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<document id="0986D9166591050AE1652E583146F3EE" ID-CLB-Dataset="27607" ID-DOI="10.11646/zootaxa.4551.2.3" ID-GBIF-Dataset="d29576b8-ec7a-4de1-a82c-f493611e681a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2622761" ID-ZooBank="23CD3A2F-8E89-4324-A81C-511BE02C0559" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554269759961" checkinUser="plazi" docAuthor="Botero-Trujillo, Ricardo, Ott, Ricardo, Mattoni, Camilo I., Nime, Mónica F. &amp; Ojanguren-Affilastro, Andrés A." docDate="2019" docId="03986B0AFFA6251CFF6322C2FF140AAB" docLanguage="en" docName="zootaxa.4551.2.3.pdf" docOrigin="Zootaxa 4551 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gaucha santana Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="192" masterDocId="FFA11372FFA02510FFF42420FFFB0D12" masterDocTitle="Two new species of the sun-spider genus Gaucha from Argentina and Brazil (Solifugae, Mummuciidae)" masterLastPageNumber="194" masterPageNumber="180" pageNumber="186" updateTime="1698716052786" updateUser="ExternalLinkService">
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<mods:title id="0278BEB37D701C6AF2CDD160A7A754BB">Two new species of the sun-spider genus Gaucha from Argentina and Brazil (Solifugae, Mummuciidae)</mods:title>
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<mods:namePart id="A4B4C722DB79437C18D47910DC629F0F">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:namePart id="E0F62B1D03D2E820D171DC670242F0D9">Ott, Ricardo</mods:namePart>
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<mods:namePart id="3E2DCB389539BB44AA09E2E8F9CD504C">Mattoni, Camilo I.</mods:namePart>
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<mods:namePart id="0131F29FA4B282D4E7BB6F6AA853A520">Nime, Mónica F.</mods:namePart>
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<mods:namePart id="68EBB0F40EBFCB6C1C27361CE2C39364">Ojanguren-Affilastro, Andrés A.</mods:namePart>
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<mods:date id="EEC8E87936AD3498D487DD2FEC2FD431">2019</mods:date>
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<subSubSection id="C32B8997FFA62516FF6322C2FE380BEE" box="[151,451,1762,1788]" pageId="6" pageNumber="186" type="nomenclature">
<paragraph id="8B8EDA1CFFA62516FF6322C2FE380BEE" blockId="6.[151,451,1762,1822]" box="[151,451,1762,1788]" pageId="6" pageNumber="186">
<heading id="D0C66D70FFA62516FF6322C2FE380BEE" bold="true" box="[151,451,1762,1788]" fontSize="11" level="1" pageId="6" pageNumber="186" reason="1">
<emphasis id="B945060EFFA62516FF6322C2FE380BEE" bold="true" box="[151,451,1762,1788]" pageId="6" pageNumber="186">
<taxonomicName id="4C31A19FFFA62516FF6322C2FEA50BEE" ID-CoL="6JXL7" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[151,350,1762,1788]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="6" pageNumber="186" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA62516FF6322C2FEA50BEE" bold="true" box="[151,350,1762,1788]" italics="true" pageId="6" pageNumber="186">Gaucha santana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A276BB75FFA62516FE9122C2FE380BEE" box="[357,451,1762,1788]" pageId="6" pageNumber="186" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C32B8997FFA62516FF632325FB310A71" pageId="6" pageNumber="186" type="description">
<paragraph id="8B8EDA1CFFA62516FF632325FE620A0C" blockId="6.[151,451,1762,1822]" box="[151,409,1797,1822]" pageId="6" pageNumber="186">
<figureCitation id="130AC699FFA62516FF632325FEFB0A0C" box="[151,256,1797,1822]" captionStart="FIGURE 1" captionStartId="1.[151,250,1813,1835]" captionTargetBox="[152,1435,1121,1801]" captionTargetId="figure@1.[152,1435,1121,1801]" captionTargetPageId="1" captionText="FIGURE 1. Map plotting known locality records of Gaucha ramirezi sp. nov. and Gaucha santana sp. nov." httpUri="https://zenodo.org/record/2622763/files/figure.png" pageId="6" pageNumber="186">Figures 1</figureCitation>
,
<figureCitation id="130AC699FFA62516FEFA2325FEC00A0C" box="[270,315,1797,1822]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="8.[151,250,1995,2017]" captionStartId-1="9.[151,250,1181,1203]" captionStartId-2="10.[151,250,1886,1908]" captionStartId-3="12.[151,250,1645,1667]" captionStartId-4="13.[151,250,1537,1559]" captionTargetBox-0="[150,1436,517,1973]" captionTargetBox-1="[150,1436,219,1160]" captionTargetBox-2="[150,1436,511,1838]" captionTargetBox-3="[150,1436,398,1586]" captionTargetBox-4="[194,1396,900,1483]" captionTargetId-0="figure@8.[150,1436,517,1973]" captionTargetId-1="figure@9.[150,1436,193,1160]" captionTargetId-2="figure@10.[150,1436,511,1838]" captionTargetId-3="figure@12.[150,1436,398,1586]" captionTargetId-4="figure@13.[150,1436,878,1515]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionTargetPageId-2="10" captionTargetPageId-3="12" captionTargetPageId-4="13" captionText-0="FIGURE 5. Live specimens of Gaucha santana sp. nov., photographed in the field. A. Male. B, C. Two different females." captionText-1="FIGURE 6. Habitat of Gaucha santana sp. nov. in Santana do Livramento—Área de Proteção Ambiental do Rio Ibirapuitã, Rio Grande do Sul, Brazil. A, B. Fazenda Rincão dos Moraes, showing typical sandy and stony soils. C, D. Large basalt outcrops, “Afloramento Passo do Ferrão”." captionText-2="FIGURE 7. Gaucha santana sp. nov. A. Male holotype, habitus, dorsal aspect (MCN-Sol-018). B. Male paratype, propeltidium and chelicerae, dorsal aspect (MCN-Sol-044). C. Ibid. for female paratype (MCN-Sol-047). D. Male paratype, opisthosoma, lateral aspect (MCN-Sol-044). E. Male paratype, opisthosoma, 1 st to 4 th post-genital sternites (MCN-Sol-044); selected ctenidia are indicated by arrows (zooming-in is recommended for proper visualization). Scale bars: 2 mm (A); 1 mm (BD); 0.25 mm (E)." captionText-3="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." captionText-4="FIGURE 9. Neighbor-Joining clustering for selected species of the fasciata species-group of Gaucha Mello-Leitão, 1924." httpUri-0="https://zenodo.org/record/2622772/files/figure.png" httpUri-1="https://zenodo.org/record/2622774/files/figure.png" httpUri-2="https://zenodo.org/record/2622776/files/figure.png" httpUri-3="https://zenodo.org/record/2622778/files/figure.png" httpUri-4="https://zenodo.org/record/2622780/files/figure.png" pageId="6" pageNumber="186">59</figureCitation>
;
<tableCitation id="C6B3EFA7FFA62516FEB22325FE620A0C" box="[326,409,1797,1822]" captionStart="TABLE 1" captionStartId="11.[151,239,152,175]" captionText="TABLE 1. Metric data for Gaucha ramirezi sp. nov. and Gaucha santana sp. nov. Measurements in millimeters for one male and one female. L = length; W = width; H = height. 1 Measured along medial axis, from the propeltidium anterior margin to the opisthosoma posterior margin. 2 Measured in dorsal view at widest point. 3 Measured in retrolateral view parallel to longitudinal axis of chelicera, from the fixed finger apex to anterolateral propeltidial lobe anterior margin. 4 Measured in retrolateral view, along vertical axis at highest part of manus. 5 Sum of individual segment lengths. 6 Maximum height. 7 Measurement excludes claws." pageId="6" pageNumber="186">Table 1</tableCitation>
</paragraph>
<paragraph id="8B8EDA1CFFA62516FF63236DFB310A71" blockId="6.[151,1226,1869,1891]" box="[151,1226,1869,1891]" pageId="6" pageNumber="186">
<treatmentCitationGroup id="AB21FD32FFA62516FF63236DFB310A71" box="[151,1226,1869,1891]" pageId="6" pageNumber="186">
<taxonomicName id="4C31A19FFFA62516FF63236DFF1C0A71" ID-CoL="4MFZ" authorityName="Mello-Leitao" authorityYear="1924" box="[151,231,1869,1891]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="6" pageNumber="186" phylum="Arthropoda" rank="genus">
<emphasis id="B945060EFFA62516FF63236DFF1C0A71" box="[151,231,1869,1891]" italics="true" pageId="6" pageNumber="186">Gaucha</emphasis>
</taxonomicName>
sp.:
<treatmentCitation id="0A90FC0DFFA62516FEE3236DFDB10A71" author="Botero-Trujillo, R. &amp; Ott, R. &amp; Carvalho, L. S." box="[279,586,1869,1891]" page="39" pageId="6" pageNumber="186" year="2017">
<bibRefCitation id="EFA0A7EDFFA62516FEE3236DFDDC0A71" author="Botero-Trujillo, R. &amp; Ott, R. &amp; Carvalho, L. S." box="[279,551,1869,1891]" pageId="6" pageNumber="186" pagination="3 - 44" refId="ref7208" refString="Botero-Trujillo, R., Ott, R. &amp; Carvalho, L. S. (2017) Systematic revision and phylogeny of the South American sun-spider genus Gaucha Mello-Leitao (Solifugae: Mummuciidae), with description of four new species and two new generic synonymies. Arthropod Systematics &amp; Phylogeny, 75 (1), 3 - 44." type="journal article" year="2017">
Botero-Trujillo
<emphasis id="B945060EFFA62516FE43236EFE1F0A71" box="[439,484,1869,1891]" italics="true" pageId="6" pageNumber="186">et al</emphasis>
. 2017
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: 39
</treatmentCitation>
[in part, records from Santana do Livramento only], figs. 3cf.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<paragraph id="8B8EDA1CFFA62516FF6323AFFA670AE2" blockId="6.[151,1436,1935,2032]" pageId="6" pageNumber="186">
<emphasis id="B945060EFFA62516FF6323AFFE400ABA" bold="true" box="[151,443,1935,1960]" pageId="6" pageNumber="186">
Type material.
