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<document id="8FA690E672441368A3FD26C6DEF3B5AB" ID-DOI="10.5281/zenodo.5429936" ID-ISSN="0253-116X" ID-Zenodo-Dep="5429936" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630422064935" checkinUser="valdenar" docAuthor="Zettel, H." docDate="2008" docId="03BA77072E2A3847A8F0E5B60C79FEC8" docLanguage="en" docName="LinzerbiolBeitr.2008.40.1.891-895.pdf" docOrigin="Linzer biologische Beiträge 40 (1)" docSource="http://antcat.org/documents/4783/Zettel_2008_Linzer_Biol_Beitr.pdf" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Recurvidris nigrans Zettel 2008, nov.sp." docType="treatment" docVersion="3" lastPageNumber="893" masterDocId="FF830F7F2E2B3844A917E13C0866FFAD" masterDocTitle="On the ants (Hymenoptera: Formicidae) of the Philippine Islands: III. The genus Recurvidris BOLTON 1992" masterLastPageNumber="895" masterPageNumber="891" pageNumber="892" updateTime="1699264472653" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="FC5F00D1CE53DE6CD9876C08A4F99AFF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BAC5B581CFEC69F3BBC6BAF470279CA3">On the ants (Hymenoptera: Formicidae) of the Philippine Islands: III. The genus Recurvidris BOLTON 1992</mods:title>
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<mods:namePart id="1E5924858FB0C1A0611E7FA1AB8DB197">Zettel, H.</mods:namePart>
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<mods:title id="98D0F568F2CF99EC2C33067262EDC19B">Linzer biologische Beiträge</mods:title>
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<mods:date id="C3B970F4DF1A60776C37FFCE7E0DD521">2008</mods:date>
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<mods:number id="9AFACF4BD5CCB8C5F4A911FF024BE62D">2008-07-10</mods:number>
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<mods:number id="AC23F95FC588E9DD146A264296E30FFC">40</mods:number>
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<mods:url id="8A155EB5BB44CD516F20D84FD425D93B">http://antcat.org/documents/4783/Zettel_2008_Linzer_Biol_Beitr.pdf</mods:url>
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<mods:identifier id="000023CB49525C28786BA3A457ACAD08" type="DOI">10.5281/zenodo.5429936</mods:identifier>
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<treatment id="03BA77072E2A3847A8F0E5B60C79FEC8" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8" httpUri="http://treatment.plazi.org/id/03BA77072E2A3847A8F0E5B60C79FEC8" lastPageId="3" lastPageNumber="893" pageId="1" pageNumber="892">
<subSubSection id="C309959A2E2A3845A8F0E5B60BA8FB5D" pageId="1" pageNumber="892" type="nomenclature">
<paragraph id="8BACC6112E2A3845A8F0E5B60B6FFB0C" blockId="1.[487,777,1162,1185]" box="[487,777,1162,1185]" pageId="1" pageNumber="892">
<heading id="D0E4717D2E2A3845A8F0E5B60B6FFB0C" bold="true" box="[487,777,1162,1185]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="892" reason="3">
<emphasis id="B9671A032E2A3845A8F0E5B60B6FFB0C" bold="true" box="[487,777,1162,1185]" pageId="1" pageNumber="892">
<taxonomicName id="4C13BD922E2A3845A8F0E5B60AD0FB0D" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[487,694,1162,1184]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="892" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E2A3845A8F0E5B60AD0FB0D" bold="true" box="[487,694,1162,1184]" italics="true" pageId="1" pageNumber="892">Recurvidris nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E2A3845ABABE5B70B6FFB0C" box="[700,777,1163,1185]" pageId="1" pageNumber="892" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BACC6112E2A3845A9A5E5810BA8FB5D" blockId="1.[178,1085,1213,1264]" pageId="1" pageNumber="892">E t y m o l o g y: The Latin adjective &quot;nigrans&quot; means dark and refers to the dark body colour of the new species that differs from all other hitherto known congeners.</paragraph>
