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52 KiB
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404 lines
52 KiB
XML
<document id="F1A0E90EFA7AC0E21BF8F79232F65134" ID-CLB-Dataset="8176" ID-DOI="10.5852/ejt.2021.734.1233" ID-GBIF-Dataset="fd8f5cb9-13e9-4c90-ac15-502fa7ee8d38" ID-ISSN="2118-9773" ID-Zenodo-Dep="4542107" ID-ZooBank="663A0C2D-E3CF-4B0C-8BA1-566B174829A2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613423098920" checkinUser="plazi" docAuthor="Montoya-Molina, Santiago & Vaz-De, Fernando Z." docDate="2021" docId="03850669FFA4FFF8FE3D24C33ABFCC55" docLanguage="en" docName="ejt-734_1-64.pdf" docOrigin="European Journal of Taxonomy 734" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius (Luederwaldtinia) validipilosus" docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="FFBC7E11FF90FFCEFFAD26453B59CB56" masterDocTitle="Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="53" updateTime="1698929631379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="0E12095BD907D85F6B10E60D14B180E1">Taxonomic review of the Dichotomius (Luederwaldtinia) agenor species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="6A667C096A6C6FA8E747BF75211F243E" type="personal">
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<mods:namePart id="A952382C3DF58F490793D11D8380B859">Montoya-Molina, Santiago</mods:namePart>
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<mods:affiliation id="7AD56934D1CF141B461E4E91E1A1F35F">Federal University of Mato Grosso, Institute of Biosciences, Programa de Pós Graduaç „ o em Ecología e Conservaç „ o da Biodiversidade, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil. Faculty of Environmental Sciences, Czech University of Life Sciences Prague, Kamýcká 129, Praha-Suchdol, 16500, Czech Republic. Federal University of Mato Grosso, Institute of Biosciences, Department of Biology and Zoology, Av. Fernando Correa da Costa, 2367, Boa Esperança, Cuiabá, MT, Brazil.</mods:affiliation>
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<mods:nameIdentifier id="3AD44DC31112516249AA7AB5DD13C513" type="email">horomo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B22EC66765C9AEA43C3DD2627629212F">Vaz-De, Fernando Z.</mods:namePart>
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<mods:title id="167EFF51CF77BC9E5740E3F01A0CA245">European Journal of Taxonomy</mods:title>
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<mods:part id="3FEFBF96CBCA88FA825A0A975DB262C3">
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<mods:date id="761583010FD61D8D195F1D2DE3B5EDE0">2021</mods:date>
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<mods:number id="193DA9E7F9547BD9BCEC32C622CD9B89">2021-02-11</mods:number>
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<mods:number id="3AD8F8FDCB5755E7D611013AB0610856">734</mods:number>
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<treatment id="03850669FFA4FFF8FE3D24C33ABFCC55" ID-DOI="http://doi.org/10.5281/zenodo.4545867" ID-GBIF-Taxon="177420807" ID-Zenodo-Dep="4545867" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55" httpUri="http://treatment.plazi.org/id/03850669FFA4FFF8FE3D24C33ABFCC55" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
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<subSubSection id="C336E4F4FFA4FFFAFE3D24C3382EC993" pageId="52" pageNumber="53" type="nomenclature">
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<paragraph id="8B93B77FFFA4FFFAFE3D24C3382EC993" blockId="52.[400,1188,646,709]" pageId="52" pageNumber="53">
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<taxonomicName id="4C2CCCFCFFA4FFFAFE3D24C33FFDC9F7" ID-CoL="35ND9" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[400,1188,646,673]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus" subGenus="Luederwaldtinia">
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<emphasis id="B9586B6DFFA4FFFAFE3D24C33972C9F7" bold="true" box="[400,555,646,673]" italics="true" pageId="52" pageNumber="53">Dichotomius</emphasis>
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(
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<emphasis id="B9586B6DFFA4FFFAFD9124C3385AC9F7" bold="true" box="[572,771,646,673]" italics="true" pageId="52" pageNumber="53">Luederwaldtinia</emphasis>
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)
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<emphasis id="B9586B6DFFA4FFFAFCB924C338F5C9F7" bold="true" box="[788,940,646,673]" italics="true" pageId="52" pageNumber="53">validipilosus</emphasis>
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(
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<bibRefCitation id="EFBDCA8EFFA4FFFAFC1724C33FC2C9F7" author="Luederwaldt H." box="[954,1179,646,673]" pageId="52" pageNumber="53" pagination="353 - 361" refId="ref30150" refString="Luederwaldt H. 1931. Tres especies novas de Pinotus, supplementos a Memoria " As especies Brasileiras do genero Pinotus ". na " Revista do Museu Paulista " Tomo XVI, 1929, P 603 SS. Revista do Museu Paulista 17: 353 - 361." type="journal article" year="1931">Luederwaldt, 1931</bibRefCitation>
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)
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</taxonomicName>
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<figureCitation id="1317ABFAFFA4FFFAFD1124EF3849C992" box="[700,784,682,708]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="52" pageNumber="53">Fig. 18</figureCitation>
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, Map 15
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</paragraph>
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</subSubSection>
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<subSubSection id="C336E4F4FFA4FFFAFF1024A83999C83A" pageId="52" pageNumber="53" type="reference_group">
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<paragraph id="8B93B77FFFA4FFFAFF1024A83885C851" blockId="52.[189,988,748,775]" box="[189,988,748,775]" pageId="52" pageNumber="53">