<materialsCitation id="3B59D041FFA62516FEBA23AFFE400ABA" ID-GBIF-Occurrence="2234218666" box="[334,443,1935,1960]" collectingDate="2013-11-30" collectorName="Fazenda Rincao dos Moraes &amp; R. Ott" country="Brazil" elevation="185" latitude="-30.48583" location="Santana do Livramento-Area de Protecao Ambiental do Rio Ibirapuita" longLatPrecision="1" longitude="-55.57653" pageId="6" pageNumber="186" specimenCount="1" stateProvince="Rio Grande do Sul" typeStatus="holotype">
<typeStatus id="548A64BEFFA62516FEBA23AFFE400ABA" box="[334,443,1935,1960]" pageId="6" pageNumber="186" type="holotype">Holotype</typeStatus>
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</emphasis>
: male from
<emphasis id="B945060EFFA62516FDBC23AFFD480ABA" bold="true" box="[584,691,1935,1960]" pageId="6" pageNumber="186">
<collectingCountry id="F3269A8CFFA62516FDBC23AFFD480ABA" box="[584,691,1935,1960]" name="Brazil" pageId="6" pageNumber="186">BRAZIL</collectingCountry>
</emphasis>
:
<emphasis id="B945060EFFA62516FD3723AFFC5E0ABA" bold="true" box="[707,933,1935,1960]" pageId="6" pageNumber="186">
<collectingRegion id="49F514FEFFA62516FD3723AFFC5E0ABA" box="[707,933,1935,1960]" country="Brazil" name="Rio Grande do Sul" pageId="6" pageNumber="186">Rio Grande do Sul</collectingRegion>
</emphasis>
:
<location id="8EEE8CC7FFA62516FC4123B0FE0B0ADE" LSID="urn:lsid:plazi:treatment:03986B0AFFA6251CFF6322C2FF140AAB:8EEE8CC7FFA62516FC4123B0FE0B0ADE" country="Brazil" latitude="-30.48583" longLatPrecision="1" longitude="-55.57653" name="Santana do Livramento-Area de Protecao Ambiental do Rio Ibirapuita" pageId="6" pageNumber="186" stateProvince="Rio Grande do Sul">Santana do Livramento—Área de Proteção Ambiental do Rio Ibirapuitã</location>
,
<collectorName id="26C4BFCAFFA62516FDF72393FCA70AD9" box="[515,860,1971,1996]" pageId="6" pageNumber="186">Fazenda Rincão dos Moraes</collectorName>
,
<quantity id="4CC977F9FFA62516FC9B2393FC450AD9" box="[879,958,1971,1996]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="186" unit="m" value="185.0">
<elevation id="001C3D2FFFA62516FC9B2393FC450AD9" box="[879,958,1971,1996]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="6" pageNumber="186" unit="m" value="185.0">185 m</elevation>
</quantity>
elev.,
<geoCoordinate id="EE05BCDBFFA62516FBEC2393FB310ADE" box="[1048,1226,1971,1996]" direction="south" orientation="latitude" pageId="6" pageNumber="186" precision="1" value="-30.48583">30°2908.99″ S</geoCoordinate>
<geoCoordinate id="EE05BCDBFFA62516FB2E2393FA630AD9" box="[1242,1432,1971,1996]" direction="west" orientation="longitude" pageId="6" pageNumber="186" precision="1" value="-55.57653">55°3435.51″ W</geoCoordinate>
,
<date id="FF8FFCDCFFA62516FF6323F7FEEA0AE2" box="[151,273,2007,2032]" pageId="6" pageNumber="186" value="2013-11-30">
<collectingDate id="EFCB0534FFA62516FF6323F7FEEA0AE2" box="[151,273,2007,2032]" pageId="6" pageNumber="186" value="2013-11-30">30.xi.2013</collectingDate>
</date>
,
<collectorName id="26C4BFCAFFA62516FEEA23F8FE9F0AE2" box="[286,356,2008,2032]" pageId="6" pageNumber="186">R. Ott</collectorName>
(MCN-Sol-018).
<materialsCitation id="3B59D041FFA62516FDC523F7FA670AE2" ID-GBIF-Occurrence="2234218738" box="[561,1436,2007,2032]" collectingDate="2013-11-30" collectorName="Fazenda Rincao dos Moraes &amp; R. Ott" country="Brazil" elevation="185" latitude="-30.48583" location="Santana do Livramento-Area de Protecao Ambiental do Rio Ibirapuita" longLatPrecision="1" longitude="-55.57653" pageId="6" pageNumber="186" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<emphasis id="B945060EFFA62516FDC523F7FD520AE2" bold="true" box="[561,681,2007,2032]" pageId="6" pageNumber="186">
<typeStatus id="548A64BEFFA62516FDC523F7FD520AE2" box="[561,681,2007,2032]" pageId="6" pageNumber="186" type="paratype">Paratypes</typeStatus>
</emphasis>
: same data of
<typeStatus id="548A64BEFFA62516FCA623F7FC4C0AE2" box="[850,951,2007,2032]" pageId="6" pageNumber="186" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D371195FFA62516FC3023F8FBEB0AE2" box="[964,1040,2007,2032]" pageId="6" pageNumber="186" type="male">1 male</specimenCount>
(MCN-Sol-016);
<specimenCount id="9D371195FFA62516FB2A23F8FABA0AE2" box="[1246,1345,2007,2032]" pageId="6" pageNumber="186" type="female">1 female</specimenCount>
(MCN-
</materialsCitation>
</paragraph>
<paragraph id="8B8EDA1CFFA72517FF6324B7FAB50C52" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="187">
Sol-017);
<specimenCount id="9D371195FFA72517FEFC24B8FE9A0DA2" box="[264,353,151,176]" pageId="7" pageNumber="187" type="male">2 males</specimenCount>
,
<specimenCount id="9D371195FFA72517FE9924B8FE2A0DA2" box="[365,465,151,176]" pageId="7" pageNumber="187" type="female">1 female</specimenCount>
,
<specimenCount id="9D371195FFA72517FE2924B8FDAC0DA2" box="[477,599,151,176]" pageId="7" pageNumber="187" type="juvenile">3 juveniles</specimenCount>
(MCN-Sol-041). Same locality and data but,
<date id="FF8FFCDCFFA72517FBAF24B7FB410DA2" box="[1115,1210,151,176]" pageId="7" pageNumber="187" value="2014-12">xii.2014</date>
, R. Ott &amp; R. Botero Trujillo,
<specimenCount id="9D371195FFA72517FF0D249CFEBD0DC7" box="[249,326,188,213]" pageId="7" pageNumber="187" type="male">1 male</specimenCount>
,
<specimenCount id="9D371195FFA72517FEA5249CFE4F0DC7" box="[337,436,188,213]" pageId="7" pageNumber="187" type="female">1 female</specimenCount>
,
<specimenCount id="9D371195FFA72517FE4B249CFDBD0DC7" box="[447,582,188,213]" pageId="7" pageNumber="187" type="juvenile">10 juveniles</specimenCount>
(MCN-Sol-047);
<materialsCitation id="3B59D041FFA72517FCFB249CFE570DEA" ID-GBIF-Occurrence="2234218727" collectionCode="BOLD" pageId="7" pageNumber="187" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D371195FFA72517FCFB249CFCA10DC7" box="[783,858,188,213]" pageId="7" pageNumber="187" type="male">1 male</specimenCount>
(MCN-Sol-046, tissue sample Sol-M00054,
<collectionCode id="ED2042D9FFA72517FAA4249CFA670DC6" box="[1360,1436,188,212]" pageId="7" pageNumber="187">BOLD</collectionCode>
process ID SSA054-16)
</materialsCitation>
;
<materialsCitation id="3B59D041FFA72517FE4F24C0FA840DEA" ID-GBIF-Occurrence="2234218669" box="[443,1407,223,248]" collectionCode="BOLD" pageId="7" pageNumber="187" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D371195FFA72517FE4F24C0FDF10DEA" box="[443,522,223,248]" pageId="7" pageNumber="187" type="male">1 male</specimenCount>
(MCN-Sol-044, tissue sample Sol-M00055,
<collectionCode id="ED2042D9FFA72517FBE724C0FBA40DEA" box="[1043,1119,224,248]" pageId="7" pageNumber="187">BOLD</collectionCode>
process ID SSA055-16)
</materialsCitation>
;
<materialsCitation id="3B59D041FFA72517FA7A24C0FCD30C52" ID-GBIF-Occurrence="2234218722" collectingDate="2013-11-29" collectionCode="BOLD" collectorName="Same &amp; Fazenda Sr. Caio &amp; R. Ott" elevation="185" pageId="7" pageNumber="187" specimenCount="1" specimenCount-subadult="1">
<specimenCount id="9D371195FFA72517FA7A24C0FF0D0C0F" pageId="7" pageNumber="187" type="subadult">1 subadult</specimenCount>
female (MCN-Sol-043, tissue sample Sol-M00056,
<collectionCode id="ED2042D9FFA72517FCA02524FC5B0C0E" box="[852,928,260,284]" pageId="7" pageNumber="187">BOLD</collectionCode>
process ID SSA056-16).
<collectorName id="26C4BFCAFFA72517FB392524FAF00C0E" box="[1229,1291,260,284]" pageId="7" pageNumber="187">Same</collectorName>
locality and data but,
<collectorName id="26C4BFCAFFA72517FF0A2507FE3B0C52" box="[254,448,295,320]" pageId="7" pageNumber="187">Fazenda Sr. Caio</collectorName>
,
<quantity id="4CC977F9FFA72517FE3F2507FDE90C2D" box="[459,530,295,320]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="187" unit="m" value="185.0">
<elevation id="001C3D2FFFA72517FE3F2507FDE90C2D" box="[459,530,295,320]" metricMagnitude="2" metricUnit="m" metricValue="1.85" pageId="7" pageNumber="187" unit="m" value="185.0">185 m</elevation>
</quantity>
elev.,
<date id="FF8FFCDCFFA72517FDAE2507FD2E0C52" box="[602,725,295,320]" pageId="7" pageNumber="187" value="2013-11-29">
<collectingDate id="EFCB0534FFA72517FDAE2507FD2E0C52" box="[602,725,295,320]" pageId="7" pageNumber="187" value="2013-11-29">29.xi.2013</collectingDate>
</date>
,
<collectorName id="26C4BFCAFFA72517FD142508FCD30C52" box="[736,808,296,320]" pageId="7" pageNumber="187">R. Ott</collectorName>
</materialsCitation>
,
<specimenCount id="9D371195FFA72517FCC72508FC770C52" box="[819,908,295,320]" pageId="7" pageNumber="187" type="male">4 males</specimenCount>
,
<specimenCount id="9D371195FFA72517FC632508FBFD0C52" box="[919,1030,295,320]" pageId="7" pageNumber="187" type="female">2 females</specimenCount>
,
<specimenCount id="9D371195FFA72517FBE52508FB700C52" box="[1041,1163,295,320]" pageId="7" pageNumber="187" type="juvenile">4 juveniles</specimenCount>
(MCN-Sol-015).
</paragraph>
<paragraph id="8B8EDA1CFFA72517FF33256CFD3D0F0A" blockId="7.[151,1436,151,536]" pageId="7" pageNumber="187">
<emphasis id="B945060EFFA72517FF33256CFD500C77" bold="true" box="[199,683,332,357]" pageId="7" pageNumber="187">
Additional material examined.
