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<document id="106707FFB3A24894B6BC8BA27059BBDC" ID-DOI="10.5281/zenodo.213946" ID-GBIF-Dataset="2a54e127-4d40-404e-ab7f-5b48f6e37e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="213946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564534622" checkinUser="plazi" docAuthor="Grisales, Diana, Wolff, Marta &amp; De, Claudio J. B." docDate="2012" docId="03BE87F8521BD403C09420A4FD8BF92F" docLanguage="en" docName="zt03591p046.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fannia trimaculata Stein" docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="FF87FF805231D429C0032033FFBBFFA9" masterDocTitle="Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia" masterLastPageNumber="46" masterPageNumber="1" pageNumber="43" updateTime="1698320797981" updateUser="plazi">
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<mods:title id="6DF66733BA26FCD4CA5F7B2288E8A4BF">Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia</mods:title>
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<mods:namePart id="FBB08FF66CD03BE7EA8167EF6C373121">Grisales, Diana</mods:namePart>
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<mods:namePart id="BF64F09E2F0956B2971376E1EE91474E">Wolff, Marta</mods:namePart>
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<mods:namePart id="FBFFB0AA2760F717B37D438BE7946DE5">De, Claudio J. B.</mods:namePart>
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<mods:title id="37FAB26014F0CD4CF83B2364B4502736">Zootaxa</mods:title>
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<mods:date id="69C83C8A90E388B3BF713F8697D1ABCF">2012</mods:date>
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<treatment id="03BE87F8521BD403C09420A4FD8BF92F" ID-DOI="http://doi.org/10.5281/zenodo.6175083" ID-GBIF-Taxon="119557561" ID-Zenodo-Dep="6175083" LSID="urn:lsid:plazi:treatment:03BE87F8521BD403C09420A4FD8BF92F" httpUri="http://treatment.plazi.org/id/03BE87F8521BD403C09420A4FD8BF92F" lastPageNumber="43" pageId="42" pageNumber="43">
<subSubSection id="C30D6565521BD403C09420A4FE4AFF7B" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BA836EE521BD403C09420A4FE67FF18" blockId="42.[151,497,151,210]" box="[151,476,151,177]" pageId="42" pageNumber="43">
<heading id="D0E08182521BD403C09420A4FE67FF18" bold="true" box="[151,476,151,177]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
<taxonomicName id="4C174D6D521BD403C09420A4FE67FF18" ID-CoL="6HPYZ" authority="Stein" authorityName="Stein" box="[151,476,151,177]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C09420A4FE67FF18" bold="true" box="[151,476,151,177]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C09420A4FE38FF18" bold="true" box="[151,387,151,177]" italics="true" pageId="42" pageNumber="43">Fannia trimaculata</emphasis>
(Stein)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA836EE521BD403C0942089FE4AFF7B" blockId="42.[151,497,151,210]" box="[151,497,186,210]" pageId="42" pageNumber="43">
(
<figureCitation id="132C2A6B521BD403C09C2089FF46FF7B" box="[159,253,186,210]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="42" pageNumber="43">Figs. 37</figureCitation>
,
<figureCitation id="132C2A6B521BD403C10B2089FE9DFF7B" box="[264,294,186,210]" captionStart="FIGURES 38 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="42" pageNumber="43">55</figureCitation>
,
<figureCitation id="132C2A6B521BD403C1312089FEEAFF7B" box="[306,337,186,210]" captionStart="FIGURES 56 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="42" pageNumber="43">73</figureCitation>
,