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<treatmentCitationGroup id="AB3C9051FFA4FFFAFF1024A83885C851" box="[189,988,748,775]" pageId="52" pageNumber="53">
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<treatmentCitation id="0A8D916EFFA4FFFAFF1024A8398DC850" author="Luederwaldt H." box="[189,724,748,774]" page="103" pageId="52" pageNumber="53" year="1931">
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<taxonomicName id="4C2CCCFCFFA4FFFAFF1024A8398DC850" ID-CoL="4HZWX" authority="Luederwaldt, 1931: 103" authorityName="Luederwaldt" authorityPageNumber="103" authorityYear="1931" box="[189,724,748,774]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus">
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<emphasis id="B9586B6DFFA4FFFAFF1024A83AEDC850" box="[189,436,748,774]" italics="true" pageId="52" pageNumber="53">Pinotus validipilosus</emphasis>
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<bibRefCitation id="EFBDCA8EFFA4FFFAFE1624A9398DC850" author="Luederwaldt H." box="[443,724,748,774]" pageId="52" pageNumber="53" pagination="353 - 361" refId="ref30150" refString="Luederwaldt H. 1931. Tres especies novas de Pinotus, supplementos a Memoria " As especies Brasileiras do genero Pinotus ". na " Revista do Museu Paulista " Tomo XVI, 1929, P 603 SS. Revista do Museu Paulista 17: 353 - 361." type="journal article" year="1931">Luederwaldt, 1931: 103</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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(original description).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF10256A39B2C81F" blockId="52.[189,747,814,877]" box="[189,747,814,841]" pageId="52" pageNumber="53">
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<treatmentCitationGroup id="AB3C9051FFA4FFFAFF10256A39B2C81F" box="[189,747,814,841]" pageId="52" pageNumber="53">
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<taxonomicName id="4C2CCCFCFFA4FFFAFF10256A3AEDC81E" ID-CoL="4HZWX" authorityName="Luederwaldt" authorityYear="1931" box="[189,436,814,840]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus">
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<emphasis id="B9586B6DFFA4FFFAFF10256A3AEDC81E" box="[189,436,814,840]" italics="true" pageId="52" pageNumber="53">Pinotus validipilosus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A8D916EFFA4FFFAFE7C256B39BEC81F" author="Blackwelder R. E." box="[465,743,814,841]" page="208" pageId="52" pageNumber="53" year="1944">
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<bibRefCitation id="EFBDCA8EFFA4FFFAFE7C256B39BEC81F" author="Blackwelder R. E." box="[465,743,814,841]" pageId="52" pageNumber="53" pagination="189 - 341" refId="ref29526" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 208</bibRefCitation>
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</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF1025173999C83A" blockId="52.[189,747,814,877]" box="[189,704,850,877]" pageId="52" pageNumber="53">
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<treatmentCitationGroup id="AB3C9051FFA4FFFAFF1025173999C83A" box="[189,704,850,877]" pageId="52" pageNumber="53">
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<taxonomicName id="4C2CCCFCFFA4FFFAFF1025173AA9C83A" ID-CoL="35ND9" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[189,496,850,876]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus">
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<emphasis id="B9586B6DFFA4FFFAFF1025173AA9C83A" box="[189,496,850,876]" italics="true" pageId="52" pageNumber="53">Dichotomius validipilosus</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0A8D916EFFA4FFFAFDA0251639E5C83A" author="Roze J. A." box="[525,700,850,877]" page="44" pageId="52" pageNumber="53" year="1955">
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<bibRefCitation id="EFBDCA8EFFA4FFFAFDA0251639E5C83A" author="Roze J. A." box="[525,700,850,877]" pageId="52" pageNumber="53" pagination="39 - 63" refId="ref30793" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C336E4F4FFA4FFFAFF1025D13F91CFE3" pageId="52" pageNumber="53" type="diagnosis">
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||
<paragraph id="8B93B77FFFA4FFFAFF1025D13A6FC8F8" blockId="52.[189,310,916,942]" box="[189,310,916,942]" pageId="52" pageNumber="53">
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<heading id="D0DB0013FFA4FFFAFF1025D13A6FC8F8" bold="true" box="[189,310,916,942]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
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<emphasis id="B9586B6DFFA4FFFAFF1025D13A6FC8F8" bold="true" box="[189,310,916,942]" pageId="52" pageNumber="53">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF1025813F91CFE3" blockId="52.[187,1399,964,1205]" pageId="52" pageNumber="53">
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<taxonomicName id="4C2CCCFCFFA4FFFAFF1025813AB6C888" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[189,495,964,990]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus">
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<emphasis id="B9586B6DFFA4FFFAFF1025813AB6C888" box="[189,495,964,990]" italics="true" pageId="52" pageNumber="53">Dichotomius validipilosus</emphasis>
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</taxonomicName>
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is separated from the other species in the group by the following combination of characters: head surface subrugose. Clypeus not rounded, clypeal margin displaying projection protruding on anterior portion of clypeo-genal suture. Cephalic carina conical, median tubercle apically emarginate, lateral tubercles slightly developed. Pronotal anterior disc with declivity, median edge deeper than lateral sides resembling two underdeveloped lobes. Pronotal median disc punctation fine. Anterior and lateral edges of metasternum with long and abundant setae. Elytral interstriae microsculpture shagreened. Sixth abdominal sternite marginally lower or depressed than other sternites.
