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<document ID-DOI="http://doi.org/10.5281/zenodo.7501705" ID-ISSN="1211-87 8 8" ID-Zenodo-Dep="7501705" checkinTime="1672754391430" checkinUser="diego" docAuthor="Löbl, Ivan" docDate="2022" docId="03BE8788FFBCC75303C76D17DDD75121" docLanguage="en" docName="ActaMusMoraviae.107.1-2.7-19.pdf" docOrigin="Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2)" docTitle="Scaphoxium sinuatum Löbl 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="FF87FFF0FFB6C75802466814DF5C556E" masterDocTitle="Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="19" masterPageNumber="7" pageNumber="17" updateTime="1672761276865" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Löbl, Ivan</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Musei Moraviae, Scientiae biologicae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>107</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1 - 2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>7</mods:start>
<mods:end>19</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.7501705</mods:identifier>
<mods:identifier type="ISSN">1211-8788</mods:identifier>
<mods:identifier type="Zenodo-Dep">7501705</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7501633" ID-Zenodo-Dep="7501633" LSID="urn:lsid:plazi:treatment:03BE8788FFBCC75303C76D17DDD75121" httpUri="http://treatment.plazi.org/id/03BE8788FFBCC75303C76D17DDD75121" lastPageId="11" lastPageNumber="18" pageId="10" pageNumber="17">
<subSubSection box="[385,729,1283,1308]" pageId="10" pageNumber="17" type="nomenclature">
<paragraph blockId="10.[385,729,1283,1308]" box="[385,729,1283,1308]" pageId="10" pageNumber="17">
<heading bold="true" box="[385,729,1283,1308]" fontSize="10" level="2" pageId="10" pageNumber="17" reason="2">
<taxonomicName authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[385,634,1283,1308]" class="Insecta" family="Staphylinidae" genus="Scaphoxium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="17" phylum="Arthropoda" rank="species" species="sinuatum" status="sp. nov.">Scaphoxium sinuatum</taxonomicName>
<taxonomicNameLabel box="[641,729,1284,1308]" pageId="10" pageNumber="17" rank="species">
<emphasis bold="true" box="[641,721,1284,1308]" pageId="10" pageNumber="17">sp. nov</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1100,1219,1284,1308]" pageId="10" pageNumber="17" type="description">
<paragraph blockId="10.[1100,1219,1284,1308]" box="[1100,1219,1284,1308]" pageId="10" pageNumber="17">
<figureCitation box="[1100,1178,1284,1308]" captionStart="Figs 8-13. 8-10" captionStartId="6.[332,370,1715,1734]" captionTargetBox="[355,1235,431,1586]" captionTargetId="figure-89@6.[345,1267,390,1595]" captionTargetPageId="6" captionText="Figs 8-13. 8-10 ̅ Pseudobironium rapax sp. nov.; 8, 9 ̅ Aedeagus in dorsal and lateral views, scale = 0.1 mm; 10 ̅ Internal sac in dorsal view, scale = 0.1 mm. 11-13 ̅ Scaphoxium sinuatum sp. nov.; 11 ̅ Aedeagus in dorsal view, scale = 0.1 mm; 12 ̅ Apical part of paramere, dorsal view, scale = 0.05 mm; 13 ̅ Internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7501709" httpUri="https://zenodo.org/record/7501709/files/figure.png" pageId="10" pageNumber="17">Figs 11</figureCitation>
̅13
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="17" type="materials_examined">
<paragraph blockId="10.[332,1279,1330,1376]" pageId="10" pageNumber="17">
<materialsCitation collectingDate="1996-07-01" collectionCode="MHNG" country="Malaysia" location="Batu Punggul Resort" pageId="10" pageNumber="17" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis bold="true" box="[332,464,1330,1350]" pageId="10" pageNumber="17">Type material.</emphasis>
<typeStatus box="[468,547,1331,1350]" pageId="10" pageNumber="17">Holotype</typeStatus>
<specimenCount box="[553,597,1331,1350]" pageId="10" pageNumber="17" type="male">male</specimenCount>
,
<collectingCountry box="[604,684,1331,1350]" name="Malaysia" pageId="10" pageNumber="17" value="Malaysia">Malaysia</collectingCountry>
,
<collectingRegion box="[692,746,1331,1350]" country="Malaysia" name="Sabah" pageId="10" pageNumber="17">Sabah</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03BE8788FFBCC75303C76D17DDD75121:8EC86045FFBCC75200B56D27DCF85028" box="[755,932,1331,1350]" country="Malaysia" name="Batu Punggul Resort" pageId="10" pageNumber="17" stateProvince="Sabah">Batu Punggul Resort</location>
env.,
<date box="[981,1037,1331,1350]" pageId="10" pageNumber="17">24.VI.</date>
̅
<date box="[1048,1142,1331,1350]" pageId="10" pageNumber="17" value="1996-07-01">
<collectingDate box="[1048,1142,1331,1350]" pageId="10" pageNumber="17" value="1996-07-01">1.VII.1996</collectingDate>
</date>
, 11c. vegetation debris and forest floor litter accumulated around large trees near river (
<collectionCode box="[942,1012,1356,1375]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="17" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="18" pageId="10" pageNumber="17" type="description">
<paragraph blockId="10.[332,1280,1397,1773]" lastBlockId="11.[304,1252,393,1104]" lastPageId="11" lastPageNumber="18" pageId="10" pageNumber="17">
<emphasis bold="true" box="[332,471,1397,1421]" pageId="10" pageNumber="17">Description.</emphasis>
Length
<quantity box="[560,658,1397,1421]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" pageId="10" pageNumber="17" unit="mm" value="1.18">1.18 mm</quantity>
, width
<quantity box="[737,835,1397,1421]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="17" unit="mm" value="0.55">0.55 mm</quantity>
, dorsoventral diameter
<quantity box="[1087,1185,1397,1421]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="10" pageNumber="17" unit="mm" value="0.62">0.62 mm</quantity>
. Thorax, elytra and most of abdomen uniformly reddish­brown, head somewhat lighter, apical abdominal segments light, nearly yellowish, femora and tibiae roughly as thorax, tarsi lighter, antennae light brown. Length/width ratios of antennomeres as follows:
<date box="[1190,1253,1493,1517]" pageId="10" pageNumber="17" value="2022-03">III 22</date>
/5:
<date box="[332,393,1525,1549]" pageId="10" pageNumber="17" value="2017-04">IV 17</date>
/4:
<date box="[426,480,1525,1549]" pageId="10" pageNumber="17" value="2022-05">V 22</date>
/4:
<date box="[511,574,1525,1549]" pageId="10" pageNumber="17" value="1923-06">VI 23</date>
/4:
<date box="[606,678,1525,1549]" pageId="10" pageNumber="17" value="1928-07">VII 28</date>
/7:
<date box="[710,791,1525,1549]" pageId="10" pageNumber="17" value="2020-08">VIII 20</date>
/7:
<date box="[823,920,1525,1549]" pageId="10" pageNumber="17" value="2011-09-28">IX 28/11</date>
: X: 28/11:
<date box="[1036,1098,1525,1549]" pageId="10" pageNumber="17" value="1938-11">XI 38</date>
/12. Pronotal and elytral punctation similar, very fine, hardly visible at 40 times magnification. Scutellum concealed. Elytra with sutural striae parallel, shortened, starting about
<quantity box="[1083,1177,1589,1613]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="17" unit="mm" value="0.15">0.15 mm</quantity>
posterior of margin of pronotal lobe, adsutural areas hardly raised. Hypomeron with scattered, very fine punctured and short oblique stria. Mesoventrite with mesal impression shallow, narrowed posteriad, lacking mesal ridge and lacking microsculpture. Margin of mesocoxal process rounded. Metaventrite lacking microsculpture, very finely punctate, convex in middle. Lateral stria oblique, nearly reaching convex, finely punctate submesocoxal line. Submesocoxal area about
<quantity box="[808,909,393,417]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="18" unit="mm" value="0.03">0.03 mm</quantity>
, as long as half of the shortest interval to margin of metacoxa. Metanepisternum flat, with suture fairly narrow, punctate, somewhat rounded. Abdomen very finely punctate, median areas of ventrites with punctulate microsculpture.
</paragraph>
<paragraph blockId="11.[304,1252,393,1104]" pageId="11" pageNumber="18">
<emphasis box="[304,359,521,545]" italics="true" pageId="11" pageNumber="18">Male</emphasis>
. Segments Ito III of protarsi hardly widened. Aedeagus (
<figureCitation box="[963,1042,521,545]" captionStart="Figs 8-13. 8-10" captionStartId="6.[332,370,1715,1734]" captionTargetBox="[355,1235,431,1586]" captionTargetId="figure-89@6.[345,1267,390,1595]" captionTargetPageId="6" captionText="Figs 8-13. 8-10 ̅ Pseudobironium rapax sp. nov.; 8, 9 ̅ Aedeagus in dorsal and lateral views, scale = 0.1 mm; 10 ̅ Internal sac in dorsal view, scale = 0.1 mm. 11-13 ̅ Scaphoxium sinuatum sp. nov.; 11 ̅ Aedeagus in dorsal view, scale = 0.1 mm; 12 ̅ Apical part of paramere, dorsal view, scale = 0.05 mm; 13 ̅ Internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.7501709" httpUri="https://zenodo.org/record/7501709/files/figure.png" pageId="11" pageNumber="18">Figs 11</figureCitation>
̅13)
<quantity box="[1097,1190,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="4.4" pageId="11" pageNumber="18" unit="mm" value="0.44">0.44 mm</quantity>
long. Parameres narrowed posterior of basal section, gradually weakly widened toward narrow apical section, not forming subapical lobe or process; outer margins of parameres concave and inner margins strongly sclerotized posterior of narrowed apical section. Internal sac bearing two pairs of sclerotized pieces, outer larger and more sclerotized than inner pair, expanded apically, inner pair widened proximally, with margins becoming indistinct apically; scale­like, spine­like or filamentous structures absent.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="18" type="etymology">
<paragraph blockId="11.[304,1252,393,1104]" pageId="11" pageNumber="18">
<emphasis bold="true" box="[304,432,753,777]" pageId="11" pageNumber="18">Etymology.</emphasis>
The species epithet is a Latin adjective meaning &quot;sinuatȅ, referring to the outer margins of the parameres.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="18" type="diagnosis">
<paragraph blockId="11.[304,1252,393,1104]" pageId="11" pageNumber="18">
<emphasis bold="true" box="[304,557,824,848]" pageId="11" pageNumber="18">Differential diagnosis.</emphasis>
The body­size of this new species is significantly smaller than that of other Bornean species and the shape of its parameres is diagnostic for it. The new species is similar to
<taxonomicName authority="Lobl, 2015" authorityName="Lobl" authorityYear="2015" box="[531,776,888,912]" class="Insecta" family="Staphylinidae" genus="Scaphoxium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="18" phylum="Arthropoda" rank="species" species="bilobum">
<emphasis box="[531,647,888,912]" italics="true" pageId="11" pageNumber="18">S. bilobum</emphasis>
<bibRefCitation author="LOBL I." box="[657,776,888,912]" journalOrPublisher="Revue suisse de zoologie" pageId="11" pageNumber="18" pagination="75" part="122" refId="ref6862" refString="LOBL I. 2015: On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands. Revue suisse de zoologie 122 (1): 75 ‾ 120." title="On the Scaphidiinae (Coleoptera: Staphylinidae) of the Lesser Sunda Islands" type="journal article" year="2015">Löbl, 2015</bibRefCitation>
</taxonomicName>
from Lombok. It shares with the latter the simple internal sac of the aedeagus lacking spine­like, denticulate or striate membranous structures in combination with the presence of two pairs of admesal sclerites. However, the outer pair of sclerites is much more expanded and the inner sclerites are much shorter and broader in
<taxonomicName authorityName="Löbl" authorityYear="2022" box="[463,588,1016,1040]" class="Insecta" family="Staphylinidae" genus="Scaphoxium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="18" phylum="Arthropoda" rank="species" species="sinuatum">
<emphasis box="[463,588,1016,1040]" italics="true" pageId="11" pageNumber="18">S. sinuatum</emphasis>
</taxonomicName>
, which may be distinguished from
<taxonomicName authorityName="Lobl" authorityYear="2015" box="[962,1078,1016,1040]" class="Insecta" family="Staphylinidae" genus="Scaphoxium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="18" phylum="Arthropoda" rank="species" species="bilobum">
<emphasis box="[962,1078,1016,1040]" italics="true" pageId="11" pageNumber="18">B. bilobum</emphasis>
</taxonomicName>
also by its wide impression of the mesoventrite, finer punctation of the lateral parts of the metaventrite and shorter submesocoxal areas.
</paragraph>
</subSubSection>
</treatment>
</document>