<figureCitation id="132C2A6B521BD403C15F2089FEC1FF7B" box="[348,378,186,210]" captionStart="FIGURES 83 91" captionStartId="17.[151,264,1695,1718]" captionTargetBox="[168,1431,195,1670]" captionTargetId="figure@17.[168,1432,195,1670]" captionTargetPageId="17" captionText="FIGURES 83 91. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (83) F. l a m o s c a sp. nov., paratype; (84) F. magdalena sp. nov., paratype; (85) F. pijao sp. nov., paratype; (86) F. porce sp. nov., holotype; (87) F. p u s i o; (88) F. quimbaya sp. nov., paratype; (89) F. sumapaz sp. nov., paratype; (90) F. sutagao sp. nov., paratype; (91) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213953/files/figure.png" pageId="42" pageNumber="43">91</figureCitation>
,
<figureCitation id="132C2A6B521BD403C1852089FE09FF7B" box="[390,434,186,210]" captionStart="FIGURES 101 109" captionStartId="19.[151,264,1921,1944]" captionTargetBox="[186,1406,193,1902]" captionTargetId="figure@19.[183,1407,193,1903]" captionTargetPageId="19" captionText="FIGURES 101 109. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (101) F. l a m os c a sp. nov., paratype; (102) F. magdalena sp. nov., paratype; (103) F. pijao sp. nov., paratype; (104) F. porce sp. nov., holotype; (105) F. p us i o; (106) F. quimbaya sp. nov., partype; (107) F. sumapaz sp. nov., paratype; (108) F. sutagao sp. nov., paratype; (109) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213955/files/figure.png" pageId="42" pageNumber="43">109</figureCitation>
,
<figureCitation id="132C2A6B521BD403C1BD2089FE52FF7B" box="[446,489,186,210]" captionStart="FIGURES 118 127" captionStartId="21.[151,264,1599,1622]" captionTargetBox="[168,1429,193,1576]" captionTargetId="figure@21.[165,1429,193,1577]" captionTargetPageId="21" captionText="FIGURES 118 127. Fannia: male, hypandrium and associated structures, dorsal view: (118) F. katios sp. nov., paratype; (119) F. l a m o s c a sp. nov., paratype; (120) F. magdalena sp. nov., paratype; (121) F. pijao sp. nov., paratype; (122) F. po rc e sp. nov., holotype; (123) F. p u s i o; (124) F. quimbaya sp. nov., paratype; (125) F. sumapaz sp. nov., paratype; (126) F. sutagao sp. nov., paratype; (127) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213957/files/figure.png" pageId="42" pageNumber="43">127</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C0942130FA26FEFF" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BA836EE521BD403C0942130FA26FEFF" blockId="42.[151,1437,258,342]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="AB0711C0521BD403C0942130FA26FEFF" infered="true" pageId="42" pageNumber="43">
<taxonomicName id="4C174D6D521BD403C0942130FD6DFEB1" ID-CoL="6HPYZ" authority="Stein, 1898: 176" authorityName="Stein" authorityPageNumber="176" authorityYear="1898" box="[151,726,258,280]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C0942130FEDDFEB1" box="[151,358,258,280]" italics="true" pageId="42" pageNumber="43">Fannia trimaculata</emphasis>
<treatmentCitation id="0AB610FF521BD403C1742131FD8AFEB1" author="Stein" box="[375,561,258,280]" page="176" pageId="42" pageNumber="43" year="1898">
<bibRefCitation id="EF864B1F521BD403C1742131FE4CFEB1" author="Stein" box="[375,503,258,280]" pageId="42" pageNumber="43" refString="Stein, P. (1898) Nordamerikanische Anthomyiden. Beitrag zur Dipterenfauna der Vereinigten Staaten. Berliner entomologische Zeitschrift, 42 [1897], 161 - 288." type="journal article" year="1898">Stein, 1898</bibRefCitation>
: 176
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(
<emphasis id="B963EAFC521BD403C2492131FD76FEB1" box="[586,717,258,280]" italics="true" pageId="42" pageNumber="43">Homalomyia</emphasis>
)
</taxonomicName>
. Male
<typeStatus id="54AC884C521BD403C3332130FC33FEB1" box="[816,904,259,280]" pageId="42" pageNumber="43" type="syntype">syntypes</typeStatus>