<materialsCitation id="3B59D041FFA72517FDB4256CFD500C77" ID-GBIF-Occurrence="2234218728" box="[576,683,332,357]" collectingDate="2011-11-14" collectingDateMax="2014-12" collectingDateMin="2011-11-14" collectorName="R. Ott &amp; R. Botero Trujillo" country="Brazil" county="Santana do Livramento-Area de Protecao Ambiental do Rio Ibirapuita" elevation="263" latitude="-30.454437" location="Area de Protecao Ambiental do Rio Ibirapuita" longLatPrecision="1" longitude="-55.72381" municipality="Area de Protecao Ambiental do Rio Ibirapuita" pageId="7" pageNumber="187" specimenCount="10" specimenCount-female="4" specimenCount-juvenile="2" specimenCount-male="4" stateProvince="Rio Grande do Sul">
<collectingCountry id="F3269A8CFFA72517FDB4256CFD500C77" box="[576,683,332,357]" name="Brazil" pageId="7" pageNumber="187">BRAZIL</collectingCountry>
</materialsCitation>
</emphasis>
:
<emphasis id="B945060EFFA72517FD48256CFC590C77" bold="true" box="[700,930,332,357]" pageId="7" pageNumber="187">
<collectingRegion id="49F514FEFFA72517FD48256CFC590C77" box="[700,930,332,357]" country="Brazil" name="Rio Grande do Sul" pageId="7" pageNumber="187">Rio Grande do Sul</collectingRegion>
</emphasis>
:
<collectingCounty id="62EFA290FFA72517FC46256CFE2E0C9A" pageId="7" pageNumber="187">Santana do Livramento—Área de Proteção Ambiental do Rio Ibirapuitã</collectingCounty>
, “Cerrito Reserva”,
<quantity id="4CC977F9FFA72517FD41254FFD000C95" box="[693,763,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="7" pageNumber="187" unit="m" value="263.0">
<elevation id="001C3D2FFFA72517FD41254FFD000C95" box="[693,763,367,392]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="7" pageNumber="187" unit="m" value="263.0">263 m</elevation>
</quantity>
elev.,
<geoCoordinate id="EE05BCDBFFA72517FCB6254FFC160C9A" box="[834,1005,367,392]" direction="south" orientation="latitude" pageId="7" pageNumber="187" precision="1" value="-30.60941">30°3633.87″ S</geoCoordinate>
<geoCoordinate id="EE05BCDBFFA72517FC07254FFB530C95" box="[1011,1192,367,392]" direction="west" orientation="longitude" pageId="7" pageNumber="187" precision="1" value="-55.64172">55°3830.18″ W</geoCoordinate>
,
<date id="FF8FFCDCFFA72517FB46254FFAE90C9A" box="[1202,1298,367,392]" pageId="7" pageNumber="187" value="2014-12">
<collectingDate id="EFCB0534FFA72517FB46254FFAE90C9A" box="[1202,1298,367,392]" pageId="7" pageNumber="187" value="2014-12">xii.2014</collectingDate>
</date>
,
<collectorName id="26C4BFCAFFA72517FAE82550FA9B0C9A" box="[1308,1376,368,392]" pageId="7" pageNumber="187">R. Ott</collectorName>
&amp;
<collectorName id="26C4BFCAFFA72517FA772550FEB30CBF" pageId="7" pageNumber="187">R. Botero Trujillo</collectorName>
,
<specimenCount id="9D371195FFA72517FEAC25B4FE510CBF" box="[344,426,404,429]" pageId="7" pageNumber="187" type="male">1 male</specimenCount>
(MCN-Sol-045); unknown date,
<collectorName id="26C4BFCAFFA72517FCC225B5FC7F0CBE" box="[822,900,404,428]" pageId="7" pageNumber="187">R. Ott</collectorName>
,
<specimenCount id="9D371195FFA72517FC6025B4FC000CBF" box="[916,1019,404,429]" pageId="7" pageNumber="187" type="female">1 female</specimenCount>
(MCN-Sol-042).
<collectingMunicipality id="6BEA4066FFA72517FB2425B4FE130CC2" pageId="7" pageNumber="187">Área de Proteção Ambiental do Rio Ibirapuitã</collectingMunicipality>
, “Afloramento Passo do Ferrão”,
<quantity id="4CC977F9FFA72517FC8B2598FC370CDD" box="[895,972,440,464]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="187" unit="m" value="174.0">
<elevation id="001C3D2FFFA72517FC8B2598FC370CDD" box="[895,972,440,464]" metricMagnitude="2" metricUnit="m" metricValue="1.7399999999999998" pageId="7" pageNumber="187" unit="m" value="174.0">174 m</elevation>
</quantity>
elev.,
<geoCoordinate id="EE05BCDBFFA72517FBD42597FB2B0CC2" box="[1056,1232,439,464]" direction="south" orientation="latitude" pageId="7" pageNumber="187" precision="1" value="-30.454437">30°2715.97″ S</geoCoordinate>
<geoCoordinate id="EE05BCDBFFA72517FB292597FA630CDD" box="[1245,1432,439,464]" direction="west" orientation="longitude" pageId="7" pageNumber="187" precision="1" value="-55.72381">55°4325.71″ W</geoCoordinate>
,
<date id="FF8FFCDCFFA72517FF6325FCFEEB0CE7" box="[151,272,476,501]" pageId="7" pageNumber="187" value="2011-11-14">
<collectingDate id="EFCB0534FFA72517FF6325FCFEEB0CE7" box="[151,272,476,501]" pageId="7" pageNumber="187" value="2011-11-14">14.xi.2011</collectingDate>
</date>
,
<specimenCount id="9D371195FFA72517FED625FCFE7A0CE7" box="[290,385,476,501]" pageId="7" pageNumber="187" type="male">2 males</specimenCount>
,
<specimenCount id="9D371195FFA72517FE6625FCFDFC0CE7" box="[402,519,476,501]" pageId="7" pageNumber="187" type="female">2 females</specimenCount>
,
<specimenCount id="9D371195FFA72517FDEC25FCFD630CE7" box="[536,664,476,501]" pageId="7" pageNumber="187" type="juvenile">2 juveniles</specimenCount>
(MCN-Sol-019).
<location id="8EEE8CC7FFA72517FC9925FCFA630CE7" LSID="urn:lsid:plazi:treatment:03986B0AFFA6251CFF6322C2FF140AAB:8EEE8CC7FFA72517FC9925FCFA630CE7" box="[877,1432,476,501]" country="Brazil" county="Santana do Livramento-Area de Protecao Ambiental do Rio Ibirapuita" latitude="-30.454437" longLatPrecision="1" longitude="-55.72381" municipality="Area de Protecao Ambiental do Rio Ibirapuita" name="Area de Protecao Ambiental do Rio Ibirapuita" pageId="7" pageNumber="187" stateProvince="Rio Grande do Sul">Área de Proteção Ambiental do Rio Ibirapuitã</location>
,
<date id="FF8FFCDCFFA72517FF6325DFFEEB0F0A" box="[151,272,511,536]" pageId="7" pageNumber="187" value="2013-11-01">
<collectingDate id="EFCB0534FFA72517FF6325DFFEEB0F0A" box="[151,272,511,536]" pageId="7" pageNumber="187" value="2013-11-01">01.xi.2013</collectingDate>
</date>
,
<collectorName id="26C4BFCAFFA72517FEE92620FE9F0F0A" box="[285,356,512,536]" pageId="7" pageNumber="187">R. Ott</collectorName>
,
<specimenCount id="9D371195FFA72517FE9B2620FE460F0A" box="[367,445,511,536]" pageId="7" pageNumber="187" type="male">1 male</specimenCount>
,
<specimenCount id="9D371195FFA72517FE3C2620FDD10F0A" box="[456,554,511,536]" pageId="7" pageNumber="187" type="female">1 female</specimenCount>
(MACN-Ar).
</paragraph>
</subSubSection>
<caption id="DF4E8A94FFA72517FF6322D2FBBC0A54" httpUri="https://zenodo.org/record/2622770/files/figure.png" pageId="7" pageNumber="187" startId="7.[151,250,1778,1800]" targetBox="[150,1436,589,1757]" targetPageId="7">
<paragraph id="8B8EDA1CFFA72517FF6322D2FBBC0A54" blockId="7.[151,1436,1778,1862]" pageId="7" pageNumber="187">
<emphasis id="B945060EFFA72517FF6322D2FEED0A15" bold="true" box="[151,278,1778,1800]" pageId="7" pageNumber="187">FIGURE 4.</emphasis>
<taxonomicName id="4C31A19FFFA72517FED422D2FE2B0A1A" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[288,464,1778,1800]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="7" pageNumber="187" phylum="Arthropoda" rank="species" species="ramirezi" status="sp. nov.">
<emphasis id="B945060EFFA72517FED422D2FE2B0A1A" box="[288,464,1778,1800]" italics="true" pageId="7" pageNumber="187">Gaucha ramirezi</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA72517FE2D22D3FDD50A1A" bold="true" box="[473,558,1779,1800]" pageId="7" pageNumber="187">
<taxonomicNameLabel id="A276BB75FFA72517FE2D22D3FDD50A1A" box="[473,558,1779,1800]" pageId="7" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, chelicerae.
<emphasis id="B945060EFFA72517FD4222D2FD330A15" bold="true" box="[694,712,1778,1799]" pageId="7" pageNumber="187">A</emphasis>
,
<emphasis id="B945060EFFA72517FD2D22D3FD110A1A" bold="true" box="[729,746,1779,1800]" pageId="7" pageNumber="187">B</emphasis>
. Right chelicera of female paratype (MACN-Ar).
<emphasis id="B945060EFFA72517FAF622D2FAEF0A15" bold="true" box="[1282,1300,1778,1799]" pageId="7" pageNumber="187">A</emphasis>
. Retrolateral aspect.
<emphasis id="B945060EFFA72517FF112331FF0D0A34" bold="true" box="[229,246,1809,1830]" pageId="7" pageNumber="187">B</emphasis>
. Prolateral aspect.
<emphasis id="B945060EFFA72517FE492331FE120A34" bold="true" box="[445,489,1809,1831]" pageId="7" pageNumber="187">CF</emphasis>
. Right chelicera of male paratype (MACN-Ar).
<emphasis id="B945060EFFA72517FC162331FC0F0A35" bold="true" box="[994,1012,1809,1831]" pageId="7" pageNumber="187">C</emphasis>
. Retrolateral aspect.
<emphasis id="B945060EFFA72517FB252331FB180A34" bold="true" box="[1233,1251,1809,1830]" pageId="7" pageNumber="187">D</emphasis>
. Prolateral aspect (under SEM).
<emphasis id="B945060EFFA72517FEDC2310FEC20A57" bold="true" box="[296,313,1840,1861]" pageId="7" pageNumber="187">E</emphasis>
. Flagellum.
<emphasis id="B945060EFFA72517FE4D2310FE3D0A57" bold="true" box="[441,454,1840,1861]" pageId="7" pageNumber="187">F</emphasis>
. Ibid. (under SEM). Scale bars: 0.5 mm (AD); 0.2 mm (E, F).
</paragraph>
</caption>
<subSubSection id="C32B8997FFA72518FF332352FC020CC2" lastPageId="8" lastPageNumber="188" pageId="7" pageNumber="187" type="etymology">
<paragraph id="8B8EDA1CFFA72518FF332352FC020CC2" blockId="7.[151,1436,1906,2003]" lastBlockId="8.[151,1436,151,464]" lastPageId="8" lastPageNumber="188" pageId="7" pageNumber="187">
<emphasis id="B945060EFFA72517FF332352FEB50A99" bold="true" box="[199,334,1906,1931]" pageId="7" pageNumber="187">Etymology.</emphasis>
Noun in apposition taken from the municipality Santana do Livramento, part of whose territory forms the Ibirapuitã Environmental Protection Area where this species was collected.