</subSubSection>
<subSubSection id="C309959A2E2A3847A9A5E43D0C79FEC8" lastPageId="3" lastPageNumber="894" pageId="1" pageNumber="892" type="description">
<paragraph id="8BACC6112E2A3845A9A5E43D0B90FA27" blockId="1.[178,1086,1281,1418]" pageId="1" pageNumber="892">
<materialsCitation id="3B7BCC4C2E2A3845A9A5E43D0B90FA27" collectingDate="2008-03-04" collectionCode="T" collectorName="H. Zettel &amp; Entomological Collection &amp; City" country="Philippines" location="Natural History Museum" municipality="University of San Carlos" pageId="1" pageNumber="892" specimenCount="10" specimenCount-worker="10" stateProvince="Wien" typeStatus="holotype">
<collectionCode id="ED025ED42E2A3845A9A5E43D08A6FAB8" box="[178,192,1281,1301]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="892">T</collectionCode>
y p e m a t e r i a l:
<typeStatus id="54A878B32E2A3845A8AFE43D0A6AFAB8" box="[440,524,1281,1301]" pageId="1" pageNumber="892" type="holotype">Holotype</typeStatus>
worker labelled &quot;
<collectingCountry id="F30486812E2A3845ABD7E43D0B4AFAB8" box="[704,812,1281,1301]" name="Philippines" pageId="1" pageNumber="892">Philippines</collectingCountry>
: Negros Or., Cuer-\ nos de Negros, Valencia\ Apolong, Casaroro Falls, 28.\ 1.2007, leg.
<collectorName id="26E6A3C72E2A3845ABF6E4220B54FA9F" box="[737,818,1310,1330]" pageId="1" pageNumber="892">H. Zettel</collectorName>
(456)&quot;, in the
<collectorName id="26E6A3C72E2A3845AAAFE4220976FAFD" pageId="1" pageNumber="892">Entomological Collection</collectorName>
of the
<collectingMunicipality id="6BC85C6B2E2A3845A840E4000A25FAFD" box="[343,579,1340,1360]" pageId="1" pageNumber="892">University of San Carlos</collectingMunicipality>
, Cebu
<collectorName id="26E6A3C72E2A3845AB91E4000AC9FAFD" box="[646,687,1340,1360]" pageId="1" pageNumber="892">City</collectorName>
,
<collectingCountry id="F30486812E2A3845ABACE4000B42FAFD" box="[699,804,1340,1360]" name="Philippines" pageId="1" pageNumber="892">Philippines</collectingCountry>
.
<specimenCount id="9D150D982E2A3845AA38E4000B8BFAFD" box="[815,1005,1340,1360]" pageId="1" pageNumber="892" type="worker" typeStatus="paratype">10 paratype workers</specimenCount>
labelled &quot;
<collectingCountry id="F30486812E2A3845A9AEE4650943FAC0" box="[185,293,1369,1389]" name="Philippines" pageId="1" pageNumber="892">Philippines</collectingCountry>
: Negros Oriental\ Cuernos d.N., Valencia\ Apolong, Casaroro Falls\ 3-
<date id="FFADE0D12E2A3845AAD3E4650C7FFAC0" box="[964,1049,1369,1389]" pageId="1" pageNumber="892" value="2008-03-04">
<collectingDate id="EFE919392E2A3845AAD3E4650C7FFAC0" box="[964,1049,1369,1389]" pageId="1" pageNumber="892" value="2008-03-04">4.3.2008</collectingDate>
</date>
, lg. Zettel (513)&quot; in the
<location id="8ECC90CA2E2A3845A871E44A0A2EFA27" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8:8ECC90CA2E2A3845A871E44A0A2EFA27" box="[358,584,1398,1418]" country="Philippines" municipality="University of San Carlos" name="Natural History Museum" pageId="1" pageNumber="892" stateProvince="Wien">Natural History Museum</location>
<collectingRegion id="49D708F32E2A3845AB59E44A0AF4FA27" box="[590,658,1398,1418]" country="Austria" name="Wien" pageId="1" pageNumber="892">Vienna</collectingRegion>
,
<collectingCountry id="F30486812E2A3845AB8CE44A0A86FA27" box="[667,736,1398,1418]" name="Austria" pageId="1" pageNumber="892">Austria</collectingCountry>
, and in the author's collection.