deposited in the Museum für Naturkunde der HumboldtUniversität (ZMHB), Museum of Comparative Zoology (MCZ), Field Museum of Natural History (FMNH) and National Museum of Natural History (USNM)(
<treatmentCitation id="0AB610FF521BD403C2C82173FC7CFEFF" author="Pont" box="[715,967,320,342]" page="15" pageId="42" pageNumber="43" year="2006">
<bibRefCitation id="EF864B1F521BD403C2C82173FC7CFEFF" author="Pont" box="[715,967,320,342]" pageId="42" pageNumber="43" refString="Pont, A. C. &amp; Werner, D. (2006) The types of Fanniidae and Muscidae (Diptera) in the Museum fur Naturkunde, Humboldt- Universitat zu Berlin, Germany. Zoosystematics and Evolution, 82, 3 - 139." type="journal article" year="2006">Pont &amp; Werner 2006: 15</bibRefCitation>
</treatmentCitation>
). Type-localities: North
<collectingCountry id="F300767E521BD403C4BF2173FAA8FEFF" box="[1212,1299,320,342]" name="United States of America" pageId="42" pageNumber="43">America</collectingCountry>
and
<collectingCountry id="F300767E521BD403C5472173FA22FEFF" box="[1348,1433,320,342]" name="Jamaica" pageId="42" pageNumber="43">Jamaica</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C09421B1FBACFC57" pageId="42" pageNumber="43" type="diagnosis">
<paragraph id="8BA836EE521BD403C09421B1FBACFC57" blockId="42.[151,1437,386,1670]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C09421B1FEAAFE32" bold="true" box="[151,273,386,411]" pageId="42" pageNumber="43">Diagnosis.</emphasis>
these characters apply only to the male sex. General coloration black; eye bare; frontal vitta black to dark brown, golden-silver pollinose, anterior margin brownish or black; 1011
<emphasis id="B963EAFC521BD403C43D2195FBEBFE17" box="[1086,1104,422,446]" italics="true" pageId="42" pageNumber="43">fr</emphasis>
; fronto-orbital plate greyish pollinose and posterior margin black; parafacial with short setae; parafacial and gena greyish pollinose; scape and pedicel dark brown greyish pollinose; arista dark brown; palpus dark brown, filiform and slender; thorax dark-grey; scutum dark brown to black up to second pair of postsutural
<emphasis id="B963EAFC521BD403C33A2220FCEFFD82" box="[825,852,531,555]" italics="true" pageId="42" pageNumber="43">dc</emphasis>
and grey up to intra-alar area and base of scutellum, sometimes with 2 grey or brown vittae, weak or visible between
<emphasis id="B963EAFC521BD403C383220BFC1EFDE6" box="[896,933,568,591]" italics="true" pageId="42" pageNumber="43">acr</emphasis>
and
<emphasis id="B963EAFC521BD403C3DC2205FC42FDE7" box="[991,1017,566,590]" italics="true" pageId="42" pageNumber="43">dc</emphasis>
;
<emphasis id="B963EAFC521BD403C408220BFB8BFDE6" box="[1035,1072,568,591]" italics="true" pageId="42" pageNumber="43">acr</emphasis>
3:34, in unordered series; 34 long
<emphasis id="B963EAFC521BD403C0D7226FFEB3FDDA" box="[212,264,604,627]" italics="true" pageId="42" pageNumber="43">pprn</emphasis>
; 2 short and strong
<emphasis id="B963EAFC521BD403C1F8226FFD99FDDA" box="[507,546,604,627]" italics="true" pageId="42" pageNumber="43">pra</emphasis>
, near
<emphasis id="B963EAFC521BD403C26D2268FD26FDDA" box="[622,669,603,627]" italics="true" pageId="42" pageNumber="43">spal</emphasis>
, disposed asymmetrically in some specimens; calypters and wing yellowish; haltere whitish to yellowish; legs dark brown; mid femur with 3 short
<emphasis id="B963EAFC521BD403C42922B3FBFEFD3E" box="[1066,1093,640,663]" italics="true" pageId="42" pageNumber="43">av</emphasis>
on basal third, 2 on mid third, and apical third with a set of short and strong setae decreasing in size towards apex, 1 row of sparse
<emphasis id="B963EAFC521BD403C4F42297FAABFD12" box="[1271,1296,676,699]" italics="true" pageId="42" pageNumber="43">pv</emphasis>