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</paragraph>
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</subSubSection>
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<subSubSection id="C336E4F4FFA4FFFAFF1022983E0ECD9F" pageId="52" pageNumber="53" type="materials_examined">
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<paragraph id="8B93B77FFFA4FFFAFF1022983AF3CFA1" blockId="52.[189,426,1245,1271]" box="[189,426,1245,1271]" pageId="52" pageNumber="53">
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<heading id="D0DB0013FFA4FFFAFF1022983AF3CFA1" bold="true" box="[189,426,1245,1271]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
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<emphasis id="B9586B6DFFA4FFFAFF1022983AF3CFA1" bold="true" box="[189,426,1245,1271]" pageId="52" pageNumber="53">Material examined</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF4723493A05CE70" blockId="52.[189,1399,1292,1426]" box="[234,348,1292,1318]" pageId="52" pageNumber="53">
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<emphasis id="B9586B6DFFA4FFFAFF4723493A05CE70" bold="true" box="[234,348,1292,1318]" pageId="52" pageNumber="53">
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<typeStatus id="549709DDFFA4FFFAFF4723493A05CE70" box="[234,348,1292,1318]" pageId="52" pageNumber="53" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF10237538D8CEC4" blockId="52.[189,1399,1292,1426]" pageId="52" pageNumber="53">
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<materialsCitation id="3B44BD22FFA4FFFAFF10237538D8CEC4" ID-GBIF-Occurrence="3037418530" collectionCode="MNHN" collectorName="Ex-Museo N. Van de Poil" country="Venezuela" location="VENEZUELA" pageId="52" pageNumber="53" specimenCount="1" specimenCount-male="1" typeStatus="typus">
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<collectingCountry id="F33BF7EFFFA4FFFAFF1023753A32CE1C" box="[189,363,1328,1354]" name="Venezuela" pageId="52" pageNumber="53">VENEZUELA</collectingCountry>
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•
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||
<specimenCount id="9D2A7CF6FFA4FFFAFED323753AF1CE1C" box="[382,424,1326,1354]" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
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; “Venézuela Caracas” [white, p]/ “Ex-Museo N. Van de Poil” [white, p]/ “MUSÉUM PARIS 1938 COLL. A. BOUCOMONT” [green, p]/ “
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<typeStatus id="549709DDFFA4FFFAFCE2231138C5CE38" box="[847,924,1364,1390]" pageId="52" pageNumber="53">Typus</typeStatus>
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” [red with black margins, p]/ “
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||
<taxonomicName id="4C2CCCFCFFA4FFFAFAB023103A0ACEC4" authority="Montoya-Molina & Vaz-De, 2021" authorityName="Montoya-Molina & Vaz-De" authorityYear="2021" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="validipilosus" status="sp. nov.">
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||
<emphasis id="B9586B6DFFA4FFFAFAB023103A0ACEC4" italics="true" pageId="52" pageNumber="53">Pinotus validipilosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26BD616FFA4FFFAFEF7233D3AC0CEC4" box="[346,409,1400,1426]" pageId="52" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
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Lueder. Det. 1931” [white, hw];
|
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<collectionCode id="ED3D2FBAFFA4FFFAFCB2233D3824CEC4" box="[799,893,1400,1426]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="52" pageNumber="53">MNHN</collectionCode>
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||
.
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||
</materialsCitation>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF4723FF3AB8CE82" blockId="52.[189,1399,1466,1600]" box="[234,481,1466,1492]" pageId="52" pageNumber="53">
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<emphasis id="B9586B6DFFA4FFFAFF4723FF3AB8CE82" bold="true" box="[234,481,1466,1492]" pageId="52" pageNumber="53">Additional material</emphasis>
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</paragraph>
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<paragraph id="8B93B77FFFA4FFFAFF10239B393CCD16" blockId="52.[189,1399,1466,1600]" pageId="52" pageNumber="53">
|
||
<materialsCitation id="3B44BD22FFA4FFFAFF10239B3AF5CD4A" ID-GBIF-Occurrence="3037418368" collectingDate="2008-06" collectionCode="CEMT" collectorName="Andrea Esparza" country="Colombia" elevation="2800" location="Rio Rancheria" municipality="Cermanke" pageId="52" pageNumber="53" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="La Guajira">
|
||
<collectingCountry id="F33BF7EFFFA4FFFAFF10239B3A01CEAE" box="[189,344,1502,1528]" name="Colombia" pageId="52" pageNumber="53">COLOMBIA</collectingCountry>
|
||
–
|
||
<emphasis id="B9586B6DFFA4FFFAFED1239B3952CEA1" bold="true" box="[380,523,1501,1528]" pageId="52" pageNumber="53">
|
||
<collectingRegion id="49E8799DFFA4FFFAFED1239B3952CEA1" box="[380,523,1501,1528]" country="Colombia" name="La Guajira" pageId="52" pageNumber="53">La Guajira</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFD87239B390FCEAE" box="[554,598,1500,1528]" pageId="52" pageNumber="53" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFDCA239B39CFCEAE" box="[615,662,1502,1528]" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