<emphasis id="B945060EFFA72517FF33239AFEBA0AC1" bold="true" box="[199,321,1978,2003]" pageId="7" pageNumber="187">Diagnosis.</emphasis>
<taxonomicName id="4C31A19FFFA72517FEBD239BFDF80AC0" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[329,515,1979,2003]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="7" pageNumber="187" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA72517FEBD239BFDF80AC0" box="[329,515,1979,2003]" italics="true" pageId="7" pageNumber="187">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA72517FDF8239AFD9C0AC1" bold="true" box="[524,615,1978,2003]" pageId="7" pageNumber="187">
<taxonomicNameLabel id="A276BB75FFA72517FDF8239AFD9C0AC1" box="[524,615,1978,2003]" pageId="7" pageNumber="187" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C31A19FFFA72517FCEC239BFC8A0AC1" authorityName="Mello-Leitao" authorityYear="1924" box="[792,881,1979,2003]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="7" pageNumber="187" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B945060EFFA72517FCEC239BFC8A0AC1" box="[792,881,1979,2003]" italics="true" pageId="7" pageNumber="187">fasciata</emphasis>
</taxonomicName>
species-group of
<taxonomicName id="4C31A19FFFA72517FBC9239BFB6D0AC1" authorityName="Mello-Leitao" authorityYear="1924" box="[1085,1174,1979,2003]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="7" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis id="B945060EFFA72517FBC9239BFB6D0AC1" box="[1085,1174,1979,2003]" italics="true" pageId="7" pageNumber="187">Gaucha</emphasis>
</taxonomicName>
, as defined by Botero- Trujillo
<emphasis id="B945060EFFA82518FF0624B9FED80DA2" box="[242,291,152,176]" italics="true" pageId="8" pageNumber="188">et al</emphasis>
. (2017: 15). In various respects, males of
<taxonomicName id="4C31A19FFFA82518FD0C24B8FC940DA2" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[760,879,152,176]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA82518FD0C24B8FC940DA2" box="[760,879,152,176]" italics="true" pageId="8" pageNumber="188">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA82518FC8224B7FC340DA2" bold="true" box="[886,975,151,176]" pageId="8" pageNumber="188">
<taxonomicNameLabel id="A276BB75FFA82518FC8224B7FC340DA2" box="[886,975,151,176]" pageId="8" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
most closely resemble those of
<taxonomicName id="4C31A19FFFA82518FAC024B8FA670DBD" authorityName="Botero-Trujillo, Ott &amp; Carvalho" authorityYear="2017" box="[1332,1436,152,175]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="curupi">
<emphasis id="B945060EFFA82518FAC024B8FA670DBD" box="[1332,1436,152,175]" italics="true" pageId="8" pageNumber="188">G. curupi</emphasis>
</taxonomicName>
among species in this group. These include that the fixed finger FP and FM teeth are elongated, whereas the FD tooth is much smaller, such that primary teeth are graded as FP=FM&gt;&gt;FD (
<figureCitation id="130AC699FFA82518FBF624FFFBA70DEA" box="[1026,1116,223,248]" captionStart="FIGURE 8" captionStartId="12.[151,250,1645,1667]" captionTargetBox="[150,1436,398,1586]" captionTargetId="figure@12.[150,1436,398,1586]" captionTargetPageId="12" captionText="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="8" pageNumber="188">Fig. 8C</figureCitation>
). Likewise, the fixed finger mucron is markedly long, similarly shaped in males of both species, and a subterminal flange (STF) is not present in either (
<figureCitation id="130AC699FFA82518FEFE2507FE730C52" box="[266,392,295,320]" captionStart="FIGURE 8" captionStartId="12.[151,250,1645,1667]" captionTargetBox="[150,1436,398,1586]" captionTargetId="figure@12.[150,1436,398,1586]" captionTargetPageId="12" captionText="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="8" pageNumber="188">Figs. 8C,E</figureCitation>
), besides both have the flagellum moderately inflated (
<figureCitation id="130AC699FFA82518FBF92507FB7C0C52" box="[1037,1159,295,320]" captionStart="FIGURE 8" captionStartId="12.[151,250,1645,1667]" captionTargetBox="[150,1436,398,1586]" captionTargetId="figure@12.[150,1436,398,1586]" captionTargetPageId="12" captionText="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="8" pageNumber="188">Figs. 8E,F</figureCitation>
) (e.g., unlike that of
<taxonomicName id="4C31A19FFFA82518FA702508FF0D0C76" authorityName="Botero-Trujillo, Ott &amp; Carvalho" authorityYear="2017" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="casuhati">
<emphasis id="B945060EFFA82518FA702508FF0D0C76" italics="true" pageId="8" pageNumber="188">G. casuhati</emphasis>
</taxonomicName>
).
<taxonomicName id="4C31A19FFFA82518FEFA256CFE3C0C76" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[270,455,332,356]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA82518FEFA256CFE3C0C76" box="[270,455,332,356]" italics="true" pageId="8" pageNumber="188">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA82518FE3A256CFDD20C77" bold="true" box="[462,553,332,357]" pageId="8" pageNumber="188">
<taxonomicNameLabel id="A276BB75FFA82518FE3A256CFDD20C77" box="[462,553,332,357]" pageId="8" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be recognized by the absence of the fixed finger FSD tooth (regardless of sex or maturity of the specimens) (
<figureCitation id="130AC699FFA82518FDEB254FFD650C9A" box="[543,670,367,392]" captionStart="FIGURE 8" captionStartId="12.[151,250,1645,1667]" captionTargetBox="[150,1436,398,1586]" captionTargetId="figure@12.[150,1436,398,1586]" captionTargetPageId="12" captionText="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="8" pageNumber="188">Figs. 8A,C</figureCitation>
); in contrast, females of
<taxonomicName id="4C31A19FFFA82518FC402550FBE60C95" authorityName="Botero-Trujillo, Ott &amp; Carvalho" authorityYear="2017" box="[948,1053,368,391]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="curupi">
<emphasis id="B945060EFFA82518FC402550FBE60C95" box="[948,1053,368,391]" italics="true" pageId="8" pageNumber="188">G. curupi</emphasis>
</taxonomicName>
always bear the FSD tooth, while FSD is most often present (though reduced) in males. Furthermore, the FD tooth, which is reduced in males of the two species, is comparatively larger in those of
<taxonomicName id="4C31A19FFFA82518FD582598FCDF0CC2" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[684,804,440,464]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA82518FD582598FCDF0CC2" box="[684,804,440,464]" italics="true" pageId="8" pageNumber="188">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA82518FCDF2597FC7F0CC2" bold="true" box="[811,900,439,464]" pageId="8" pageNumber="188">
<taxonomicNameLabel id="A276BB75FFA82518FCDF2597FC7F0CC2" box="[811,900,439,464]" pageId="8" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="130AC699FFA82518FC602597FC170CC2" box="[916,1004,439,464]" captionStart="FIGURE 8" captionStartId="12.[151,250,1645,1667]" captionTargetBox="[150,1436,398,1586]" captionTargetId="figure@12.[150,1436,398,1586]" captionTargetPageId="12" captionText="FIGURE 8. Gaucha santana sp. nov., chelicerae. A, B. Right chelicera of female paratype (MCN-Sol-015). A. Retrolateral aspect. B. Prolateral aspect. CF. Right chelicera of male paratype (MCN-Sol-041). C. Retrolateral aspect. D. Prolateral aspect (under SEM). E. Flagellum. F. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F)." httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="8" pageNumber="188">Fig. 8C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF4E8A94FFA82518FF6323EBFA8A0AF3" box="[151,1393,1995,2017]" httpUri="https://zenodo.org/record/2622772/files/figure.png" pageId="8" pageNumber="188" startId="8.[151,250,1995,2017]" targetBox="[150,1436,517,1973]" targetPageId="8">
<paragraph id="8B8EDA1CFFA82518FF6323EBFA8A0AF3" blockId="8.[151,1393,1995,2017]" box="[151,1393,1995,2017]" pageId="8" pageNumber="188">
<emphasis id="B945060EFFA82518FF6323EBFEE90AF2" bold="true" box="[151,274,1995,2017]" pageId="8" pageNumber="188">FIGURE 5.</emphasis>
Live specimens of
<taxonomicName id="4C31A19FFFA82518FE2D23EBFD840AF3" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[473,639,1995,2017]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="8" pageNumber="188" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA82518FE2D23EBFD840AF3" box="[473,639,1995,2017]" italics="true" pageId="8" pageNumber="188">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA82518FD7223EBFD2D0AF2" bold="true" box="[646,726,1995,2016]" pageId="8" pageNumber="188">
<taxonomicNameLabel id="A276BB75FFA82518FD7223EBFD2D0AF2" box="[646,726,1995,2016]" pageId="8" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, photographed in the field.
<emphasis id="B945060EFFA82518FC1B23EBFBFA0AF2" bold="true" box="[1007,1025,1995,2016]" pageId="8" pageNumber="188">A</emphasis>
. Male.
<emphasis id="B945060EFFA82518FBBB23EBFB9B0AF2" bold="true" box="[1103,1120,1995,2016]" pageId="8" pageNumber="188">B</emphasis>
,
<emphasis id="B945060EFFA82518FB9823EBFB850AF3" bold="true" box="[1132,1150,1995,2017]" pageId="8" pageNumber="188">C</emphasis>
. Two different females.
</paragraph>
</caption>
<caption id="DF4E8A94FFA92519FF6320BDFDB909E2" httpUri="https://zenodo.org/record/2622774/files/figure.png" pageId="9" pageNumber="189" startId="9.[151,250,1181,1203]" targetBox="[150,1436,219,1160]" targetPageId="9">
<paragraph id="8B8EDA1CFFA92519FF6320BDFDB909E2" blockId="9.[151,1436,1181,1264]" pageId="9" pageNumber="189">
<emphasis id="B945060EFFA92519FF6320BDFEEF09A0" bold="true" box="[151,276,1181,1203]" pageId="9" pageNumber="189">FIGURE 6.</emphasis>
Habitat of
<taxonomicName id="4C31A19FFFA92519FE7F20BDFDC809A1" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[395,563,1181,1203]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="9" pageNumber="189" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFA92519FE7F20BDFDC809A1" box="[395,563,1181,1203]" italics="true" pageId="9" pageNumber="189">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFA92519FDCF20BEFD7609A1" bold="true" box="[571,653,1182,1203]" pageId="9" pageNumber="189">
<taxonomicNameLabel id="A276BB75FFA92519FDCF20BEFD7609A1" box="[571,653,1182,1203]" pageId="9" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in Santana do Livramento—Área de Proteção Ambiental do Rio Ibirapuitã, Rio Grande do Sul, Brazil.
<emphasis id="B945060EFFA92519FE48209CFE3509C3" bold="true" box="[444,462,1212,1233]" pageId="9" pageNumber="189">A</emphasis>
,
<emphasis id="B945060EFFA92519FE2A209CFE1409C3" bold="true" box="[478,495,1212,1233]" pageId="9" pageNumber="189">B</emphasis>
. Fazenda Rincão dos Moraes, showing typical sandy and stony soils.
<emphasis id="B945060EFFA92519FB2C209CFB1109C0" bold="true" box="[1240,1258,1212,1234]" pageId="9" pageNumber="189">C</emphasis>
,
<emphasis id="B945060EFFA92519FB0E209CFAF709C3" bold="true" box="[1274,1292,1212,1233]" pageId="9" pageNumber="189">D</emphasis>
. Large basalt outcrops, “Afloramento Passo do Ferrão”.
</paragraph>
</caption>
<subSubSection id="C32B8997FFA9251AFF33213DFC7F0CBF" lastPageId="10" lastPageNumber="190" pageId="9" pageNumber="189" type="description">
<paragraph id="8B8EDA1CFFA9251AFF33213DFC7F0CBF" blockId="9.[151,1437,1309,2021]" lastBlockId="10.[151,1437,151,429]" lastPageId="10" lastPageNumber="190" pageId="9" pageNumber="189">
<emphasis id="B945060EFFA92519FF33213DFEA20824" bold="true" box="[199,345,1309,1334]" pageId="9" pageNumber="189">Description.</emphasis>
For most aspects the general morphology fits that described for the genus and the
<taxonomicName id="4C31A19FFFA92519FAB7213DFA670827" authorityName="Mello-Leitao" authorityYear="1924" box="[1347,1436,1309,1333]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="9" pageNumber="189" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B945060EFFA92519FAB7213DFA670827" box="[1347,1436,1309,1333]" italics="true" pageId="9" pageNumber="189">fasciata</emphasis>
</taxonomicName>
species-group by
<bibRefCitation id="EFA0A7EDFFA92519FEA92161FD670848" author="Botero-Trujillo, R. &amp; Ott, R. &amp; Carvalho, L. S." box="[349,668,1345,1370]" pageId="9" pageNumber="189" pagination="3 - 44" refId="ref7208" refString="Botero-Trujillo, R., Ott, R. &amp; Carvalho, L. S. (2017) Systematic revision and phylogeny of the South American sun-spider genus Gaucha Mello-Leitao (Solifugae: Mummuciidae), with description of four new species and two new generic synonymies. Arthropod Systematics &amp; Phylogeny, 75 (1), 3 - 44." type="journal article" year="2017">
Botero-Trujillo
<emphasis id="B945060EFFA92519FDFA2162FDBB084B" box="[526,576,1345,1369]" italics="true" pageId="9" pageNumber="189">et al</emphasis>
. (2017)
</bibRefCitation>
.
<emphasis id="B945060EFFA92519FD532161FD130848" bold="true" box="[679,744,1345,1370]" italics="true" pageId="9" pageNumber="189">Color</emphasis>
: Propeltidium brown, with some slightly lighter areas; ocular tubercle brown to black. Chelicerae with manus predominantly brown and fingers reddish. Meso-, metapeltidium and dorsal surface of opisthosoma with a three-dark-band design typical of the family: tergites with broad, median, longitudinal brown band, and paired, thinner lateral whitish bands; pleural membranes with sub-dorsal black and sub-ventral white bands; white band of opisthosomal pleural membrane with black marks surrounding the socket of most setae, and black pigment along the inter-segmental transversal vertices especially on posterior half; sternites yellowish brown, with lateral margins conspicuously darkened, especially on two/three posteriormost sternites. Ventral surface of prosoma uniformly yellowish; sternum lighter than coxae. Pedipalps and legs dark brown especially on dorsal and prolateral surfaces, lighter on ventral and retrolateral surfaces; pedipalp with telotarsus darker than the rest of pedipalp. Malleoli whitish, often with distal margin darkened.
<emphasis id="B945060EFFA92519FB9E22A4FB0C0B8F" bold="true" box="[1130,1271,1668,1693]" italics="true" pageId="9" pageNumber="189">Morphology</emphasis>
: Opisthosoma with scarce ctenidia on 1
<superScript id="7C447754FFA92519FE432288FE3A0BA4" attach="left" box="[439,449,1704,1718]" fontSize="6" pageId="9" pageNumber="189">st</superScript>
and 2
<superScript id="7C447754FFA92519FDFF2288FDE00BA4" attach="left" box="[523,539,1704,1718]" fontSize="6" pageId="9" pageNumber="189">nd</superScript>
post-genital sternites (spiracular sternites), more abundant on 3
<superScript id="7C447754FFA92519FB0C2288FAFE0BA4" attach="left" box="[1272,1285,1704,1718]" fontSize="6" pageId="9" pageNumber="189">rd</superScript>
and 4
<superScript id="7C447754FFA92519FABA2288FAA10BA4" attach="left" box="[1358,1370,1704,1718]" fontSize="6" pageId="9" pageNumber="189">th</superScript>
postgenital sternites (post-spiracular sternites III); ctenidia filiform and setiform, similar in thickness to the bifid setae (in male and female). Chelicerae, fixed finger with median teeth series comprising all primary teeth, i.e., FP, FM, FD; with one (
<collectingCountry id="F3269A8CFFA92519FEB42338FE800A22" box="[320,379,1816,1840]" name="Micronesia" pageId="9" pageNumber="189">FSM</collectingCountry>
) secondary teeth series and without FSD teeth. Legs II and III: basitarsus with row of three proventral, row of three retroventral, and one distal subventral spiniform setae, in a 2.2.3 rather staggered pattern; telotarsus bi-segmented with pro- and retroventral rows of five and four spiniform setae respectively, in a 1.2.2/2.2 pattern. Leg IV: basitarsus with row of four proventral and one distal retroventral spiniform setae, in a 1.1.1.2 pattern; telotarsus bi-segmented with incomplete (ventral) segmentation on first (basal) tarsomere, with pro- and retroventral rows of six spiniform setae each, in a 2.2.2-2/2.2 pattern.
<emphasis id="B945060EFFA92519FC4623ECFC150AF7" bold="true" box="[946,1006,1996,2021]" pageId="9" pageNumber="189">Male</emphasis>
: Metric data as in
<tableCitation id="C6B3EFA7FFA92519FB3123ECFAE10AF7" box="[1221,1306,1996,2021]" captionStart="TABLE 1" captionStartId="11.[151,239,152,175]" captionText="TABLE 1. Metric data for Gaucha ramirezi sp. nov. and Gaucha santana sp. nov. Measurements in millimeters for one male and one female. L = length; W = width; H = height. 1 Measured along medial axis, from the propeltidium anterior margin to the opisthosoma posterior margin. 2 Measured in dorsal view at widest point. 3 Measured in retrolateral view parallel to longitudinal axis of chelicera, from the fixed finger apex to anterolateral propeltidial lobe anterior margin. 4 Measured in retrolateral view, along vertical axis at highest part of manus. 5 Sum of individual segment lengths. 6 Maximum height. 7 Measurement excludes claws." pageId="9" pageNumber="189">Table 1</tableCitation>
. Cheliceral fixed finger with well-developed FP and FM primary teeth, FD tooth much smaller, graded as FP=FM&gt;&gt;FD; mucron markedly long, without subterminal flange (STF). Movable finger MP tooth pronounced, markedly taller than MM; mucron short, typical of the
<taxonomicName id="4C31A19FFFAA251AFDBD24C0FD590DEA" authorityName="Mello-Leitao" authorityYear="1924" box="[585,674,224,248]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="10" pageNumber="190" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B945060EFFAA251AFDBD24C0FD590DEA" box="[585,674,224,248]" italics="true" pageId="10" pageNumber="190">fasciata</emphasis>
</taxonomicName>
species-group, with gnathal edge carina very prominent and convex on lateral aspect. Flagellum inflated and narrowing anteriorly, with minute spicules along prodorsal and proventral margins.
<emphasis id="B945060EFFAA251AFF0A2507FEAE0C52" bold="true" box="[254,341,295,320]" pageId="10" pageNumber="190">Female</emphasis>
: Metric data as in
<tableCitation id="C6B3EFA7FFAA251AFDD12507FD820C52" box="[549,633,295,320]" captionStart="TABLE 1" captionStartId="11.[151,239,152,175]" captionText="TABLE 1. Metric data for Gaucha ramirezi sp. nov. and Gaucha santana sp. nov. Measurements in millimeters for one male and one female. L = length; W = width; H = height. 1 Measured along medial axis, from the propeltidium anterior margin to the opisthosoma posterior margin. 2 Measured in dorsal view at widest point. 3 Measured in retrolateral view parallel to longitudinal axis of chelicera, from the fixed finger apex to anterolateral propeltidial lobe anterior margin. 4 Measured in retrolateral view, along vertical axis at highest part of manus. 5 Sum of individual segment lengths. 6 Maximum height. 7 Measurement excludes claws." pageId="10" pageNumber="190">Table 1</tableCitation>
. Similar to male but without male-specific secondary sexual characters. Chelicera on lateral aspect, fixed finger dorsal margin moderately curved and without angular dorsal crest, with highest elevation at level between FM and FD teeth. Fixed finger robust, with mucron distinctly curved towards the venter. Movable finger MP tooth moderately taller than MM tooth.
</paragraph>
</subSubSection>
<caption id="DF4E8A94FFAA251AFF63237EFE970AE0" httpUri="https://zenodo.org/record/2622776/files/figure.png" pageId="10" pageNumber="190" startId="10.[151,250,1886,1908]" targetBox="[150,1436,511,1838]" targetPageId="10">
<paragraph id="8B8EDA1CFFAA251AFF63237EFE970AE0" blockId="10.[151,1436,1886,2034]" pageId="10" pageNumber="190">
<emphasis id="B945060EFFAA251AFF63237EFEE00A66" bold="true" box="[151,283,1886,1908]" pageId="10" pageNumber="190">FIGURE 7.</emphasis>
<taxonomicName id="4C31A19FFFAA251AFEDD237EFE230A67" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[297,472,1886,1909]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="10" pageNumber="190" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAA251AFEDD237EFE230A67" box="[297,472,1886,1909]" italics="true" pageId="10" pageNumber="190">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAA251AFE13237FFD9B0A61" bold="true" box="[487,608,1886,1908]" pageId="10" pageNumber="190">
<taxonomicNameLabel id="A276BB75FFAA251AFE13237FFDBB0A66" box="[487,576,1887,1908]" pageId="10" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. Male holotype, habitus, dorsal aspect (MCN-Sol-018).
<emphasis id="B945060EFFAA251AFB22237FFB1C0A66" bold="true" box="[1238,1255,1887,1908]" pageId="10" pageNumber="190">B</emphasis>
. Male paratype, propeltidium and chelicerae, dorsal aspect (MCN-Sol-044).
<emphasis id="B945060EFFAA251AFCF0235DFCED0A81" bold="true" box="[772,790,1917,1939]" pageId="10" pageNumber="190">C</emphasis>
. Ibid. for female paratype (MCN-Sol-047).
<emphasis id="B945060EFFAA251AFB15235EFB080A81" bold="true" box="[1249,1267,1918,1939]" pageId="10" pageNumber="190">D</emphasis>
. Male paratype, opisthosoma, lateral aspect (MCN-Sol-044).
<emphasis id="B945060EFFAA251AFD9C23BFFD820AA6" bold="true" box="[616,633,1951,1972]" pageId="10" pageNumber="190">E</emphasis>
. Male paratype, opisthosoma, 1
<superScript id="7C447754FFAA251AFC3223BCFC340ABA" attach="left" box="[966,975,1948,1960]" fontSize="5" pageId="10" pageNumber="190">st</superScript>
to 4
<superScript id="7C447754FFAA251AFBF523BBFBF70ABA" attach="left" box="[1025,1036,1947,1960]" fontSize="5" pageId="10" pageNumber="190">th</superScript>
post-genital sternites (MCN-Sol-044); selected ctenidia are indicated by arrows (zooming-in is recommended for proper visualization). Scale bars: 2 mm (A); 1 mm (BD); 0.25 mm (E).
</paragraph>
</caption>
<caption id="DF4E8A94FFAB251BFF6324B8FD510C4E" pageId="11" pageNumber="191">
<paragraph id="8B8EDA1CFFAB251BFF6324B8FD510C4E" blockId="11.[151,1436,151,348]" pageId="11" pageNumber="191">
<emphasis id="B945060EFFAB251BFF6324B8FEF10DBD" bold="true" box="[151,266,152,175]" pageId="11" pageNumber="191">TABLE 1.</emphasis>
Metric data for
<taxonomicName id="4C31A19FFFAB251BFE4C24B7FD960DBD" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[440,621,151,175]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="ramirezi" status="sp. nov.">
<emphasis id="B945060EFFAB251BFE4C24B7FD960DBD" box="[440,621,151,175]" italics="true" pageId="11" pageNumber="191">Gaucha ramirezi</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFD8024B8FD310DBD" bold="true" box="[628,714,152,175]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFD8024B8FD310DBD" box="[628,714,152,175]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C31A19FFFAB251BFD0A24B7FC560DBD" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[766,941,151,175]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAB251BFD0A24B7FC560DBD" box="[766,941,151,175]" italics="true" pageId="11" pageNumber="191">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFC4024B8FBF10DBD" bold="true" box="[948,1034,152,175]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFC4024B8FBF10DBD" box="[948,1034,152,175]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Measurements in millimeters for one male and one female. L = length; W = width; H = height.
<superScript id="7C447754FFAB251BFCFD2498FCEB0DD6" attach="right" box="[777,784,184,196]" fontSize="5" pageId="11" pageNumber="191">1</superScript>
Measured along medial axis, from the propeltidium anterior margin to the opisthosoma posterior margin.
<superScript id="7C447754FFAB251BFD8A24FBFD7E0DF5" attach="right" box="[638,645,219,231]" fontSize="5" pageId="11" pageNumber="191">2</superScript>
Measured in dorsal view at widest point.
<superScript id="7C447754FFAB251BFBBA24FBFBAE0DF5" attach="right" box="[1102,1109,219,231]" fontSize="5" pageId="11" pageNumber="191">3</superScript>
Measured in retrolateral view parallel to longitudinal axis of chelicera, from the fixed finger apex to anterolateral propeltidial lobe anterior margin.
<superScript id="7C447754FFAB251BFA6024DDFA600C1B" attach="right" box="[1428,1435,253,265]" fontSize="5" pageId="11" pageNumber="191">4</superScript>
Measured in retrolateral view, along vertical axis at highest part of manus.
<superScript id="7C447754FFAB251BFC1F2500FC090C3E" attach="right" box="[1003,1010,288,300]" fontSize="5" pageId="11" pageNumber="191">5</superScript>
Sum of individual segment lengths.
<superScript id="7C447754FFAB251BFA602500FA600C3E" attach="right" box="[1428,1435,288,300]" fontSize="5" pageId="11" pageNumber="191">6</superScript>
Maximum height.
<superScript id="7C447754FFAB251BFEA82563FE980C5D" attach="left" box="[348,355,323,335]" fontSize="5" pageId="11" pageNumber="191">7</superScript>
Measurement excludes claws.
</paragraph>
</caption>
<paragraph id="8B8EDA1CFFAB251BFF6A2558FB73086E" pageId="11" pageNumber="191">
<table id="F93128BCFFABDAEFFF6A2558FA8A086D" box="[158,1393,376,1407]" gridcols="4" gridrows="24" pageId="11" pageNumber="191">
<tr id="3501D85EFFABDAEFFF6A2558FA8A0C9C" box="[158,1393,376,398]" gridrow="0" pageId="11" pageNumber="191" rowspan-1="1">
<th id="76D0B122FFABDAEFFF6A2558FEBC0C9C" box="[158,327,376,398]" gridcol="0" gridrow="0" pageId="11" pageNumber="191">Species</th>
<th id="76D0B122FFABDAEFFD6A2558FC170C9C" box="[670,1004,376,398]" gridcol="2" gridrow="0" pageId="11" pageNumber="191">
<taxonomicName id="4C31A19FFFAB251BFD6A2558FCB20C9C" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[670,841,376,398]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="ramirezi" status="sp. nov.">
<emphasis id="B945060EFFAB251BFD6A2558FCB20C9C" box="[670,841,376,398]" italics="true" pageId="11" pageNumber="191">Gaucha ramirezi</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFCA42558FC5B0C9F" bold="true" box="[848,928,376,397]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFCA42558FC5B0C9F" box="[848,928,376,397]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76D0B122FFABDAEFFBEB2558FA8A0C9C" box="[1055,1393,376,398]" gridcol="3" gridrow="0" pageId="11" pageNumber="191">
<taxonomicName id="4C31A19FFFAB251BFBEB2558FB3E0C9C" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[1055,1221,376,398]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAB251BFBEB2558FB3E0C9C" box="[1055,1221,376,398]" italics="true" pageId="11" pageNumber="191">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFB382558FAE70C9F" bold="true" box="[1228,1308,376,397]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFB382558FAE70C9F" box="[1228,1308,376,397]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="3501D85EFFABDAEFFF6A2581FA8A0CC4" box="[158,1393,417,470]" gridrow="1" pageId="11" pageNumber="191" rowspan-1="1">
<th id="76D0B122FFABDAEFFF6A2581FEBC0CC4" box="[158,327,417,470]" gridcol="0" gridrow="1" pageId="11" pageNumber="191">Voucher</th>
<td id="76D0B122FFABDAEFFD6A2581FC170CC4" box="[670,1004,417,470]" gridcol="2" gridrow="1" pageId="11" pageNumber="191">Male holotype MACN-Ar 39101 [Female paratype MACN-Ar]</td>
<td id="76D0B122FFABDAEFFBEB2581FA8A0CC4" box="[1055,1393,417,470]" gridcol="3" gridrow="1" pageId="11" pageNumber="191">Male holotype MCN-Sol-018 [Female paratype MCN-Sol-041]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A25C8FA8A0F10" box="[158,1393,488,514]" gridrow="2" pageId="11" pageNumber="191">
<th id="76D0B122FFABDAEFFF6A25C8FD8A0F10" box="[158,625,488,514]" colspan="2" colspanRight="1" gridcol="0" gridrow="2" pageId="11" pageNumber="191">Total body L (w/o chelicerae)1</th>
<td id="76D0B122FFABDAEFFD6A25C8FC170F10" box="[670,1004,488,514]" gridcol="2" gridrow="2" pageId="11" pageNumber="191">8.11[9.58]</td>
<td id="76D0B122FFABDAEFFBEB25C8FA8A0F10" box="[1055,1393,488,514]" gridcol="3" gridrow="2" pageId="11" pageNumber="191">8.51[15.16]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2635FA8A0F39" box="[158,1393,533,555]" gridrow="3" pageId="11" pageNumber="191">
<th id="76D0B122FFABDAEFFF6A2635FEBC0F39" box="[158,327,533,555]" gridcol="0" gridrow="3" pageId="11" pageNumber="191">Propeltidium</th>
<td id="76D0B122FFABDAEFFE832635FD8A0F39" box="[375,625,533,555]" gridcol="1" gridrow="3" pageId="11" pageNumber="191">L</td>
<td id="76D0B122FFABDAEFFD6A2635FC170F39" box="[670,1004,533,555]" gridcol="2" gridrow="3" pageId="11" pageNumber="191">1.57[1.77]</td>
<td id="76D0B122FFABDAEFFBEB2635FA8A0F39" box="[1055,1393,533,555]" gridcol="3" gridrow="3" pageId="11" pageNumber="191">1.97[2.50]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A261DFA8A0F45" box="[158,1393,573,599]" gridrow="4" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE83261DFD8A0F45" box="[375,625,573,599]" gridcol="1" gridrow="4" pageId="11" pageNumber="191">W2</td>
<td id="76D0B122FFABDAEFFD6A261DFC170F45" box="[670,1004,573,599]" gridcol="2" gridrow="4" pageId="11" pageNumber="191">1.67[2.17]</td>
<td id="76D0B122FFABDAEFFBEB261DFA8A0F45" box="[1055,1393,573,599]" gridcol="3" gridrow="4" pageId="11" pageNumber="191">2.37[3.59]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A264AFA8A0F90" box="[158,1393,618,642]" gridrow="5" pageId="11" pageNumber="191">
<th id="76D0B122FFABDAEFFF6A264AFEBC0F90" box="[158,327,618,642]" gridcol="0" gridrow="5" pageId="11" pageNumber="191">Chelicera</th>
<td id="76D0B122FFABDAEFFE83264AFD8A0F90" box="[375,625,618,642]" gridcol="1" gridrow="5" pageId="11" pageNumber="191">L3</td>
<td id="76D0B122FFABDAEFFD6A264AFC170F90" box="[670,1004,618,642]" gridcol="2" gridrow="5" pageId="11" pageNumber="191">2.10[2.80]</td>
<td id="76D0B122FFABDAEFFBEB264AFA8A0F90" box="[1055,1393,618,642]" gridcol="3" gridrow="5" pageId="11" pageNumber="191">2.50[4.52]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A26B5FA8A0FBC" box="[158,1393,661,686]" gridrow="6" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE8326B5FD8A0FBC" box="[375,625,661,686]" gridcol="1" gridrow="6" pageId="11" pageNumber="191">W2</td>
<td id="76D0B122FFABDAEFFD6A26B5FC170FBC" box="[670,1004,661,686]" gridcol="2" gridrow="6" pageId="11" pageNumber="191">0.75[1.00]</td>
<td id="76D0B122FFABDAEFFBEB26B5FA8A0FBC" box="[1055,1393,661,686]" gridcol="3" gridrow="6" pageId="11" pageNumber="191">1.13[1.63]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A26E1FA8A0FC8" box="[158,1393,705,730]" gridrow="7" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE8326E1FD8A0FC8" box="[375,625,705,730]" gridcol="1" gridrow="7" pageId="11" pageNumber="191">H4</td>
<td id="76D0B122FFABDAEFFD6A26E1FC170FC8" box="[670,1004,705,730]" gridcol="2" gridrow="7" pageId="11" pageNumber="191">0.77[1.00]</td>
<td id="76D0B122FFABDAEFFBEB26E1FA8A0FC8" box="[1055,1393,705,730]" gridcol="3" gridrow="7" pageId="11" pageNumber="191">1.17[1.67]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A26CDFA8A0E14" box="[158,1393,749,774]" gridrow="8" pageId="11" pageNumber="191" rowspan-1="1">
<th id="76D0B122FFABDAEFFF6A26CDFEBC0E14" box="[158,327,749,774]" gridcol="0" gridrow="8" pageId="11" pageNumber="191">Pedipalp total L5</th>
<td id="76D0B122FFABDAEFFD6A26CDFC170E14" box="[670,1004,749,774]" gridcol="2" gridrow="8" pageId="11" pageNumber="191">4.90[5.33]</td>
<td id="76D0B122FFABDAEFFBEB26CDFA8A0E14" box="[1055,1393,749,774]" gridcol="3" gridrow="8" pageId="11" pageNumber="191">6.59[7.87]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A273AFA8A0E22" box="[158,1393,794,816]" gridrow="9" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE83273AFD8A0E22" box="[375,625,794,816]" gridcol="1" gridrow="9" pageId="11" pageNumber="191">Femur L</td>
<td id="76D0B122FFABDAEFFD6A273AFC170E22" box="[670,1004,794,816]" gridcol="2" gridrow="9" pageId="11" pageNumber="191">1.80[1.83]</td>
<td id="76D0B122FFABDAEFFBEB273AFA8A0E22" box="[1055,1393,794,816]" gridcol="3" gridrow="9" pageId="11" pageNumber="191">2.53[2.87]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2763FA8A0E4B" box="[158,1393,835,857]" gridrow="10" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832763FD8A0E4B" box="[375,625,835,857]" gridcol="1" gridrow="10" pageId="11" pageNumber="191">Tibia L</td>
<td id="76D0B122FFABDAEFFD6A2763FC170E4B" box="[670,1004,835,857]" gridcol="2" gridrow="10" pageId="11" pageNumber="191">1.43[1.57]</td>
<td id="76D0B122FFABDAEFFBEB2763FA8A0E4B" box="[1055,1393,835,857]" gridcol="3" gridrow="10" pageId="11" pageNumber="191">1.93[2.33]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A274CFA8A0E97" box="[158,1393,876,901]" gridrow="11" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE83274CFD8A0E97" box="[375,625,876,901]" gridcol="1" gridrow="11" pageId="11" pageNumber="191">Tibia W2</td>
<td id="76D0B122FFABDAEFFD6A274CFC170E97" box="[670,1004,876,901]" gridcol="2" gridrow="11" pageId="11" pageNumber="191">0.42[0.48]</td>
<td id="76D0B122FFABDAEFFBEB274CFA8A0E97" box="[1055,1393,876,901]" gridcol="3" gridrow="11" pageId="11" pageNumber="191">0.55[0.67]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A27B8FA8A0EBC" box="[158,1393,920,942]" gridrow="12" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE8327B8FD8A0EBC" box="[375,625,920,942]" gridcol="1" gridrow="12" pageId="11" pageNumber="191">Basitarsus + telotarsus L</td>
<td id="76D0B122FFABDAEFFD6A27B8FC170EBC" box="[670,1004,920,942]" gridcol="2" gridrow="12" pageId="11" pageNumber="191">1.67[1.93]</td>
<td id="76D0B122FFABDAEFFBEB27B8FA8A0EBC" box="[1055,1393,920,942]" gridcol="3" gridrow="12" pageId="11" pageNumber="191">2.13[2.67]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A27E1FA8A0EC8" box="[158,1393,961,986]" gridrow="13" pageId="11" pageNumber="191" rowspan-1="1">
<th id="76D0B122FFABDAEFFF6A27E1FEBC0EC8" box="[158,327,961,986]" gridcol="0" gridrow="13" pageId="11" pageNumber="191">Leg I total L5</th>
<td id="76D0B122FFABDAEFFD6A27E1FC170EC8" box="[670,1004,961,986]" gridcol="2" gridrow="13" pageId="11" pageNumber="191">3.73[4.00]</td>
<td id="76D0B122FFABDAEFFBEB27E1FA8A0EC8" box="[1055,1393,961,986]" gridcol="3" gridrow="13" pageId="11" pageNumber="191">5.31[5.93]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A27CDFA8A0911" box="[158,1393,1005,1027]" gridrow="14" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE8327CDFD8A0911" box="[375,625,1005,1027]" gridcol="1" gridrow="14" pageId="11" pageNumber="191">Patella L</td>
<td id="76D0B122FFABDAEFFD6A27CDFC170911" box="[670,1004,1005,1027]" gridcol="2" gridrow="14" pageId="11" pageNumber="191">1.10[1.17]</td>
<td id="76D0B122FFABDAEFFBEB27CDFA8A0911" box="[1055,1393,1005,1027]" gridcol="3" gridrow="14" pageId="11" pageNumber="191">1.67[1.60]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2036FA8A093E" box="[158,1393,1046,1068]" gridrow="15" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832036FD8A093E" box="[375,625,1046,1068]" gridcol="1" gridrow="15" pageId="11" pageNumber="191">Tibia L</td>
<td id="76D0B122FFABDAEFFD6A2036FC17093E" box="[670,1004,1046,1068]" gridcol="2" gridrow="15" pageId="11" pageNumber="191">1.23[1.33]</td>
<td id="76D0B122FFABDAEFFBEB2036FA8A093E" box="[1055,1393,1046,1068]" gridcol="3" gridrow="15" pageId="11" pageNumber="191">1.67[2.10]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2060FA8A0944" box="[158,1393,1088,1110]" gridrow="16" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832060FD8A0944" box="[375,625,1088,1110]" gridcol="1" gridrow="16" pageId="11" pageNumber="191">Basitarsus L</td>
<td id="76D0B122FFABDAEFFD6A2060FC170944" box="[670,1004,1088,1110]" gridcol="2" gridrow="16" pageId="11" pageNumber="191">0.77[0.87]</td>
<td id="76D0B122FFABDAEFFBEB2060FA8A0944" box="[1055,1393,1088,1110]" gridcol="3" gridrow="16" pageId="11" pageNumber="191">1.07[1.33]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2049FA8A096D" box="[158,1393,1129,1151]" gridrow="17" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832049FD8A096D" box="[375,625,1129,1151]" gridcol="1" gridrow="17" pageId="11" pageNumber="191">Telotarsus L</td>
<td id="76D0B122FFABDAEFFD6A2049FC17096D" box="[670,1004,1129,1151]" gridcol="2" gridrow="17" pageId="11" pageNumber="191">0.63[0.63]</td>
<td id="76D0B122FFABDAEFFBEB2049FA8A096D" box="[1055,1393,1129,1151]" gridcol="3" gridrow="17" pageId="11" pageNumber="191">0.90[0.90]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A20B2FA8A09B9" box="[158,1393,1170,1195]" gridrow="18" pageId="11" pageNumber="191">
<th id="76D0B122FFABDAEFFF6A20B2FD8A09B9" box="[158,625,1170,1195]" colspan="2" colspanRight="1" gridcol="0" gridrow="18" pageId="11" pageNumber="191">Leg IV total L (w/o claws)5</th>
<td id="76D0B122FFABDAEFFD6A20B2FC1709B9" box="[670,1004,1170,1195]" gridcol="2" gridrow="18" pageId="11" pageNumber="191">6.24[5.90]</td>
<td id="76D0B122FFABDAEFFBEB20B2FA8A09B9" box="[1055,1393,1170,1195]" gridcol="3" gridrow="18" pageId="11" pageNumber="191">8.67[9.84]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A209EFA8A09C6" box="[158,1393,1214,1236]" gridrow="19" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE83209EFD8A09C6" box="[375,625,1214,1236]" gridcol="1" gridrow="19" pageId="11" pageNumber="191">Patella L</td>
<td id="76D0B122FFABDAEFFD6A209EFC1709C6" box="[670,1004,1214,1236]" gridcol="2" gridrow="19" pageId="11" pageNumber="191">2.07[1.97]</td>
<td id="76D0B122FFABDAEFFBEB209EFA8A09C6" box="[1055,1393,1214,1236]" gridcol="3" gridrow="19" pageId="11" pageNumber="191">2.87[3.10]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A20C7FA8A0812" box="[158,1393,1255,1280]" gridrow="20" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE8320C7FD8A0812" box="[375,625,1255,1280]" gridcol="1" gridrow="20" pageId="11" pageNumber="191">Patella H6</td>
<td id="76D0B122FFABDAEFFD6A20C7FC170812" box="[670,1004,1255,1280]" gridcol="2" gridrow="20" pageId="11" pageNumber="191">0.57[0.65]</td>
<td id="76D0B122FFABDAEFFBEB20C7FA8A0812" box="[1055,1393,1255,1280]" gridcol="3" gridrow="20" pageId="11" pageNumber="191">0.80[0.97]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2134FA8A0838" box="[158,1393,1300,1322]" gridrow="21" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832134FD8A0838" box="[375,625,1300,1322]" gridcol="1" gridrow="21" pageId="11" pageNumber="191">Tibia L</td>
<td id="76D0B122FFABDAEFFD6A2134FC170838" box="[670,1004,1300,1322]" gridcol="2" gridrow="21" pageId="11" pageNumber="191">1.83[1.83]</td>
<td id="76D0B122FFABDAEFFBEB2134FA8A0838" box="[1055,1393,1300,1322]" gridcol="3" gridrow="21" pageId="11" pageNumber="191">2.63[3.07]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A211DFA8A0841" box="[158,1393,1341,1363]" gridrow="22" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE83211DFD8A0841" box="[375,625,1341,1363]" gridcol="1" gridrow="22" pageId="11" pageNumber="191">Basitarsus L</td>
<td id="76D0B122FFABDAEFFD6A211DFC170841" box="[670,1004,1341,1363]" gridcol="2" gridrow="22" pageId="11" pageNumber="191">1.47[1.37]</td>
<td id="76D0B122FFABDAEFFBEB211DFA8A0841" box="[1055,1393,1341,1363]" gridcol="3" gridrow="22" pageId="11" pageNumber="191">2.00[2.30]</td>
</tr>
<tr id="3501D85EFFABDAEFFF6A2146FA8A086D" box="[158,1393,1382,1407]" gridrow="23" pageId="11" pageNumber="191" rowspan-0="1">
<td id="76D0B122FFABDAEFFE832146FD8A086D" box="[375,625,1382,1407]" gridcol="1" gridrow="23" pageId="11" pageNumber="191">Telotarsus L 7</td>
<td id="76D0B122FFABDAEFFD6A2146FC17086D" box="[670,1004,1382,1407]" gridcol="2" gridrow="23" pageId="11" pageNumber="191">0.87[0.73]</td>
<td id="76D0B122FFABDAEFFBEB2146FA8A086D" box="[1055,1393,1382,1407]" gridcol="3" gridrow="23" pageId="11" pageNumber="191">1.17[1.37]</td>
</tr>
</table>
</paragraph>
<subSubSection id="C32B8997FFAB251CFF33219FFB0C0C52" lastPageId="12" lastPageNumber="192" pageId="11" pageNumber="191" type="distribution">
<paragraph id="8B8EDA1CFFAB251BFF33219FFD7A0A0E" blockId="11.[151,1437,1471,2037]" pageId="11" pageNumber="191">
<emphasis id="B945060EFFAB251BFF33219FFCE708CA" bold="true" box="[199,796,1471,1496]" pageId="11" pageNumber="191">Distribution, habitat and biological observations.</emphasis>
<taxonomicName id="4C31A19FFFAB251BFCD121E0FC1A08CA" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[805,993,1472,1496]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAB251BFCD121E0FC1A08CA" box="[805,993,1472,1496]" italics="true" pageId="11" pageNumber="191">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFC1E219FFBBD08CA" bold="true" box="[1002,1094,1471,1496]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFC1E219FFBBD08CA" box="[1002,1094,1471,1496]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known only from the
<typeStatus id="548A64BEFFAB251BFA9821E0FA6708CA" box="[1388,1436,1472,1496]" pageId="11" pageNumber="191">type</typeStatus>
locality, Santana do Livramento—Área de Proteção Ambiental do Rio Ibirapuitã, in the state of
<collectingRegion id="49F514FEFFAB251BFB3121C4FA6308EF" box="[1221,1432,1508,1533]" country="Brazil" name="Rio Grande do Sul" pageId="11" pageNumber="191">Rio Grande do Sul</collectingRegion>
, southern
<collectingCountry id="F3269A8CFFAB251BFEF42227FEB30B32" box="[256,328,1543,1568]" name="Brazil" pageId="11" pageNumber="191">Brazil</collectingCountry>
. The Ibirapuitã Environmental Protection Area is a conservation unit located in the northernmost portion of the Uruguayan Savanna ecoregion (NT0710 in
<bibRefCitation id="EFA0A7EDFFAB251BFCB1220CFBE30B56" author="Olson, D. M. &amp; Dinerstein, E. &amp; Wikramanayake, E. D. &amp; Burgess, N. D. &amp; Powell, G. V. N. &amp; Underwood, E. C. &amp; D'Amico, J. A. &amp; Itoua, I. &amp; Strand, H. E. &amp; Morrison, J. C. &amp; Loucks, C. J. &amp; Alltutt, T. F. &amp; Ricketts, T. H. &amp; Kura, Y. &amp; Lamoreux, J. F. &amp; Wettengel, W. W. &amp; Hedao, P. &amp; Kassem, K. R." box="[837,1048,1580,1605]" pageId="11" pageNumber="191" pagination="933 - 938" refId="ref7637" refString="Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Alltutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P. &amp; Kassem, K. R. (2001) Terrestrial ecoregions of the world: a new map of life on earth. BioScience, 51, 933 - 938. https: // doi. org / 10.1641 / 0006 - 3568 (2001) 051 [0933: TEOTWA] 2.0. CO; 2" type="journal article" year="2001">
Olson
<emphasis id="B945060EFFAB251BFC60220DFC320B56" box="[916,969,1580,1604]" italics="true" pageId="11" pageNumber="191">et al</emphasis>
. 2001
</bibRefCitation>
), into the biome of Tropical and Subtropical Grasslands, Savannas, and Shrublands. Grasslands dominate the area, and local geology is characterized by shallow, sandy or stony soils with low capacity for water retention (
<bibRefCitation id="EFA0A7EDFFAB251BFBBA2254FABB0B9F" author="Overbeck, G. E. &amp; Boldrini, I. I. &amp; Carmo, M. R. B. &amp; Garcia, E. N. &amp; Moro, R. S. &amp; Pinto, C. E. &amp; Trevisan, R. &amp; Zannin, A." box="[1102,1344,1652,1677]" pageId="11" pageNumber="191" pagination="31 - 42" refId="ref7807" refString="Overbeck, G. E., Boldrini, I. I., Carmo, M. R. B., Garcia, E. N., Moro, R. S., Pinto, C. E., Trevisan, R. &amp; Zannin, A. (2015) Fisionomia dos campos. In: Pillar, V. D. &amp; Lange, O. (Org.), Campos do Sul. 1 st Edition. Rede Campos Sulinos, Porto Alegre, pp. v + 31 - 42." type="book chapter" year="2015">
Overbeck
<emphasis id="B945060EFFAB251BFB302255FB0F0B9E" box="[1220,1268,1652,1676]" italics="true" pageId="11" pageNumber="191">et al</emphasis>
. 2015
</bibRefCitation>
). Rocky outcrops of basalt are notable formations; in the easternmost part of the protected area there is a place where sandstone emerges below the basalt cover, in a formation known as &quot;Costa do Haedo&quot;. Specimens of
<taxonomicName id="4C31A19FFFAB251BFB09229CFA8F0BC6" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[1277,1396,1724,1748]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAB251BFB09229CFA8F0BC6" box="[1277,1396,1724,1748]" italics="true" pageId="11" pageNumber="191">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFA8E229CFF330BEA" bold="true" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFA8E229CFF330BEA" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have been observed in open fields of stony soils with little grassy vegetation, often resembling small rocky islands, or running on large basalt outcrops.
