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<mods:title id="C747E732B1A1F0CFC48DBB3D4C538D05">A preliminary review of the fossil species of Ranina Lamarck, 1801 (Decapoda, Brachyura, Raninidae), with systematic remarks</mods:title>
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?
<taxonomicName id="4C64499A7B116B7FFCDDF95C8096A7B8" ID-CoL="BTSJG" authority="Lorenthey, 1898" authorityName="Lorenthey" authorityYear="1898" box="[844,1239,1732,1758]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B116B7FFCDDF95C8058A7B8" box="[844,1049,1732,1758]" italics="true" pageId="5" pageNumber="48">Ranina brevispina</emphasis>
Lőrenthey, 1898
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<figureCitation id="135F2E9C7B116B7FFCAEF97B87C0A79A" box="[831,897,1762,1788]" captionStart="Fig" captionStartId="6.[113,146,1919,1941]" captionTargetBox="[154,709,1212,1890]" captionTargetId="figure-829@6.[154,716,1209,1894]" captionTargetPageId="6" captionText="Fig. 4 -?Ranina brevispina Lőrenthey, 1898; Lőrentheys original figure (Pl. 10, fig. 2)." figureDoi="http://doi.org/10.5281/zenodo.12753813" httpUri="https://zenodo.org/record/12753813/files/figure.png" pageId="5" pageNumber="48">Fig. 4</figureCitation>
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<treatmentCitation id="0AC514087B116B7FFCAEF8868120A65F" author="Lorenthey E." box="[831,1377,1822,1849]" page="138" pageId="5" pageNumber="48" year="1898">
<taxonomicName id="4C64499A7B116B7FFCAEF8868120A65F" ID-CoL="BTSJG" authority="Lorenthey, 1898 b: 135 - 138" authorityName="Lorenthey" authorityPageNumber="135 - 138" authorityYear="1898" box="[831,1377,1822,1849]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B116B7FFCAEF8868052A65E" box="[831,1043,1822,1848]" italics="true" pageId="5" pageNumber="48">Ranina brevispina</emphasis>
<bibRefCitation id="EFF54FE87B116B7FFBB1F8878120A65F" author="Lorenthey E." box="[1056,1377,1822,1849]" pageId="5" pageNumber="48" pagination="134 - 152" refId="ref24292" refString="Lorenthey E., 1898 b - Uber die Brachyuren der palaeon- tologischen sammlung des Bayrischen Staates. Termeszetrajzi Fuzetek, 21: 134 - 152." type="journal article" year="1898">Lőrenthey, 1898b: 135-138</bibRefCitation>
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, Pl. 10, figs. 2, 3, Pl. 11, fig. 1.
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<taxonomicName id="4C64499A7B116B7FFCAEF8E38050A6F5" ID-CoL="BTSJG" authorityName="Lorenthey" authorityYear="1898" box="[831,1041,1913,1939]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B116B7FFCAEF8E38050A6F5" box="[831,1041,1913,1939]" italics="true" pageId="5" pageNumber="48">Ranina brevispina</emphasis>
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-
<treatmentCitation id="0AC514087B116B7FFBA3F8E080BAA6F5" author="Fabiani R." box="[1074,1275,1913,1939]" page="91" pageId="5" pageNumber="48" year="1910">
<bibRefCitation id="EFF54FE87B116B7FFBA3F8E080BAA6F5" author="Fabiani R." box="[1074,1275,1913,1939]" pageId="5" pageNumber="48" pagination="85 - 102" refId="ref23776" refString="Fabiani R., 1910 - Sulle specie di Ranina finora note ed in particolare sulla Ranina Aldrovandii. Atti della Accademia Scientifica Veneto-Trentino-Istriana, 3 (3): 85 - 102." type="journal article" year="1910">Fabiani 1910: 91</bibRefCitation>
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, 92.