|
||
; Cermanke,
|
||
<location id="8EF3E1A4FFA4FFFAFC9C239B38B9CEAE" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55:8EF3E1A4FFA4FFFAFC9C239B38B9CEAE" box="[817,992,1502,1528]" country="Colombia" municipality="Cermanke" name="Rio Rancheria" pageId="52" pageNumber="53" stateProvince="La Guajira">Rio Rancheria</location>
|
||
;
|
||
<elevation id="0001504CFFA4FFFAFC43239B3FD2CEAE" box="[1006,1163,1502,1528]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="52" pageNumber="53" unit="m" value="2800.0">
|
||
<quantity id="4CD41A9AFFA4FFFAFC43239B3F10CEAE" box="[1006,1097,1502,1528]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="52" pageNumber="53" unit="m" value="2800.0">2800 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FF9291BFFFA4FFFAFB36239B3E4ACEAE" box="[1179,1299,1502,1528]" pageId="52" pageNumber="53" value="2008-06">
|
||
<collectingDate id="EFD66857FFA4FFFAFB36239B3E4ACEAE" box="[1179,1299,1502,1528]" pageId="52" pageNumber="53" value="2008-06">Jun. 2008</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FFA4FFFAFA8D239B3A43CD4A" pageId="52" pageNumber="53">Andrea Esparza</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3D2FBAFFA4FFFAFEF420473AF5CD4A" box="[345,428,1538,1564]" collectionName="CEMT" pageId="52" pageNumber="53">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B44BD22FFA4FFFAFE6720473939CD16" ID-GBIF-Occurrence="3037418389" collectingDate="1973-05-14" collectionCode="CEMT" collectorName="Campbell & Howden" country="Colombia" elevation="914" location="Campana." pageId="52" pageNumber="53" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena">
|
||
<emphasis id="B9586B6DFFA4FFFAFE672047390FCD4A" bold="true" box="[458,598,1538,1564]" pageId="52" pageNumber="53">
|
||
<collectingRegion id="49E8799DFFA4FFFAFE672047390FCD4A" box="[458,598,1538,1564]" country="Colombia" name="Magdalena" pageId="52" pageNumber="53">Magdalena</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFDC3204739C3CD4A" box="[622,666,1538,1564]" pageId="52" pageNumber="53" type="female">1 ♀</specimenCount>
|
||
;
|
||
<locationDeviation id="9A21AE5DFFA4FFFAFD0A204738FECD4A" box="[679,935,1538,1564]" location="Sta. Marta" pageId="52" pageNumber="53">
|
||
<quantity id="4CD41A9AFFA4FFFAFD0A204739A9CD4A" box="[679,752,1538,1564]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="52" pageNumber="53" unit="km" value="24.0">24 km</quantity>
|
||
S of Sta. Marta
|
||
</locationDeviation>
|
||
,
|
||
<location id="8EF3E1A4FFA4FFFAFC1F20473F75CD4A" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55:8EF3E1A4FFA4FFFAFC1F20473F75CD4A" box="[946,1068,1538,1564]" country="Colombia" name="Campana." pageId="52" pageNumber="53" stateProvince="Magdalena">Campana.</location>
|
||
;
|
||
<elevation id="0001504CFFA4FFFAFB9B20473F91CD4A" box="[1078,1224,1538,1564]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="52" pageNumber="53" unit="ft" value="3000.0">
|
||
<quantity id="4CD41A9AFFA4FFFAFB9B20473FD0CD4A" box="[1078,1161,1538,1564]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="52" pageNumber="53" unit="ft" value="3000.0">3000 ft</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FF9291BFFFA4FFFAFB7920473E2BCD4A" box="[1236,1394,1538,1564]" pageId="52" pageNumber="53" value="1973-05-14">
|
||
<collectingDate id="EFD66857FFA4FFFAFB7920473E2BCD4A" box="[1236,1394,1538,1564]" pageId="52" pageNumber="53" value="1973-05-14">14 May 1973</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FFA4FFFAFF1020603A76CD69" box="[189,303,1573,1599]" pageId="52" pageNumber="53">Campbell</collectorName>
|
||
and
|
||
<collectorName id="26D9D2A9FFA4FFFAFEC520603A92CD69" box="[360,459,1573,1599]" pageId="52" pageNumber="53">Howden</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3D2FBAFFA4FFFAFDA020633939CD16" box="[525,608,1574,1600]" collectionName="CEMT" pageId="52" pageNumber="53">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA4FFFAFF10202D3E0ECD9F" blockId="52.[189,1399,1639,1738]" pageId="52" pageNumber="53">
|
||
<materialsCitation id="3B44BD22FFA4FFFAFF10202D3FBECDD4" ID-GBIF-Occurrence="3037418491" box="[189,1255,1639,1666]" collectingDate="2004-07-06" collectionCode="CEMT" collectorName="D. Garcia" country="Venezuela" elevation="50" location="Cuyagua" pageId="52" pageNumber="53" specimenCount="2" specimenCount-female="2" stateProvince="Aragua">
|
||
<collectingCountry id="F33BF7EFFFA4FFFAFF10202D3A33CDD4" box="[189,362,1640,1666]" name="Venezuela" pageId="52" pageNumber="53">VENEZUELA</collectingCountry>
|
||
–
|
||
<emphasis id="B9586B6DFFA4FFFAFE2A20223ABFCDD7" bold="true" box="[391,486,1639,1665]" pageId="52" pageNumber="53">
|
||
<collectingRegion id="49E8799DFFA4FFFAFE2A20223ABFCDD7" box="[391,486,1639,1665]" country="Venezuela" name="Aragua" pageId="52" pageNumber="53">Aragua</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFE53202D3918CDD4" box="[510,577,1640,1666]" pageId="52" pageNumber="53" type="female">2 ♀♀</specimenCount>
|
||
;
|
||