, apical third with a set of short strong setae; hind coxa with 23
<emphasis id="B963EAFC521BD403C2D222FBFD64FD76" box="[721,735,712,735]" italics="true" pageId="42" pageNumber="43">p</emphasis>
; hind femur with 1 strong
<emphasis id="B963EAFC521BD403C40422FBFB9AFD76" box="[1031,1057,712,735]" italics="true" pageId="42" pageNumber="43">av</emphasis>
on apical third, median third with 2 rows of short and straight setae (
<figureCitation id="132C2A6B521BD403C22522D9FDC0FCAB" box="[550,635,746,770]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="42" pageNumber="43">Fig. 37</figureCitation>
), posteroventral surface on apical third with weak protuberance and 1 row of
<emphasis id="B963EAFC521BD403C0E92323FEBEFC8E" box="[234,261,784,807]" italics="true" pageId="42" pageNumber="43">pv</emphasis>
that increase in length towards the apex, 34 setae inserted on protuberance (
<figureCitation id="132C2A6B521BD403C475233DFB71FC8F" box="[1142,1226,782,806]" captionStart="FIGURES 38 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="42" pageNumber="43">Fig. 55</figureCitation>
); hind tibia with 1 median
<emphasis id="B963EAFC521BD403C0F12307FEB0FCE2" box="[242,267,820,843]" italics="true" pageId="42" pageNumber="43">av</emphasis>
, 1 row of short
<emphasis id="B963EAFC521BD403C1C72307FE69FCE2" box="[452,466,820,843]" italics="true" pageId="42" pageNumber="43">a</emphasis>
ending in 1 longer pre-apical (
<figureCitation id="132C2A6B521BD403C3362301FC32FCE3" box="[821,905,818,842]" captionStart="FIGURES 20 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="42" pageNumber="43">Fig. 37</figureCitation>
), 1 median
<emphasis id="B963EAFC521BD403C4122300FBA4FCE2" box="[1041,1055,819,843]" italics="true" pageId="42" pageNumber="43">d</emphasis>
and 1 developed pre-apical; hind tarsus on first and second tarsomeres with a strong ventral seta; abdomen trimaculate; sternite 1 bare; sternite 5 as in
<figureCitation id="132C2A6B521BD403C0B02349FEBDFC3B" box="[179,262,890,914]" captionStart="FIGURES 56 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="42" pageNumber="43">Fig. 73</figureCitation>
; terminalia (
<figureCitation id="132C2A6B521BD403C1912349FE54FC3B" box="[402,495,890,914]" captionStart="FIGURES 83 91" captionStartId="17.[151,264,1695,1718]" captionTargetBox="[168,1431,195,1670]" captionTargetId="figure@17.[168,1432,195,1670]" captionTargetPageId="17" captionText="FIGURES 83 91. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (83) F. l a m o s c a sp. nov., paratype; (84) F. magdalena sp. nov., paratype; (85) F. pijao sp. nov., paratype; (86) F. porce sp. nov., holotype; (87) F. p u s i o; (88) F. quimbaya sp. nov., paratype; (89) F. sumapaz sp. nov., paratype; (90) F. sutagao sp. nov., paratype; (91) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213953/files/figure.png" pageId="42" pageNumber="43">Figs. 91</figureCitation>
,
<figureCitation id="132C2A6B521BD403C1F82349FD92FC3B" box="[507,553,890,914]" captionStart="FIGURES 101 109" captionStartId="19.[151,264,1921,1944]" captionTargetBox="[186,1406,193,1902]" captionTargetId="figure@19.[183,1407,193,1903]" captionTargetPageId="19" captionText="FIGURES 101 109. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (101) F. l a m os c a sp. nov., paratype; (102) F. magdalena sp. nov., paratype; (103) F. pijao sp. nov., paratype; (104) F. porce sp. nov., holotype; (105) F. p us i o; (106) F. quimbaya sp. nov., partype; (107) F. sumapaz sp. nov., paratype; (108) F. sutagao sp. nov., paratype; (109) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213955/files/figure.png" pageId="42" pageNumber="43">109</figureCitation>