<treatmentCitation id="0AC514087B116B7FFAC5F8E08798A6D7" author="Glaessner M. F." page="361" pageId="5" pageNumber="48" year="1929">
<bibRefCitation id="EFF54FE87B116B7FFAC5F8E08798A6D7" author="Glaessner M. F." pageId="5" pageNumber="48" pagination="1 - 464" refId="ref23938" refString="Glaessner M. F., 1929 - Crustacea Decapoda. In: Fossilium Catalogus. I: Animalia. Pompeckj F. J. (ed.). W. Junk, Berlin, 41: 1 - 464." type="journal article" year="1929">Glaessner 1929: 361</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B116B7FFC68F80E8164A6D7" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." box="[1017,1317,1943,1969]" page="120" pageId="5" pageNumber="48" year="2009">
<bibRefCitation id="EFF54FE87B116B7FFC68F80E8164A6D7" author="De Angeli A. &amp; Dall'Igna G. L. &amp; Ceccon L." box="[1017,1317,1943,1969]" pageId="5" pageNumber="48" pagination="119 - 122" refId="ref23670" refString="De Angeli A., Dall'Igna G. L. &amp; Ceccon L., 2009 - Segnalazione di Ranina sp. (Decapoda, Brachyura, Raninidae) dell'Eocene superiore di Monte Malo (Vicenza, Italia settentrionale). Lavori - Societa Veneziana di Scienze Naturali, 34: 119 - 122." type="journal article" year="2009">
De Angeli
<emphasis id="B910EE0B7B116B7FFBE2F80180EAA6D7" box="[1139,1195,1943,1969]" italics="true" pageId="5" pageNumber="48">et al.</emphasis>
2009: 120
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B116B7CFAD7F80E8501A1C4" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." lastPageId="6" lastPageNumber="49" page="74" pageId="5" pageNumber="48" year="2010">
<bibRefCitation id="EFF54FE87B116B7CFAD7F80E8501A1C4" author="Schweitzer C. &amp; Feldmann R. M. &amp; Garassino A. &amp; Karasawa H. &amp; Schweigert G." lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48" pagination="1 - 222" refId="ref24966" refString="Schweitzer C. E, Feldmann R. M., Garassino A., Karasawa H. &amp; Schweigert G., 2010 - Systematic list of fossil decapod crustacean species. Crustaceana Monographs, 10: 1 - 222." type="journal article" year="2010">
Schweitzer
<emphasis id="B910EE0B7B126B7CFF06FF118490A1C7" box="[151,209,135,161]" italics="true" pageId="6" pageNumber="49">et al.</emphasis>
2010: 74
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B126B7CFEF3FF118685A1C4" author="De Angeli A. &amp; Beschin C." box="[354,708,135,162]" page="13" pageId="6" pageNumber="49" year="2011">
<bibRefCitation id="EFF54FE87B126B7CFEF3FF118685A1C4" author="De Angeli A. &amp; Beschin C." box="[354,708,135,162]" pageId="6" pageNumber="49" pagination="11 - 20" refId="ref23611" refString="De Angeli A. &amp; Beschin C., 2011 - Il genere Ranina Lamarck, 1801 (Crustacea, Decapoda, Raninidae) nel Terziario del Vicentino, con descrizione di due nuove specie. Studi e Ricerche - Associazione Amici del Museo - Museo Civico &quot; G. Zannato &quot;, 18: 11 - 20." type="book chapter" year="2011">De Angeli &amp; Beschin 2011: 13</bibRefCitation>
</treatmentCitation>
, 15.