<location id="8EF3E1A4FFA4FFFAFDE2202D39E2CDD4" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55:8EF3E1A4FFA4FFFAFDE2202D39E2CDD4" box="[591,699,1640,1666]" country="Venezuela" name="Cuyagua" pageId="52" pageNumber="53" stateProvince="Aragua">Cuyagua</location>
|
||
;
|
||
<elevation id="0001504CFFA4FFFAFD6A2022381BCDD7" box="[711,834,1639,1666]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="53" unit="m" value="50.0">
|
||
<quantity id="4CD41A9AFFA4FFFAFD6A2022385ACDD4" box="[711,771,1639,1666]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="52" pageNumber="53" unit="m" value="50.0">50 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="FF9291BFFFA4FFFAFCFD2022388DCDD4" box="[848,980,1639,1666]" pageId="52" pageNumber="53" value="2004-07-06">
|
||
<collectingDate id="EFD66857FFA4FFFAFCFD2022388DCDD4" box="[848,980,1639,1666]" pageId="52" pageNumber="53" value="2004-07-06">6 Jul. 2004</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FFA4FFFAFC4C202D3F0BCDD4" box="[993,1106,1640,1666]" pageId="52" pageNumber="53">D. García</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3D2FBAFFA4FFFAFB39202D3FBECDD4" box="[1172,1255,1640,1666]" collectionName="CEMT" pageId="52" pageNumber="53">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B44BD22FFA4FFFAFAA520223A95CD9F" ID-GBIF-Occurrence="3037418570" collectingDate="2015-03-13" collectionCode="CEMT" collectorName="Racconcia" country="Venezuela" elevation="1380" latitude="8.238334" location="Mutucuy" longLatPrecision="21" longitude="-71.28667" municipality="Arzobispo Chacon" pageId="52" pageNumber="53" specimenCount="6" specimenCount-female="2" specimenCount-male="4" stateProvince="Merida">
|
||
<emphasis id="B9586B6DFFA4FFFAFAA520223E3CCDD7" bold="true" box="[1288,1381,1639,1665]" pageId="52" pageNumber="53">
|
||
<collectingRegion id="49E8799DFFA4FFFAFAA520223E3CCDD7" box="[1288,1381,1639,1665]" country="Venezuela" name="Merida" pageId="52" pageNumber="53">Merida</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFF1020C93A5ACDF3" box="[189,259,1673,1702]" pageId="52" pageNumber="53" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFEB920C93A02CDF0" box="[276,347,1676,1702]" pageId="52" pageNumber="53" type="female">2 ♀♀</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BF72D05FFA4FFFAFEC620CE3914CDF3" box="[363,589,1675,1701]" pageId="52" pageNumber="53">Arzobispo Chacón</collectingMunicipality>
|
||
,
|
||
<location id="8EF3E1A4FFA4FFFAFDF620C93990CDF0" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55:8EF3E1A4FFA4FFFAFDF620C93990CDF0" box="[603,713,1676,1702]" country="Venezuela" latitude="8.238334" longLatPrecision="21" longitude="-71.28667" municipality="Arzobispo Chacon" name="Mutucuy" pageId="52" pageNumber="53" stateProvince="Merida">Mutucuy</location>
|
||
;
|
||
<elevation id="0001504CFFA4FFFAFD7520C9382ECDF3" box="[728,887,1675,1702]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="52" pageNumber="53" unit="m" value="1380.0">
|
||
<quantity id="4CD41A9AFFA4FFFAFD7520C9386DCDF0" box="[728,820,1676,1702]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="52" pageNumber="53" unit="m" value="1380.0">1380 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<geoCoordinate id="EE18D1B8FFA4FFFAFC2520CF3F49CDF0" box="[904,1040,1674,1702]" degrees="8" direction="north" minutes="14" orientation="latitude" pageId="52" pageNumber="53" precision="15" seconds="18" value="8.238334">8º14′18″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE18D1B8FFA4FFFAFBB220CF3FE2CDF0" box="[1055,1211,1674,1702]" degrees="71" direction="west" minutes="17" orientation="longitude" pageId="52" pageNumber="53" precision="15" seconds="12" value="-71.28667">71º17′12″ W</geoCoordinate>
|
||
;
|
||
<date id="FF9291BFFFA4FFFAFB6720C93E2BCDF0" box="[1226,1394,1675,1702]" pageId="52" pageNumber="53" value="2015-03-13">
|
||
<collectingDate id="EFD66857FFA4FFFAFB6720C93E2BCDF0" box="[1226,1394,1675,1702]" pageId="52" pageNumber="53" value="2015-03-13">13 Mar. 2015</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FFA4FFFAFF1020EA3A6ECD9F" box="[189,311,1711,1737]" pageId="52" pageNumber="53">Racconcia</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3D2FBAFFA4FFFAFED420EA3A95CD9F" box="[377,460,1711,1737]" collectionName="CEMT" pageId="52" pageNumber="53">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B44BD22FFA4FFFAFE4020EA3E0ACD9F" ID-GBIF-Occurrence="3037418357" box="[493,1363,1709,1738]" collectingDate="1985-07-15" collectionCode="CEMT" collectorName="B. Joffre" country="Venezuela" location="La Flautera" municipality="Palmira" pageId="52" pageNumber="53" specimenCount="2" specimenCount-male="2" stateProvince="Tachira">
|
||
<emphasis id="B9586B6DFFA4FFFAFE4020EA3917CD9F" bold="true" box="[493,590,1711,1737]" pageId="52" pageNumber="53">
|
||
<collectingRegion id="49E8799DFFA4FFFAFE4020EA3917CD9F" box="[493,590,1711,1737]" country="Venezuela" name="Tachira" pageId="52" pageNumber="53">Tachira</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D2A7CF6FFA4FFFAFDCA20EA39F3CD9F" box="[615,682,1709,1737]" pageId="52" pageNumber="53" type="male">2 ♂♂</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BF72D05FFA4FFFAFD1520EA384FCD9F" box="[696,790,1711,1737]" pageId="52" pageNumber="53">Palmira</collectingMunicipality>
|
||
,
|
||