</materialsCitation>
</paragraph>
<paragraph id="8BACC6112E2A3845A9A5E4A60B54FA47" blockId="1.[178,1086,1434,1514]" pageId="1" pageNumber="892">
<materialsCitation id="3B7BCC4C2E2A3845A9A5E4A6092EFA47" collectionCode="T" country="Philippines" county="Negros Island" location="Mount Talinis" municipality="Cuernos de Negros Mountains" pageId="1" pageNumber="892" specimenCount="1" stateProvince="Negros Oriental">
<collectionCode id="ED025ED42E2A3845A9A5E4A608A7FA02" box="[178,193,1434,1455]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="892">T</collectionCode>
y p e l o c a l i t y
<collectingCountry id="F30486812E2A3845A8A4E4A60A46FA1D" box="[435,544,1434,1456]" name="Philippines" pageId="1" pageNumber="892">Philippines</collectingCountry>
,
<collectingCounty id="62CDBE9D2E2A3845AB23E4A60AD9FA1D" box="[564,703,1434,1456]" pageId="1" pageNumber="892">Negros Island</collectingCounty>
,
<collectingRegion id="49D708F32E2A3845ABC4E4A60BB3FA1D" box="[723,981,1434,1456]" country="Philippines" name="Negros Oriental" pageId="1" pageNumber="892">Negros Oriental Province</collectingRegion>
,
<collectingMunicipality id="6BC85C6B2E2A3845AAFEE4A609F4FA60" pageId="1" pageNumber="892">Cuernos de Negros Mountains</collectingMunicipality>
(=
<location id="8ECC90CA2E2A3845A8ACE4840A34FA60" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8:8ECC90CA2E2A3845A8ACE4840A34FA60" box="[443,594,1463,1485]" country="Philippines" county="Negros Island" municipality="Cuernos de Negros Mountains" name="Mount Talinis" pageId="1" pageNumber="892" stateProvince="Negros Oriental">Mount Talinis</location>
), municipality of
<location id="8ECC90CA2E2A3845AA19E48B0B0CFA60" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8:8ECC90CA2E2A3845AA19E48B0B0CFA60" box="[782,874,1463,1485]" country="Philippines" county="Negros Island" municipality="Cuernos de Negros Mountains" name="Valencia" pageId="1" pageNumber="892" stateProvince="Negros Oriental">Valencia</location>
, barangay
<location id="8ECC90CA2E2A3845AAC9E48B0C5FFA60" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8:8ECC90CA2E2A3845AAC9E48B0C5FFA60" box="[990,1081,1463,1485]" country="Philippines" county="Negros Island" municipality="Cuernos de Negros Mountains" name="Apolong" pageId="1" pageNumber="892" stateProvince="Negros Oriental">Apolong</location>
,
<location id="8ECC90CA2E2A3845A9A5E4E8092EFA47" LSID="urn:lsid:plazi:treatment:03BA77072E2A3847A8F0E5B60C79FEC8:8ECC90CA2E2A3845A9A5E4E8092EFA47" box="[178,328,1492,1514]" country="Philippines" county="Negros Island" municipality="Cuernos de Negros Mountains" name="Casaroro Falls" pageId="1" pageNumber="892" stateProvince="Negros Oriental">Casaroro Falls</location>
</materialsCitation>
;
<materialsCitation id="3B7BCC4C2E2A3845A845E4E80B49FA47" box="[338,815,1492,1514]" collectionCode="GPS" country="Philippines" elevation="525" latitude="9.266666" longLatPrecision="1300" longitude="123.2" pageId="1" pageNumber="892" specimenCount="1" stateProvince="Negros Oriental">
<geoCoordinate id="EE27A0D62E2A3845A845E4E809CEFA47" box="[338,424,1492,1514]" degrees="9" direction="north" minutes="16" orientation="latitude" pageId="1" pageNumber="892" precision="925" value="9.266666">9° 16' N</geoCoordinate>
,
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, c.
