318 lines
36 KiB
XML
318 lines
36 KiB
XML
<document id="5FE9754896534F858F27F0D9C59BD672" ID-DOI="10.4081/nhs.2017.310" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1721184724886" checkinUser="felipe" docAuthor="Pasini, Giovanni & Garassino, Alessandro" docDate="2017" docId="03CD830F7B176B7EFFE0FF1185EBA0D7" docLanguage="en" docName="NatHistSci.4.1.43-72.pdf" docOrigin="Natural History Sciences 4 (1)" docSource="http://dx.doi.org/10.4081/nhs.2017.310" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Ranina berglundi Squires & Demetrion 1992" docType="treatment" docVersion="1" lastPageNumber="47" masterDocId="FFF4FB777B146B7AFF91FF998441A166" masterDocTitle="A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks" masterLastPageNumber="72" masterPageNumber="43" pageNumber="46" updateTime="1726747683517" updateUser="guilherme" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="1B4C2DFA243DE056A3C7E05BE606640C">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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<mods:namePart id="7BF692BB20498C2EDD5C81F01C8DE7F3">Pasini, Giovanni</mods:namePart>
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<mods:namePart id="6819C0B4A9A6B13148F144F1F04AFECE">Garassino, Alessandro</mods:namePart>
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<mods:title id="CE1692C99874CEFCA020235AB02DF445">Natural History Sciences</mods:title>
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<mods:date id="43ACF61938A05414370DF57F6A282DC7">2017</mods:date>
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<treatment id="03CD830F7B176B7EFFE0FF1185EBA0D7" LSID="urn:lsid:plazi:treatment:03CD830F7B176B7EFFE0FF1185EBA0D7" httpUri="http://treatment.plazi.org/id/03CD830F7B176B7EFFE0FF1185EBA0D7" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">
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<subSubSection id="C37E61927B176B79FFE0FF11863FA1C4" box="[113,638,135,162]" pageId="3" pageNumber="46" type="nomenclature">
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<paragraph id="8BDB32197B176B79FFE0FF11863FA1C4" blockId="3.[113,638,135,192]" box="[113,638,135,162]" pageId="3" pageNumber="46">
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?
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<taxonomicName id="4C64499A7B176B79FFEFFF11863FA1C4" ID-CoL="BDDGD" authority="Squires & Demetrion, 1992" authorityName="Squires & Demetrion" authorityPageNumber="43" authorityYear="1992" box="[126,638,135,162]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="berglundi">
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<emphasis id="B910EE0B7B176B79FFEFFF118503A1C7" box="[126,322,135,161]" italics="true" pageId="3" pageNumber="46">Ranina berglundi</emphasis>
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<bibRefCitation id="EFF54FE87B176B79FED8FF1E863FA1C4" author="Squires R. L. & Demetrion R. A." box="[329,638,135,162]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion, 1992</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<subSubSection id="C37E61927B176B79FFE0FF3F84F2A1A6" box="[113,179,166,192]" pageId="3" pageNumber="46" type="description">
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<paragraph id="8BDB32197B176B79FFE0FF3F84F2A1A6" blockId="3.[113,638,135,192]" box="[113,179,166,192]" pageId="3" pageNumber="46">
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<figureCitation id="135F2E9C7B176B79FFE0FF3F84F2A1A6" box="[113,179,166,192]" captionStart="Fig" captionStartId="3.[113,146,1918,1940]" captionTargetBox="[113,756,744,1894]" captionTargetId="figure-657@3.[113,756,744,1355]" captionTargetPageId="3" captionText="Fig. 2 -?Ranina berglundi Squires & Demetrion, 1992. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5)." figureDoi="http://doi.org/10.5281/zenodo.12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="3" pageNumber="46">Fig. 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E61927B176B79FFE0FF7A8650A0A9" pageId="3" pageNumber="46" type="reference_group">
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<paragraph id="8BDB32197B176B79FFE0FF7A8576A07C" blockId="3.[113,755,226,282]" pageId="3" pageNumber="46">
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<heading id="D09385757B176B79FFE0FF7A8576A07C" centered="true" fontSize="10" level="2" pageId="3" pageNumber="46" reason="3">
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<treatmentCitation id="0AC514087B176B79FFE0FF7A86FEA19A" author="Squires R. L. & Demetrion R. A." box="[113,703,226,252]" page="43" pageId="3" pageNumber="46" year="1992">
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<taxonomicName id="4C64499A7B176B79FFE0FF7A86FEA19A" ID-CoL="BDDGD" authority="Squires & Demetrion, 1992: 43" authorityName="Squires & Demetrion" authorityPageNumber="43" authorityYear="1992" box="[113,703,226,252]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="berglundi">
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<emphasis id="B910EE0B7B176B79FFE0FF7A857AA19A" box="[113,315,226,252]" italics="true" pageId="3" pageNumber="46">Ranina berglundi</emphasis>
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<bibRefCitation id="EFF54FE87B176B79FED6FF7B86FEA19A" author="Squires R. L. & Demetrion R. A." box="[327,703,226,252]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion, 1992: 43</bibRefCitation>
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</taxonomicName>
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, 44, figs. 128, 129.
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</heading>
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</paragraph>
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<paragraph id="8BDB32197B176B79FFE0FEA48650A0A9" blockId="3.[113,756,316,463]" pageId="3" pageNumber="46">
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<taxonomicName id="4C64499A7B176B79FFE0FEA4857BA030" ID-CoL="BDDGD" authorityName="Squires & Demetrion" authorityYear="1992" box="[113,314,316,342]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="berglundi">
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||
<emphasis id="B910EE0B7B176B79FFE0FEA4857BA030" box="[113,314,316,342]" italics="true" pageId="3" pageNumber="46">Ranina berglundi</emphasis>
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</taxonomicName>
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||
-
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<treatmentCitation id="0AC514087B176B79FECBFEA586D3A030" author="Schweitzer C. E. & Feldmann R. M. & Gonzalez-Barba G. & Cosovic V." box="[346,658,316,342]" page="24" pageId="3" pageNumber="46" year="2006">
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<bibRefCitation id="EFF54FE87B176B79FECBFEA586D3A030" author="Schweitzer C. E. & Feldmann R. M. & Gonzalez-Barba G. & Cosovic V." box="[346,658,316,342]" pageId="3" pageNumber="46" pagination="26 - 45" refId="ref25008" refString="Schweitzer C. E., Feldmann R. M., Gonzalez-Barba G. & Cosovic V., 2006 - New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico. Bulletin of the Mizunami Fossil Museum, 33: 26 - 45." type="journal article" year="2006">
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Schweitzer
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<emphasis id="B910EE0B7B176B79FE70FEA4865FA030" box="[481,542,316,342]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
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2006: 24
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||
</bibRefCitation>
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</treatmentCitation>
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||
, 27, 28, figs. 2.5, 2.6. –
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||
<treatmentCitation id="0AC514087B176B79FEC1FEC2867EA013" author="Vega F. J. & Nyborg T. & Coutino M. A. & Hernandez-Monzon O." box="[336,575,346,373]" page="54" pageId="3" pageNumber="46" year="2008">
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<bibRefCitation id="EFF54FE87B176B79FEC1FEC2867EA013" author="Vega F. J. & Nyborg T. & Coutino M. A. & Hernandez-Monzon O." box="[336,575,346,373]" pageId="3" pageNumber="46" pagination="51 - 71" refId="ref25274" refString="Vega F. J., Nyborg T., Coutino M. A. & Hernandez-Monzon O., 2008 - Review and additions to the Eocene decapod Crustacea from Chiapas, Mexico. Bulletin of the Mizunami Fossil Museum, 34: 51 - 71." type="journal article" year="2008">
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||
Vega
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||
<emphasis id="B910EE0B7B176B79FE03FEC2858CA012" box="[402,461,346,372]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
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||
2008: 54
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
, Pl. 1, figs. 13- 15, Pl. 2, figs. 1, 2. –
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||
<treatmentCitation id="0AC514087B176B79FE18FEE186F3A0F5" author="Schweitzer C. & Feldmann R. M. & Garassino A. & Karasawa H. & Schweigert G." box="[393,690,376,403]" page="74" pageId="3" pageNumber="46" year="2010">
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<bibRefCitation id="EFF54FE87B176B79FE18FEE186F3A0F5" author="Schweitzer C. & Feldmann R. M. & Garassino A. & Karasawa H. & Schweigert G." box="[393,690,376,403]" pageId="3" pageNumber="46" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. & Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
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||
Schweitzer
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||
<emphasis id="B910EE0B7B176B79FD9CFEE08604A0F4" box="[525,581,376,402]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
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||
2010: 74
|
||
</bibRefCitation>
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||
</treatmentCitation>
|
||
. –
|
||
<treatmentCitation id="0AC514087B176B79FD42FEE08592A0D7" author="De Angeli A. & Beschin C." page="13" pageId="3" pageNumber="46" year="2011">
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<bibRefCitation id="EFF54FE87B176B79FD42FEE08592A0D7" author="De Angeli A. & Beschin C." pageId="3" pageNumber="46" pagination="11 - 20" refId="ref23611" refString="De Angeli A. & Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico " G. Zannato ", 18: 11 - 20." type="book chapter" year="2011">De Angeli & Beschin 2011: 13</bibRefCitation>
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</treatmentCitation>
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||
. –
|
||
<treatmentCitation id="0AC514087B176B79FE64FE0E8485A0A9" author="Van Bakel W. M. & Guinot D. & Artal P. & Fraaije R. H. B. & Jagt J. W. M." page="208" pageId="3" pageNumber="46" year="2012">
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<bibRefCitation id="EFF54FE87B176B79FE64FE0E8485A0A9" author="Van Bakel W. M. & Guinot D. & Artal P. & Fraaije R. H. B. & Jagt J. W. M." pageId="3" pageNumber="46" pagination="1 - 216" refId="ref25151" refString="Van Bakel W. M., Guinot D., Artal P., Fraaije R. H. B. & Jagt J. W. M., 2012 - A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata). Zootaxa, 3215: 1 - 216." type="journal article" year="2012">
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||
Van Bakel
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||
<emphasis id="B910EE0B7B176B79FDE3FE0E86EDA0D7" box="[626,684,407,433]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
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||
2012: 208
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||
</bibRefCitation>
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||
</treatmentCitation>
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||
. –
|
||
<treatmentCitation id="0AC514087B176B79FF75FE2C864CA0A9" author="Karasawa H. & Schweitzer C. E. & Feldmann R. M. & Luque J." box="[228,525,437,463]" page="260" pageId="3" pageNumber="46" year="2014">
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<bibRefCitation id="EFF54FE87B176B79FF75FE2C864CA0A9" author="Karasawa H. & Schweitzer C. E. & Feldmann R. M. & Luque J." box="[228,525,437,463]" pageId="3" pageNumber="46" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. & Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
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||
Karasawa
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||
<emphasis id="B910EE0B7B176B79FECBFE2F85D3A0A9" box="[346,402,437,463]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
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2014: 260
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||
</bibRefCitation>
|
||
</treatmentCitation>
|
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.
|
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</treatmentCitationGroup>
|
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</heading>
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E61927B176B79FF06FE68803AA05F" pageId="3" pageNumber="46" type="materials_examined">
|
||
<paragraph id="8BDB32197B176B79FF06FE6884F0A34C" blockId="3.[113,756,497,705]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FF06FE688501A36D" bold="true" box="[151,320,497,523]" pageId="3" pageNumber="46">Type material</emphasis>
|
||
:
|
||
<typeStatus id="54DF8CBB7B176B79FEC0FE6885F7A36D" box="[337,438,497,523]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
,
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||
<collectionCode id="ED75AADC7B176B79FE55FE6885BBA36D" box="[452,506,497,523]" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="3" pageNumber="46">IGM</collectionCode>
|
||
5913;
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||
<typeStatus id="54DF8CBB7B176B79FDDCFE6B86F1A36A" box="[589,688,498,524]" pageId="3" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
,
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||
<collectionCode id="ED75AADC7B176B79FD2CFE6886B2A36D" box="[701,755,497,523]" country="Mongolia" httpUri="http://grbio.org/cool/0h5a-qjh4" name="Geological Institute, Mongolian Academy of Sciences" pageId="3" pageNumber="46">IGM</collectionCode>
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||
5914.
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||
</paragraph>
|
||
<paragraph id="8BDB32197B176B79FF06FDB485B4A303" blockId="3.[113,756,497,705]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FF06FDB48570A321" bold="true" box="[151,305,557,583]" pageId="3" pageNumber="46">Type locality</emphasis>
|
||
: locality
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||
<collectionCode id="ED75AADC7B176B79FE0FFDB485A8A321" box="[414,489,557,583]" country="USA" httpUri="http://grbio.org/cool/sgeh-wrgd" name="California State University, Northridge" pageId="3" pageNumber="46" type="University or college">CSUN</collectionCode>
|
||
1220b;
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||
<collectingRegion id="49A0FCFB7B176B79FDD6FDB484D9A300" country="Mexico" name="Baja California Sur" pageId="3" pageNumber="46">Baja California Sur</collectingRegion>
|
||
(
|
||
<collectingCountry id="F37372897B176B79FF36FDD28543A300" box="[167,258,587,614]" name="Mexico" pageId="3" pageNumber="46">Mexico</collectingCountry>
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||
), Bateque Formation.
|
||
</paragraph>
|
||
<paragraph id="8BDB32197B176B79FF06FDF085ADA3E2" blockId="3.[113,756,497,705]" box="[151,492,617,644]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FF06FDF08505A3E2" bold="true" box="[151,324,617,644]" pageId="3" pageNumber="46">Geological age</emphasis>
|
||
: early Eocene.
|
||
</paragraph>
|
||
<paragraph id="8BDB32197B176B79FF06FD1E803AA05F" blockId="3.[113,756,497,705]" lastBlockId="3.[832,1475,135,1970]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FF06FD1E85C7A3C7" bold="true" box="[151,390,647,673]" pageId="3" pageNumber="46">Examined material</emphasis>
|
||
:
|
||
<typeStatus id="54DF8CBB7B176B79FE0DFD1185BEA3C4" box="[412,511,648,674]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54DF8CBB7B176B79FDD5FD1186E8A3C4" box="[580,681,648,674]" pageId="3" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
;
|
||
<specimenCount id="9D62F9907B176B79FD2AFD1184A6A3A6" count="3" pageId="3" pageNumber="46" type="generic">three specimens</specimenCount>
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||
(IHNFG-3013-3015) from the early Eocene (El Bosque Formation), locality El Veinte (
|
||
<collectingRegion id="49A0FCFB7B176B79FACFFF1E81FFA1C7" box="[1374,1470,135,161]" country="Mexico" name="Chiapas" pageId="3" pageNumber="46">Chiapas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37372897B176B79FCD1FF3C87DAA1A6" box="[832,923,165,192]" name="Mexico" pageId="3" pageNumber="46">Mexico</collectingCountry>
|
||
), reported by
|
||
<bibRefCitation id="EFF54FE87B176B79FBAEFF3F8154A1A6" author="Vega F. J. & Nyborg T. & Coutino M. A. & Hernandez-Monzon O." box="[1087,1301,165,192]" pageId="3" pageNumber="46" pagination="51 - 71" refId="ref25274" refString="Vega F. J., Nyborg T., Coutino M. A. & Hernandez-Monzon O., 2008 - Review and additions to the Eocene decapod Crustacea from Chiapas, Mexico. Bulletin of the Mizunami Fossil Museum, 34: 51 - 71." type="journal article" year="2008">
|
||
Vega
|
||
<emphasis id="B910EE0B7B176B79FB10FF3F80FCA1D9" box="[1153,1213,165,191]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
;
|
||
<specimenCount id="9D62F9907B176B79FAB5FF3F8183A1D9" box="[1316,1474,165,192]" count="1" pageId="3" pageNumber="46" type="generic">one specimen</specimenCount>
|
||
(
|
||
<collectionCode id="ED75AADC7B176B79FCD6FF5D87B9A1B8" box="[839,1016,196,222]" pageId="3" pageNumber="46">MHN-UABCS</collectionCode>
|
||
/Ba7-3) from the middle Eocene (Bateque Formation), locality Waypoint 70 (
|
||
<geoCoordinate id="EE5054DE7B176B79FAB6FF7B81FCA19B" box="[1319,1469,226,253]" degrees="26" direction="north" minutes="45.918" orientation="latitude" pageId="3" pageNumber="46" precision="1" value="26.7653">26°45.918’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE5054DE7B176B79FCD1FE99879BA07D" box="[832,986,256,283]" degrees="113" direction="west" minutes="0.953" orientation="longitude" pageId="3" pageNumber="46" precision="1" value="-113.015884">113°0.953’W</geoCoordinate>
|
||
) (
|
||
<collectingRegion id="49A0FCFB7B176B79FC64FE9980A1A07C" box="[1013,1248,256,282]" country="Mexico" name="Baja California Sur" pageId="3" pageNumber="46">Baja California Sur</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37372897B176B79FB61FE99810AA07D" box="[1264,1355,256,283]" name="Mexico" pageId="3" pageNumber="46">Mexico</collectingCountry>
|
||
), reported by
|
||
<bibRefCitation id="EFF54FE87B176B79FCF2FE878036A05F" author="Schweitzer C. E. & Feldmann R. M. & Gonzalez-Barba G. & Cosovic V." box="[867,1143,286,313]" pageId="3" pageNumber="46" pagination="26 - 45" refId="ref25008" refString="Schweitzer C. E., Feldmann R. M., Gonzalez-Barba G. & Cosovic V., 2006 - New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico. Bulletin of the Mizunami Fossil Museum, 33: 26 - 45." type="journal article" year="2006">
|
||
Schweitzer
|
||
<emphasis id="B910EE0B7B176B79FC79FE868060A05E" box="[1000,1057,286,312]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E61927B176B79FCF4FEA5800AA5A6" pageId="3" pageNumber="46" type="description">
|
||
<paragraph id="8BDB32197B176B79FCF4FEA5800AA5A6" blockId="3.[832,1475,135,1970]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FCF4FEA58137A030" bold="true" box="[869,1398,316,342]" pageId="3" pageNumber="46">
|
||
Description by
|
||
<bibRefCitation id="EFF54FE87B176B79FB8EFEA58137A030" author="Squires R. L. & Demetrion R. A." box="[1055,1398,316,342]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion (1992)</bibRefCitation>
|
||
</emphasis>
|
||
: “
|
||
<emphasis id="B910EE0B7B176B79FA02FEA4807EA5A6" italics="true" pageId="3" pageNumber="46">
|
||
Moderate-sized raninid with ovate, moderately convex carapace, very broad in proportion to its length; widest along anterior one-fifth at outermost anterolateral spine area; carapace maximum width (including outermost anterolateral spines) equals total length; carapace width (excluding outermost anterolateral spines) 80 percent of total length; lateral borders curve inward, posterior margin very narrow; surface with small pits, sparse on anterior middle part, closer posteriorly and toward lateral and anterior borders; posterolaterally, pits in transverse rows of about 10; rostrum trifid, small, and pointed; fronto-orbital margin curved with three spines on each side; first (innermost) fronto-orbital spine small, outward-pointing, and separated from wider second spine by a short furrow; second fronto-orbital spine separated from forward-pointing and more prominent third spine by a very short and narrow furrow; length of fronto-orbital area about 60 percent of width of carapace (excluding outermost anterolateral spines); two anterolateral spines, equally spaced; first anterolateral spine forward-pointing, about equal in size to outermost fronto-orbital spine, but more pointed; second (outermost) anterolateral spine widest and strongest of all spines, extended at a 45-degree angle to a blunt point, with four serrations on anterior edge of spine (the middle two the strongest); posteriorly to second anterolateral spine, a fine raised rim extends around the carapace;
|
||
<typeStatus id="54DF8CBB7B176B79FAF1FBF38183A5E2" box="[1376,1474,1130,1156]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
<quantity id="4C9C9FFC7B176B79FCD1FB1187CFA5C4" box="[832,910,1160,1186]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="46" unit="mm" value="39.0">39 mm</quantity>
|
||
in width (including outermost anterolateral spines),
|
||
<quantity id="4C9C9FFC7B176B79FCEFFB3F878AA5A6" box="[894,971,1190,1216]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="46" unit="mm" value="38.0">38 mm</quantity>
|
||
in length.
|
||
</emphasis>
|
||
”
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E61927B176B7EFCF4FB5D85EBA0D7" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46" type="discussion">
|
||
<paragraph id="8BDB32197B176B79FCF4FB5D81DCA610" blockId="3.[832,1475,135,1970]" pageId="3" pageNumber="46">
|
||
<emphasis id="B910EE0B7B176B79FCF4FB5D87A5A5B8" bold="true" box="[869,996,1220,1246]" pageId="3" pageNumber="46">Discussion</emphasis>
|
||
. The type material described by
|
||
<bibRefCitation id="EFF54FE87B176B79FAFCFB5D806FA59A" author="Squires R. L. & Demetrion R. A." pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion (1992)</bibRefCitation>
|
||
has trifid rostrum, first anterolateral spine single and second anterolateral spine, strongly directed outward, with four spinules on anterior edge.
|
||
<bibRefCitation id="EFF54FE87B176B79FCD1FAA5800EA431" author="Schweitzer C. E. & Feldmann R. M. & Gonzalez-Barba G. & Cosovic V." box="[832,1103,1340,1367]" pageId="3" pageNumber="46" pagination="26 - 45" refId="ref25008" refString="Schweitzer C. E., Feldmann R. M., Gonzalez-Barba G. & Cosovic V., 2006 - New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico. Bulletin of the Mizunami Fossil Museum, 33: 26 - 45." type="journal article" year="2006">
|
||
Schweitzer
|
||
<emphasis id="B910EE0B7B176B79FC55FAA487BDA430" box="[964,1020,1340,1366]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
assigned
|
||
<specimenCount id="9D62F9907B176B79FB2CFAA5811AA430" box="[1213,1371,1340,1366]" count="6" pageId="3" pageNumber="46" type="generic">six specimens</specimenCount>
|
||
from the Bateque Formation (
|
||
<collectingRegion id="49A0FCFB7B176B79FBB8FAC2814AA413" box="[1065,1291,1371,1397]" country="Mexico" name="Baja California Sur" pageId="3" pageNumber="46">Baja California Sur</collectingRegion>
|
||
) to this species, without adding additional information appreciably to the description of the carapace made by
|
||
<bibRefCitation id="EFF54FE87B176B79FB64FA0E8788A4A9" author="Squires R. L. & Demetrion R. A." pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion (1992)</bibRefCitation>
|
||
. Later
|
||
<bibRefCitation id="EFF54FE87B176B79FB84FA2F809CA4A9" author="Vega F. J. & Nyborg T. & Coutino M. A. & Hernandez-Monzon O." box="[1045,1245,1461,1488]" pageId="3" pageNumber="46" pagination="51 - 71" refId="ref25274" refString="Vega F. J., Nyborg T., Coutino M. A. & Hernandez-Monzon O., 2008 - Review and additions to the Eocene decapod Crustacea from Chiapas, Mexico. Bulletin of the Mizunami Fossil Museum, 34: 51 - 71." type="journal article" year="2008">
|
||
Vega
|
||
<emphasis id="B910EE0B7B176B79FBC5FA2F80CAA4A9" box="[1108,1163,1461,1487]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
(2008)
|
||
</bibRefCitation>
|
||
assigned tentatively
|
||
<specimenCount id="9D62F9907B176B79FCD1FA4A87B6A48B" box="[832,1015,1491,1517]" count="3" pageId="3" pageNumber="46" type="generic">three specimens</specimenCount>
|
||
from
|
||
<collectingRegion id="49A0FCFB7B176B79FBD0FA4A80DDA48B" box="[1089,1180,1491,1517]" country="Mexico" name="Chiapas" pageId="3" pageNumber="46">Chiapas</collectingRegion>
|
||
(
|
||
<collectingCountry id="F37372897B176B79FB3CFA4A8146A488" box="[1197,1287,1491,1518]" name="Mexico" pageId="3" pageNumber="46">Mexico</collectingCountry>
|
||
) to this species, pointing out, as difference from the type material, the first anterolateral spine bifid, small size, and distribution of the carapace dorsal pits. However, the review of the type material described by
|
||
<bibRefCitation id="EFF54FE87B176B79FBE8F9D58183A700" author="Squires R. L. & Demetrion R. A." box="[1145,1474,1612,1638]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion (1992)</bibRefCitation>
|
||
allow us to establish that the first anterolateral spine is also bifid, well preserved in the left side of the
|
||
<typeStatus id="54DF8CBB7B176B79FACEF9118183A7C4" box="[1375,1474,1672,1698]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
and slightly broken in the
|
||
<typeStatus id="54DF8CBB7B176B79FBF5F93E8089A7A7" box="[1124,1224,1703,1729]" pageId="3" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
, attesting that the specimens from
|
||
<collectingRegion id="49A0FCFB7B176B79FC47F95D80F5A7B8" box="[982,1204,1732,1758]" country="Mexico" name="Baja California Sur" pageId="3" pageNumber="46">Baja California Sur</collectingRegion>
|
||
(
|
||
<typeStatus id="54DF8CBB7B176B79FB52F95D8169A7B8" box="[1219,1320,1732,1758]" pageId="3" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
and
|
||
<typeStatus id="54DF8CBB7B176B79FAF0F95C8183A7B9" box="[1377,1474,1733,1759]" pageId="3" pageNumber="46" type="paratype">paratype</typeStatus>
|
||
described by
|
||
<bibRefCitation id="EFF54FE87B176B79FC4CF97B8160A79A" author="Squires R. L. & Demetrion R. A." box="[989,1313,1762,1788]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion, 1992</bibRefCitation>
|
||
) and
|
||
<collectingRegion id="49A0FCFB7B176B79FAF6F97B8183A79A" box="[1383,1474,1762,1788]" country="Mexico" name="Chiapas" pageId="3" pageNumber="46">Chiapas</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EFF54FE87B176B79FCD6F8998014A67D" author="Schweitzer C. E. & Feldmann R. M. & Gonzalez-Barba G. & Cosovic V." box="[839,1109,1792,1819]" pageId="3" pageNumber="46" pagination="26 - 45" refId="ref25008" refString="Schweitzer C. E., Feldmann R. M., Gonzalez-Barba G. & Cosovic V., 2006 - New Decapoda (Anomura, Brachyura) from the Eocene Bateque and Tepetate Formations, Baja California Sur, Mexico. Bulletin of the Mizunami Fossil Museum, 33: 26 - 45." type="journal article" year="2006">
|
||
Schweitzer
|
||
<emphasis id="B910EE0B7B176B79FC41F898804AA67C" box="[976,1035,1792,1818]" italics="true" pageId="3" pageNumber="46">et al.</emphasis>
|
||
, 2006
|
||
</bibRefCitation>
|
||
) could belong to the same species, though the smaller size of
|
||
<collectingRegion id="49A0FCFB7B176B79FB35F88680BEA65F" box="[1188,1279,1823,1849]" country="Mexico" name="Chiapas" pageId="3" pageNumber="46">Chiapas</collectingRegion>
|
||
specimens could be a diagnostic characters to distinguish these specimens from the type material (F.J. Vega pers. comm., 2016).
|
||
</paragraph>
|
||
<caption id="DF1B62917B176B79FFE0F8E7867AA6C9" ID-DOI="http://doi.org/10.5281/zenodo.12753809" ID-Zenodo-Dep="12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="3" pageNumber="46" startId="3.[113,146,1918,1940]" targetBox="[113,756,744,1894]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BDB32197B176B79FFE0F8E7867AA6C9" blockId="3.[113,757,1918,1967]" pageId="3" pageNumber="46">
|
||
Fig. 2 -?
|
||
<taxonomicName id="4C64499A7B176B79FF5BF8E68637A6F2" authority="Squires & Demetrion, 1992" authorityName="Squires & Demetrion" authorityYear="1992" box="[202,630,1918,1940]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="berglundi">
|
||
<emphasis id="B910EE0B7B176B79FF5BF8E6852BA6F5" box="[202,362,1918,1940]" italics="true" pageId="3" pageNumber="46">Ranina berglundi</emphasis>
|
||
<bibRefCitation id="EFF54FE87B176B79FEE3F8E78637A6F2" author="Squires R. L. & Demetrion R. A." box="[370,630,1918,1940]" pageId="3" pageNumber="46" pagination="1 - 55" refId="ref25064" refString="Squires R. L. & Demetrion R. A., 1992 - Paleontology of the Eocene Bateque Formation, Baja California Sur, Mexico. Los Angeles County Museum, Contributions in Science, 434: 1 - 55." type="journal article" year="1992">Squires & Demetrion, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB32197B176B7EFCF4F8E08514A19A" blockId="3.[832,1475,135,1970]" lastBlockId="4.[113,756,135,433]" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">
|
||
Based upon the main morphological characters of
|
||
<taxonomicName id="4C64499A7B176B79FA3BF8E387EDA6D7" authorityName="Squires & Demetrion" authorityYear="1992" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="berglundi">
|
||
<emphasis id="B910EE0B7B176B79FA3BF8E387EDA6D7" italics="true" pageId="3" pageNumber="46">R. berglundi</emphasis>
|
||
</taxonomicName>
|
||
, such as the trifid rostrum and the peculiar shape and arrangement of the anterolateral spines (above all the second one), we can attest that these characters do not fit those of
|
||
<taxonomicName id="4C64499A7B106B7EFF7FFF5C857EA1B8" authorityName="Lamark" authorityYear="1810" box="[238,319,197,222]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910EE0B7B106B7EFF7FFF5C857EA1B8" box="[238,319,197,222]" italics="true" pageId="4" pageNumber="47">Ranina</emphasis>
|
||
</taxonomicName>
|
||
, questioning the placement of
|
||
<taxonomicName id="4C64499A7B106B7EFD04FF5D84C6A19A" authorityName="Squires & Demetrion" authorityYear="1992" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="berglundi">
|
||
<emphasis id="B910EE0B7B106B7EFD04FF5D84C6A19A" italics="true" pageId="4" pageNumber="47">berglundi</emphasis>
|
||
</taxonomicName>
|
||
within this genus.
|
||
</paragraph>
|
||
<paragraph id="8BDB32197B106B7EFF06FE9985EBA0D7" blockId="4.[113,756,135,433]" pageId="4" pageNumber="47">
|
||
Indeed
|
||
<taxonomicName id="4C64499A7B106B7EFF7AFE988536A07C" authorityName="Squires & Demetrion" authorityYear="1992" box="[235,375,256,282]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="berglundi">
|
||
<emphasis id="B910EE0B7B106B7EFF7AFE988536A07C" box="[235,375,256,282]" italics="true" pageId="4" pageNumber="47">R. berglundi</emphasis>
|
||
</taxonomicName>
|
||
from the early Eocene of Mexico and the rich sample of specimens from the late Oligocene (Rupelian) of the Ligure-Piemontese Basin (
|
||
<collectionCode id="ED75AADC7B106B7EFD10FEA586F0A030" box="[641,689,316,342]" pageId="4" pageNumber="47">NW</collectionCode>
|
||
<collectingCountry id="F37372897B106B7EFD26FEA586ADA031" box="[695,748,316,343]" name="Italy" pageId="4" pageNumber="47">Italy</collectingCountry>
|
||
) ascribed to
|
||
<taxonomicName id="4C64499A7B106B7EFF63FEC28530A012" baseAuthorityName="von Munster" baseAuthorityYear="1840" box="[242,369,347,372]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="speciosa">
|
||
<emphasis id="B910EE0B7B106B7EFF63FEC28530A012" box="[242,369,347,372]" italics="true" pageId="4" pageNumber="47">R. speciosa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFF54FE87B106B7EFE0BFEC38614A013" author="Allasinaz A." box="[410,597,346,373]" pageId="4" pageNumber="47" pagination="509 - 566" refId="ref23326" refString="Allasinaz A., 1987 - Brachyura Decapoda oligocenici (Rupeliano) del Bacino Ligure Piemontese. Bollettino del Museo regionale di Scienze naturali di Torino, 5 (2): 509 - 566." type="journal article" year="1987">Allasinaz (1987)</bibRefCitation>
|
||
have been assigned to a new genus within the Raninindae
|
||
<emphasis id="B910EE0B7B106B7EFDF9FEE086B5A0F4" box="[616,756,377,402]" italics="true" pageId="4" pageNumber="47">sensu stricto</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF54FE87B106B7EFFE8FE0E85DFA0D7" author="Pasini G. & Garassino A." box="[121,414,407,433]" pageId="4" pageNumber="47" pagination="87 - 96" refId="ref24487" refString="Pasini G. & Garassino A., 2017 - A new raninoid crab (Crustacea, Decapoda, Raninidae) from the early Oligocene (late Rupelian) of Italy: systematics and palaeogeographic evidences. Natural History Sciences Atti della Societa italiana di Scienze naturali e del Museo civico di Storia naturale in Milano, 4 (1): 87 - 96. doi: 10.4081 / nhs. 2017.323" type="journal article" year="2017">Pasini & Garassino, 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |