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<document id="01BC15AE40C836C540926AAE0E348BD8" ID-DOI="10.4081/nhs.2017.310" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753805" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch,operationResults" IM.taxonomicNames_approvedBy="guilherme" IM.treatmentCitations_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1721184724886" checkinUser="felipe" docAuthor="Pasini, Giovanni &amp; Garassino, Alessandro" docDate="2017" docId="03CD830F7B156B78FCD1FDF08183A6D5" docLanguage="en" docName="NatHistSci.4.1.43-72.pdf" docOrigin="Natural History Sciences 4 (1)" docSource="http://dx.doi.org/10.4081/nhs.2017.310" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Ranina americana Withers 1924" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="FFF4FB777B146B7AFF91FF998441A166" masterDocTitle="A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks" masterLastPageNumber="72" masterPageNumber="43" pageNumber="44" updateTime="1726747683517" updateUser="guilherme" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="D2D032D52E4FED559F680B5F87B74F82">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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<treatment id="03CD830F7B156B78FCD1FDF08183A6D5" LSID="urn:lsid:plazi:treatment:03CD830F7B156B78FCD1FDF08183A6D5" httpUri="http://treatment.plazi.org/id/03CD830F7B156B78FCD1FDF08183A6D5" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<subSubSection id="C37E61927B156B7BFCD1FDF08081A3E5" box="[832,1216,617,643]" pageId="1" pageNumber="44" type="nomenclature">
<paragraph id="8BDB32197B156B7BFCD1FDF08081A3E5" blockId="1.[832,1216,617,674]" box="[832,1216,617,643]" pageId="1" pageNumber="44">
?
<taxonomicName id="4C64499A7B156B7BFCDDFDF38081A3E5" ID-CoL="9WX4B" authority="Withers, 1924" authorityName="Withers" authorityYear="1924" box="[844,1216,617,643]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B156B7BFCDDFDF3805DA3E5" box="[844,1052,618,643]" italics="true" pageId="1" pageNumber="44">Ranina americana</emphasis>
Withers, 1924
</taxonomicName>
</paragraph>
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<subSubSection id="C37E61927B156B7BFCD1FD1E87C0A3C4" box="[832,897,647,674]" pageId="1" pageNumber="44" type="description">
<paragraph id="8BDB32197B156B7BFCD1FD1E87C0A3C4" blockId="1.[832,1216,617,674]" box="[832,897,647,674]" pageId="1" pageNumber="44">
<figureCitation id="135F2E9C7B156B7BFCD1FD1E87C0A3C4" box="[832,897,647,674]" captionStart="Fig" captionStartId="2.[189,222,839,861]" captionTargetBox="[171,1421,140,817]" captionTargetPageId="2" captionText="Fig. 1 -?Ranina americana Whiters, 1924; holotype, NHM 23798. A) Dorsal view. B) Ventral view. (x 1.3)." figureDoi="http://doi.org/10.5281/zenodo.12753807" httpUri="https://zenodo.org/record/12753807/files/figure.png" pageId="1" pageNumber="44">Fig. 1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B156B7BFCD1FD5D879CA56D" pageId="1" pageNumber="44" type="reference_group">
<paragraph id="8BDB32197B156B7BFCD1FD5D87D3A39A" blockId="1.[832,1475,707,764]" pageId="1" pageNumber="44">
<heading id="D09385757B156B7BFCD1FD5D87D3A39A" centered="true" fontSize="10" level="2" pageId="1" pageNumber="44" reason="3">
<taxonomicName id="4C64499A7B156B7BFCD1FD5D8177A3B8" authority="Withers, 1924: 125 - 127" authorityName="Withers" authorityPageNumber="125 - 127" authorityYear="1924" box="[832,1334,707,734]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B156B7BFCD1FD5D8054A3BB" box="[832,1045,708,733]" italics="true" pageId="1" pageNumber="44">Ranina americana</emphasis>
Withers, 1924: 125-127
</taxonomicName>
, Pl. 4, figs. 1-3.
</heading>
</paragraph>
<paragraph id="8BDB32197B156B7BFCD1FC8681CFA2D6" blockId="1.[832,1474,798,945]" pageId="1" pageNumber="44">
<heading id="D09385757B156B7BFCD1FC8681CFA2D6" centered="true" fontSize="10" level="2" pageId="1" pageNumber="44" reason="3">
<treatmentCitationGroup id="AB7415377B156B7BFCD1FC8681CFA2D6" pageId="1" pageNumber="44">
<taxonomicName id="4C64499A7B156B7BFCD1FC868053A25E" ID-CoL="9WX4B" authorityName="Withers" authorityYear="1924" box="[832,1042,799,824]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B156B7BFCD1FC868053A25E" box="[832,1042,799,824]" italics="true" pageId="1" pageNumber="44">Ranina americana</emphasis>
</taxonomicName>
-
<treatmentCitation id="0AC514087B156B7BFBBCFC8780BDA25E" author="Rathbun M. J." box="[1069,1276,798,824]" page="91" pageId="1" pageNumber="44" year="1926">
<bibRefCitation id="EFF54FE87B156B7BFBBCFC8780BDA25E" author="Rathbun M. J." box="[1069,1276,798,824]" pageId="1" pageNumber="44" pagination="1 - 155" refId="ref24728" refString="Rathbun M. J., 1926 - The fossil stalk-eyed Crustacea of the Pacific slope of North America. Bulletin of the United States National Museum, 138: 1 - 155." type="journal article" year="1926">Rathbun 1926: 91</bibRefCitation>
</treatmentCitation>
, Pl. 23, figs. 1-4.