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<quantity id="4CEB6BF42E2A3845AB55E4E80AD6FA47" box="[578,688,1492,1514]" metricMagnitude="2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="1" pageNumber="892" unit="m" value="525.0" valueMax="550.0" valueMin="500.0">500-550 m</quantity>
a.s.l.
</elevation>
(
<collectionCode id="ED025ED42E2A3845ABE3E4E80B41FA47" box="[756,807,1492,1514]" pageId="1" pageNumber="892">GPS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BACC6112E2A3846A9A5E4C60AA1FE2E" blockId="1.[178,1086,1530,1581]" lastBlockId="2.[178,1086,219,387]" lastPageId="2" lastPageNumber="893" pageId="1" pageNumber="892">
D i a g n o s i s (worker):Body chiefly blackish brown, funiculusof antenna and tarsi yellowish. Body surface smooth and shiny. Head broad, HW 0.61-0.65, CI 99-103. Cly- peus without distinct carinae. Mandible with four acute teeth on apical margin and one prominent, acute tooth at basal margin (
<figureCitation id="1328DA942E293846AB4EE1C40AFDFEA3" box="[601,667,248,270]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 4</figureCitation>
). Propodeum dorsally with 1-2 pairs of short decumbent setae, with long, slender, recurved spines (
<figureCitation id="1328DA942E293846AA34E0290B06FE86" box="[803,864,277,299]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 1</figureCitation>
), without infradental lamellae between spines and metapleural lobes. Petiole (
<figureCitation id="1328DA942E293846AA1FE00E0B20FEE5" box="[776,838,306,328]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 1</figureCitation>
) with very slender peduncle ventrally bearing a spiniform subpetiolar process; node in lateral view forming an acute angle anteriorly (at insertion of anterior setae).
</paragraph>
<paragraph id="8BACC6112E293846A9A5E0AF0981FE05" blockId="2.[178,487,402,424]" box="[178,487,402,424]" pageId="2" pageNumber="893">D e s c r i p t i o n (worker):</paragraph>
<paragraph id="8BACC6112E293846A9A5E0840BDFFDA5" blockId="2.[178,1085,439,520]" pageId="2" pageNumber="893">
Measurements of
<typeStatus id="54A878B32E293846A879E08B09ADFE60" box="[366,459,439,461]" pageId="2" pageNumber="893" type="holotype">holotype</typeStatus>
: TL 3.2; HL 0.63; HW 0.65; CI 103; SL 0.57; SI 89; PW 0.35; AL 0.86. Measurements of
<typeStatus id="54A878B32E293846AB00E0E90A1EFE47" box="[535,632,469,490]" pageId="2" pageNumber="893" type="paratype">paratypes</typeStatus>
(n = 10): TL 2.9-3.2; HL 0.61-0.64; HW 0.61-0.65; CI 99-102; SL 0.56-0.59; SI 89-94; PW 0.34-0.35; AL 0.81-0.86.
</paragraph>
<paragraph id="8BACC6112E293846A9A5E32B0BCAFBF7" blockId="2.[178,1088,535,1114]" pageId="2" pageNumber="893">
Head (
<figureCitation id="1328DA942E293846A9E1E32B0937FD80" box="[246,337,535,557]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Figs 2, 3</figureCitation>
) broad and smooth, lacking sculpture except hair pits, setiferous pits and some rugae near mandible base. Mandible (
<figureCitation id="1328DA942E293846AB68E3090AD9FDE6" box="[639,703,565,587]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 4</figureCitation>
) smooth and shiny, with some relatively large hair pits, apical margin with four teeth, forth tooth acute, basal margin with one acute tooth almost as large as forth tooth of apical margin. Clypeus (
<figureCitation id="1328DA942E293846AAB5E3530B87FD28" box="[930,993,623,645]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 4</figureCitation>
) without paired carinae, but with indistinct paired elongate swellings. Eyes (
<figureCitation id="1328DA942E293846AA76E3B00BFAFD0F" box="[865,924,652,674]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 1</figureCitation>
) comparatively large, containing 8 ommatidia in longest row. Profile shape of alitrunk, petiole and postpetiole as in
<figureCitation id="1328DA942E293846A822E3FB09EAFD70" box="[309,396,711,733]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Figure 1</figureCitation>
. Alitrunk smooth and polished, with some very fine setiferous pits, only part of mesopleuron with some fine rugae. Promesonotum with 4 pairs of setae plus one median seta posteriorly. Impression behind mesonotum appearing angular in profile. Propodeum comparatively high, dorsal surface with 1-2 pairs of short subcumbent setae in front of spiracle; spines very high, slender, divergent, and in caudal view very narrow (blade-like). Petiole (
<figureCitation id="1328DA942E293846A89AE26509AFFCC2" box="[397,457,857,879]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 1</figureCitation>
) with 2-3 pairs of dorsal setae; peduncle in profile long and very slender, its dorsal outline distinctly concave and ending posteriorly in a sharp angle at insertions of anterior pair of setae (insertions separated from each other by shallow impression), its ventral outline with long, spiniform subpetiolar process; node low. Postpetiole with 3 dorsal pairs and 1 ventrolateral pair of setae, low and with deep helcium, as typical for the genus. Gaster moderately depressed. Petiole, postpetiole, and gaster polished except for very fine setiferous pits. Colour blackish brown, alitrunk, petiole and postpetiole slightly lighter. Scape of antenna brown, funiculus yellowish. Mandibles brown, distally yellowish. Legs dark brown, but tarsi yellowish.
</paragraph>
<paragraph id="8BACC6112E293847A9A5E5550C79FEC8" blockId="2.[178,1087,1129,1590]" lastBlockId="3.[178,1086,219,358]" lastPageId="3" lastPageNumber="894" pageId="2" pageNumber="893">
C o m p a r a t i v e n o t e s:
<taxonomicName id="4C13BD922E293846AB02E5550A85FBD2" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[533,739,1129,1151]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846AB02E5550A85FBD2" box="[533,739,1129,1151]" italics="true" pageId="2" pageNumber="893">Recurvidris nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846ABFAE5560B51FBD2" box="[749,823,1130,1151]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
meets all morphological criteria of the
<taxonomicName id="4C13BD922E293846A854E5BB09D7FB31" baseAuthorityName="WHEELER &amp; WHEELER" baseAuthorityYear="1954" box="[323,433,1158,1180]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="kemneri">
<emphasis id="B9671A032E293846A854E5BB0934FB31" box="[323,338,1159,1180]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A877E5BA09D7FB31" box="[352,433,1158,1180]" italics="true" pageId="2" pageNumber="893">kemneri</emphasis>
</taxonomicName>
group listed by
<bibRefCitation id="EF82BBE02E293846AB4FE5BB0A91FB31" author="BOLTON B" box="[600,759,1158,1180]" pageId="2" pageNumber="893" pagination="35 - 48" refId="ref2308" refString="BOLTON B. (1992): A review of the ant genus Recurvidris (Hym.: Formicidae), a new name for Trigonogaster FOREL. - Psyche 99: 35 - 48." type="journal article" year="1992">BOLTON (1992)</bibRefCitation>
