-08-02" collectionCode="CEMT" collectorName="Fco. E. Figueroa" location="Tetipac" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Guerrero" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FBD6F084FBE1FC31" box="[1089,1138,940,966]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FB18F084FB6DFC31" box="[1167,1278,940,966]" pageId="13" pageNumber="14">Guerrero</collectingRegion>
|
||
,
|
||
<location id="8EB9D580047B1F42FA9AF084FAF0FC31" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FA9AF084FAF0FC31" box="[1293,1379,940,966]" country="Mexico" municipality="Guerrero" name="Tetipac" pageId="13" pageNumber="14" stateProvince="Guerrero">Tetipac</location>
|
||
/
|
||
<date id="FFD8A59B047B1F42FF2AF0E7FEDFFC1E" box="[189,332,975,1001]" pageId="13" pageNumber="14" value="1990-08-02">
|
||
<collectingDate id="EF9C5C73047B1F42FF2AF0E7FEDFFC1E" box="[189,332,975,1001]" pageId="13" pageNumber="14" value="1990-08-02">2 Aug. 1990</collectingDate>
|
||
</date>
|
||
/ excrement burro /
|
||
<collectorName id="2693E68D047B1F42FDA0F0E7FD6CFC1E" box="[567,767,975,1001]" pageId="13" pageNumber="14">Fco. E. Figueroa</collectorName>
|
||
”;
|
||
<collectionCode id="ED771B9E047B1F42FC82F0E7FCFBFC1E" box="[789,872,975,1001]" pageId="13" pageNumber="14">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B0E8906047B1F42FC10F0E7FABFFBFB" ID-GBIF-Occurrence="4164095336" collectingDate="1997-08" collectionCode="CEMT" collectorName="M. Lopez" elevation="1900" location="Estacion Cientifica Las Joyas" pageId="13" pageNumber="14" specimenCount="1" specimenCount-male="1" stateProvince="Jalisco" typeStatus="paratype">
|
||
<emphasis id="B9125F49047B1F42FC10F0E7FC4FFC1E" bold="true" box="[903,988,975,1001]" pageId="13" pageNumber="14">
|
||
<collectingRegion id="49A24DB9047B1F42FC10F0E7FC4FFC1E" box="[903,988,975,1001]" country="Mexico" name="Jalisco" pageId="13" pageNumber="14">Jalisco</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D6048D2047B1F42FC63F0E7FBB1FC1E" box="[1012,1058,975,1001]" pageId="13" pageNumber="14" type="male">1 ♂</specimenCount>
|
||
; “Jalisco,
|
||
<location id="8EB9D580047B1F42FB0EF0E7FEBCFBFA" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FB0EF0E7FEBCFBFA" country="Mexico" municipality="Jalisco" name="Estacion Cientifica Las Joyas" pageId="13" pageNumber="14" stateProvince="Jalisco">Estación Cientifica Las Joyas</location>
|
||
/
|
||
<elevation id="004B6468047B1F42FED3F0DAFE4AFBFB" box="[324,473,1010,1037]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="14" unit="m" value="1900.0">
|
||
<quantity id="4C9E2EBE047B1F42FED3F0DAFE0FFBFA" box="[324,412,1010,1037]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="14" unit="m" value="1900.0">1900 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
/
|
||
<date id="FFD8A59B047B1F42FE7BF0DAFDF3FBFB" box="[492,608,1010,1036]" pageId="13" pageNumber="14" value="1997-08">
|
||
<collectingDate id="EF9C5C73047B1F42FE7BF0DAFDF3FBFB" box="[492,608,1010,1036]" pageId="13" pageNumber="14" value="1997-08">Aug 1997</collectingDate>
|
||
</date>
|
||
/ Sancampús Reg. Pino, necrotrampa 8F /
|
||
<collectorName id="2693E68D047B1F42FBD9F0DBFB56FBFB" box="[1102,1221,1010,1037]" pageId="13" pageNumber="14">M. Lopez</collectorName>
|
||
”;
|
||
<collectionCode id="ED771B9E047B1F42FB4CF0DAFABFFBFB" box="[1243,1324,1010,1036]" pageId="13" pageNumber="14">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0E8906047B1F42FAD4F0DAFE4EFBA4" ID-GBIF-Occurrence="4164095340" collectingDate="1997-08" collectionCode="CEMT" collectorName="M. Lopez" elevation="1900" location="Estacion Cientifica Las Joyas" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Jalisco" typeStatus="paratype">
|
||
<specimenCount id="9D6048D2047B1F42FAD4F0DAFAFCFBFA" box="[1347,1391,1010,1037]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
|
||
; “
|
||
<collectingRegion id="49A24DB9047B1F42FF5FF73DFE8EFBD8" box="[200,285,1045,1071]" pageId="13" pageNumber="14">Jalisco</collectingRegion>
|
||
,
|
||
<location id="8EB9D580047B1F42FEBEF73DFD11FBC7" LSID="urn:lsid:plazi:treatment:03CF324D047D1F40FDF3F7EAFB67F9A0:8EB9D580047B1F42FEBEF73DFD11FBC7" box="[297,642,1045,1072]" country="Mexico" municipality="Jalisco" name="Estacion Cientifica Las Joyas" pageId="13" pageNumber="14" stateProvince="Jalisco">Estación Científica Las Joyas</location>
|
||
/
|
||
<elevation id="004B6468047B1F42FD0DF73DFCA1FBD8" box="[666,818,1045,1072]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="14" unit="m" value="1900.0">
|
||
<quantity id="4C9E2EBE047B1F42FD0DF73DFD60FBC7" box="[666,755,1045,1072]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="14" unit="m" value="1900.0">1900 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
/
|
||
<date id="FFD8A59B047B1F42FCD0F73DFC2FFBD8" box="[839,956,1045,1071]" pageId="13" pageNumber="14" value="1997-08">
|
||
<collectingDate id="EF9C5C73047B1F42FCD0F73DFC2FFBD8" box="[839,956,1045,1071]" pageId="13" pageNumber="14" value="1997-08">Aug 1997</collectingDate>
|
||
</date>
|
||
/ Sancampús Reg. Pino, necrotrampa 7D /
|
||
<collectorName id="2693E68D047B1F42FF6FF711FEFCFBA4" box="[248,367,1081,1107]" pageId="13" pageNumber="14">M. Lopez</collectorName>
|
||
”;
|
||
<collectionCode id="ED771B9E047B1F42FE11F711FE4AFBA4" box="[390,473,1081,1107]" pageId="13" pageNumber="14">CEMT</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37CD0D0047B1F41FF2AF752FCFAFE7B" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BD9835B047B1F42FF2AF752FE73FB63" blockId="13.[189,480,1146,1172]" box="[189,480,1146,1172]" pageId="13" pageNumber="14">
|
||
<heading id="D0913437047B1F42FF2AF752FE73FB63" bold="true" box="[189,480,1146,1172]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B9125F49047B1F42FF2AF752FE73FB63" bold="true" box="[189,480,1146,1172]" pageId="13" pageNumber="14">Description of the male</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF2AF781FAD1FB11" blockId="13.[189,1398,1192,1254]" pageId="13" pageNumber="14">
|
||
<smallCapsWord id="8D3F1587047B1F42FF2AF781FE8BFB36" baselines="1212,1212" box="[189,280,1193,1219]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="13" pageNumber="14">COLOUR</smallCapsWord>
|
||
. Body black with strong purple to greenish iridescence on pronotum and elytra; dorsally, tegument shining, head and pronotal disc silky; antennal segments brown, antennal club brownish.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF2AF626FCFCFA66" blockId="13.[187,1399,1293,1425]" pageId="13" pageNumber="14">
|
||
<smallCapsWord id="8D3F1587047B1F42FF2AF626FF6FFAD1" baselines="1313,1313" box="[189,252,1294,1320]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="13" pageNumber="14">HEAD</smallCapsWord>
|
||
. Clypeus obtusely triangular and elongated, with a medial, high triangular horn; from posterior side, horn wide and longitudinally sulcate, tip slightly to distinctly depressed at middle; head punctation very obsolete and superficial; genae squared, with elevated and acuminate angle; frons smooth, unpunctured, with two weak and oblique ridges along inner side of eyes.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF2AF691FD19F93E" blockId="13.[188,1400,1464,1737]" pageId="13" pageNumber="14">
|
||
<smallCapsWord id="8D3F1587047B1F42FF2AF691FE8AFA26" baselines="1484,1484" box="[189,281,1465,1491]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="13" pageNumber="14">THORAX</smallCapsWord>
|
||
. In lateral view, pronotum weakly convex; in dorsal view, posterior edge of pronotum without margin at middle, lateral edges weakly curved from posterior angles to midline, straight and convergent from midline to anterior angles; anterior angles obtuse; pronotal disc either unpunctured or with a few, shallow punctures across midline, sides with strong and coarse punctures sparsely distributed from posterior to anterior angles; in lateral view, anteromedial projection curved upward apically; in dorsal view, projection with sides parallel, tip widely emarginated at middle. Elytral interstriae slightly convex, very finely and densely punctate. Ventral side of thoracic sclerites (i.e., pro-, meso- and metasternum) with long, yellowish to brownish setae.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF2AF5D8FD4DF8A6" blockId="13.[189,1398,1776,1873]" pageId="13" pageNumber="14">