<treatmentCitation id="0AC514087B126B7CFF3DFF3F859CA1A6" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." box="[172,477,166,192]" page="209" pageId="6" pageNumber="49" year="2012">
<bibRefCitation id="EFF54FE87B126B7CFF3DFF3F859CA1A6" author="Van Bakel W. M. &amp; Guinot D. &amp; Artal P. &amp; Fraaije R. H. B. &amp; Jagt J. W. M." box="[172,477,166,192]" pageId="6" pageNumber="49" pagination="1 - 216" refId="ref25151" refString="Van Bakel W. M., Guinot D., Artal P., Fraaije R. H. B. &amp; Jagt J. W. M., 2012 - A revision of the Palaeocorystoidea and the phylogeny of raninoidian crabs (Crustacea, Decapoda, Brachyura, Podotremata). Zootaxa, 3215: 1 - 216." type="journal article" year="2012">
Van Bakel
<emphasis id="B910EE0B7B126B7CFEB9FF3F8520A1A6" box="[296,353,166,192]" italics="true" pageId="6" pageNumber="49">et al.</emphasis>
2012: 209
</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AC514087B126B7CFE6FFF3F8484A1B8" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." page="260" pageId="6" pageNumber="49" year="2014">
<bibRefCitation id="EFF54FE87B126B7CFE6FFF3F8484A1B8" author="Karasawa H. &amp; Schweitzer C. E. &amp; Feldmann R. M. &amp; Luque J." pageId="6" pageNumber="49" pagination="216 - 272" refId="ref24222" refString="Karasawa H., Schweitzer C. E., Feldmann R. M. &amp; Luque J., 2014 - Phylogeny and classification of Raninoida (Decapoda: Brachyura). Journal of Crustacean Biology, 34 (2): 216 - 272." type="journal article" year="2014">
Karasawa
<emphasis id="B910EE0B7B126B7CFDE5FF3F86ECA1A6" box="[628,685,166,192]" italics="true" pageId="6" pageNumber="49">et al.</emphasis>
2014: 260
</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BDB32197B126B7CFFE0FE99854EA05E" blockId="6.[113,756,256,312]" pageId="6" pageNumber="49">
<heading id="D09385757B126B7CFFE0FE99854EA05E" centered="true" fontSize="10" level="2" pageId="6" pageNumber="49" reason="3">
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<taxonomicName id="4C64499A7B126B7CFFE0FE99856AA07C" authority="Lőrenthey in Lőrenthey &amp; Beurlen 1929: 109" box="[113,299,256,282]" class="Malacostraca" family="Unciolidae" genus="Hela" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B126B7CFFE0FE99856AA07C" box="[113,299,256,282]" italics="true" pageId="6" pageNumber="49">Hela brevispina</emphasis>
</taxonomicName>
- Lőrenthey
<emphasis id="B910EE0B7B126B7CFE43FE9885A9A07C" box="[466,488,257,282]" italics="true" pageId="6" pageNumber="49">in</emphasis>
<treatmentCitation id="0AC514087B126B7CFE67FE99854AA05E" author="Lorenthey E. &amp; Beurlen K." page="109" pageId="6" pageNumber="49" year="1929">
<bibRefCitation id="EFF54FE87B126B7CFE67FE99854AA05E" author="Lorenthey E. &amp; Beurlen K." pageId="6" pageNumber="49" pagination="1 - 421" refId="ref24319" refString="Lorenthey E. &amp; Beurlen K., 1929 - Die Fossilen Deka- poden der Lander der Ungarischen Krone. Geologica Hungarica (Palaeontologica), 3: 1 - 421." type="journal article" year="1929">Lőrenthey &amp; Beurlen 1929: 109</bibRefCitation>
</treatmentCitation>
.
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</heading>
</paragraph>
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<subSubSection id="C37E61927B126B7CFF06FEC286FEA0A9" pageId="6" pageNumber="49" type="materials_examined">
<paragraph id="8BDB32197B126B7CFF06FEC2868FA012" blockId="6.[113,756,346,1156]" box="[151,718,346,373]" pageId="6" pageNumber="49">
<emphasis id="B910EE0B7B126B7CFF06FEC2857FA012" bold="true" box="[151,318,346,373]" pageId="6" pageNumber="49">Type material</emphasis>
:
<typeStatus id="54DF8CBB7B126B7CFEDCFEC385EEA012" box="[333,431,346,372]" pageId="6" pageNumber="49" type="holotype">holotype</typeStatus>
and repository unknown.
</paragraph>
<paragraph id="8BDB32197B126B7CFF06FEE08620A0F4" blockId="6.[113,756,346,1156]" box="[151,609,376,403]" pageId="6" pageNumber="49">
<materialsCitation id="3B0C38447B126B7CFF06FEE0861CA0F4" box="[151,605,376,403]" country="Algeria" location="Beni Chennadha" pageId="6" pageNumber="49" specimenCount="1" typeStatus="holotype">
<emphasis id="B910EE0B7B126B7CFF06FEE08571A0F4" bold="true" box="[151,304,376,403]" pageId="6" pageNumber="49">
<typeStatus id="54DF8CBB7B126B7CFF06FEE08493A0F5" box="[151,210,377,403]" pageId="6" pageNumber="49">Type</typeStatus>
locality
</emphasis>
:
<location id="8EBB64C27B126B7CFEAEFEE185BDA0F4" LSID="urn:lsid:plazi:treatment:03CD830F7B116B7DFCAEF95D80E2A34F:8EBB64C27B126B7CFEAEFEE185BDA0F4" box="[319,508,376,402]" country="Algeria" name="Beni Chennadha" pageId="6" pageNumber="49">Beni Chennadha</location>
,
<collectingCountry id="F37372897B126B7CFD94FEE1861CA0F4" box="[517,605,376,402]" name="Algeria" pageId="6" pageNumber="49">Algeria</collectingCountry>
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.