, i.e., the characteristic dentition of the mandible (
<figureCitation id="1328DA942E293846A873E59F09C2FB14" box="[356,420,1187,1209]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 4</figureCitation>
), the absence of infradental lamellae on the propodeal declivity, and a broad head (
<figureCitation id="1328DA942E293846A894E5FD09A3FB7A" box="[387,453,1217,1239]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 2</figureCitation>
) (range of CI
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the group,
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<taxonomicName id="4C13BD922E293846AAD9E5FE0C5BFB7A" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[974,1085,1217,1239]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846AAD9E5FE0BBBFB7A" box="[974,989,1218,1239]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AAE7E5FE0C5BFB7A" box="[1008,1085,1218,1239]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846A9A5E5E30899FB59" box="[178,255,1247,1268]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
). This species group contains only two further taxa,
<taxonomicName id="4C13BD922E293846AA21E5E3093BFABC" authority="(WHEELER &amp; WHEELER 1954)" baseAuthorityName="WHEELER &amp; WHEELER" baseAuthorityYear="1954" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="kemneri">
<emphasis id="B9671A032E293846AA21E5E30B23FB59" box="[822,837,1247,1268]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AA41E5E20BC1FB59" box="[854,935,1246,1268]" italics="true" pageId="2" pageNumber="893">kemneri</emphasis>
(WHEELER &amp; WHEELER 1954)
</taxonomicName>
from Borneo and Java and
<taxonomicName id="4C13BD922E293846AB83E5C00BF2FABC" authority="BOLTON 1992" authorityName="BOLTON" authorityYear="1992" box="[660,916,1275,1297]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846AB83E5C00AC5FABC" box="[660,675,1276,1297]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846ABA1E5C70A93FABC" box="[694,757,1275,1297]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
<bibRefCitation id="EF82BBE02E293846AA16E5C00BF2FABC" author="BOLTON B" box="[769,916,1275,1297]" pageId="2" pageNumber="893" pagination="35 - 48" refId="ref2308" refString="BOLTON B. (1992): A review of the ant genus Recurvidris (Hym.: Formicidae), a new name for Trigonogaster FOREL. - Psyche 99: 35 - 48." type="journal article" year="1992">BOLTON 1992</bibRefCitation>
</taxonomicName>
from Sulawesi. Comparing the key characteristics which distinguish these two species,
<taxonomicName id="4C13BD922E293846AABEE4250C70FA83" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[937,1046,1305,1326]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846AABEE4250BDEFA83" box="[937,952,1305,1326]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AADEE4250C70FA83" box="[969,1046,1305,1326]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846AD37E42508B9FAE1" pageId="2" pageNumber="893" rank="species">sp. nov.</taxonomicNameLabel>
is more similar to
<taxonomicName id="4C13BD922E293846A8A3E40B0A72FAE1" authorityName="BOLTON" authorityYear="1992" box="[436,532,1334,1356]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846A8A3E40B09A5FAE1" box="[436,451,1335,1356]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A8C2E40A0A72FAE1" box="[469,532,1334,1356]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
</taxonomicName>
, because it is relatively large in size, lacks clypeal carinae and bears 1-2 pairs of setae on the propodeal dorsum. The dark body colour of
<taxonomicName id="4C13BD922E293846AD3FE4680899FA2B" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846AD3FE4680C51FAC4" box="[1064,1079,1364,1385]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A9A5E44D0899FA2B" box="[178,255,1393,1414]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846A81CE44D0933FA2B" box="[267,341,1393,1414]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
differs from both species (as well as from all other congeners!);
<taxonomicName id="4C13BD922E293846AD30E44D0965FA09" baseAuthorityName="WHEELER &amp; WHEELER" baseAuthorityYear="1954" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="kemneri">
<emphasis id="B9671A032E293846AD30E44D0C50FA2B" box="[1063,1078,1393,1414]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A9A5E4B20965FA09" box="[178,259,1422,1444]" italics="true" pageId="2" pageNumber="893">kemneri</emphasis>
</taxonomicName>
is uniformly yellow,
<taxonomicName id="4C13BD922E293846A8FBE4B30A2CFA09" authorityName="BOLTON" authorityYear="1992" box="[492,586,1422,1444]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846A8FBE4B3099DFA09" box="[492,507,1423,1444]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AB1CE4B20A2CFA09" box="[523,586,1422,1444]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
</taxonomicName>
is yellow with brown head and gaster. Further differences between
<taxonomicName id="4C13BD922E293846A891E4900989FA6C" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[390,495,1452,1473]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="nov.sp. and R.">
<emphasis id="B9671A032E293846A891E49009F3FA6C" box="[390,405,1452,1473]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A8B5E4900989FA6C" box="[418,495,1452,1473]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846A8E0E4900AEEFA6C" box="[503,648,1451,1473]" pageId="2" pageNumber="893" rank="subSpecies">
nov.sp. and
<taxonomicName id="4C13BD922E293846AB63E4900AEEFA6C" authorityName="BOLTON" authorityYear="1992" box="[628,648,1452,1473]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846AB63E4900AE4FA6C" box="[628,642,1452,1473]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="B9671A032E293846AB87E4970AA9FA6C" box="[656,719,1451,1473]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
are the following: The tooth on the basal margin of the mandible is strongly developed in
<taxonomicName id="4C13BD922E293846ABCBE4F50B23FA73" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[732,837,1481,1502]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846ABCBE4F50A8DFA73" box="[732,747,1481,1502]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846ABEFE4F50B23FA73" box="[760,837,1481,1502]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846AA5AE4F50BF1FA73" box="[845,919,1481,1502]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
(almost as large as the forth tooth,
<figureCitation id="1328DA942E293846A861E4DA09D0FA51" box="[374,438,1510,1532]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="2" pageNumber="893">Fig. 4</figureCitation>
), but small in
<taxonomicName id="4C13BD922E293846AB45E4DA0AD6FA51" authorityName="BOLTON" authorityYear="1992" box="[594,688,1510,1532]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846AB45E4DA0A07FA56" box="[594,609,1510,1531]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AB66E4DA0AD6FA51" box="[625,688,1510,1532]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
</taxonomicName>
. The dorsum of the propodeum bears short, decumbent setae in
<taxonomicName id="4C13BD922E293846A8ABE7380A40F9B4" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[444,550,1540,1561]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846A8ABE73809ADF9B4" box="[444,459,1540,1561]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A8CEE7380A40F9B4" box="[473,550,1540,1561]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846AB39E7380A1FF9B4" box="[558,633,1540,1561]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
, but comparatively long standing setae in
<taxonomicName id="4C13BD922E293846AD3FE7380897F99B" authorityName="BOLTON" authorityYear="1992" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E293846AD3FE7380C51F9B4" box="[1064,1079,1540,1561]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846A9A5E71C0897F99B" box="[178,241,1568,1590]" italics="true" pageId="2" pageNumber="893">proles</emphasis>
</taxonomicName>
. The subpetiolar process is narrow, relatively long and acute in
<taxonomicName id="4C13BD922E293846AA93E71D0B8BF99B" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[900,1005,1569,1590]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="893" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E293846AA93E71D0BF5F99B" box="[900,915,1569,1590]" italics="true" pageId="2" pageNumber="893">R</emphasis>
.
<emphasis id="B9671A032E293846AAB7E71D0B8BF99B" box="[928,1005,1569,1590]" italics="true" pageId="2" pageNumber="893">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E293846AAE2E71D0C59F99B" box="[1013,1087,1569,1590]" pageId="2" pageNumber="893" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="1328DA942E283847A9ADE1E7089CFF5C" box="[186,250,219,241]" captionStart="Figs 1-4" captionStartId="4.[178,218,1474,1494]" captionTargetBox="[178,1085,732,1458]" captionTargetId="figure-138@4.[178,1085,732,1458]" captionTargetPageId="4" captionText="Figs 1-4: Recurvidris nigrans nov.sp., paratype worker (HW 0.63 mm, AL 0.82 mm) in the Natural History Museum Vienna. (1) Habitus, lateral view; (2) Habitus, dorsal view; (3) Head, full face view; (4) Head in oblique view showing dentition of mandibles." figureDoi="http://doi.org/10.5281/zenodo.5429938" httpUri="https://zenodo.org/record/5429938/files/figure.png" pageId="3" pageNumber="894">Fig. 1</figureCitation>
), but small and triangular in
<taxonomicName id="4C13BD922E283847AB53E1E70AC3FF5C" authorityName="BOLTON" authorityYear="1992" box="[580,677,219,241]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="894" phylum="Arthropoda" rank="species" species="proles">
<emphasis id="B9671A032E283847AB53E1E70A35FF5D" box="[580,595,219,240]" italics="true" pageId="3" pageNumber="894">R</emphasis>
.
<emphasis id="B9671A032E283847AB71E1E70AC3FF5C" box="[614,677,219,241]" italics="true" pageId="3" pageNumber="894">proles</emphasis>
</taxonomicName>
(for the latter two characteristics see
<bibRefCitation id="EF82BBE02E283847A9A5E1C40921FEA3" author="BOLTON B" box="[178,327,248,270]" pageId="3" pageNumber="894" pagination="35 - 48" refId="ref2308" refString="BOLTON B. (1992): A review of the ant genus Recurvidris (Hym.: Formicidae), a new name for Trigonogaster FOREL. - Psyche 99: 35 - 48." type="journal article" year="1992">BOLTON 1992</bibRefCitation>
: fig. 8). &quot;
<emphasis id="B9671A032E283847A8A3E1C40A4CFEA3" box="[436,554,248,270]" italics="true" pageId="3" pageNumber="894">Recurvidris</emphasis>
sp_phi1&quot; from
<collectingRegion id="49D708F32E283847ABC1E1C40B16FEA3" box="[726,880,248,270]" country="Philippines" name="Camarines Sur" pageId="3" pageNumber="894">Camarines Sur</collectingRegion>
in southern Luzon (
<bibRefCitation id="EF82BBE02E283847A9AEE02909E4FE86" author="ALPERT G &amp; GENERAL D" box="[185,386,277,299]" pageId="3" pageNumber="894" refId="ref2205" refString="ALPERT G. D., GENERAL D. M. &amp; V. SAMARITA (2008): Ants of the Philippines. &lt;http: // pick 4. pick. uga. edu / mp / 20 q? act = x _ checklist &amp; list = Ants _ Philippines&gt;, retrieved on 25 March 2008." type="url" year="2008">ALPERT &amp; al. 2008</bibRefCitation>
: see figures) differs from the new species by much smaller size (TL c. 2.1, HW c. 0.4), uniformly yellow colour, and a relatively stout petiolar peduncle in lateral aspect. On average,
<taxonomicName id="4C13BD922E283847A8ACE06D0A47FEC8" authority="Zettel, 2008" authorityName="Zettel" authorityYear="2008" box="[443,545,336,358]" class="Insecta" family="Formicidae" genus="Recurvidris" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="894" phylum="Arthropoda" rank="species" species="nigrans" status="sp. nov.">
<emphasis id="B9671A032E283847A8ACE06D09ACFECB" box="[443,458,337,358]" italics="true" pageId="3" pageNumber="894">R</emphasis>
.
<emphasis id="B9671A032E283847A8C1E06C0A47FEC8" box="[470,545,336,357]" italics="true" pageId="3" pageNumber="894">nigrans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A254A7782E283847AB30E06C0A09FEC8" box="[551,623,336,357]" pageId="3" pageNumber="894" rank="species">nov.sp.</taxonomicNameLabel>
is the largest known species of
<emphasis id="B9671A032E283847AAB1E06C0C7EFECB" box="[934,1048,336,358]" italics="true" pageId="3" pageNumber="894">Recurvidris</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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