<location id="8EF3E1A4FFA4FFFAFC8F20F538F5CD9F" LSID="urn:lsid:plazi:treatment:03850669FFA4FFF8FE3D24C33ABFCC55:8EF3E1A4FFA4FFFAFC8F20F538F5CD9F" box="[802,940,1711,1738]" country="Venezuela" municipality="Palmira" name="La Flautera" pageId="52" pageNumber="53" stateProvince="Tachira">La Flautera</location>
|
||
;
|
||
<date id="FF9291BFFFA4FFFAFC1520EA3F15CD9C" box="[952,1100,1711,1738]" pageId="52" pageNumber="53" value="1985-07-15">
|
||
<collectingDate id="EFD66857FFA4FFFAFC1520EA3F15CD9C" box="[952,1100,1711,1738]" pageId="52" pageNumber="53" value="1985-07-15">15 Jul. 1985</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26D9D2A9FFA4FFFAFBF520F53FE7CD9F" box="[1112,1214,1711,1738]" pageId="52" pageNumber="53">B. Joffre</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED3D2FBAFFA4FFFAFAAD20EA3E0ACD9F" box="[1280,1363,1711,1737]" collectionName="CEMT" pageId="52" pageNumber="53">CEMT</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FFA4FFF8FF1020B439B7CD71" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53" type="description">
|
||
<paragraph id="8B93B77FFFA4FFFAFF1020B43A32CC5D" blockId="52.[189,363,1777,1803]" box="[189,363,1777,1803]" pageId="52" pageNumber="53">
|
||
<heading id="D0DB0013FFA4FFFAFF1020B43A32CC5D" bold="true" box="[189,363,1777,1803]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
|
||
<emphasis id="B9586B6DFFA4FFFAFF1020B43A32CC5D" bold="true" box="[189,363,1777,1803]" pageId="52" pageNumber="53">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA4FFFAFF4721643A73CC6D" blockId="52.[189,590,1825,1887]" box="[234,298,1825,1851]" pageId="52" pageNumber="53">
|
||
<emphasis id="B9586B6DFFA4FFFAFF4721643A73CC6D" bold="true" box="[234,298,1825,1851]" pageId="52" pageNumber="53">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA4FFFAFF1021003917CC09" blockId="52.[189,590,1825,1887]" box="[189,590,1860,1887]" pageId="52" pageNumber="53">
|
||
MEASUREMENTS. Length:
|
||
<quantity id="4CD41A9AFFA4FFFAFE7321013910CC09" box="[478,585,1860,1887]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="52" pageNumber="53" unit="mm" value="21.6">21.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA4FFF8FF1021C23860CA3C" blockId="52.[188,1399,1926,2025]" lastBlockId="54.[189,1399,265,363]" lastPageId="54" lastPageNumber="55" pageId="52" pageNumber="53">
|
||
HEAD. Anterior surface subrugose (
|
||
<figureCitation id="1317ABFAFFA4FFFAFDFE21C339E5CCF7" box="[595,700,1926,1953]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="52" pageNumber="53">Fig. 18C</figureCitation>
|
||
), extending posteriorly not reaching base of cephalic carina. Clypeus not rounded, with large central emargination separating two rounded clypeal teeth, feebly angled upwards (
|
||
<figureCitation id="1317ABFAFFA4FFFAFE20218B397ACCBE" box="[397,547,1998,2024]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="52" pageNumber="53">Fig. 18A, C</figureCitation>
|
||
). Clypeal margin distinct, projection protruding on anterior portion of clypeo-genal suture. Cephalic carina, as wide as high, conical in shape. Median tubercle three times as high as lateral tubercles, apically emarginate. Lateral tubercles slightly developed. Ventral clypeal process truncate and broadened apically (see
|
||
<figureCitation id="1317ABFAFFA6FFF8FD7D27153872CA3C" box="[720,811,336,362]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="54" pageNumber="55">Fig. 2B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF53E7F7FFA5FFFBFF1021203935CCBF" ID-DOI="http://doi.org/10.5281/zenodo.4542143" ID-Zenodo-Dep="4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="53" pageNumber="54" startId="53.[189,232,1893,1919]" targetBox="[241,1349,296,1842]" targetPageId="53">
|
||
<paragraph id="8B93B77FFFA5FFFBFF1021203935CCBF" blockId="53.[188,1398,1891,2025]" pageId="53" pageNumber="54">
|
||
<emphasis id="B9586B6DFFA5FFFBFF1021203A42CC29" bold="true" box="[189,283,1893,1919]" pageId="53" pageNumber="54">Fig. 18.</emphasis>
|
||
<taxonomicName id="4C2CCCFCFFA5FFFBFE8821203800CCD6" authority="(Luederwaldt, 1931)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[293,857,1893,1920]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="validipilosus">
|
||
<emphasis id="B9586B6DFFA5FFFBFE8821203903CC29" box="[293,602,1893,1919]" italics="true" pageId="53" pageNumber="54">Dichotomius validipilosus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFBDCA8EFFA5FFFBFDC621203809CCD6" author="Luederwaldt H." box="[619,848,1893,1920]" pageId="53" pageNumber="54" pagination="353 - 361" refId="ref30150" refString="Luederwaldt H. 1931. Tres especies novas de Pinotus, supplementos a Memoria " As especies Brasileiras do genero Pinotus ". na " Revista do Museu Paulista " Tomo XVI, 1929, P 603 SS. Revista do Museu Paulista 17: 353 - 361." type="journal article" year="1931">Luederwaldt, 1931</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B9586B6DFFA5FFFBFCC9212038C2CC29" bold="true" box="[868,923,1893,1919]" pageId="53" pageNumber="54">A–B</emphasis>
|
||
. Dorsal habitus.
|
||
<emphasis id="B9586B6DFFA5FFFBFBC721203F27CC29" bold="true" box="[1130,1150,1893,1919]" pageId="53" pageNumber="54">A</emphasis>
|
||
. ♂.
|
||
<emphasis id="B9586B6DFFA5FFFBFB1921233F91CCD6" bold="true" box="[1204,1224,1894,1920]" pageId="53" pageNumber="54">B</emphasis>
|
||
. ♀.