): epandrium wider than long, with setae of differing lengths; cercal plate fused only apically, with short and strong setae; surstylus straight and slightly pointed apically, with 2 strong pre-apical and setulose setae on inner basal region, with a short latero-ventral projection and with short strong setae; bacilliform process absent; hypandrium and associated structures as in
<figureCitation id="132C2A6B521BD403C3B123D5FBA9FC57" box="[946,1042,998,1022]" captionStart="FIGURES 118 127" captionStartId="21.[151,264,1599,1622]" captionTargetBox="[168,1429,193,1576]" captionTargetId="figure@21.[165,1429,193,1577]" captionTargetPageId="21" captionText="FIGURES 118 127. Fannia: male, hypandrium and associated structures, dorsal view: (118) F. katios sp. nov., paratype; (119) F. l a m o s c a sp. nov., paratype; (120) F. magdalena sp. nov., paratype; (121) F. pijao sp. nov., paratype; (122) F. po rc e sp. nov., holotype; (123) F. p u s i o; (124) F. quimbaya sp. nov., paratype; (125) F. sumapaz sp. nov., paratype; (126) F. sutagao sp. nov., paratype; (127) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213957/files/figure.png" pageId="42" pageNumber="43">Fig. 127</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C0C42439FB1BFB1B" pageId="42" pageNumber="43" type="biology_ecology">
<paragraph id="8BA836EE521BD403C0C42439FB1BFB1B" blockId="42.[151,1437,386,1670]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C0C42439FE9CFB8A" bold="true" box="[199,295,1034,1059]" pageId="42" pageNumber="43">Biology.</emphasis>
<taxonomicName id="4C174D6D521BD403C12D243FFE76FB8A" box="[302,461,1035,1059]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C12D243FFE76FB8A" box="[302,461,1035,1059]" italics="true" pageId="42" pageNumber="43">F. trimaculata</emphasis>
</taxonomicName>
is associated with forest areas (de
<bibRefCitation id="EF864B1F521BD403C3542439FBD8FB8B" author="Carvalho" box="[855,1123,1034,1058]" pageId="42" pageNumber="43" refString="de Carvalho, C. J. B. &amp; Couri, M. S (1991) Muscidae, Fanniidae, e Calliphoridae (Diptera) do Projeto Marac, Roraima, Brazil. Acta Amazonica, 21, 35 - 43." type="journal article" year="1991">Carvalho &amp; Couri 1991</bibRefCitation>
) and is found in abundance on the dung of confined birds. For this reason, research on possible
<taxonomicName id="4C174D6D521BD403C39E2403FB86FBEF" box="[925,1085,1070,1095]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C39E2403FB86FBEF" box="[925,1085,1070,1095]" italics="true" pageId="42" pageNumber="43">F. trimaculata</emphasis>
</taxonomicName>
predators, such as histerid and staphilinid beetles that develop in the same substrate (
<bibRefCitation id="EF864B1F521BD403C3092461FC60FBC3" author="Bruno" box="[778,987,1106,1131]" pageId="42" pageNumber="43" refString="Bruno, T. V., Guimaraes, J. H., dos Santos,. M. M. &amp; Tucci, E. C. (1993) Moscas sinantropicas (Diptera) e seus predadores que se criam em esterco de aves poedeiras confinadas, no estado de Sao Paulo, Brazil. Revista Brasileira de Entomologia, 37, 577 - 590." type="journal article" year="1993">
Bruno
<emphasis id="B963EAFC521BD403C3582467FC2CFBC2" box="[859,919,1107,1131]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
1993
</bibRefCitation>
;
<bibRefCitation id="EF864B1F521BD403C3E92460FAA8FBC3" author="Lomonaco" box="[1002,1299,1106,1131]" pageId="42" pageNumber="43" refString="Lomonaco, C. &amp; Prado, A. P. (1994) Estrutura comunitaria e dinamica populacional da fauna de dipteros e seus inimigos naturais em granjas avicolas. Anais da Sociedade Entomolgica Brasileira, 23, 71 - 79." type="journal article" year="1994">Lomônaco &amp; Prado 1994</bibRefCitation>
), is needed.