</paragraph>
<paragraph id="8B8EDA1CFFAB251BFF332308FD550ABE" blockId="11.[151,1437,1471,2037]" pageId="11" pageNumber="191">
No specimens of
<taxonomicName id="4C31A19FFFAB251BFE6D2308FDEE0A52" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[409,533,1832,1856]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="11" pageNumber="191" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAB251BFE6D2308FDEE0A52" box="[409,533,1832,1856]" italics="true" pageId="11" pageNumber="191">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAB251BFDD52307FD840A52" bold="true" box="[545,639,1831,1856]" pageId="11" pageNumber="191">
<taxonomicNameLabel id="A276BB75FFAB251BFDD52307FD840A52" box="[545,639,1831,1856]" pageId="11" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were found in morning hours. The peak of activity of this species seemingly starts at noon, and activity can be noticed until right before twilight begins. With respect to seasonality, the species exhibits the greatest activity in spring and summer, especially from November to January, and this presumably coincides with the breeding season.
</paragraph>
<paragraph id="8B8EDA1CFFAB251CFF332398FDFB0C0E" blockId="11.[151,1437,1471,2037]" lastBlockId="12.[151,1437,151,320]" lastPageId="12" lastPageNumber="192" pageId="11" pageNumber="191">
In the field, adult females were sometimes observed digging burrows in the late afternoon. In early
<date id="FF8FFCDCFFAB251BFADD2397FF290AE6" pageId="11" pageNumber="191" value="2012-12">December 2012</date>
, one of these specimens was taken to the laboratory where shortly afterwards, in its new place over a litter layer, laid
<specimenCount id="9D371195FFAC251CFEE724B8FE940DA2" box="[275,367,152,176]" pageId="12" pageNumber="192" type="egg">73 eggs</specimenCount>
, but none hatched. A year later, in
<date id="FF8FFCDCFFAC251CFCF924B7FC3E0DA2" box="[781,965,151,176]" pageId="12" pageNumber="192" value="2013-12">December 2013</date>
, another female was captured in digging behavior and taken to the laboratory; there she laid
<specimenCount id="9D371195FFAC251CFD26249CFCD10DC7" box="[722,810,188,213]" pageId="12" pageNumber="192" type="egg">25 eggs</specimenCount>
, 23 from which larvae emerged after 37 days. All larvae died and no other observations could be performed. Images of the cited egg-laying females are available in Botero- Trujillo
<emphasis id="B945060EFFAC251CFF062525FED80C0E" box="[242,291,260,284]" italics="true" pageId="12" pageNumber="192">et al</emphasis>
. (2017: figs. 3E,F).
</paragraph>
<paragraph id="8B8EDA1CFFAC251CFF332507FB0C0C52" blockId="12.[151,1437,151,320]" box="[199,1271,295,320]" pageId="12" pageNumber="192">
<emphasis id="B945060EFFAC251CFF332507FEEA0C52" bold="true" box="[199,273,295,320]" pageId="12" pageNumber="192">Notes.</emphasis>
No any other mummuciid species was found in the area inhabited by
<taxonomicName id="4C31A19FFFAC251CFBD42508FB6C0C52" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[1056,1175,296,320]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="192" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAC251CFBD42508FB6C0C52" box="[1056,1175,296,320]" italics="true" pageId="12" pageNumber="192">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAC251CFB6A2507FB0C0C52" bold="true" box="[1182,1271,295,320]" pageId="12" pageNumber="192">
<taxonomicNameLabel id="A276BB75FFAC251CFB6A2507FB0C0C52" box="[1182,1271,295,320]" pageId="12" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<caption id="DF4E8A94FFAC251CFF63224DFB3A0BD2" httpUri="https://zenodo.org/record/2622778/files/figure.png" pageId="12" pageNumber="192" startId="12.[151,250,1645,1667]" targetBox="[150,1436,398,1586]" targetPageId="12">
<paragraph id="8B8EDA1CFFAC251CFF63224DFB3A0BD2" blockId="12.[151,1435,1644,1728]" pageId="12" pageNumber="192">
<emphasis id="B945060EFFAC251CFF63224DFEEF0B90" bold="true" box="[151,276,1645,1667]" pageId="12" pageNumber="192">FIGURE 8.</emphasis>
<taxonomicName id="4C31A19FFFAC251CFEE8224CFE3E0B91" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[284,453,1644,1667]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="192" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAC251CFEE8224CFE3E0B91" box="[284,453,1644,1667]" italics="true" pageId="12" pageNumber="192">Gaucha santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAC251CFE39224DFDDB0B90" bold="true" box="[461,544,1645,1666]" pageId="12" pageNumber="192">
<taxonomicNameLabel id="A276BB75FFAC251CFE39224DFDDB0B90" box="[461,544,1645,1666]" pageId="12" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, chelicerae.
<emphasis id="B945060EFFAC251CFD50224DFD4D0B90" bold="true" box="[676,694,1645,1666]" pageId="12" pageNumber="192">A</emphasis>
,
<emphasis id="B945060EFFAC251CFD30224DFD2E0B90" bold="true" box="[708,725,1645,1666]" pageId="12" pageNumber="192">B</emphasis>
. Right chelicera of female paratype (MCN-Sol-015).
<emphasis id="B945060EFFAC251CFAF0224DFAED0B90" bold="true" box="[1284,1302,1645,1666]" pageId="12" pageNumber="192">A</emphasis>
. Retrolateral aspect.
<emphasis id="B945060EFFAC251CFF1722ACFF0F0BB3" bold="true" box="[227,244,1676,1697]" pageId="12" pageNumber="192">B</emphasis>
. Prolateral aspect.
<emphasis id="B945060EFFAC251CFE4022ABFE1B0BB3" bold="true" box="[436,480,1675,1697]" pageId="12" pageNumber="192">CF</emphasis>
. Right chelicera of male paratype (MCN-Sol-041).
<emphasis id="B945060EFFAC251CFC1922ABFC040BB3" bold="true" box="[1005,1023,1675,1697]" pageId="12" pageNumber="192">C</emphasis>
. Retrolateral aspect.
<emphasis id="B945060EFFAC251CFB2122ACFB1C0BB3" bold="true" box="[1237,1255,1676,1697]" pageId="12" pageNumber="192">D</emphasis>
. Prolateral aspect (under SEM).
<emphasis id="B945060EFFAC251CFEDC228AFEC20BAD" bold="true" box="[296,313,1706,1727]" pageId="12" pageNumber="192">E</emphasis>
. Flagellum.
<emphasis id="B945060EFFAC251CFE4D228AFE3D0BAD" bold="true" box="[441,454,1706,1727]" pageId="12" pageNumber="192">F</emphasis>
. Ibid. (under SEM). Scale bars: 1 mm (A,B); 0.5 mm (C,D); 0.2 mm (E,F).
</paragraph>
</caption>
<subSubSection id="C32B8997FFAC251CFF3322CCFF140AAB" pageId="12" pageNumber="192" type="description">
<paragraph id="8B8EDA1CFFAC251CFF3322CCFF140AAB" blockId="12.[151,1436,1772,1977]" pageId="12" pageNumber="192">
<emphasis id="B945060EFFAC251CFF3322CCFECA0A17" bold="true" box="[199,305,1772,1797]" pageId="12" pageNumber="192">Barcode.</emphasis>
Three COI sequences (657 bp) obtained from different specimens of
<taxonomicName id="4C31A19FFFAC251CFB8922CDFB000A17" authority="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro, 2019" authorityName="Botero-Trujillo &amp; Ott &amp; Mattoni &amp; Nime &amp; Ojanguren-Affilastro" authorityYear="2019" box="[1149,1275,1773,1797]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="192" phylum="Arthropoda" rank="species" species="santana" status="sp. nov.">
<emphasis id="B945060EFFAC251CFB8922CDFB000A17" box="[1149,1275,1773,1797]" italics="true" pageId="12" pageNumber="192">G. santana</emphasis>
</taxonomicName>
<emphasis id="B945060EFFAC251CFAF322CCFA9C0A17" bold="true" box="[1287,1383,1772,1797]" pageId="12" pageNumber="192">
<taxonomicNameLabel id="A276BB75FFAC251CFAF322CCFA9C0A17" box="[1287,1383,1772,1797]" pageId="12" pageNumber="192" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
had identical nucleotide composition (0% intra-specific divergence). An identification search in BOLD Systems for this sequence (Barcode Index Number: BOLD:ADD5797) indicated closer match with
<taxonomicName id="4C31A19FFFAC251CFB2C2315FA670A5F" authorityName="Mello-Leitao" authorityYear="1924" box="[1240,1436,1845,1869]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="192" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="B945060EFFAC251CFB2C2315FA670A5F" box="[1240,1436,1845,1869]" italics="true" pageId="12" pageNumber="192">Gaucha fasciata</emphasis>
</taxonomicName>
(BIN:BOLD:ADD5134) followed by
<taxonomicName id="4C31A19FFFAC251CFDB92379FD000A62" authorityName="Botero-Trujillo, Ott &amp; Carvalho" authorityYear="2017" box="[589,763,1881,1905]" class="Arachnida" family="Mummuciidae" genus="Gaucha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Solifugae" pageId="12" pageNumber="192" phylum="Arthropoda" rank="species" species="curupi">
<emphasis id="B945060EFFAC251CFDB92379FD000A62" box="[589,763,1881,1905]" italics="true" pageId="12" pageNumber="192">Gaucha curupi</emphasis>
</taxonomicName>
(BIN:BOLD:ADD8503), with 90.6690.67% and 89.76 90.21% similarity range values, respectively.
<figureCitation id="130AC699FFAC251CFD5A235DFCEA0A87" box="[686,785,1917,1942]" captionStart="FIGURE 9" captionStartId="13.[151,250,1537,1559]" captionTargetBox="[194,1396,900,1483]" captionTargetId="figure@13.[150,1436,878,1515]" captionTargetPageId="13" captionText="FIGURE 9. Neighbor-Joining clustering for selected species of the fasciata species-group of Gaucha Mello-Leitão, 1924." httpUri="https://zenodo.org/record/2622780/files/figure.png" pageId="12" pageNumber="192">Figure 9</figureCitation>
shows the clustering pattern of sequences of the three species.
</paragraph>
</subSubSection>
</treatment>
</document>