</paragraph>
<paragraph id="8BDB32197B126B7CFF06FE0F8678A0D4" blockId="6.[113,756,346,1156]" box="[151,569,406,434]" pageId="6" pageNumber="49">
<emphasis id="B910EE0B7B126B7CFF06FE0F857DA0D7" bold="true" box="[151,316,406,433]" pageId="6" pageNumber="49">Geologcal age</emphasis>
: Miocene (Langhian).
</paragraph>
<paragraph id="8BDB32197B126B7CFF06FE2C86FEA0A9" blockId="6.[113,756,346,1156]" box="[151,703,437,463]" pageId="6" pageNumber="49">
<emphasis id="B910EE0B7B126B7CFF06FE2C853CA0A9" bold="true" box="[151,381,437,463]" pageId="6" pageNumber="49">Examined material</emphasis>
: Lőrentheys original plates.
</paragraph>
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<subSubSection id="C37E61927B126B7DFF06FE4A813EA1A6" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="description">
<paragraph id="8BDB32197B126B7CFF06FE4A815FA79A" blockId="6.[113,756,346,1156]" lastBlockId="6.[831,1475,135,1970]" pageId="6" pageNumber="49">
<emphasis id="B910EE0B7B126B7CFF06FE4A8654A08B" bold="true" box="[151,533,466,493]" pageId="6" pageNumber="49">Description by Lőrenthey (1898)</emphasis>
: “
<emphasis id="B910EE0B7B126B7CFDBEFE4D8510A34F" italics="true" pageId="6" pageNumber="49">
In dem genannten Musem finden sich aus
<collectingCountry id="F37372897B126B7CFEEFFE688585A36D" box="[382,452,497,523]" name="Algeria" pageId="6" pageNumber="49">Algier</collectingCountry>
stammene drei Exemplare einer unbestimmten
</emphasis>
<taxonomicName id="4C64499A7B126B7CFECBFD9685E8A34F" authorityName="Lamark" authorityYear="1810" box="[346,425,527,553]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">Ranina</taxonomicName>
<emphasis id="B910EE0B7B126B7CFE3BFD898183A7A6" italics="true" pageId="6" pageNumber="49">
-Art, welche in den Formenkreis von
<taxonomicName id="4C64499A7B126B7CFF75FDB785B9A321" authority="Munst." authorityName="Munst." box="[228,504,558,583]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="speciosa">Ranina speciosa Münst.</taxonomicName>
gehört, doch wie wir sogleich sehen werden, von allen bekannten verwandten Formen, wesentlich abweicht und so eine gut charakterisirte neue Form ist; die Charaktere derselben lassen sich in Folgendes zusammenfassen; des Cephalothorax ist birnförmig, der grösste Breitendurchmesser befindet sich in dem vorderen Drittel der Längslinie und verschmälert sich sowohl gegen vorn als auch hinten nur schwach; von dem grössten Breitenpunkte erfolgt die Verschmälerung nach rückwärts zu plötzlicher als nach vorne; der Rand des Cephalothorax verläuft von hier S-förmig nach hinten, indem er anfangs im zweiten Drittel der Länge schwach convex, im letzten Drittel aber concav ist, wodurch sich das Schild gegen hinten plötzlicher verschmälert, so dass der Durchmesser im vordren Drittel noch
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, der Hinterland aber schon nur mehr