<bibRefCitation id="EF864B1F521BD403C0942445FEC6FB27" author="Albuquerque" box="[151,381,1142,1166]" pageId="42" pageNumber="43" refString="Albuquerque, D. de O. (1945) Sobre Fannia trimaculata (Stein, 1897) Malloch, 1913 (Diptera, Muscidae). Boletim do Museu Nacional Zoologia, 34, 1 - 16." type="journal article" year="1945">Albuquerque (1945)</bibRefCitation>
presented data on the biology, morphology and distribution of the species from specimens reared in the laboratory, but the data he presented was for
<taxonomicName id="4C174D6D521BD403C32324AFFCC2FB1A" box="[800,889,1180,1203]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="B963EAFC521BD403C32324AFFCC2FB1A" box="[800,889,1180,1203]" italics="true" pageId="42" pageNumber="43">F. pusio</emphasis>
</taxonomicName>
(de
<bibRefCitation id="EF864B1F521BD403C3A924A9FB2FFB1B" author="Carvalho" box="[938,1172,1178,1203]" pageId="42" pageNumber="43" refString="de Carvalho, C. J. B., Pont A. C., Couri, M. S. &amp; Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32." type="journal article" year="2003">
Carvalho
<emphasis id="B963EAFC521BD403C41B24AFFBEBFB1A" box="[1048,1104,1179,1203]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2003
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C0C4248DFE8FFACF" pageId="42" pageNumber="43" type="discussion">
<paragraph id="8BA836EE521BD403C0C4248DFE8FFACF" blockId="42.[151,1437,386,1670]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C0C4248DFEF4FB7E" bold="true" box="[199,335,1214,1239]" pageId="42" pageNumber="43">Comments.</emphasis>
<taxonomicName id="4C174D6D521BD403C155248CFD89FB7F" box="[342,562,1214,1238]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C155248CFD89FB7F" box="[342,562,1214,1238]" italics="true" pageId="42" pageNumber="43">Fannia trimaculata</emphasis>
</taxonomicName>
is newly recorded in
<collectingCountry id="F300767E521BD403C324248DFC22FB7F" box="[807,921,1214,1238]" name="Colombia" pageId="42" pageNumber="43">Colombia</collectingCountry>
. It is close to
<taxonomicName id="4C174D6D521BD403C43624F3FAB9FB7F" box="[1077,1282,1214,1239]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculatoides">
<emphasis id="B963EAFC521BD403C43624F3FAB9FB7F" box="[1077,1282,1214,1239]" italics="true" pageId="42" pageNumber="43">F. trimaculatoides</emphasis>
</taxonomicName>
, but males of both species are distinguished by the hind femur, setulose in the middle of ventral surface in
<taxonomicName id="4C174D6D521BD403C49C24D7FA86FB52" box="[1183,1341,1251,1275]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C49C24D7FA86FB52" box="[1183,1341,1251,1275]" italics="true" pageId="42" pageNumber="43">F. trimaculata</emphasis>
</taxonomicName>
.
<taxonomicName id="4C174D6D521BD403C54824D7FEA1FAB7" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="trimaculata">
<emphasis id="B963EAFC521BD403C54824D7FEA1FAB7" italics="true" pageId="42" pageNumber="43">Fannia trimaculata</emphasis>
</taxonomicName>
is close to
<taxonomicName id="4C174D6D521BD403C19A253BFDAAFAB7" box="[409,529,1286,1311]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="howardi">
<emphasis id="B963EAFC521BD403C19A253BFDAAFAB7" box="[409,529,1286,1311]" italics="true" pageId="42" pageNumber="43">F. howardi</emphasis>
</taxonomicName>
, but can be easily recognized by the hind femur with one row of 7-8
<emphasis id="B963EAFC521BD403C51E253BFA83FAB6" box="[1309,1336,1288,1311]" italics="true" pageId="42" pageNumber="43">av</emphasis>
on basal third and apical third, bare basally, except for one strong seta on apex (
<bibRefCitation id="EF864B1F521BD403C3B32519FB8DFAEB" author="Seago" box="[944,1078,1322,1346]" pageId="42" pageNumber="43" refString="Seago, J. M. (1954) The Pusio Group of the genus Fannia Robineau - Desvoidy, with descriptions of the new species. American Museum Novittates, 1699, 1 - 14." type="journal article" year="1954">Seago 1954</bibRefCitation>
) (these characters apply only to the male sex).
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C0C42540FD53F9B3" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BA836EE521BD403C0C42540FD9AFA23" blockId="42.[151,1437,386,1670]" box="[199,545,1394,1419]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C0C42540FED6FA22" bold="true" box="[199,365,1394,1419]" pageId="42" pageNumber="43">
<typeStatus id="54AC884C521BD403C0C42540FEB9FA22" box="[199,258,1395,1419]" pageId="42" pageNumber="43">Type</typeStatus>
material
</emphasis>
. Not examined.
</paragraph>
<paragraph id="8BA836EE521BD403C0C425A5FD53F9B3" blockId="42.[151,1437,386,1670]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C0C425A5FE0AFA06" bold="true" box="[199,433,1430,1455]" pageId="42" pageNumber="43">Material examined.</emphasis>
<collectingCountry id="F300767E521BD403C1BB25A5FDEAFA07" box="[440,593,1430,1454]" name="Colombia" pageId="42" pageNumber="43">COLOMBIA</collectingCountry>
. Antioquia:
<specimenCount id="9D11FD67521BD403C2D925A5FC8BFA07" box="[730,816,1430,1454]" pageId="42" pageNumber="43" type="male">8 males</specimenCount>
Medellín, U de A, En cadáver Curi, 9:00,
<date id="FFA9102E521BD403C51225A5FA23FA07" box="[1297,1432,1430,1454]" pageId="42" pageNumber="43" value="2000-12-01">1 Dic. 2000</date>
, P. Duque,
<taxonomicName id="4C174D6D521BD403C10C2589FE3BFA7B" box="[271,384,1466,1490]" class="Insecta" family="Fanniidae" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Fanniidae</taxonomicName>
,
<taxonomicName id="4C174D6D521BD403C18D258FFE64FA7A" box="[398,479,1468,1491]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC521BD403C18D258FFE64FA7A" box="[398,479,1468,1491]" italics="true" pageId="42" pageNumber="43">Fannia</emphasis>
</taxonomicName>
sp., M. Wolff (
<collectionCode id="ED06AE2B521BD403C2972589FD5DFA7B" box="[660,742,1466,1490]" pageId="42" pageNumber="43">CEUA</collectionCode>
),
<specimenCount id="9D11FD67521BD403C2F82589FCF2FA7B" box="[763,841,1466,1490]" pageId="42" pageNumber="43" type="male">1 male</specimenCount>
Medellín, Universidad de Antioquia, Jameo, Cerdo forense,
<date id="FFA9102E521BD403C0F825EDFEEBFA5F" box="[251,336,1502,1526]" pageId="42" pageNumber="43">Junio 9</date>
de 1999, A. Uribe Toro,
<taxonomicName id="4C174D6D521BD403C28325EDFD56FA5F" box="[640,749,1502,1526]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="family">Muscidae</taxonomicName>
543 (
<collectionCode id="ED06AE2B521BD403C33025EDFC3EFA5F" box="[819,901,1502,1526]" pageId="42" pageNumber="43">CEUA</collectionCode>
),
<specimenCount id="9D11FD67521BD403C39F25EDFC50FA5F" box="[924,1003,1502,1526]" pageId="42" pageNumber="43" type="male">1 male</specimenCount>
Medellín, RamaJama, 1.480 msnm, Fecha
<date id="FFA9102E521BD403C0E22631FEFCF9B3" box="[225,327,1538,1562]" pageId="42" pageNumber="43" value="2002-01-25">02/25/01</date>
,
<collectionCode id="ED06AE2B521BD403C1512631FE25F9B3" box="[338,414,1538,1562]" pageId="42" pageNumber="43">CEUA</collectionCode>
13341, M. Castaño (
<collectionCode id="ED06AE2B521BD403C28A2631FD67F9B3" box="[649,732,1538,1562]" pageId="42" pageNumber="43">CEUA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D6565521BD403C0C42615FD8BF92F" pageId="42" pageNumber="43" type="distribution">