<treatmentCitation id="0AC514087B156B7BFCEDFCA58029A230" author="Glaessner M. F." box="[892,1128,828,854]" page="360" pageId="1" pageNumber="44" year="1929">
<bibRefCitation id="EFF54FE87B156B7BFCEDFCA58029A230" author="Glaessner M. F." box="[892,1128,828,854]" pageId="1" pageNumber="44" pagination="1 - 464" refId="ref23938" refString="Glaessner M. F., 1929 - Crustacea Decapoda. In: Fossilium Catalogus. I: Animalia. Pompeckj F. J. (ed.). W. Junk, Berlin, 41: 1 - 464." type="journal article" year="1929">Glaessner 1929: 360</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B156B7BFB1AFCA481FFA230" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." box="[1163,1470,828,855]" page="120" pageId="1" pageNumber="44" year="2009">
<bibRefCitation id="EFF54FE87B156B7BFB1AFCA481FFA230" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." box="[1163,1470,828,855]" pageId="1" pageNumber="44" pagination="119 - 122" refId="ref23670" refString="De Angeli A., Dall'Igna G. L. &amp; Ceccon L., 2009 - Segnalazione di Ranina sp. (Decapoda, Brachyura, Raninidae) dell'Eocene superiore di Monte Malo (Vicenza, Italia settentrionale). Lavori - Societa Veneziana di Scienze Naturali, 34: 119 - 122." type="journal article" year="2009">
De Angeli
<emphasis id="B910EE0B7B156B7BFA96FCA48100A230" box="[1287,1345,828,854]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
2009: 120
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B156B7BFCECFCC380C2A212" author="Waugh D. A. &amp; Feldmann R. M. &amp; Schweitzer C. E." box="[893,1155,858,884]" page="20" pageId="1" pageNumber="44" year="2009">
<bibRefCitation id="EFF54FE87B156B7BFCECFCC380C2A212" author="Waugh D. A. &amp; Feldmann R. M. &amp; Schweitzer C. E." box="[893,1155,858,884]" pageId="1" pageNumber="44" pagination="15 - 41" refId="ref25322" refString="Waugh D. A., Feldmann R. M. &amp; Schweitzer C. E., 2009 - Systematic evaluation of raninid cuticle microstructure. Bulletin of the Mizunami Fossil Museum, 35: 15 - 41." type="journal article" year="2009">
Waugh
<emphasis id="B910EE0B7B156B7BFC44FCC28051A212" box="[981,1040,858,884]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
2009: 20
</bibRefCitation>
</treatmentCitation>
,
<tableCitation id="C6E607A27B156B7BFB01FCC38098A212" box="[1168,1241,858,884]" captionStart="Tab" captionStartId="27.[190,235,370,396]" captionTargetId="graphics-17@27.[113,1474,430,1966]" captionTargetPageId="27" captionText="Tab. 1 - List of the Ranina fossil species as revised in this paper." httpUri="http://table.plazi.org/id/DF1B62917B0F6B61FF2FFEEB87CEA0EA" pageId="1" pageNumber="44" tableUuid="DF1B62917B0F6B61FF2FFEEB87CEA0EA">Tab. 1</tableCitation>
.