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zählt; der Cephalothorax ist in der Richtung des Breitedurchmessers convex, in der Linie der Längenaxe dagegen fällt er langsam nach rückwärts ab, ohne jedoch convex zu werden; der Stirnrand ist in der Mitte breit und zeigt einen, ein Drittel der Frontalrandbreite einnehmenden Lobus, welcher in drei Spitzen endigt und zwar so, dass die mittlere kürzer und die seitlichen länger sind, die Randkante dagegen ist zwischen den drei Spitzen stark concav; diesel dreispitzige Lobus wird durch je einen schmalen, doch genug tiefen Einschnitt von dem schmalen, zahnartigen, ersten Seitenlobus getrennt, an welchem man, obwohl die Spitze abgebroehen ist, doch gut sehen kann, dass er spitz und mit seiner Spitze gerade nach vorne gerichtet ist; der durch einen viel schwächeren Einschnitt abgetrennte zweite Seitenlobus ist an seinem Ende zweigetheilt und zwar so, dass der innere der beiden Zähne der stärkere ist, während übrigens beide mit ihrer Spitze schwach gegen die Mittellinie zu neigen; nach diesel Lobus folgt ein breiter, und tiefer als der erste reichender Einschnitt, (welcher auf
<figureCitation id="135F2E9C7B126B7CFBC1FDD580D3A303" box="[1104,1170,587,613]" captionStart="Fig" captionStartId="3.[113,146,1918,1940]" captionTargetBox="[113,756,744,1894]" captionTargetId="figure-657@3.[113,756,744,1355]" captionTargetPageId="3" captionText="Fig. 2 -?Ranina berglundi Squires &amp; Demetrion, 1992. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5)." figureDoi="http://doi.org/10.5281/zenodo.12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="6" pageNumber="49">Fig. 2</figureCitation>
nicht genug gut dargestellt ist); nach demselben kommt wieder ein, in zwei Zähnen endender Lobus, der dritte Seitenlappen, welcher ähnlich entwickelt, wie der vorige, nur schmäler, kürzer, im Ganzen schwächer ist; der vierte Seitenlobus ist ebenfalls breit und wird durch einen noch weiter nach hinten reicheden Einschnitt von dem Dritten getrennt; er weicht aber in seiner Ausbildung von den anderei dreien ab, da er sich in drei Stacheln theilt, so wie dies auf
<figureCitation id="135F2E9C7B126B7CFAABFCA4813CA230" box="[1338,1405,828,854]" captionStart="Fig" captionStartId="3.[113,146,1918,1940]" captionTargetBox="[113,756,744,1894]" captionTargetId="figure-657@3.[113,756,744,1355]" captionTargetPageId="3" captionText="Fig. 2 -?Ranina berglundi Squires &amp; Demetrion, 1992. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5)." figureDoi="http://doi.org/10.5281/zenodo.12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="6" pageNumber="49">Fig. 2</figureCitation>
und
<figureCitation id="135F2E9C7B126B7CFA25FCA58183A230" box="[1460,1474,828,854]" captionStart="Fig" captionStartId="4.[189,222,1915,1937]" captionTargetBox="[189,1394,486,1879]" captionTargetId="figure-197@4.[189,1398,478,1891]" captionTargetPageId="4" captionText="Fig. 3 -?Ranina bouilleana A. Milne Edwards, 1872.A) Milne Edwardss original figures (Pl. 8, figs. 2, 2a-c). B-C) Two specimens, MNHM R03540, collected from the same locality (Phare de Biarritz) of the type series (natural size)." figureDoi="http://doi.org/10.5281/zenodo.12753811" httpUri="https://zenodo.org/record/12753811/files/figure.png" pageId="6" pageNumber="49">3</figureCitation>
theilweise dargestellt ist, nä mlich der innere Stachel ist stark entwickelt und scharf von dem anderei zweispitzigen Stachel abgesetzt; der innere Stachel dieses zweiendigen Lobus ist schwächer als der äussere; am meisten streckt sich der innere Zahn des zweiten Seitenlobus nach vorne; von hier lässt sich eine gerade Linie bis zu dem mittleren Dorn des mittleren Lappens über den ersten Seitenzahn und die Spitze des Seitenstachels des Mittellobus ziehen; eine andere von dem inneren Zahn des zweiten Seitenlobus über die Spitzen des Zähne bis zur Spitze des äusseren Zahnes des vierten Seitenlobus; der ganze Cephalothorax ist von dem vierten Seitenlobus angefangen nach hinten zu mit einer, mit kleinen Tuberkeln geschmückten Leiste begrenzt, während der mordere Rand vollkommen glatt erscheint; die Oberfläche des Cephalothorax mit Ausnahme des Frontalrandes ist mit länglichen, grossen Tuberkeln bedeckt, zwischen denen hier und dort zerstreut sich kleinere, runde Tuberkel beobachten lassen, deren Zahl gegen die Ränder zu zunimmt; dieselben herrschen dann über der mit Tuberkeln geschmückten Leiste am Rande des Cephalothoax vor; der ganze Stirnrand und dessen Lobi sind mit ebensolchen runden Tuberkeln bedeckt; der untere Theil des Cephalothorax, das Sternum und die Extremitäten konnten wegen ihres mangelhaften Erhaltungszustandes nicht studirt werden; um aber einigermaassen einen Begriff von denselben zu geben, publicire ich in
<figureCitation id="135F2E9C7B126B7CFA05F9D5870CA7E2" captionStart="Fig" captionStartId="2.[189,222,839,861]" captionTargetBox="[171,1421,140,817]" captionTargetPageId="2" captionText="Fig. 1 -?Ranina americana Whiters, 1924; holotype, NHM 23798. A) Dorsal view. B) Ventral view. (x 1.3)." figureDoi="http://doi.org/10.5281/zenodo.12753807" httpUri="https://zenodo.org/record/12753807/files/figure.png" pageId="6" pageNumber="49">Fig. 1</figureCitation>
der Tab. XI. die mangelhafte Unterseite der in
<figureCitation id="135F2E9C7B126B7CFAEDF9F28183A7E2" box="[1404,1474,1642,1668]" captionStart="Fig" captionStartId="3.[113,146,1918,1940]" captionTargetBox="[113,756,744,1894]" captionTargetId="figure-657@3.[113,756,744,1355]" captionTargetPageId="3" captionText="Fig. 2 -?Ranina berglundi Squires &amp; Demetrion, 1992. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5)." figureDoi="http://doi.org/10.5281/zenodo.12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="6" pageNumber="49">Fig. 2</figureCitation>
der Tab. X. dargestellten Form, an welcher auch die Eindrücke der Scheerenhände sichtbar sind; die Grösse von
</emphasis>
<taxonomicName id="4C64499A7B126B7CFCAEF95D804EA7B8" authorityName="Lorenthey" authorityYear="1898" box="[831,1039,1732,1758]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="brevispina">Ranina brevispina</taxonomicName>
<emphasis id="B910EE0B7B126B7CFB8BF95C8153A79A" italics="true" pageId="6" pageNumber="49">
ist recht variabel; das vollständigste Exemplar (Tab. X.
<figureCitation id="135F2E9C7B126B7CFB82F97A8017A79A" box="[1043,1110,1762,1788]" captionStart="Fig" captionStartId="3.[113,146,1918,1940]" captionTargetBox="[113,756,744,1894]" captionTargetId="figure-657@3.[113,756,744,1355]" captionTargetPageId="3" captionText="Fig. 2 -?Ranina berglundi Squires &amp; Demetrion, 1992. A) holotype, IGM 5913 (x 1.8). B) paratype, IGM 5914 (x 4.5)." figureDoi="http://doi.org/10.5281/zenodo.12753809" httpUri="https://zenodo.org/record/12753809/files/figure.png" pageId="6" pageNumber="49">Fig. 2</figureCitation>
) ist das kleinste.
</emphasis>
</paragraph>
<paragraph id="8BDB32197B126B7CFCF4F8998180A6D7" blockId="6.[831,1475,135,1970]" pageId="6" pageNumber="49">
Literal translation: in the mentioned museum (the Bavarian State Collection in Munich) there are
<specimenCount id="9D62F9907B126B7CFAAEF887873BA631" count="3" pageId="6" pageNumber="49" type="generic">three specimens</specimenCount>
of an undetermined species of
<taxonomicName id="4C64499A7B126B7CFB4EF8A48171A630" authorityName="Lamark" authorityYear="1810" box="[1247,1328,1853,1878]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B126B7CFB4EF8A48171A630" box="[1247,1328,1853,1878]" italics="true" pageId="6" pageNumber="49">Ranina</emphasis>
</taxonomicName>
from
<collectingCountry id="F37372897B126B7CFAE4F8A481FFA631" box="[1397,1470,1853,1879]" name="Algeria" pageId="6" pageNumber="49">Algier</collectingCountry>
, which belongs to the group of
<taxonomicName id="4C64499A7B126B7CFB18F8C581CFA613" authority="Munst" authorityName="Munst" box="[1161,1422,1883,1909]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="B910EE0B7B126B7CFB18F8C5817EA613" box="[1161,1343,1884,1909]" italics="true" pageId="6" pageNumber="49">Ranina speciosa</emphasis>
Münst
</taxonomicName>
, but, as we will see below, differs from all previously known related forms and therefore it is a well-defined new form.
</paragraph>
<caption id="DF1B62917B126B7CFFE0F8E68571A6C8" ID-DOI="http://doi.org/10.5281/zenodo.12753813" ID-Zenodo-Dep="12753813" httpUri="https://zenodo.org/record/12753813/files/figure.png" pageId="6" pageNumber="49" startId="6.[113,146,1919,1941]" targetBox="[154,709,1212,1890]" targetPageId="6" targetType="figure">
<paragraph id="8BDB32197B126B7CFFE0F8E68571A6C8" blockId="6.[113,756,1919,1966]" pageId="6" pageNumber="49">
Fig. 4 -?
<taxonomicName id="4C64499A7B126B7CFF45F8198666A6F2" authority="Lorenthey, 1898" authorityName="Lorenthey" authorityYear="1898" box="[212,551,1919,1941]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B126B7CFF45F81985C1A6F2" box="[212,384,1919,1941]" italics="true" pageId="6" pageNumber="49">Ranina brevispina</emphasis>
Lőrenthey, 1898
</taxonomicName>
; Lőrentheys original figure (Pl. 10, fig. 2).
</paragraph>
</caption>
<paragraph id="8BDB32197B136B7DFF06FF1E813EA1A6" blockId="7.[113,756,135,1970]" lastBlockId="7.[831,1475,135,553]" pageId="7" pageNumber="50">
The features of this form are as follows: the cephalothorax is pear-shaped; the maximum width is located in the anterior third and it becomes only gradually taller towards the anterior and posterior parts; from the point of maximum width this narrowing is brusquer towards the posterior part than towards the anterior one; form here the margin is S-shaped; at the beginning in the second third of the carapace´s length it is weekly convex, in the last third it becomes concave, so that the narrowing of the carapace is brusquer, so that its diameter in the anterior third is
<quantity id="4C9C9FFC7B136B7DFE9CFE2C851EA0A9" box="[269,351,437,463]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" pageId="7" pageNumber="50" unit="mm" value="42.0">42 mm</quantity>
, but at the posterior margin only
<quantity id="4C9C9FFC7B136B7DFD49FE2C84E1A08B" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="50" unit="mm" value="19.0">19 mm</quantity>
; the (section of the) cephalothorax is convex in the direction of the width diameter, in its longitudinal axis, however, it gradually slopes down without becoming convex; the frontal margin is broad in the middle and exhibits a lobe, which is one third of the entire frontal margin wide and ends in three spines in that way that the central spine is shorten, whereas the lateral ones are longer, and the marginal edge between these three spines is strongly concave; this three-spined lobe is separated by a narrow, moderately deep incision from the narrow, tooth-like first lateral lobe, at which it is still discernible that it was pointed and the tip was foreward-directed, despite of its broken state; the second lateral lobe, which is separated by a much weaker incision, is bisected at its ending in that way, that the inner one of the two teeth is the stronger one, whereas both are curved pointing with their tips towards the median axis; on the far side of this lobe follows an incision, which is wider and first one (which is not very well illustrated in fig. 2); behind follows again a lobe ending in two teeth, the third lateral lobe, which is similar to the previous one, but narrower, shorter and generally weaker developed; the fourth lateral lobe is broad as well and is separated from the third one by a still deeper incision; it differs from the other three ones by subdividing in three spines, as it is partly shown in figs. 2 and 3, the inner spine is strong and sharply separated from the other bicuspid spine; the inner spine of this twice-ending lobe is weaker than the outer one; the inner tooth of the second lateral lobe is the longest projecting one; it is possible to draw a straight line from here (? the inner tooth) to the middle spine of the middle lappet and the first lateral tooth and the tip of the lateral spine of the median lobe; another one connecting the inner tooth of the second lobe, the tips of the teth and the tip of the outer marginal tooth of the forth lateral lobe; the entire cephalothorax is separated posteriorly by a finely tuberculate crest starting from the forth lateral lobe, whereas the fronal margin appears to be completely smooth; the entire surface of the cephalothorax except of the frontal margin is covered with elongate, large tubercles, between the latter small round tubercles are scattered; their number becomes greater towards the lateral margins, the same tubercles predominate in front of the tuberculate crest at the margin of the cephalothorax; the entire frontal margin and its lobes are covered with the same round tubercles; the ventral part of the cephaolothorax, the sternum and the appendages could not be studied due to the poor preservation state; but to give an impression how they look like I show in fig. 1 of Plate 11 the poor ventral part of the specimen illustrated in fig 2 of Plate 10, in which the imprints of the chelae are visible; the size of
<taxonomicName id="4C64499A7B136B7DFC2EFF1180CFA1C7" authorityName="Lorenthey" authorityYear="1898" box="[959,1166,135,161]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="brevispina">
<emphasis id="B910EE0B7B136B7DFC2EFF1180CFA1C7" box="[959,1166,135,161]" italics="true" pageId="7" pageNumber="50">Ranina brevispina</emphasis>
</taxonomicName>
is variable; the most complete specimen (Plate 10, fig. 2) is the smallest one.
</paragraph>
</subSubSection>
<subSubSection id="C37E61927B136B7DFCF4FF5A80E2A34F" pageId="7" pageNumber="50" type="discussion">
<paragraph id="8BDB32197B136B7DFCF4FF5A80E2A34F" blockId="7.[831,1475,135,553]" pageId="7" pageNumber="50">
<emphasis id="B910EE0B7B136B7DFCF4FF5A87A2A1BB" bold="true" box="[869,995,195,221]" pageId="7" pageNumber="50">Discussion</emphasis>
. The
<typeStatus id="54DF8CBB7B136B7DFBBFFF5D801FA1B8" box="[1070,1118,196,222]" pageId="7" pageNumber="50">type</typeStatus>
material, as described and figured by Lőrenthey (1898: Pl. 10, fig. 2; Pl. 11, fig. 1) preserves the complete frontal part with the orbital and postorbital spines, triangular rostrum, first anterolateral spine bifid (second one not preserved), typical raninid pleonal sternum shield-like, and shapped and flattened palms of the chelae. Moreover, the general shape and the dorsal granulate ornamentation of the carapace seems typical of
<emphasis id="B910EE0B7B136B7DFCF0FE2C87F8A0A8" box="[865,953,437,462]" italics="true" pageId="7" pageNumber="50">
<taxonomicName id="4C64499A7B136B7DFCF0FE2C87F7A0A8" authorityName="Lamark" authorityYear="1810" box="[865,950,437,462]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="50" phylum="Arthropoda" rank="genus">Ranina</taxonomicName>
.
</emphasis>
Though the specimen can be assigned to the
<taxonomicName id="4C64499A7B136B7DFCAEFE4A87F0A08B" authorityName="De Haan" authorityYear="1839" box="[831,945,467,493]" class="Malacostraca" family="Raninidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="50" phylum="Arthropoda" rank="family">Raninidae</taxonomicName>
<emphasis id="B910EE0B7B136B7DFC28FE4A8006A08A" box="[953,1095,467,492]" italics="true" pageId="7" pageNumber="50">sensu stricto</emphasis>
based upon the above-mentioned characters, the lack of the second anterolateral spine questions its assignment to
<taxonomicName id="4C64499A7B136B7DFBDAFD8980DDA34F" authorityName="Lamark" authorityYear="1810" box="[1099,1180,528,553]" class="Malacostraca" family="Raninidae" genus="Ranina" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B910EE0B7B136B7DFBDAFD8980DDA34F" box="[1099,1180,528,553]" italics="true" pageId="7" pageNumber="50">Ranina</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>