<paragraph id="8BA836EE521BD403C0C42615FD8BF92F" blockId="42.[151,1437,386,1670]" pageId="42" pageNumber="43">
<emphasis id="B963EAFC521BD403C0C42615FEDBF996" bold="true" box="[199,352,1574,1599]" pageId="42" pageNumber="43">Distribution.</emphasis>
<collectingCountry id="F300767E521BD403C1692615FE0FF997" box="[362,436,1574,1598]" name="Belize" pageId="42" pageNumber="43">Belize</collectingCountry>
,
<collectingCountry id="F300767E521BD403C1C12614FDA5F997" box="[450,542,1575,1598]" name="Panama" pageId="42" pageNumber="43">Panama</collectingCountry>
,
<collectingCountry id="F300767E521BD403C22F2614FD0CF997" box="[556,695,1574,1598]" name="Puerto Rico" pageId="42" pageNumber="43">Puerto Rico</collectingCountry>
,
<collectingCountry id="F300767E521BD403C2C62615FC99F997" box="[709,802,1574,1598]" name="Jamaica" pageId="42" pageNumber="43">Jamaica</collectingCountry>
,
<collectingCountry id="F300767E521BD403C3332615FBA5F997" box="[816,1054,1574,1598]" name="Dominican Republic" pageId="42" pageNumber="43">Dominican Republic</collectingCountry>
,
<collectingCountry id="F300767E521BD403C42F2615FBDCF997" box="[1068,1127,1574,1598]" name="Haiti" pageId="42" pageNumber="43">Haiti</collectingCountry>
, Santo
<collectingCountry id="F300767E521BD403C4BD2615FA90F997" box="[1214,1323,1574,1598]" name="Dominican Republic" pageId="42" pageNumber="43">Domingo</collectingCountry>
,
<collectingCountry id="F300767E521BD403C53A2615FA23F997" box="[1337,1432,1574,1598]" name="Ecuador" pageId="42" pageNumber="43">Ecuador</collectingCountry>
,
<collectingCountry id="F300767E521BD403C0942678FF76F9CB" box="[151,205,1611,1634]" name="Peru" pageId="42" pageNumber="43">Peru</collectingCountry>
,
<collectingCountry id="F300767E521BD403C0DC2679FEEEF9CB" box="[223,341,1610,1634]" name="Venezuela" pageId="42" pageNumber="43">Venezuela</collectingCountry>
,
<collectingCountry id="F300767E521BD403C1652679FE16F9CB" box="[358,429,1610,1634]" name="Brazil" pageId="42" pageNumber="43">Brazil</collectingCountry>
,
<collectingCountry id="F300767E521BD403C1BC2678FD99F9CA" box="[447,546,1611,1635]" name="Uruguay" pageId="42" pageNumber="43">Uruguay</collectingCountry>
,
<collectingCountry id="F300767E521BD403C2372679FD18F9CB" box="[564,675,1610,1634]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
(de
<bibRefCitation id="EF864B1F521BD403C2E12679FC64F9CB" author="Carvalho" box="[738,991,1610,1635]" pageId="42" pageNumber="43" refString="de Carvalho, C. J. B., Pont A. C., Couri, M. S. &amp; Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32." type="journal article" year="2003">
Carvalho
<emphasis id="B963EAFC521BD403C355267FFC2EF9CA" box="[854,917,1611,1635]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
2003
</bibRefCitation>
).
<collectingCountry id="F300767E521BD403C3FB2679FBD0F9CB" box="[1016,1131,1610,1634]" name="Colombia" pageId="42" pageNumber="43">Colombia</collectingCountry>
: department of Antioquia (Andean region, Central Cordillera).
</paragraph>
</subSubSection>
</treatment>
</document>