|
||
<smallCapsWord id="8D3F1587047B1F42FF2AF5D8FF65F8FF" baselines="1795,1795" box="[189,246,1776,1802]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Legs" pageId="13" pageNumber="14">LEGS</smallCapsWord>
|
||
. Protibiae robust, with six external teeth, ventral side with a longitudinal, crenulated carina; apical spur long and slender; meso- and metatibiae each with four entire, transversal carinae on posterior side; femora unmodified, with long yellowish setae.
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F42FF2AF45FFE23F866" blockId="13.[189,432,1911,1937]" box="[189,432,1911,1937]" pageId="13" pageNumber="14">
|
||
<heading id="D0913437047B1F42FF2AF45FFE23F866" bold="true" box="[189,432,1911,1937]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="B9125F49047B1F42FF2AF45FFE23F866" bold="true" box="[189,432,1911,1937]" pageId="13" pageNumber="14">Sexual dimorphism</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B047B1F41FF2AF48EFCFAFE7B" blockId="13.[189,1399,1958,2020]" lastBlockId="14.[188,1400,265,397]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Female specimens have the clypeal horn distinctly shorter and slightly depressed posteriorly; from above, the clypeal horn is triangular-shaped, apex almost reaching clypeal edge; anteromedial pronotal protuberance callus-like and transverse, depressed on superior side and close to anterior pronotal edge. Female specimens of
|
||
<taxonomicName id="4C66F8D804781F41FD90F205FD1CFEB1" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[519,655,300,327]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F4904781F41FD90F205FD1CFEB1" box="[519,655,300,327]" italics="true" pageId="14" pageNumber="15">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E23204781F41FD0FF205FD60FEB0" box="[664,755,301,327]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are very similar to those of
|
||
<taxonomicName id="4C66F8D804781F41FBD3F205FB22FEB1" baseAuthorityName="Jekel" baseAuthorityYear="1866" box="[1092,1201,301,327]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F4904781F41FBD3F205FB22FEB1" box="[1092,1201,301,327]" italics="true" pageId="14" pageNumber="15">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
, but they can be distinguished by the posterior edge of the pronotum not having a margin at the middle, and the presence of very fine and dense punctation on the elytral interstriae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF19D3D304781F41FF2AF4A0FD1CF81F" ID-DOI="http://doi.org/10.5281/zenodo.8177552" ID-Zenodo-Dep="8177552" httpUri="https://zenodo.org/record/8177552/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1928,1954]" targetBox="[209,1382,582,1879]" targetPageId="14">
|
||
<paragraph id="8BD9835B04781F41FF2AF4A0FD1CF81F" blockId="14.[189,1398,1928,2025]" pageId="14" pageNumber="15">
|
||
<emphasis id="B9125F4904781F41FF2AF4A0FE82F855" bold="true" box="[189,273,1928,1954]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
|
||
Type specimens and original labels.
|
||
<emphasis id="B9125F4904781F41FD71F4A0FC8EF855" bold="true" box="[742,797,1928,1954]" pageId="14" pageNumber="15">A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C66F8D804781F41FCA5F4A0FB47F854" authority="(Jekel, 1865)" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[818,1236,1928,1955]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F4904781F41FCA5F4A0FBB6F855" box="[818,1061,1928,1954]" italics="true" pageId="14" pageNumber="15">Ceratotrupes sturmi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF7FEAA04781F41FBADF4A0FB59F854" author="Jekel H." box="[1082,1226,1928,1955]" pageId="14" pageNumber="15" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, holotype, ♂ (NHML).
|
||
<emphasis id="B9125F4904781F41FEABF483FEC2F832" bold="true" box="[316,337,1963,1989]" pageId="14" pageNumber="15">A</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="B9125F4904781F41FDBBF483FDD3F832" bold="true" box="[556,576,1963,1989]" pageId="14" pageNumber="15">B</emphasis>
|
||
. Labels.
|
||
<emphasis id="B9125F4904781F41FD2FF483FD62F832" bold="true" box="[696,753,1963,1989]" pageId="14" pageNumber="15">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C66F8D804781F41FC91F484FB41F831" authority="(Jekel, 1865)" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[774,1234,1963,1990]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mniszechi">
|
||
<emphasis id="B9125F4904781F41FC91F484FBB0F832" box="[774,1059,1963,1990]" italics="true" pageId="14" pageNumber="15">Ceratotrupes mniszechi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF7FEAA04781F41FBAFF483FB5AF831" author="Jekel H." box="[1080,1225,1963,1990]" pageId="14" pageNumber="15" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, holotype, ♂ (MNHN).
|
||
<emphasis id="B9125F4904781F41FEAEF4E6FEDDF81F" bold="true" box="[313,334,1998,2024]" pageId="14" pageNumber="15">C</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="B9125F4904781F41FD81F4E6FDB8F81F" bold="true" box="[534,555,1998,2024]" pageId="14" pageNumber="15">D</emphasis>
|
||
. Labels.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF19D3D304791F40FF2AF7E8FC64FB2D" ID-DOI="http://doi.org/10.5281/zenodo.8177554" ID-Zenodo-Dep="8177554" box="[189,1015,1216,1243]" httpUri="https://zenodo.org/record/8177554/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1216,1242]" targetBox="[208,1383,278,1155]" targetPageId="15">
|
||
<paragraph id="8BD9835B04791F40FF2AF7E8FC64FB2D" blockId="15.[189,1015,1216,1243]" box="[189,1015,1216,1243]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9125F4904791F40FF2AF7E8FE98FB2D" bold="true" box="[189,267,1216,1242]" pageId="15" pageNumber="16">Fig. 6.</emphasis>
|
||
Geographic distribution of the genus
|
||
<taxonomicName id="4C66F8D804791F40FD5FF7E8FC63FB2C" authority="Jekel, 1865" authorityName="Jekel" authorityYear="1865" box="[712,1008,1216,1243]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9125F4904791F40FD5FF7E8FCF0FB2D" box="[712,867,1216,1242]" italics="true" pageId="15" pageNumber="16">Ceratotrupes</emphasis>
|
||
<bibRefCitation id="EFF7FEAA04791F40FCFDF7E8FC63FB2C" author="Jekel H." box="[874,1008,1216,1243]" pageId="15" pageNumber="16" pagination="513 - 618" refId="ref10191" refString="Jekel H. 1865. Essai sur la classification naturelle des Geotrupes Latreille et description d'especes nouvelles. Annales de la Societe entomologique de France, 4 e serie 5: 513 - 618. Available from https: // www. biodiversitylibrary. org / page / 14221920 [accessed 12 May 2023]." type="journal article" year="1865">Jekel, 1865</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37CD0D004791F40FF2AF630FB67F9A0" pageId="15" pageNumber="16" type="biology_ecology">
|
||
<paragraph id="8BD9835B04791F40FF2AF630FD9AFAC5" blockId="15.[189,521,1304,1330]" box="[189,521,1304,1330]" pageId="15" pageNumber="16">
|
||
<heading id="D091343704791F40FF2AF630FD9AFAC5" bold="true" box="[189,521,1304,1330]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="B9125F4904791F40FF2AF630FD9AFAC5" bold="true" box="[189,521,1304,1330]" pageId="15" pageNumber="16">Ecology and biogeography</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BD9835B04791F40FF2AF66EFB67F9A0" blockId="15.[187,1399,1350,1623]" pageId="15" pageNumber="16">
|
||
This species occurs in the Sierra Madre del Sur and it seems to be restricted to the surrounding mountains of Omiltemi,
|
||
<collectingRegion id="49A24DB904791F40FECAF642FE59FA73" box="[349,458,1386,1412]" country="Mexico" name="Guerrero" pageId="15" pageNumber="16">Guerrero</collectingRegion>
|
||
, where it occurs exclusively in pine forest. Many of the specimens historically identified as
|
||
<taxonomicName id="4C66F8D804791F40FEC3F6A5FE52FA50" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[340,449,1421,1447]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F4904791F40FEC3F6A5FE52FA50" box="[340,449,1421,1447]" italics="true" pageId="15" pageNumber="16">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
in natural history collections instead belong to this new species.
|
||
<bibRefCitation id="EFF7FEAA04791F40FB57F6A5FAE0FA50" author="Howden H. F." box="[1216,1395,1421,1447]" pageId="15" pageNumber="16" pagination="567 - 575" refId="ref9986" refString="Howden H. F. 1974. Additional records and descriptions of North and Central American Geotrupinae (Coleoptera, Scarabaeidae). Canadian Journal of Zoology 52 (5): 567 - 575. https: // doi. org / 10.1139 / z 74 - 073" type="journal article" year="1974">Howden (1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFF7FEAA04791F40FF2AF698FF6AFA3D" author="Howden H. F." box="[189,249,1456,1482]" pageId="15" pageNumber="16" pagination="95 - 108" refId="ref10098" refString="Howden H. F. 2003. Subfamilia Geotrupinae. In: Moron M. A. (ed.) Atlas de los Escarabajos de Mexico. Coleoptera: Lamellicornia Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 95 - 108. Argania, Barcelona." type="book chapter" year="2003">2003</bibRefCitation>
|
||
) noticed that specimens identified as
|
||
<taxonomicName id="4C66F8D804791F40FD3AF698FC8AFA3D" authorityName="Jekel" baseAuthorityName="Jekel" baseAuthorityYear="1865" box="[685,793,1456,1482]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F4904791F40FD3AF698FC8AFA3D" box="[685,793,1456,1482]" italics="true" pageId="15" pageNumber="16">C. sturmi</emphasis>
|
||
</taxonomicName>
|
||
showed some geographically restricted differences, such as shape of cephalic horn and genitalia in males, elytral punctation and posterior edge of pronotum with or without margin at middle. During this research we examined the specimens mentioned by Howden (Juquila and Juchatengo,
|
||
<collectingRegion id="49A24DB904791F40FDF4F532FD2DF9C3" box="[611,702,1562,1588]" country="Mexico" name="Oaxaca" pageId="15" pageNumber="16">Oaxaca</collectingRegion>
|
||
, and Omiltemi,
|
||
<collectingRegion id="49A24DB904791F40FC10F532FC66F9C3" box="[903,1013,1562,1588]" country="Mexico" name="Guerrero" pageId="15" pageNumber="16">Guerrero</collectingRegion>
|
||
; CMNC) and confirm that they belong to different species, namely
|
||
<taxonomicName id="4C66F8D804791F40FDC8F515FD75F9A0" authority="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini, 2023" authorityName="Arriaga-Jiménez & Zunino & Halffter & Escobar-Hernández & Rossini" authorityYear="2023" box="[607,742,1597,1623]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gonzaloi" status="sp. nov.">
|
||
<emphasis id="B9125F4904791F40FDC8F515FD75F9A0" box="[607,742,1597,1623]" italics="true" pageId="15" pageNumber="16">C. gonzaloi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A221E23204791F40FD79F515FCDBF9A0" box="[750,840,1597,1623]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(Omiltemi) and
|
||
<taxonomicName id="4C66F8D804791F40FB9BF515FB63F9A0" authority="(Oaxaca)" authorityName="Jekel" baseAuthorityName="Oaxaca" baseAuthorityYear="1865" box="[1036,1264,1597,1623]" class="Insecta" family="Geotrupidae" genus="Ceratotrupes" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="sturmi">
|
||
<emphasis id="B9125F4904791F40FB9BF515FBEAF9A0" box="[1036,1145,1597,1623]" italics="true" pageId="15" pageNumber="16">C. sturmi</emphasis>
|
||
(
|
||
<collectingRegion id="49A24DB904791F40FB1EF515FB7BF9A0" box="[1161,1256,1597,1623]" country="Mexico" name="Oaxaca" pageId="15" pageNumber="16">Oaxaca</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |