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<document id="8F8C18705DB32892A0DDF17056F42AFE" ID-DOI="10.5281/zenodo.208740" ID-GBIF-Dataset="219d1ad7-0bc2-4982-b619-951254c1291b" ID-ISSN="1175-5326" ID-Zenodo-Dep="208740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460551926673" checkinUser="plazi" docAuthor="Van, Barry W. M., Guinot, Danièle, Corral, José Carmelo &amp; Artal, Pedro" docDate="2012" docId="E552DD5FFF8B0B6BA7D8FCEA279B3AFA" docLanguage="en" docName="zt03534p052.pdf" docOrigin="Zootaxa 3534" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Graptocarcinus Roemer 1887" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="196BA527FF8D0B6CA74FFFF5236A3815" masterDocTitle="Graptocarcininae n. subfam., an extinct subfamily of Dynomenidae Ortmann, 1892 (Crustacea, Brachyura, Podotremata)" masterLastPageNumber="52" masterPageNumber="40" pageNumber="46" updateTime="1698318472179" updateUser="plazi">
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<mods:title id="1729D3656D35F2B9546E6FF10114D00C">Graptocarcininae n. subfam., an extinct subfamily of Dynomenidae Ortmann, 1892 (Crustacea, Brachyura, Podotremata)</mods:title>
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<mods:namePart id="0D6F8890FBC0C5013332A2C6F34CF4C1">Van, Barry W. M.</mods:namePart>
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<mods:namePart id="C89AEC0C09D1C6B5C12BDADE1916DD0D">Guinot, Danièle</mods:namePart>
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<mods:namePart id="D302DFFA3126B2AA4E6E2744F511829F">Corral, José Carmelo</mods:namePart>
</mods:name>
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<mods:namePart id="5B6C12894EA2F7338F52A85F4FF6F96D">Artal, Pedro</mods:namePart>
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<mods:date id="351FED60A98F4684A7F8CE93F7431191">2012</mods:date>
<mods:detail id="74261637AF6040111461620E86B5E87F" type="volume">
<mods:number id="85A03271B0B94E90E6889DABA1270873">3534</mods:number>
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<mods:classification id="59C7C2CBB68015B5FA26F7E7C0C2933F">journal article</mods:classification>
<mods:identifier id="C378C9A8A90272DBD6DF1D9E848F6B3E" type="DOI">10.5281/zenodo.208740</mods:identifier>
<mods:identifier id="43A4536DB64259865B6589A3B7EA3EE0" type="GBIF-Dataset">219d1ad7-0bc2-4982-b619-951254c1291b</mods:identifier>
<mods:identifier id="BDD1108B3004088D155BE80666EB78F9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="17616A6AFC942B54F9F1EBCFA2FA71F4" type="Zenodo-Dep">208740</mods:identifier>
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<treatment id="E552DD5FFF8B0B6BA7D8FCEA279B3AFA" ID-DOI="http://doi.org/10.5281/zenodo.6173702" ID-GBIF-Taxon="119530296" ID-Zenodo-Dep="6173702" LSID="urn:lsid:plazi:treatment:E552DD5FFF8B0B6BA7D8FCEA279B3AFA" httpUri="http://treatment.plazi.org/id/E552DD5FFF8B0B6BA7D8FCEA279B3AFA" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46">
<subSubSection id="25E13FC2FF8B0B6AA7D8FCEA21313B2C" box="[151,603,799,825]" pageId="6" pageNumber="46" type="nomenclature">
<paragraph id="6D446C49FF8B0B6AA7D8FCEA21313B2C" blockId="6.[151,603,799,825]" box="[151,603,799,825]" pageId="6" pageNumber="46">
<heading id="360CDB25FF8B0B6AA7D8FCEA21313B2C" bold="true" box="[151,603,799,825]" fontSize="11" level="1" pageId="6" pageNumber="46" reason="1">
<emphasis id="5F8FB05BFF8B0B6AA7D8FCEA21313B2C" bold="true" box="[151,603,799,825]" pageId="6" pageNumber="46">
Genus
<taxonomicName id="AAFB17CAFF8B0B6AA7A1FCEA21313B2C" ID-CoL="759a060e-c8cd-4b10-9643-118b071434a0" authority="Roemer, 1887" authorityName="Roemer" authorityYear="1887" box="[238,603,799,825]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8B0B6AA7A1FCEA22C13B2C" bold="true" box="[238,427,799,825]" italics="true" pageId="6" pageNumber="46">Graptocarcinus</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA6FDFCEA21313B2C" author="Roemer" box="[434,603,799,825]" pageId="6" pageNumber="46" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer, 1887</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25E13FC2FF8B0B6AA7D8FC9227DB3C8B" pageId="6" pageNumber="46" type="materials_examined">
<paragraph id="6D446C49FF8B0B6AA7D8FC9222643BB6" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<emphasis id="5F8FB05BFF8B0B6AA7D8FC9222583B6A" bold="true" box="[151,306,870,895]" pageId="6" pageNumber="46">
<typeStatus id="B240D2EBFF8B0B6AA7D8FC9223BA3B6A" box="[151,208,871,895]" pageId="6" pageNumber="46">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="AAFB17CAFF8B0B6AA677FC9221823B6B" authority="Roemer, 1887" authorityName="Roemer" authorityYear="1887" box="[312,744,870,895]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8B0B6AA677FC92212F3B6A" box="[312,581,871,895]" italics="true" pageId="6" pageNumber="46">Graptocarcinus texanus</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA504FC9221823B6B" author="Roemer" box="[587,744,870,894]" pageId="6" pageNumber="46" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer, 1887</bibRefCitation>
</taxonomicName>
(=
<taxonomicName id="AAFB17CAFF8B0B6AA447FC9326073B6B" authority="Wright &amp; Wright, 1950" authorityName="Wright &amp; Wright" authorityYear="1950" box="[776,1389,870,894]" class="Insecta" family="Melolonthidae" genus="Cyphonotus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="integrimarginatus">
<emphasis id="5F8FB05BFF8B0B6AA447FC9327373B6B" box="[776,1117,870,894]" italics="true" pageId="6" pageNumber="46">Cyphonotus integrimarginatus</emphasis>
Wright &amp; Wright, 1950
</taxonomicName>
), by monotypy.
</paragraph>
<paragraph id="6D446C49FF8B0B6AA788FC5A21C53BFF" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<emphasis id="5F8FB05BFF8B0B6AA788FC5A22F83BD2" bold="true" box="[199,402,942,967]" pageId="6" pageNumber="46">Included species.</emphasis>
<taxonomicName id="AAFB17CAFF8B0B6AA6D6FC5A207E3BD3" authority="Collins &amp; Dieni, 1995" authorityName="Collins &amp; Dieni" authorityYear="1995" box="[409,788,942,966]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="bellonii">
<emphasis id="5F8FB05BFF8B0B6AA6D6FC5A21653BD3" box="[409,527,942,966]" italics="true" pageId="6" pageNumber="46">G. bellonii</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA559FC5B207E3BD3" author="Collins" box="[534,788,942,966]" pageId="6" pageNumber="46" refString="Collins, J. S. H. &amp; Dieni, I. (1995) New decapod crustaceans from the Cenomanian Rudist Limestones of NE Italy. Bulletin of the Mizunami Fossil Museum, 22, 67 - 72." type="journal article" year="1995">Collins &amp; Dieni, 1995</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AAFB17CAFF8B0B6AA46FFC5A27E73BD3" authority="Fraaye, 1996" authorityName="Fraaye" authorityYear="1996" box="[800,1165,942,967]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="maastrichtensis">
<emphasis id="5F8FB05BFF8B0B6AA46FFC5A20853BD3" box="[800,1007,942,966]" italics="true" pageId="6" pageNumber="46">G. maastrichtensis</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA4B8FC5A27E73BD3" author="Fraaye" box="[1015,1165,942,967]" pageId="6" pageNumber="46" refString="Fraaye, R. H. B. (1996) Two new crabs, Graptocarcinus maastrichtensis and Caloxanthus kuypersi (Crustacea, Decapoda), from the type Maastrichtian of the Netherlands. Journal of Paleontology, 70, 463 - 465." type="journal article" year="1996">Fraaye, 1996</bibRefCitation>
</taxonomicName>
,
<emphasis id="5F8FB05BFF8B0B6AA3D7FC5A279B3BD3" box="[1176,1265,943,967]" italics="true" pageId="6" pageNumber="46">G. m u i r i</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA3B3FC5B26F23BD3" author="Stenzel" box="[1276,1432,942,966]" pageId="6" pageNumber="46" refString="Stenzel, H. B. (1944) A new Cretaceous crab, Graptocarcinus muiri, from Mexico. Journal of Paleontology, 18, 550 - 552." type="journal article" year="1944">Stenzel, 1944</bibRefCitation>
,
<taxonomicName id="AAFB17CAFF8B0B6AA7D8FC2622D83BFF" authority="Roemer, 1887" authorityName="Roemer" authorityYear="1887" box="[151,434,978,1003]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8B0B6AA7D8FC2622603BFE" box="[151,266,979,1003]" italics="true" pageId="6" pageNumber="46">G. texanus</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA65DFC2622D83BFF" author="Roemer" box="[274,434,978,1002]" pageId="6" pageNumber="46" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer, 1887</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="AAFB17CAFF8B0B6AA6F2FC2621083BFE" box="[445,610,979,1003]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
<emphasis id="5F8FB05BFF8B0B6AA6F2FC2621083BFE" box="[445,610,979,1003]" italics="true" pageId="6" pageNumber="46">G. urbasaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44BC0D20FF8B0B6AA526FC2621CD3BFE" box="[617,679,979,1003]" pageId="6" pageNumber="46" rank="species">
<emphasis id="5F8FB05BFF8B0B6AA526FC2621CD3BFE" bold="true" box="[617,679,979,1003]" pageId="6" pageNumber="46">n. sp.</emphasis>
</taxonomicNameLabel>
.
</paragraph>
<paragraph id="6D446C49FF8B0B6AA788FC0327DB3C8B" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<emphasis id="5F8FB05BFF8B0B6AA788FC0322DE3C1A" bold="true" box="[199,436,1014,1039]" pageId="6" pageNumber="46">Material examined.</emphasis>
<taxonomicName id="AAFB17CAFF8B0B6AA6F0FC02206A3C1B" box="[447,768,1014,1038]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
<emphasis id="5F8FB05BFF8B0B6AA6F0FC02206A3C1B" box="[447,768,1014,1038]" italics="true" pageId="6" pageNumber="46">Graptocarcinus urbasaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44BC0D20FF8B0B6AA444FC0220273C1A" box="[779,845,1015,1039]" pageId="6" pageNumber="46" rank="species">
<emphasis id="5F8FB05BFF8B0B6AA444FC0220273C1A" bold="true" box="[779,845,1015,1039]" pageId="6" pageNumber="46">n. sp.</emphasis>
</taxonomicNameLabel>
:
<typeStatus id="B240D2EBFF8B0B6AA42FFC0220FA3C1A" box="[864,912,1015,1039]" pageId="6" pageNumber="46">type</typeStatus>
series, see below.
<taxonomicName id="AAFB17CAFF8B0B6AA33FFC0226F23C1B" authority="Roemer, 1887" authorityName="Roemer" authorityYear="1887" box="[1136,1432,1014,1039]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8B0B6AA33FFC0227823C1A" box="[1136,1256,1015,1039]" italics="true" pageId="6" pageNumber="46">G. texanus</emphasis>
<bibRefCitation id="096A11B8FF8B0B6AA3BCFC0226F23C1B" author="Roemer" box="[1267,1432,1014,1038]" pageId="6" pageNumber="46" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer, 1887</bibRefCitation>
</taxonomicName>
: MAB k.3208–3213 (ex collection K. Durney, Austin, Texas).
<taxonomicName id="AAFB17CAFF8B0B6AA408FBEE20923C27" box="[839,1016,1051,1074]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8B0B6AA408FBEE20923C27" box="[839,1016,1051,1074]" italics="true" pageId="6" pageNumber="46">Graptocarcinus</emphasis>
</taxonomicName>
sp. (as
<taxonomicName id="AAFB17CAFF8B0B6AA304FBEE27D43C26" box="[1099,1214,1051,1075]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8B0B6AA304FBEE27D43C26" box="[1099,1214,1051,1075]" italics="true" pageId="6" pageNumber="46">G. texanus</emphasis>
</taxonomicName>
): MNHN-R.03275, cast of large carapace, original in Musée de Cherbourg, Upper Cretaceous, Maastrichtian, “craie à
<taxonomicName id="AAFB17CAFF8B0B6AA26FFBCB26E23C43" authority="Fresnelles" authorityName="Fresnelles" box="[1312,1416,1086,1110]" class="Cephalopoda" family="Baculitidae" genus="Baculites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ammonitida" pageId="6" pageNumber="46" phylum="Mollusca" rank="genus">
<emphasis id="5F8FB05BFF8B0B6AA26FFBCB26E23C43" box="[1312,1416,1086,1110]" italics="true" pageId="6" pageNumber="46">Baculites</emphasis>
</taxonomicName>
”, Fresnelles (Manche),
<collectingCountry id="15EC2CD9FF8B0B6AA6C4FB9622B13C6F" box="[395,475,1123,1146]" name="France" pageId="6" pageNumber="46">France</collectingCountry>
. MNHN-A.32427 (coll. Milne-Edwards), small, mostly decorticated carapace, Upper Cretaceous, Cenomanian, Rouen (Ste-Catherine), Seine-Maritime, Haute-Normandie,
<collectingCountry id="15EC2CD9FF8B0B6AA312FB7227C73C8B" box="[1117,1197,1159,1182]" name="France" pageId="6" pageNumber="46">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="25E13FC2FF8B0B6AA788FB5E20CE3E67" pageId="6" pageNumber="46" type="discussion">
<paragraph id="6D446C49FF8B0B6AA788FB5E26F73D1F" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<emphasis id="5F8FB05BFF8B0B6AA788FB5E22533CD6" bold="true" box="[199,313,1195,1219]" pageId="6" pageNumber="46">Remarks.</emphasis>
<taxonomicName id="AAFB17CAFF8B0B6AA60BFB5E229A3CD7" box="[324,496,1195,1218]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8B0B6AA60BFB5E229A3CD7" box="[324,496,1195,1218]" italics="true" pageId="6" pageNumber="46">Graptocarcinus</emphasis>
</taxonomicName>
is traditionally assigned to
<taxonomicName id="AAFB17CAFF8B0B6AA479FB5F20BA3CD7" box="[822,976,1194,1218]" class="Malacostraca" family="Dynomenidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="family">Dynomenidae</taxonomicName>
(
<bibRefCitation id="096A11B8FF8B0B6AA4ACFB5F27E23CD7" author="Rathbun" box="[995,1160,1194,1218]" pageId="6" pageNumber="46" refString="Rathbun, M. J. (1935) Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America, Special Papers, 2, viii + 160 pp., pls. 1 - 26." type="book chapter" year="1935">Rathbun 1935</bibRefCitation>
: 41;
<bibRefCitation id="096A11B8FF8B0B6AA389FB5F26363CD7" author="Stenzel" box="[1222,1372,1194,1218]" pageId="6" pageNumber="46" refString="Stenzel, H. B. (1944) A new Cretaceous crab, Graptocarcinus muiri, from Mexico. Journal of Paleontology, 18, 550 - 552." type="journal article" year="1944">Stenzel 1944</bibRefCitation>
: 550;
<bibRefCitation id="096A11B8FF8B0B6AA7D8FB3A22763CF3" author="Remy" box="[151,284,1230,1255]" pageId="6" pageNumber="46" refString="Remy, J. M. (1955) Pagurus bathonicus nov. sp. et Graptocarcinus texanus Roemer. Crustaces Decapodes du Secondaire de la France. Bulletin du Museum National d'Histoire Naturelle, s e r. 2, 27 (2), 160 - 163." type="journal article" year="1955">Remy 1955</bibRefCitation>
: 162;
<bibRefCitation id="096A11B8FF8B0B6AA62DFB3B21333CF3" author="Collins" box="[354,601,1230,1254]" pageId="6" pageNumber="46" refString="Collins, J. S. H. &amp; Dieni, I. (1995) New decapod crustaceans from the Cenomanian Rudist Limestones of NE Italy. Bulletin of the Mizunami Fossil Museum, 22, 67 - 72." type="journal article" year="1995">Collins &amp; Dieni 1995</bibRefCitation>
: 70;
<bibRefCitation id="096A11B8FF8B0B6AA5DEFB3A204F3CF3" author="McLay" box="[657,805,1230,1255]" pageId="6" pageNumber="46" refString="McLay, C. L. (1999) Crustacea Decapoda: Revision of the family Dynomenidae. In: Crosnier, A. (Ed.), Resultats des Campagnes MUSORSTOM, volume 20. Memoires du Museum national d'Histoire naturelle (Paris), 180, 427 - 569." type="journal article" year="1999">McLay 1999</bibRefCitation>
: 434;
<bibRefCitation id="096A11B8FF8B0B6AA425FB3A27DC3CF3" author="De" box="[874,1206,1230,1254]" pageId="6" pageNumber="46" refString="De Angeli, A. &amp; Garassino, A. (2007) Decapod crustaceans from the Mesozoic and Cenozoic of Friuli-Venezia Giulia (NE Italy). Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35, 24 - 27." type="journal article" year="2007">
<collectingCountry id="15EC2CD9FF8B0B6AA425FB3A20E03CF3" box="[874,906,1231,1254]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="46">De</collectingCountry>
Angeli &amp; Garassino 2007
</bibRefCitation>
: 24;
<bibRefCitation id="096A11B8FF8B0B6AA3A0FB3A23B83D1F" author="De" pageId="6" pageNumber="46" refString="De Grave, S., Pentcheff, N. D., Ahyong, S. T., Chan, T. - Y., Crandall, K. A., Dworschak, P. C., Felder, D. L., Feldmann, R. M., Fransen, C. H. J. M., Goulding, L. Y. D., Lemaitre, R., Low, M. E. Y., Martin, J. W., Ng, P. K. L., Schweitzer, C. E., Tan, S. H., Tshudy, D. &amp; Wetzer, R. (2009) A classification of living and fossil genera of decapod crustaceans. Raffles Bulletin of Zoology, Supplement no. 21, 1 - 109." type="journal article" year="2009">
<collectingCountry id="15EC2CD9FF8B0B6AA3A0FB3A26653CF3" box="[1263,1295,1231,1254]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="46">De</collectingCountry>
Grave
<emphasis id="5F8FB05BFF8B0B6AA22CFB2526FF3CF3" box="[1379,1429,1230,1255]" italics="true" pageId="6" pageNumber="46">et al</emphasis>
. 2009
</bibRefCitation>
: 27;
<bibRefCitation id="096A11B8FF8B0B6AA64BFB0622C73D1F" author="Jagt" box="[260,429,1266,1291]" pageId="6" pageNumber="46" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. &amp; Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye &amp; Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. &amp; Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
Jagt
<emphasis id="5F8FB05BFF8B0B6AA679FB0122073D1E" box="[310,365,1267,1291]" italics="true" pageId="6" pageNumber="46">et al.</emphasis>
2010
</bibRefCitation>
: table 2; Schweitzer
<emphasis id="5F8FB05BFF8B0B6AA5C2FB0121D13D1E" box="[653,699,1267,1291]" italics="true" pageId="6" pageNumber="46">et al</emphasis>
. 2010: 66; Breton &amp; Collins 2011: 142;
<bibRefCitation id="096A11B8FF8B0B6AA33DFB0626303D1F" author="Karasawa" box="[1138,1370,1266,1291]" pageId="6" pageNumber="46" refString="Karasawa, H., Schweitzer, C. E. &amp; Feldmann, R. M. (2011) Phylogenetic analysis and revised classification of podotrematous Brachyura (Decapoda) including extinct and extant families. Journal of Crustacean Biology, 31, 523 - 565." type="journal article" year="2011">
Karasawa
<emphasis id="5F8FB05BFF8B0B6AA3ABFB0126793D1E" box="[1252,1299,1267,1291]" italics="true" pageId="6" pageNumber="46">et al</emphasis>
. 2011
</bibRefCitation>
: 542).
</paragraph>
<paragraph id="6D446C49FF8B0B6AA788FAE322873DF7" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
The Italian species
<taxonomicName id="AAFB17CAFF8B0B6AA6EFFAE221C73D3B" box="[416,685,1302,1326]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="bellonii">
<emphasis id="5F8FB05BFF8B0B6AA6EFFAE221C73D3B" box="[416,685,1302,1326]" italics="true" pageId="6" pageNumber="46">Graptocarcinus bellonii</emphasis>
</taxonomicName>
is known from the Cenomanian
<typeStatus id="B240D2EBFF8B0B6AA354FAE227213D3A" box="[1051,1099,1303,1327]" pageId="6" pageNumber="46">type</typeStatus>
series (
<bibRefCitation id="096A11B8FF8B0B6AA3EEFAE322403D47" author="Collins" pageId="6" pageNumber="46" refString="Collins, J. S. H. &amp; Dieni, I. (1995) New decapod crustaceans from the Cenomanian Rudist Limestones of NE Italy. Bulletin of the Mizunami Fossil Museum, 22, 67 - 72." type="journal article" year="1995">Collins &amp; Dieni 1995: figs. 2.1, 2.2</bibRefCitation>
), as well as from a better preserved Maastrichtian specimen (
<bibRefCitation id="096A11B8FF8B0B6AA4AAFACF265D3D47" author="De" box="[997,1335,1338,1362]" pageId="6" pageNumber="46" refString="De Angeli, A. &amp; Garassino, A. (2007) Decapod crustaceans from the Mesozoic and Cenozoic of Friuli-Venezia Giulia (NE Italy). Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35, 24 - 27." type="journal article" year="2007">
<collectingCountry id="15EC2CD9FF8B0B6AA4AAFACF276D3D47" box="[997,1031,1338,1362]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="46">De</collectingCountry>
Angeli &amp; Garassino 2007
</bibRefCitation>
: fig. 1e). Differences in ornamentation between the
<typeStatus id="B240D2EBFF8B0B6AA5DFFAAA21AA3D62" box="[656,704,1375,1399]" pageId="6" pageNumber="46">type</typeStatus>
series and the additional specimen were ascribed to possible ontogenetic differences, the size difference being considerable (
<bibRefCitation id="096A11B8FF8B0B6AA414FA7727CF3D8F" author="De" box="[859,1189,1410,1434]" pageId="6" pageNumber="46" refString="De Angeli, A. &amp; Garassino, A. (2007) Decapod crustaceans from the Mesozoic and Cenozoic of Friuli-Venezia Giulia (NE Italy). Memorie della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano, 35, 24 - 27." type="journal article" year="2007">
<collectingCountry id="15EC2CD9FF8B0B6AA414FA7720173D8F" box="[859,893,1410,1434]" name="South Georgia and the South Sandwich Islands" pageId="6" pageNumber="46">De</collectingCountry>
Angeli &amp; Garassino 2007
</bibRefCitation>
: 24).
<bibRefCitation id="096A11B8FF8B0B6AA3ADFA7626F23D8F" author="Jagt" box="[1250,1432,1410,1435]" pageId="6" pageNumber="46" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. &amp; Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye &amp; Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. &amp; Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
Jagt
<emphasis id="5F8FB05BFF8B0B6AA25AFA71262C3D8E" box="[1301,1350,1411,1435]" italics="true" pageId="6" pageNumber="46">et al</emphasis>
. (2010
</bibRefCitation>
: 186) assumed that the claw figured by
<bibRefCitation id="096A11B8FF8B0B6AA515FA5327503DAB" author="Collins" box="[602,1082,1446,1470]" pageId="6" pageNumber="46" refString="Collins, J. S. H. &amp; Dieni, I. (1995) New decapod crustaceans from the Cenomanian Rudist Limestones of NE Italy. Bulletin of the Mizunami Fossil Museum, 22, 67 - 72." type="journal article" year="1995">Collins &amp; Dieni (1995: figs. 2.3, 2.5, 2.6)</bibRefCitation>
as “
<taxonomicName id="AAFB17CAFF8B0B6AA321FA53278B3DAB" box="[1134,1249,1446,1470]" class="Malacostraca" family="Paguridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="subFamily" subFamily="Pagurinae">Pagurinae</taxonomicName>
genus indet.” is actually the claw of
<emphasis id="5F8FB05BFF8B0B6AA635FA3E22883DF6" box="[378,482,1483,1507]" italics="true" pageId="6" pageNumber="46">G. b e l l o n i</emphasis>
.
</paragraph>
<paragraph id="6D446C49FF8B0B6AA788FA1A20CE3E67" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<taxonomicName id="AAFB17CAFF8B0B6AA788FA1A215B3E13" box="[199,561,1518,1542]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="maastrichtensis">
<emphasis id="5F8FB05BFF8B0B6AA788FA1A215B3E13" box="[199,561,1518,1542]" italics="true" pageId="6" pageNumber="46">Graptocarcinus maastrichtensis</emphasis>
</taxonomicName>
was described from a single, moderately preserved carapace from the
<typeStatus id="B240D2EBFF8B0B6AA224FA1A26F63E12" box="[1387,1436,1519,1543]" pageId="6" pageNumber="46">type</typeStatus>
Maastrichtian of
<collectingCountry id="15EC2CD9FF8B0B6AA62FF9E721703E3F" box="[352,538,1554,1578]" name="Netherlands" pageId="6" pageNumber="46">the Netherlands</collectingCountry>
. It shows large orbits, a broadly triangular front, and a rounded pentagonal carapace outline (
<bibRefCitation id="096A11B8FF8B0B6AA613F9C3215B3E5B" author="Fraaye" box="[348,561,1590,1614]" pageId="6" pageNumber="46" refString="Fraaye, R. H. B. (1996) Two new crabs, Graptocarcinus maastrichtensis and Caloxanthus kuypersi (Crustacea, Decapoda), from the type Maastrichtian of the Netherlands. Journal of Paleontology, 70, 463 - 465." type="journal article" year="1996">Fraaye 1996: fig. 1</bibRefCitation>
). It may represent a juvenile specimen of the poorly known
<taxonomicName id="AAFB17CAFF8B0B6AA39BF9C322AF3E67" authority="Bosquet, 1854" authorityName="Bosquet" authorityYear="1854" class="Malacostraca" family="Dynomenidae" genus="Stephanometopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="granulatum">
<emphasis id="5F8FB05BFF8B0B6AA39BF9C322703E66" italics="true" pageId="6" pageNumber="46">Stephanometopon granulatum</emphasis>
Bosquet, 1854
</taxonomicName>
, but more material is needed to verify this.
</paragraph>
</subSubSection>
<subSubSection id="25E13FC2FF8B0B6BA788F98A279B3AFA" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46" type="reference_group">
<paragraph id="6D446C49FF8B0B6AA788F98A27BE3F87" blockId="6.[151,1437,870,2011]" pageId="6" pageNumber="46">
<taxonomicName id="AAFB17CAFF8B0B6AA788F98A22BA3E82" ID-CoL="c45db722-9a8c-41f2-b4bc-d36cd9ace904" box="[199,464,1663,1687]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8B0B6AA788F98A22BA3E82" box="[199,464,1663,1687]" italics="true" pageId="6" pageNumber="46">Graptocarcinus texanus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="096A11B8FF8B0B6AA5ECF98A20263E83" author="Roemer" box="[675,844,1662,1686]" pageId="6" pageNumber="46" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer (1887)</bibRefCitation>
from four well preserved carapaces from the “upper Turonian” from Austin, Texas. The
<typeStatus id="B240D2EBFF8B0B6AA56FF95621253EAE" box="[544,591,1699,1723]" pageId="6" pageNumber="46">type</typeStatus>
series could not be located in Bonn (despite of a visit of first author together with J.W.M Jagt, NHMM, and R.H.B. Fraaije, MAB, in
<date id="19454A89FF8B0B6AA45FF93320CD3ECB" box="[784,935,1734,1758]" pageId="6" pageNumber="46" value="2011-10">October 2011</date>
) and most probably is deposited in Wroclaw (formerly Breslau),
<collectingCountry id="15EC2CD9FF8B0B6AA63FF91F22AA3F17" box="[368,448,1770,1794]" name="Poland" pageId="6" pageNumber="46">Poland</collectingCountry>
.
<bibRefCitation id="096A11B8FF8B0B6AA686F91F20183F17" author="Rathbun" box="[457,882,1770,1794]" pageId="6" pageNumber="46" refString="Rathbun, M. J. (1935) Fossil Crustacea of the Atlantic and Gulf Coastal Plain. Geological Society of America, Special Papers, 2, viii + 160 pp., pls. 1 - 26." type="book chapter" year="1935">Rathbun (1935: 41; pl. 10, figs. 13–15)</bibRefCitation>
figured an additional well preserved carapace, and
<bibRefCitation id="096A11B8FF8B0B6AA7D8F8FB221A3F33" author="Stenzel" box="[151,368,1806,1830]" pageId="6" pageNumber="46" refString="Stenzel, H. B. (1944) A new Cretaceous crab, Graptocarcinus muiri, from Mexico. Journal of Paleontology, 18, 550 - 552." type="journal article" year="1944">Stenzel (1944: 551)</bibRefCitation>
translated and emended the original description by reporting the age of the
<typeStatus id="B240D2EBFF8B0B6AA3F5F8FA27833F32" box="[1210,1257,1807,1831]" pageId="6" pageNumber="46">type</typeStatus>
series as “Buda limestone of Comanche series, Cretaceous (lower Cenomanian)”. The species is reported from
<collectingCountry id="15EC2CD9FF8B0B6AA38CF8C726743F5F" box="[1219,1310,1842,1866]" name="United Kingdom" pageId="6" pageNumber="46">England</collectingCountry>
(
<bibRefCitation id="096A11B8FF8B0B6AA260F8C7224D3F7B" author="Wright" pageId="6" pageNumber="46" refString="Wright, C. W. &amp; Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of the Palaeontographical Society (London), 126 (533), 1 - 114, 22 pls." type="journal article" year="1972">Wright &amp; Collins 1972</bibRefCitation>
),
<collectingCountry id="15EC2CD9FF8B0B6AA674F8A222EC3F7B" box="[315,390,1879,1902]" name="France" pageId="6" pageNumber="46">France</collectingCountry>
(
<bibRefCitation id="096A11B8FF8B0B6AA6DAF8A321733F7B" author="Remy" box="[405,537,1878,1902]" pageId="6" pageNumber="46" refString="Remy, J. M. (1955) Pagurus bathonicus nov. sp. et Graptocarcinus texanus Roemer. Crustaces Decapodes du Secondaire de la France. Bulletin du Museum National d'Histoire Naturelle, s e r. 2, 27 (2), 160 - 163." type="journal article" year="1955">Remy 1955</bibRefCitation>
), and
<collectingCountry id="15EC2CD9FF8B0B6AA512F8A321F13F7B" box="[605,667,1878,1902]" name="Spain" pageId="6" pageNumber="46">Spain</collectingCountry>
(
<bibRefCitation id="096A11B8FF8B0B6AA5E6F8A320123F7B" author="Van" box="[681,888,1878,1902]" pageId="6" pageNumber="46" refString="Van Straelen, V. (1940) Crustaces decapodes nouveaux du Cretacique de la Navarre. Bulletin du Musee Royal d'Histoire Naturelle de Belgique, 16, 1 - 5." type="journal article" year="1940">Van Straelen 1940</bibRefCitation>
), but well preserved material should be examined to verify if this material belongs to
<taxonomicName id="AAFB17CAFF8B0B6AA557F88E21153F86" box="[536,639,1915,1939]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="texanu">
<emphasis id="5F8FB05BFF8B0B6AA557F88E21153F86" box="[536,639,1915,1939]" italics="true" pageId="6" pageNumber="46">G. texanu</emphasis>
</taxonomicName>
s.
<bibRefCitation id="096A11B8FF8B0B6AA5D8F88E20033F87" author="Fraaye" box="[663,873,1914,1939]" pageId="6" pageNumber="46" refString="Fraaye, R. H. B. (1996) Two new crabs, Graptocarcinus maastrichtensis and Caloxanthus kuypersi (Crustacea, Decapoda), from the type Maastrichtian of the Netherlands. Journal of Paleontology, 70, 463 - 465." type="journal article" year="1996">Fraaye (1996: 463)</bibRefCitation>
provided additional information.
</paragraph>
<paragraph id="6D446C49FF8B0B6BA788F86B277238C1" blockId="6.[151,1437,870,2011]" lastBlockId="7.[151,1437,151,752]" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46">
The other American species,
<taxonomicName id="AAFB17CAFF8B0B6AA558F86A20663FA3" ID-CoL="e9d5d8d6-dc25-46fe-934a-9943ace4ed5f" box="[535,780,1951,1974]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="muiri">
<emphasis id="5F8FB05BFF8B0B6AA558F86A20663FA3" box="[535,780,1951,1974]" italics="true" pageId="6" pageNumber="46">Graptocarcinus muiri</emphasis>
</taxonomicName>
, was described from a single, isolated carapace from the Albian (middle Cretaceous) Taninul limestone of San Luis Potosí,
<collectingCountry id="15EC2CD9FF8B0B6AA4E4F837276A3FCF" box="[939,1024,1986,2010]" name="Mexico" pageId="6" pageNumber="46">Mexico</collectingCountry>
(
<bibRefCitation id="096A11B8FF8B0B6AA35BF83727C53FCF" author="Stenzel" box="[1044,1199,1986,2010]" pageId="6" pageNumber="46" refString="Stenzel, H. B. (1944) A new Cretaceous crab, Graptocarcinus muiri, from Mexico. Journal of Paleontology, 18, 550 - 552." type="journal article" year="1944">Stenzel 1944</bibRefCitation>
).
<bibRefCitation id="096A11B8FF8B0B6BA388F836238B38BA" author="Vega" lastPageId="7" lastPageNumber="47" pageId="6" pageNumber="46" refString="Vega, F. J. &amp; Feldmann, R. M. (1992) Occurence of Costacopluma (Decapoda: Brachyura: Retroplumidae) in the Maastrichtian of Southern Mexico and its paleobiogeographic implications. Annals of Carnegie Museum, 61, 133 - 152." type="journal article" year="1992">Vega &amp; Feldmann (1992)</bibRefCitation>
stated this to be of Aptian age. It was synonymised with
<taxonomicName id="AAFB17CAFF8A0B6BA4CFFF6D209D38A5" ID-CoL="c45db722-9a8c-41f2-b4bc-d36cd9ace904" box="[896,1015,152,176]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8A0B6BA4CFFF6D209D38A5" box="[896,1015,152,176]" italics="true" pageId="7" pageNumber="47">G. texanus</emphasis>
</taxonomicName>
by
<bibRefCitation id="096A11B8FF8A0B6BA369FF62260538BA" author="Wright" box="[1062,1391,151,175]" pageId="7" pageNumber="47" refString="Wright, C. W. &amp; Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of the Palaeontographical Society (London), 126 (533), 1 - 114, 22 pls." type="journal article" year="1972">Wright &amp; Collins (1972: 54)</bibRefCitation>
but subsequently
<bibRefCitation id="096A11B8FF8A0B6BA67FFF4922BB38C1" author="Bishop" box="[304,465,188,212]" pageId="7" pageNumber="47" refString="Bishop, G. A. (1986) Occurrence, preservation, and biogeography of the Cretaceous crabs of North America. In: Gore, R. H. &amp; Heck, K. L. (Eds), Crustacean Biogeography. A. A. Balkema, Rotterdam / Boston, pp. 111 - 142." type="book chapter" year="1986">Bishop (1986)</bibRefCitation>
and
<bibRefCitation id="096A11B8FF8A0B6BA548FF4821CF38C1" author="Fraaye" box="[519,677,188,213]" pageId="7" pageNumber="47" refString="Fraaye, R. H. B. (1996) Two new crabs, Graptocarcinus maastrichtensis and Caloxanthus kuypersi (Crustacea, Decapoda), from the type Maastrichtian of the Netherlands. Journal of Paleontology, 70, 463 - 465." type="journal article" year="1996">Fraaye (1996)</bibRefCitation>
treated them as separate species.
</paragraph>
<paragraph id="6D446C49FF8A0B6BA788FF2A21DE3A4A" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="47">
<bibRefCitation id="096A11B8FF8A0B6BA788FF2A217438E2" author="Guinot" box="[199,542,223,247]" pageId="7" pageNumber="47" refString="Guinot, D. &amp; Breton, G. (2006) Lithophylax trigeri A. Milne-Edwards &amp; Brocchi, 1879 from the French Cretaceous (Cenomanian) and placement of the family Lithophylacidae Van Straelen, 1936 (Crustacea, Decapoda, Brachyura). Geodiversitas, 28 (4), 591 - 633." type="journal article" year="2006">Guinot &amp; Breton (2006: 610)</bibRefCitation>
commented that “the lower Cenomanian
<taxonomicName id="AAFB17CAFF8A0B6BA349FF14261638E2" authority="Fritsch" authorityName="Fritsch" box="[1030,1404,223,248]" class="Malacostraca" family="Necrocarcinidae" genus="Necrocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="avicularis">
<emphasis id="5F8FB05BFF8A0B6BA349FF14264938ED" box="[1030,1315,224,248]" italics="true" pageId="7" pageNumber="47">Necrocarcinus avicularis</emphasis>
Fritsch
</taxonomicName>
<emphasis id="5F8FB05BFF8A0B6BA2C9FF1426F638ED" box="[1414,1436,225,248]" italics="true" pageId="7" pageNumber="47">in</emphasis>
<bibRefCitation id="096A11B8FF8A0B6BA7D8FEF122C03909" author="Fritsch" box="[151,426,260,284]" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">Fritsch &amp; Kafka, 1887</bibRefCitation>
(
<bibRefCitation id="096A11B8FF8A0B6BA68EFEF127ED3909" author="Fritsch" box="[449,1159,260,284]" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">
Fritsch &amp; Kafka 1887: 47
<emphasis id="5F8FB05BFF8A0B6BA45FFEF020EA3909" box="[784,896,261,284]" italics="true" pageId="7" pageNumber="47">pro parte</emphasis>
, pl. 10, fig. 12 only
</bibRefCitation>
) might be included in
<taxonomicName id="AAFB17CAFF8A0B6BA7D8FEDD2299392A" ID-CoL="759a060e-c8cd-4b10-9643-118b071434a0" authority="Roemer, 1887" authorityName="Roemer" authorityYear="1887" box="[151,499,295,319]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA7D8FEDD222D392A" box="[151,327,296,319]" italics="true" pageId="7" pageNumber="47">Graptocarcinus</emphasis>
<bibRefCitation id="096A11B8FF8A0B6BA600FEDD2299392A" author="Roemer" box="[335,499,295,319]" pageId="7" pageNumber="47" refString="Roemer, F. A. (1887) Graptocarcinus texanus, ein Brachyure aus der oberen Kreide von Texas. Neues Jahrbuch fur Mineralogie, Geologie und Palaontologie, 1887, 173 - 176." type="journal article" year="1887">Roemer, 1887</bibRefCitation>
</taxonomicName>
”. This species is based on claws (
<bibRefCitation id="096A11B8FF8A0B6BA43BFED22719392A" author="Fritsch" box="[884,1139,295,319]" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">Fritsch &amp; Kafka 1887</bibRefCitation>
: 47;
<bibRefCitation id="096A11B8FF8A0B6BA3E4FEDD2635392A" author="Jagt" box="[1195,1375,295,320]" pageId="7" pageNumber="47" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. &amp; Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye &amp; Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. &amp; Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
Jagt
<emphasis id="5F8FB05BFF8A0B6BA3AFFEDC26703955" box="[1248,1306,296,320]" italics="true" pageId="7" pageNumber="47">et al.</emphasis>
2010
</bibRefCitation>
: 185; table I) but two fragments of carapaces, clearly belonging to different genera, were figured (
<bibRefCitation id="096A11B8FF8A0B6BA3D2FEB9220D3992" author="Fritsch" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">Fritsch &amp; Kafka 1887: pl. 10, figs. 12, 13</bibRefCitation>
). One of these fragments is described as (
<taxonomicName id="AAFB17CAFF8A0B6BA472FE8520E53992" ID-CoL="7NRS7" box="[829,911,368,391]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA472FE8520E53992" box="[829,911,368,391]" italics="true" pageId="7" pageNumber="47">Cancer</emphasis>
</taxonomicName>
?)
<emphasis id="5F8FB05BFF8A0B6BA4E3FE85277E399D" box="[940,1044,368,392]" italics="true" pageId="7" pageNumber="47">modestus</emphasis>
(
<bibRefCitation id="096A11B8FF8A0B6BA36CFE9A238B39B9" author="Fritsch" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">Fritsch &amp; Kafka 1887: 49, pl. 10, fig. 12</bibRefCitation>
) which
<bibRefCitation id="096A11B8FF8A0B6BA674FE61211039B9" author="Wright" box="[315,634,404,428]" pageId="7" pageNumber="47" refString="Wright, C. W. &amp; Collins, J. S. H. (1972) British Cretaceous crabs. Monograph of the Palaeontographical Society (London), 126 (533), 1 - 114, 22 pls." type="journal article" year="1972">Wright &amp; Collins (1972: 54)</bibRefCitation>
questionably synonymised with
<taxonomicName id="AAFB17CAFF8A0B6BA4A5FE60279C39B9" ID-CoL="c45db722-9a8c-41f2-b4bc-d36cd9ace904" box="[1002,1270,405,428]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="texanus">
<emphasis id="5F8FB05BFF8A0B6BA4A5FE60279C39B9" box="[1002,1270,405,428]" italics="true" pageId="7" pageNumber="47">Graptocarcinus texanus</emphasis>
</taxonomicName>
. Schweitzer
<emphasis id="5F8FB05BFF8A0B6BA2C7FE6023DE39C5" italics="true" pageId="7" pageNumber="47">et al.</emphasis>
(2010: 80, 101) included
<taxonomicName id="AAFB17CAFF8A0B6BA6AAFE4C211239C5" box="[485,632,440,464]" class="Malacostraca" family="Necrocarcinidae" genus="Necrocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="avicularis">
<emphasis id="5F8FB05BFF8A0B6BA6AAFE4C229239C5" box="[485,504,441,464]" italics="true" pageId="7" pageNumber="47">N</emphasis>
.
<emphasis id="5F8FB05BFF8A0B6BA547FE4D211239C5" box="[520,632,440,464]" italics="true" pageId="7" pageNumber="47">avicularis</emphasis>
</taxonomicName>
in
<taxonomicName id="AAFB17CAFF8A0B6BA5EDFE4C202F39C5" ID-CoL="9G98G" box="[674,837,441,464]" class="Malacostraca" family="Necrocarcinidae" genus="Necrocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA5EDFE4C202F39C5" box="[674,837,441,464]" italics="true" pageId="7" pageNumber="47">Necrocarcinus</emphasis>
</taxonomicName>
, while (
<emphasis id="5F8FB05BFF8A0B6BA4E8FE4D20AA39DA" box="[935,960,440,463]" italics="true" pageId="7" pageNumber="47">C.</emphasis>
?)
<emphasis id="5F8FB05BFF8A0B6BA4AFFE4D272239C5" box="[992,1096,440,464]" italics="true" pageId="7" pageNumber="47">modestus</emphasis>
was erroneously listed under
<taxonomicName id="AAFB17CAFF8A0B6BA7D8FE28238239E1" ID-CoL="7NRS7" box="[151,232,477,500]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA7D8FE28238239E1" box="[151,232,477,500]" italics="true" pageId="7" pageNumber="47">Cancer</emphasis>
</taxonomicName>
.
<bibRefCitation id="096A11B8FF8A0B6BA7B4FE2822A839E1" author="Jagt" box="[251,450,476,501]" pageId="7" pageNumber="47" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. &amp; Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye &amp; Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. &amp; Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
Jagt
<emphasis id="5F8FB05BFF8A0B6BA67BFE28221B39E1" box="[308,369,476,500]" italics="true" pageId="7" pageNumber="47">et al.</emphasis>
(2010
</bibRefCitation>
: 185) assigned the chelae to the “form genus”
<taxonomicName id="AAFB17CAFF8A0B6BA4B8FE28267A39E1" ID-CoL="0b6c7dbf-868d-4cb9-884a-092de3f35842" authority="Bishop, 1983" authorityName="Bishop" authorityYear="1983" box="[1015,1296,476,500]" class="Malacostraca" family="Diogenidae" genus="Roemerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA4B8FE28270D39E1" box="[1015,1127,477,500]" italics="true" pageId="7" pageNumber="47">Roemerus</emphasis>
<bibRefCitation id="096A11B8FF8A0B6BA33CFE29267A39E1" author="Bishop" box="[1139,1296,476,500]" pageId="7" pageNumber="47" refString="Bishop, G. A. (1983) Fossil decapod crustaceans from the Lower Cretaceous, Glen Rose Limestone of Central Texas. Transactions of the San Diego Society of Natural History, 20, 27 - 55, pls. 1 - 3." type="journal article" year="1983">Bishop, 1983</bibRefCitation>
</taxonomicName>
, which was considered to be a dynomenid. Both the claws (
<bibRefCitation id="096A11B8FF8A0B6BA58EFE0A27A23A02" author="Fritsch" box="[705,1224,511,535]" pageId="7" pageNumber="47" refString="Fritsch, A. &amp; Kafka, J. (1887) Die Crustaceen der bohmischen Kreideformation, 53 pp., 10 pls. Fr. Rivnac, Praha." type="book" year="1887">Fritsch &amp; Kafka 1887: pl. 10, figs. 2, 10, 11</bibRefCitation>
) and the carapace fragment in figure 10, could belong to
<taxonomicName id="AAFB17CAFF8A0B6BA51BFDD0206E3A29" ID-CoL="759a060e-c8cd-4b10-9643-118b071434a0" box="[596,772,549,572]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA51BFDD0206E3A29" box="[596,772,549,572]" italics="true" pageId="7" pageNumber="47">Graptocarcinus</emphasis>
</taxonomicName>
. The material, however, is fragmentary and disassociated, and no final conclusion can be drawn at present.
</paragraph>
<paragraph id="6D446C49FF8A0B6BA788FD99279B3AFA" blockId="7.[151,1437,151,752]" pageId="7" pageNumber="47">
Claws of
<taxonomicName id="AAFB17CAFF8A0B6BA67AFD98228F3A91" ID-CoL="759a060e-c8cd-4b10-9643-118b071434a0" box="[309,485,621,644]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA67AFD98228F3A91" box="[309,485,621,644]" italics="true" pageId="7" pageNumber="47">Graptocarcinus</emphasis>
</taxonomicName>
spp. may be differentiated from other fossil dynomenid claws by the presence of a distinct articulating bulge.
<bibRefCitation id="096A11B8FF8A0B6BA688FD6521E93AB2" author="Jagt" box="[455,643,655,680]" pageId="7" pageNumber="47" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. &amp; Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye &amp; Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. &amp; Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
Jagt
<emphasis id="5F8FB05BFF8A0B6BA6B2FD64215C3ABD" box="[509,566,656,680]" italics="true" pageId="7" pageNumber="47">et al.</emphasis>
(2010
</bibRefCitation>
: 179) stated that claw forms with oval depressions in both fingers for setae can be linked to “
<taxonomicName id="AAFB17CAFF8A0B6BA687FD4121123AD9" ID-CoL="3e7d5083-3906-451e-9f99-3a7f4a0a939a" box="[456,632,692,716]" class="Malacostraca" family="Goniodromitidae" genus="Palaeodromites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA687FD4121123AD9" box="[456,632,692,716]" italics="true" pageId="7" pageNumber="47">Palaeodromites</emphasis>
</taxonomicName>
” [to
<taxonomicName id="AAFB17CAFF8A0B6BA58BFD4120523AD9" ID-CoL="eae76844-d176-495f-82f1-7b6dabddc122" box="[708,824,692,716]" class="Malacostraca" family="Goniodromitidae" genus="Distefania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA58BFD4120523AD9" box="[708,824,692,716]" italics="true" pageId="7" pageNumber="47">Distefania</emphasis>
</taxonomicName>
] while forms lacking such depressions belong to
<taxonomicName id="AAFB17CAFF8A0B6BA7D8FD2D222D3AFA" ID-CoL="759a060e-c8cd-4b10-9643-118b071434a0" box="[151,327,728,751]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="5F8FB05BFF8A0B6BA7D8FD2D222D3AFA" box="[151,327,728,751]" italics="true" pageId="7" pageNumber="47">Graptocarcinus</emphasis>
</taxonomicName>
or are closely allied dynomenids. The material described below confirms this view.
</paragraph>
</subSubSection>
</treatment>
</document>
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