<treatmentCitation id="0AC514087B156B7BFA90FCC3878AA2F5" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." page="74" pageId="1" pageNumber="44" year="2010">
<bibRefCitation id="EFF54FE87B156B7BFA90FCC3878AA2F5" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." pageId="1" pageNumber="44" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="B910EE0B7B156B7BFA16FCC28183A212" box="[1415,1474,858,884]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
2010: 74
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B156B7BFC7AFCE08106A2F5" author="De Angeli A. &amp; Beschin C." box="[1003,1351,888,915]" page="13" pageId="1" pageNumber="44" year="2011">
<bibRefCitation id="EFF54FE87B156B7BFC7AFCE08106A2F5" author="De Angeli A. &amp; Beschin C." box="[1003,1351,888,915]" pageId="1" pageNumber="44" pagination="11 - 20" refId="ref23611" refString="De Angeli A. &amp; Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico &quot; G. Zannato &quot;, 18: 11 - 20." type="book chapter" year="2011">De Angeli &amp; Beschin 2011: 13</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B156B7BFAF6FCE08000A2D7" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." page="208" pageId="1" pageNumber="44" year="2012">
<bibRefCitation id="EFF54FE87B156B7BFAF6FCE08000A2D7" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." pageId="1" pageNumber="44" pagination="1 - 216" refId="ref25151" refString="Van Bakel W. M., Guinot D., Artal P., Fraaije R. H. B. &amp; Jagt J. W. M., 2012 - A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata). Zootaxa, 3215: 1 - 216." type="journal article" year="2012">
Van Bakel
<emphasis id="B910EE0B7B156B7BFC1EFC0E8786A2D7" box="[911,967,919,945]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
2012: 208
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B156B7BFBF0FC0E81CBA2D6" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[1121,1418,918,945]" page="260" pageId="1" pageNumber="44" year="2014">
<bibRefCitation id="EFF54FE87B156B7BFBF0FC0E81CBA2D6" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[1121,1418,918,945]" pageId="1" pageNumber="44" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B156B7BFB46FC0E814EA2D7" box="[1239,1295,919,945]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
2014: 260
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</heading>
</paragraph>
<paragraph id="8BDB32197B156B7BFCD1FC4A879CA56D" blockId="1.[832,1474,979,1035]" pageId="1" pageNumber="44">
<heading id="D09385757B156B7BFCD1FC4A879CA56D" centered="true" fontSize="10" level="2" pageId="1" pageNumber="44" reason="3">
<treatmentCitationGroup id="AB7415377B156B7BFCD1FC4A879CA56D" pageId="1" pageNumber="44">
<taxonomicName id="4C64499A7B156B7BFCD1FC4A87BDA28B" baseAuthorityName="Olsson" baseAuthorityYear="1922" box="[832,1020,979,1005]" class="Malacostraca" family="Unciolidae" genus="Hela" kingdom="Animalia" order="Amphipoda" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B156B7BFCD1FC4A87BDA28B" box="[832,1020,979,1005]" italics="true" pageId="1" pageNumber="44">Hela americana</emphasis>
</taxonomicName>
- Lőrenthey
<emphasis id="B910EE0B7B156B7BFB30FC4D80F6A28B" box="[1185,1207,980,1005]" italics="true" pageId="1" pageNumber="44">in</emphasis>
<treatmentCitation id="0AC514087B156B7BFB54FC4A8798A56D" author="Lorenthey E. &amp; Beurlen K." page="109" pageId="1" pageNumber="44" year="1929">
<bibRefCitation id="EFF54FE87B156B7BFB54FC4A8798A56D" author="Lorenthey E. &amp; Beurlen K." pageId="1" pageNumber="44" pagination="1 - 421" refId="ref24319" refString="Lorenthey E. &amp; Beurlen K., 1929 - Die Fossilen Deka- poden der Lander der Ungarischen Krone. Geologica Hungarica (Palaeontologica), 3: 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen 1929: 109</bibRefCitation>
</treatmentCitation>
.
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B156B7BFCF4FBB4817DA413" pageId="1" pageNumber="44" type="materials_examined">
<paragraph id="8BDB32197B156B7BFCF4FBB48160A521" blockId="1.[832,1475,1069,1969]" box="[869,1313,1069,1096]" pageId="1" pageNumber="44">
<emphasis id="B910EE0B7B156B7BFCF4FBB4804DA521" bold="true" box="[869,1036,1069,1095]" pageId="1" pageNumber="44">Type material</emphasis>
:
<typeStatus id="54DF8CBB7B156B7BFB8AFBB480C1A521" box="[1051,1152,1069,1095]" pageId="1" pageNumber="44" type="holotype">holotype</typeStatus>
, NHM 23798.
</paragraph>
<paragraph id="8BDB32197B156B7BFCF4FBD2806CA5C4" blockId="1.[832,1475,1069,1969]" pageId="1" pageNumber="44">
<materialsCitation id="3B0C38447B156B7BFCF4FBD28068A5C4" country="United States of America" county="Late Oligocene" location="Clallam Bay" municipality="Blakeley Formation" pageId="1" pageNumber="44" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="B910EE0B7B156B7BFCF4FBD28040A503" bold="true" box="[869,1025,1099,1125]" pageId="1" pageNumber="44">
<typeStatus id="54DF8CBB7B156B7BFCF4FBD287E1A503" box="[869,928,1099,1125]" pageId="1" pageNumber="44">Type</typeStatus>
locality
</emphasis>
:
<location id="8EBB64C27B156B7BFB85FBD280E4A500" LSID="urn:lsid:plazi:treatment:03CD830F7B156B78FCD1FDF08183A6D5:8EBB64C27B156B7BFB85FBD280E4A500" box="[1044,1189,1099,1126]" country="United States of America" county="Late Oligocene" municipality="Blakeley Formation" name="Clallam Bay" pageId="1" pageNumber="44" stateProvince="Washington">Clallam Bay</location>
, south shore of
<location id="8EBB64C27B156B7BFAF5FBD28799A5E2" LSID="urn:lsid:plazi:treatment:03CD830F7B156B78FCD1FDF08183A6D5:8EBB64C27B156B7BFAF5FBD28799A5E2" country="United States of America" county="Late Oligocene" municipality="Blakeley Formation" name="Strait of Juan de Fuca" pageId="1" pageNumber="44" stateProvince="Washington">Strait of Juan de Fuca</location>
,
<collectingRegion id="49A0FCFB7B156B7BFC74FBF080EFA5E2" box="[997,1198,1129,1156]" country="United States of America" name="Washington" pageId="1" pageNumber="44">Washington State</collectingRegion>
,
<collectingCountry id="F37372897B156B7BFB2AFBF380B6A5E2" box="[1211,1271,1130,1156]" name="United States of America" pageId="1" pageNumber="44">USA</collectingCountry>
;
<collectingMunicipality id="6BBFA8637B156B7BFA92FBF0872FA5C4" pageId="1" pageNumber="44">Blakeley Formation</collectingMunicipality>
,
<collectingCounty id="62BA4A957B156B7BFCE8FB118068A5C4" box="[889,1065,1160,1186]" pageId="1" pageNumber="44">Late Oligocene</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8BDB32197B156B7BFCF4FB3C8094A5A6" blockId="1.[832,1475,1069,1969]" box="[869,1237,1189,1216]" pageId="1" pageNumber="44">
<emphasis id="B910EE0B7B156B7BFCF4FB3C8053A5A6" bold="true" box="[869,1042,1189,1216]" pageId="1" pageNumber="44">Geological age</emphasis>
: Late Oligocene.
</paragraph>
<paragraph id="8BDB32197B156B7BFCF4FB5D817DA413" blockId="1.[832,1475,1069,1969]" pageId="1" pageNumber="44">
<emphasis id="B910EE0B7B156B7BFCF4FB5D800CA5B8" bold="true" box="[869,1101,1220,1246]" pageId="1" pageNumber="44">Examined material</emphasis>
:
<materialsCitation id="3B0C38447B156B7BFBCCFB5D8797A45F" collectorName="Eocene Hoko River Formation &amp; Stanford University, N." country="United States of America" location="Eocene Hoko River" municipality="River Formation" pageId="1" pageNumber="44" specimenCount="4" stateProvince="Washington" typeStatus="holotype">
<typeStatus id="54DF8CBB7B156B7BFBCCFB5D8082A5B8" box="[1117,1219,1220,1246]" pageId="1" pageNumber="44" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D62F9907B156B7BFB41FB5D811DA5B8" box="[1232,1372,1220,1246]" count="4" pageId="1" pageNumber="44" type="generic">4 specimens</specimenCount>
reported by
<bibRefCitation id="EFF54FE87B156B7BFCF5FB7B8056A59B" author="Rathbun M. J." box="[868,1047,1250,1277]" pageId="1" pageNumber="44" pagination="1 - 155" refId="ref24728" refString="Rathbun M. J., 1926 - The fossil stalk-eyed Crustacea of the Pacific slope of North America. Bulletin of the United States National Museum, 138: 1 - 155." type="journal article" year="1926">Rathbun (1926)</bibRefCitation>
from the middle
<collectorName id="269157CF7B156B7BFB70FB7A87F9A47C" pageId="1" pageNumber="44">
<location id="8EBB64C27B156B7BFB70FB7A8183A59A" LSID="urn:lsid:plazi:treatment:03CD830F7B156B78FCD1FDF08183A6D5:8EBB64C27B156B7BFB70FB7A8183A59A" box="[1249,1474,1250,1277]" country="United States of America" municipality="River Formation" name="Eocene Hoko River" pageId="1" pageNumber="44" stateProvince="Washington">
Eocene Hoko
<collectingMunicipality id="6BBFA8637B156B7BFA15FB7B87F9A47C" pageId="1" pageNumber="44">River Formation</collectingMunicipality>
</location>
</collectorName>
,
<collectingRegion id="49A0FCFB7B156B7BFC52FA9980CAA47C" box="[963,1163,1280,1306]" country="United States of America" name="Washington" pageId="1" pageNumber="44">Washington State</collectingRegion>
,
<collectingCountry id="F37372897B156B7BFB06FA99808CA47C" box="[1175,1229,1280,1306]" name="United States of America" pageId="1" pageNumber="44">USA</collectingCountry>
(
<collectorName id="269157CF7B156B7BFB4DFA99871AA45F" pageId="1" pageNumber="44">Stanford University, N.</collectorName>
Pac. 167)
</materialsCitation>
;
<materialsCitation id="3B0C38447B156B7BFC77FA878179A413" country="United States of America" location="Eocene Hoko River" municipality="River Formation" pageId="1" pageNumber="44" specimenCount="3" stateProvince="Washington" typeStatus="holotype">
<specimenCount id="9D62F9907B156B7BFC77FA878035A45E" box="[998,1140,1310,1336]" count="3" pageId="1" pageNumber="44" type="generic">3 specimens</specimenCount>
reported by
<bibRefCitation id="EFF54FE87B156B7BFA9CFA8687CCA431" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." pageId="1" pageNumber="44" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B156B7BFA17FA868183A45E" box="[1414,1474,1310,1336]" italics="true" pageId="1" pageNumber="44">et al.</emphasis>
(2014)
</bibRefCitation>
, KSU D675-678, from the the middle
<location id="8EBB64C27B156B7BFAACFAA487E0A412" LSID="urn:lsid:plazi:treatment:03CD830F7B156B78FCD1FDF08183A6D5:8EBB64C27B156B7BFAACFAA487E0A412" country="United States of America" municipality="River Formation" name="Eocene Hoko River" pageId="1" pageNumber="44" stateProvince="Washington">
Eocene Hoko
<collectingMunicipality id="6BBFA8637B156B7BFCF2FAC38061A412" box="[867,1056,1370,1396]" pageId="1" pageNumber="44">River Formation</collectingMunicipality>
</location>
,
<collectingRegion id="49A0FCFB7B156B7BFBBBFAC380B0A413" box="[1066,1265,1370,1397]" country="United States of America" name="Washington" pageId="1" pageNumber="44">Washington State</collectingRegion>
,
<collectingCountry id="F37372897B156B7BFB6DFAC28179A413" box="[1276,1336,1371,1397]" name="United States of America" pageId="1" pageNumber="44">USA</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B156B78FCF4FAE18567A7E3" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44" type="description">
<paragraph id="8BDB32197B156B78FCF4FAE18567A7E3" blockId="1.[832,1475,1069,1969]" lastBlockId="2.[113,756,920,1971]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<emphasis id="B910EE0B7B156B7BFCF4FAE180B3A4F4" bold="true" box="[869,1266,1400,1427]" pageId="1" pageNumber="44">
Description by
<bibRefCitation id="EFF54FE87B156B7BFBA5FAE180B3A4F4" author="Whiters T. H." box="[1076,1266,1400,1426]" pageId="1" pageNumber="44" pagination="121 - 127" refId="ref25362" refString="Whiters T. H., 1924 - Some Decapod Crustaceans (Callianassa and Ranina) from the Oligocene of Washington, State, U. S. A. The Annals and Magazine of Natural History, 14 (9): 121 - 127." type="journal article" year="1924">Whiters (1924)</bibRefCitation>
</emphasis>
: “
<emphasis id="B910EE0B7B156B78FA8BFAE0855AA7E3" italics="true" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
Cephalothorax elongate-ovoid, moderately convex transversely, width of orbito-frontal margin about equal to half that of the carapace; length, imperfect,
<quantity id="4C9C9FFC7B156B7BFBC4FA4A80E5A48B" box="[1109,1188,1491,1517]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="44" unit="mm" value="60.0">60 mm</quantity>
; breadth
<quantity id="4C9C9FFC7B156B7BFA83FA4A8120A48B" box="[1298,1377,1491,1517]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="44" unit="mm" value="48.0">48 mm</quantity>
; anterolateral margins with three large lobes, very narrowly and deeply separated; the anterior lobe on the left side is the only one preserved entire, and this has a broadly rounded margin; the middle lobe is of about the same width, but evidently projected beyond the others; the third and posterior lobe being small and projected only to a small extent; the postero-lateral margins converge posteriorly, are slightly raised to form a rim, which is weakly serrated above; the orbito-frontal margins broken away, but on the right side is an impression of the eye-stalk and on the left the eye-stalk itself; outer surface smooth except on the sides of the anterior half of the carapace, where there are semicircular incisione, behind which the surface is raised and the edges slightly toothed; these markings are somewhat coarser anteriorly; on the inner surface only the anterior half of the specimen is preserved, and the pterygostomian plates, the third maxillipedes, and the first sternal plate are displaced; pterygostomian plates extremely convex, with the sides bordering the buccal cavità slightly excavated, and below there is a small rounded excavation for the basal joint of the third maxillipedes; the inner half of the surface is smooth and minutely pitted, and the outer half is coarsely ornamented with raised scale-like markings, toothed on the anterior edge, and these markings become more closely disposed towards the outer margin; third maxillipedes much elongated; first sternite widely trifid, with the outer lobes widely rounded, wider than the middle lobeor episternum; the sides curve downward and become subparallel; surface of first scernite marked by a number of pits, forming an almost symmetrical pattern; episternum with the anterior margins forming an obtuse angle and the posterior margins slightly inclined outwards; branchiostergite with the buccal margin sinuous, obliquely inclined outwards, and the margin adjoining the pterygostomian plate bordered by a close line of pits; the middle of the plate is raised to form a ridge, steep-sided and somewhat concave on the outer side, where there are a number of corse but small scale-like markings; a few similar, but smaller, markings are situated near to the buccal margin.
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF1B62917B166B78FF2CFCDE80D3A23B" ID-DOI="http://doi.org/10.5281/zenodo.12753807" ID-Zenodo-Dep="12753807" box="[189,1170,839,861]" httpUri="https://zenodo.org/record/12753807/files/figure.png" pageId="2" pageNumber="45" startId="2.[189,222,839,861]" targetBox="[171,1421,140,817]" targetPageId="2" targetType="figure">
<paragraph id="8BDB32197B166B78FF2CFCDE80D3A23B" blockId="2.[189,1170,839,861]" box="[189,1170,839,861]" pageId="2" pageNumber="45">
Fig. 1 -?
<taxonomicName id="4C64499A7B166B78FE9EFCDE867EA23A" authority="Whiters, 1924" authorityName="Whiters" authorityYear="1924" box="[271,575,839,861]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FE9EFCDE85F6A23A" box="[271,439,839,860]" italics="true" pageId="2" pageNumber="45">Ranina americana</emphasis>
<bibRefCitation id="EFF54FE87B166B78FE2DFCDE867EA23A" author="Whiters T. H." box="[444,575,839,861]" pageId="2" pageNumber="45" pagination="121 - 127" refId="ref25362" refString="Whiters T. H., 1924 - Some Decapod Crustaceans (Callianassa and Ranina) from the Oligocene of Washington, State, U. S. A. The Annals and Magazine of Natural History, 14 (9): 121 - 127." type="journal article" year="1924">Whiters, 1924</bibRefCitation>
</taxonomicName>
; holotype, NHM 23798. A) Dorsal view. B) Ventral view. (x 1.3).
</paragraph>
</caption>
<subSubSection id="C37E61927B166B78FF06F9108183A6D5" pageId="2" pageNumber="45" type="discussion">
<paragraph id="8BDB32197B166B78FF06F9108017A501" blockId="2.[113,756,920,1971]" lastBlockId="2.[831,1474,920,1971]" pageId="2" pageNumber="45">
<emphasis id="B910EE0B7B166B78FF06F9108554A7C5" bold="true" box="[151,277,1673,1699]" pageId="2" pageNumber="45">Discussion</emphasis>
. Withers (1924) described
<taxonomicName id="4C64499A7B166B78FDC9F91386B5A7C5" authorityName="Whiters" authorityYear="1924" box="[600,756,1674,1700]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FDC9F9138628A7C5" box="[600,617,1674,1699]" italics="true" pageId="2" pageNumber="45">R</emphasis>
.
<emphasis id="B910EE0B7B166B78FDEDF91386B5A7C5" box="[636,756,1674,1699]" italics="true" pageId="2" pageNumber="45">americana</emphasis>
</taxonomicName>
from the late Oligocene Blakeley Formation of Clallam Bay,
<collectingRegion id="49A0FCFB7B166B78FF3AF95C8532A7B9" box="[171,371,1733,1759]" country="United States of America" name="Washington" pageId="2" pageNumber="45">Washington State</collectingRegion>
,
<collectingCountry id="F37372897B166B78FEEFF95C85F8A7B9" box="[382,441,1733,1759]" name="United States of America" pageId="2" pageNumber="45">USA</collectingCountry>
, based upon a single poorly preserved specimen lacking the frontal margin and the anterolateral spines, the main characters useful for a specific assignment. The correct locality where this specimen was collected is unknown except for Clallam Bay which has exposures from Upper Oligocene to Lower Miocene Clallam Formation (
<bibRefCitation id="EFF54FE87B166B78FEBDF8E38588A6F2" author="Arnold R." box="[300,457,1914,1940]" pageId="2" pageNumber="45" pagination="1 - 461" refId="ref23362" refString="Arnold R., 1906 - Geologic reconnaissance of the coast of the Olympic Peninsula. Geological Society of America Bulletin, 17, 1 - 461." type="journal article" year="1906">Arnold, 1906</bibRefCitation>
;
<bibRefCitation id="EFF54FE87B166B78FE47F8E386C4A6F2" author="Addicott W. O." box="[470,645,1914,1940]" pageId="2" pageNumber="45" pagination="1 - 44" refId="ref23221" refString="Addicott W. O., 1976 - Molluscan paleontology of the lower Miocene Clallam Formation, northwestern Washington. U. S. Geological Survey Professional Paper, 976: 1 - 44." type="journal article" year="1976">Addicott, 1976</bibRefCitation>
;
<bibRefCitation id="EFF54FE87B166B78FD02F8E3855BA6D4" author="Prothero D. R. &amp; Burns C." pageId="2" pageNumber="45" pagination="234 - 241" refId="ref24560" refString="Prothero D. R. &amp; Burns C., 2001 - Magnetic stratigraphy and tectonic rotation of the upper Oligocene-? lower Miocene (type Pillarian stage) Clallam Formation, Clallam County, Washington. Pacific Section SEPM, 91: 234 - 241." type="journal article" year="2001">Prothero &amp; Burns, 2001</bibRefCitation>
). The Blakeley Formation is limited to exposures west of Seattle,
<collectingRegion id="49A0FCFB7B166B78FBC1FC018094A2D4" box="[1104,1237,920,946]" country="United States of America" name="Washington" pageId="2" pageNumber="45">Washington</collectingRegion>
(
<bibRefCitation id="EFF54FE87B166B78FB72FC01873DA2B6" author="Prothero D. R. &amp; Nesbitt E. A." pageId="2" pageNumber="45" pagination="315 - 322" refId="ref24607" refString="Prothero D. R. &amp; Nesbitt E. A., 2008 - Paleomagentism and tectonic rotation of the Restoration Point Member of the Blakeley Formation (Type Blakeley Stage), Bainbridge Island, Washington, and the Pacific Coast Oligocene-Miocene Boundary. In: Neogene Mammals. Lucas S. G., Morgan G. S., Spielmann J. A. &amp; Prothero D. R., (eds.). New Mexico Museum of Natural History and Science Bulletin, 44: 315 - 322." type="journal article" year="2008">Prothero &amp; Nesbitt, 2008</bibRefCitation>
). This is the only report of a
<taxonomicName id="4C64499A7B166B78FB49FC2E8168A2B6" authorityName="Lamark" authorityYear="1810" box="[1240,1321,951,976]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B166B78FB49FC2E8168A2B6" box="[1240,1321,951,976]" italics="true" pageId="2" pageNumber="45">Ranina</emphasis>
</taxonomicName>
crab coming from this formation and area. The sole specimen lacks enough diagnostic characters for assignment to specific level. In conclusion the inability to recognize the diagnostic characters of
<taxonomicName id="4C64499A7B166B78FB92FBB68015A52E" authorityName="Lamark" authorityYear="1810" box="[1027,1108,1071,1096]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B166B78FB92FBB68015A52E" box="[1027,1108,1071,1096]" italics="true" pageId="2" pageNumber="45">Ranina</emphasis>
</taxonomicName>
questions the placement of
<taxonomicName id="4C64499A7B166B78FA1BFBB687C9A500" authorityName="Whiters" authorityYear="1924" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FA1BFBB687C9A500" italics="true" pageId="2" pageNumber="45">americana</emphasis>
</taxonomicName>
within this genus.
</paragraph>
<paragraph id="8BDB32197B166B78FCF4FBF281EFA598" blockId="2.[831,1474,920,1971]" pageId="2" pageNumber="45">
<bibRefCitation id="EFF54FE87B166B78FCF4FBF28076A5E3" author="Rathbun M. J." box="[869,1079,1131,1157]" pageId="2" pageNumber="45" pagination="1 - 155" refId="ref24728" refString="Rathbun M. J., 1926 - The fossil stalk-eyed Crustacea of the Pacific slope of North America. Bulletin of the United States National Museum, 138: 1 - 155." type="journal article" year="1926">Rathbun (1926: 91</bibRefCitation>
, Pl. 23, figs. 1-4) described four poorly preserved specimens from the Hoko River Formation referring them to
<taxonomicName id="4C64499A7B166B78FB91FB3180D7A5A7" authorityName="Whiters" authorityYear="1924" box="[1024,1174,1191,1217]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FB91FB318050A5A7" box="[1024,1041,1192,1217]" italics="true" pageId="2" pageNumber="45">R</emphasis>
.
<emphasis id="B910EE0B7B166B78FB8FFB3180D7A5A7" box="[1054,1174,1192,1217]" italics="true" pageId="2" pageNumber="45">americana</emphasis>
</taxonomicName>
. However, as stated above, the
<typeStatus id="54DF8CBB7B166B78FCF9FB5F87D9A586" box="[872,920,1222,1248]" pageId="2" pageNumber="45">type</typeStatus>
species does not have enough diagnostic features to make a comparison with the other species of
<taxonomicName id="4C64499A7B166B78FAC7FB7D81E6A59B" authorityName="Lamark" authorityYear="1810" box="[1366,1447,1252,1277]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B166B78FAC7FB7D81E6A59B" box="[1366,1447,1252,1277]" italics="true" pageId="2" pageNumber="45">Ranina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB32197B166B78FCF4FA9B800FA786" blockId="2.[831,1474,920,1971]" pageId="2" pageNumber="45">
<bibRefCitation id="EFF54FE87B166B78FCF4FA9B80DAA47A" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[869,1179,1281,1308]" pageId="2" pageNumber="45" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B166B78FC4DFA9B8054A47D" box="[988,1045,1281,1307]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
(2014: 250
</bibRefCitation>
, fig. 18C) figured a specimen of
<taxonomicName id="4C64499A7B166B78FC0AFAB98077A45F" authorityName="Whiters" authorityYear="1924" box="[923,1078,1312,1337]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FC0AFAB98077A45F" box="[923,1078,1312,1337]" italics="true" pageId="2" pageNumber="45">R. americana</emphasis>
</taxonomicName>
from the late Eocene Hoko River Formation (
<collectionCode id="ED75AADC7B166B78FC54FAA787BEA43E" box="[965,1023,1342,1368]" country="Saudi Arabia" lsid="urn:lsid:biocol.org:col:13432" name="King Saud University" pageId="2" pageNumber="45" type="Herbarium">KSU</collectionCode>
D677). This specimen is better preserved than the ones reported by
<bibRefCitation id="EFF54FE87B166B78FB1EFAC5817EA411" author="Rathbun M. J." box="[1167,1343,1372,1399]" pageId="2" pageNumber="45" pagination="1 - 155" refId="ref24728" refString="Rathbun M. J., 1926 - The fossil stalk-eyed Crustacea of the Pacific slope of North America. Bulletin of the United States National Museum, 138: 1 - 155." type="journal article" year="1926">Rathbun (1926)</bibRefCitation>
in having a wide, subtriangular first anterolateral spine, a second, directed outward, anterolateral spine with accessorial denticles on anterior edge of the spine, and a dorsal surface ornated by scabrous granules arranged in different ways along the carapace but never forming transverse rims. In addition,
<bibRefCitation id="EFF54FE87B166B78FC39F98880E5A74D" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[936,1188,1552,1579]" pageId="2" pageNumber="45" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B166B78FB8DF9888015A74C" box="[1052,1108,1552,1578]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
(2014)
</bibRefCitation>
also lists
<specimenCount id="9D62F9907B166B78FA81F9898183A74C" box="[1296,1474,1552,1578]" count="3" pageId="2" pageNumber="45" type="generic">three specimens</specimenCount>
of
<taxonomicName id="4C64499A7B166B78FCF1F9B687B8A72E" authorityName="Whiters" authorityYear="1924" box="[864,1017,1583,1608]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FCF1F9B687B8A72E" box="[864,1017,1583,1608]" italics="true" pageId="2" pageNumber="45">R. americana</emphasis>
</taxonomicName>
(
<collectionCode id="ED75AADC7B166B78FB9BF9B68005A72F" box="[1034,1092,1583,1609]" country="Saudi Arabia" lsid="urn:lsid:biocol.org:col:13432" name="King Saud University" pageId="2" pageNumber="45" type="Herbarium">KSU</collectionCode>
D675, D676, D678) without any comparison with the
<typeStatus id="54DF8CBB7B166B78FBA8F9D48028A701" box="[1081,1129,1613,1639]" pageId="2" pageNumber="45">type</typeStatus>
material. Indeed these specimens cannot be compared with the
<typeStatus id="54DF8CBB7B166B78FB5EF9F280BEA7E3" box="[1231,1279,1643,1669]" pageId="2" pageNumber="45">type</typeStatus>
material because they do not preserve the anterolateral spines and the frontal margin, main characters necessary to comparisons at specific level in
<taxonomicName id="4C64499A7B166B78FC67F95F8006A7B9" authorityName="Lamark" authorityYear="1810" box="[1014,1095,1734,1759]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B166B78FC67F95F8006A7B9" box="[1014,1095,1734,1759]" italics="true" pageId="2" pageNumber="45">Ranina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDB32197B166B78FCF4F97A8183A6D5" blockId="2.[831,1474,920,1971]" pageId="2" pageNumber="45">
Both specimens referred to
<taxonomicName id="4C64499A7B166B78FB0EF97D8176A79B" authorityName="Whiters" authorityYear="1924" box="[1183,1335,1764,1789]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="americana">
<emphasis id="B910EE0B7B166B78FB0EF97D8176A79B" box="[1183,1335,1764,1789]" italics="true" pageId="2" pageNumber="45">R. americana</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF54FE87B166B78FAF2F97A87CBA67A" author="Rathbun M. J." pageId="2" pageNumber="45" pagination="1 - 155" refId="ref24728" refString="Rathbun M. J., 1926 - The fossil stalk-eyed Crustacea of the Pacific slope of North America. Bulletin of the United States National Museum, 138: 1 - 155." type="journal article" year="1926">Rathbun (1926)</bibRefCitation>
and
<bibRefCitation id="EFF54FE87B166B78FC57F89B808CA67A" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." box="[966,1229,1793,1820]" pageId="2" pageNumber="45" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B166B78FBAFF89B8038A67D" box="[1086,1145,1793,1819]" italics="true" pageId="2" pageNumber="45">et al.</emphasis>
(2014)
</bibRefCitation>
could be assigned to
<taxonomicName id="4C64499A7B166B78FCD1F8B987F3A65C" authorityName="De Haan" authorityYear="1839" box="[832,946,1824,1850]" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="family">Raninidae</taxonomicName>
<emphasis id="B910EE0B7B166B78FC28F8B98006A65F" box="[953,1095,1824,1849]" italics="true" pageId="2" pageNumber="45">sensu stricto</emphasis>
. However, previous authors have not compared these specimens with the
<typeStatus id="54DF8CBB7B166B78FB69F8A7811BA63E" box="[1272,1370,1854,1880]" pageId="2" pageNumber="45" type="holotype">holotype</typeStatus>
or considered the differences in age and formations. Therefore, until new specimens are found preserving specific level characters these specimens should be referred as?
<taxonomicName id="4C64499A7B166B78FADBF8008183A6D5" box="[1354,1474,1945,1971]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B910EE0B7B166B78FADBF80081DAA6D4" box="[1354,1435,1945,1970]" italics="true" pageId="2" pageNumber="45">Ranina</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>