|
||
<emphasis id="B9586B6DFFA5FFFBFB5321203E4ACC29" bold="true" box="[1278,1299,1893,1919]" pageId="53" pageNumber="54">C</emphasis>
|
||
. Lateral view of the aedeagus.
|
||
<emphasis id="B9586B6DFFA5FFFBFE1321CC3A8ACCF5" bold="true" box="[446,467,1929,1955]" pageId="53" pageNumber="54">D</emphasis>
|
||
. View from above of the head, basal third surface subrugose.
|
||
<emphasis id="B9586B6DFFA5FFFBFB0C21CC3FECCCF5" bold="true" box="[1185,1205,1929,1955]" pageId="53" pageNumber="54">E</emphasis>
|
||
. Ventral view of the aedeagus.
|
||
<emphasis id="B9586B6DFFA5FFFBFECF21E93AC0CC90" bold="true" box="[354,409,1964,1990]" pageId="53" pageNumber="54">F–G</emphasis>
|
||
. Lamella copulatrix.
|
||
<emphasis id="B9586B6DFFA5FFFBFD3921E939F2CC90" bold="true" box="[660,683,1964,1990]" pageId="53" pageNumber="54">H</emphasis>
|
||
. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B93B77FFFA6FFF8FF1027D73A7CC96C" blockId="54.[189,1399,402,570]" pageId="54" pageNumber="55">
|
||
PRONOTUM. Twice as wide as long and as wide as elytra. Pronotal median disc shiny and with fine punctures, ocellate punctures near posterior basal central margin. Anterior disc with sharp declivity, medial edge deeper than lateral sides, resembling two underdeveloped lobes (
|
||
<figureCitation id="1317ABFAFFA6FFF8FBCE279C3F8DCAA5" box="[1123,1236,473,499]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="54" pageNumber="55">Fig. 18A</figureCitation>
|
||
). Hypomeron external margin bearing rows of disorganized setigerous punctures, median disc smooth and shiny (see
|
||
<figureCitation id="1317ABFAFFA6FFF8FF1024653A41C96C" box="[189,280,544,570]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="54" pageNumber="55">Fig. 2E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF102427397EC823" blockId="54.[188,1399,610,885]" pageId="54" pageNumber="55">
|
||
ABDOMEN. Mesosternum and metepisternum surface microsculpture shagreened, ocellate setigerous punctures separated by once diameter of a puncture. Metasternum anterior portion next to basal margin of mesosternum and margins of mesocoxa microsculpture shagreened, with dense pilosity surrounding margins of mesocoxa joining medial and lateral portions of metasternum (see
|
||
<figureCitation id="1317ABFAFFA6FFF8FB2B24893FBEC9B1" box="[1158,1255,716,743]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="54" pageNumber="55">Fig. 2G</figureCitation>
|
||
,
|
||
<figureCitation id="1317ABFAFFA6FFF8FB5524893E30C9B1" box="[1272,1385,716,743]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="54" pageNumber="55">Fig. 18D</figureCitation>
|
||
). Longitudinal sulcus distinctly sculpted, ending as shallow mid-fossa. Sternites ocellate punctures along anterior margins. Sixth abdominal sternite marginally lower or depressed than other sternites (
|
||
<figureCitation id="1317ABFAFFA6FFF8FB5025513E30C878" box="[1277,1385,788,814]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="54" pageNumber="55">Fig. 18D</figureCitation>
|
||
). Pygidium completely delimited by margin. Large punctation restricted to basal margin. Medial and apical surfaces finely punctate.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF1025D83BACC8A8" blockId="54.[189,1399,925,1022]" pageId="54" pageNumber="55">
|
||
ELYTRA. Wider than long. Interstriae microsculpture shagreened (see
|
||
<figureCitation id="1317ABFAFFA6FFF8FC4725D83F62C8E1" box="[1002,1083,925,951]" captionStart="Fig" captionStartId="5.[189,232,1679,1705]" captionTargetBox="[278,1310,266,1653]" captionTargetId="figure-184@5.[278,1310,266,1653]" captionTargetPageId="5" captionText="Fig. 2. General characters that help differentiate the species of the Dichotomius (Luederwaldtinia) agenor species group. A–D. Ventral clypeal process. A. Cordate as in D. agenor (Harold, 1869). B. Truncate as in D. sagittarius (Harold, 1869) stat. rev. C. Bifurcate as in D. belus (Harold, 1880). D. Cylindrical as in D. henripittieri sp. nov. E–F. Hypomeral characters. E. Medial disc smooth, lateral margin with multiple rows of setae as in D. amplicollis (Harold, 1869). F. Medial disc punctate, lateral margin with single row of setae as in D. tristis (Luederwaldt, 1923). G–H. Metasternum characters. G. Setigerous punctures from the medial and lateral edges joining throughout the margin of the mesocaxa as in D. enioi sp. nov. H. Setigerous punctures from the medial and lateral edges not joining throughout the margin of the mesocaxa as in D. centralis (Harold, 1869). I–J. Elytra interstriae. I. Smooth as in D. amplicollis. J. Shagreened as in D. agenor." figureDoi="http://doi.org/10.5281/zenodo.4542111" httpUri="https://zenodo.org/record/4542111/files/figure.png" pageId="54" pageNumber="55">Fig. 2J</figureCitation>
|
||
) with fine shiny punctures arranged randomly. Striae with ocellate punctures separated by once diameter of a puncture on elytral disc.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF1022633F14CF32" blockId="54.[187,1399,1061,1124]" pageId="54" pageNumber="55">LEGS. Metacoxa surface microsculpture shagreened, posterior margin covered by row of dense setigerous punctures close together; setae dense, half the length of setae of metasternum.</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF1022CE3858CFF3" blockId="54.[189,769,1163,1189]" box="[189,769,1163,1189]" pageId="54" pageNumber="55">
|
||
PARAMERES AND LAMELLA COPULATRIX.
|
||
<figureCitation id="1317ABFAFFA6FFF8FDDD22CE3998CFF3" box="[624,705,1163,1189]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="54" pageNumber="55">Fig. 18</figureCitation>
|
||
E–G.
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF1022893AB7CFB0" blockId="54.[189,494,1228,1254]" box="[189,494,1228,1254]" pageId="54" pageNumber="55">
|
||
<heading id="D0DB0013FFA6FFF8FF1022893AB7CFB0" bold="true" box="[189,494,1228,1254]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
|
||
<emphasis id="B9586B6DFFA6FFF8FF1022893AB7CFB0" bold="true" box="[189,494,1228,1254]" pageId="54" pageNumber="55">Morphological variation</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF4722B93A8ACE40" blockId="54.[189,722,1275,1338]" box="[234,467,1275,1302]" pageId="54" pageNumber="55">
|
||
<emphasis id="B9586B6DFFA6FFF8FF4722B93A08CE40" bold="true" box="[234,337,1276,1302]" pageId="54" pageNumber="55">Females</emphasis>
|
||
(
|
||
<figureCitation id="1317ABFAFFA6FFF8FECC22BE3A90CE40" box="[353,457,1275,1302]" captionStart="Fig" captionStartId="53.[189,232,1893,1919]" captionTargetBox="[241,1349,296,1842]" captionTargetId="figure-73@53.[214,1373,259,1851]" captionTargetPageId="53" captionText="Fig. 18. Dichotomius validipilosus (Luederwaldt, 1931). A–B. Dorsal habitus. A. ♂. B. ♀. C. Lateral view of the aedeagus.D. View from above of the head, basal third surface subrugose. E. Ventral view of the aedeagus. F–G. Lamella copulatrix. H. Punctures along the anterior margin of the punctuated, sixth sternite marginally lower, depressed." figureDoi="http://doi.org/10.5281/zenodo.4542143" httpUri="https://zenodo.org/record/4542143/files/figure.png" pageId="54" pageNumber="55">Fig. 18B</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF10235A398BCE6F" blockId="54.[189,722,1275,1338]" box="[189,722,1311,1338]" pageId="54" pageNumber="55">As males except for the following characters:</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF10232439FECEC9" blockId="54.[189,1398,1377,1439]" pageId="54" pageNumber="55">HEAD. Anterior surface rugose, less dense on interocular space. Cephalic carina quadrituberculate; two median tubercles higher than lateral ones.</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF10238239B7CD71" blockId="54.[189,1399,1478,1575]" pageId="54" pageNumber="55">PRONOTUM. Anterior disc declivity weak, not modified. Surface finely punctate, punctures firmly impressed, denser at anterolateral margins next to fossae. Medial sulcus conspicuous. Pygidium delimited by incomplete margin, faded apically.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C336E4F4FFA6FFF8FF10200A3ABFCC55" pageId="54" pageNumber="55" type="discussion">
|
||
<paragraph id="8B93B77FFFA6FFF8FF10200A3A76CD3F" blockId="54.[189,303,1615,1641]" box="[189,303,1615,1641]" pageId="54" pageNumber="55">
|
||
<heading id="D0DB0013FFA6FFF8FF10200A3A76CD3F" bold="true" box="[189,303,1615,1641]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
|
||
<emphasis id="B9586B6DFFA6FFF8FF10200A3A76CD3F" bold="true" box="[189,303,1615,1641]" pageId="54" pageNumber="55">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B93B77FFFA6FFF8FF10203B3ABFCC55" blockId="54.[187,1399,1662,1795]" pageId="54" pageNumber="55">
|
||
The distribution of
|
||
<taxonomicName id="4C2CCCFCFFA6FFF8FE19203A392DCDCE" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[436,628,1662,1688]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="validipilosus">
|
||
<emphasis id="B9586B6DFFA6FFF8FE19203A392DCDCE" box="[436,628,1662,1688]" italics="true" pageId="54" pageNumber="55">D. validipilosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2CCCFCFFA6FFF8FD16203A3815CDCE" baseAuthorityName="Harold" baseAuthorityYear="1869" box="[699,844,1662,1688]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="deyrollei">
|
||
<emphasis id="B9586B6DFFA6FFF8FD16203A3815CDCE" box="[699,844,1662,1688]" italics="true" pageId="54" pageNumber="55">D. deyrollei</emphasis>
|
||
</taxonomicName>
|
||
overlaps and, although males can easily be differentiated, the females are quite similar and can only be distinguished by the pronotal punctation, which is lacking in
|
||
<taxonomicName id="4C2CCCFCFFA6FFF8FE3420833909CD89" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[409,592,1733,1759]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="validipilosus">
|
||
<emphasis id="B9586B6DFFA6FFF8FE3420833909CD89" box="[409,592,1733,1759]" italics="true" pageId="54" pageNumber="55">D. validipilosus</emphasis>
|
||
</taxonomicName>
|
||
, and the character of the sixth abdominal sternite, which is marginally lower in
|
||
<taxonomicName id="4C2CCCFCFFA6FFF8FE8820AF3A86CC55" baseAuthorityName="Luederwaldt" baseAuthorityYear="1931" box="[293,479,1769,1795]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="validipilosus">
|
||
<emphasis id="B9586B6DFFA6FFF8FE8820AF3A86CC55" box="[293,479,1769,1795]" italics="true" pageId="54" pageNumber="55">D. validipilosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |