304 lines
34 KiB
XML
304 lines
34 KiB
XML
<document id="F898CC5E9D6280D7FCF8FFBC61FB9B1B" ID-DOI="10.5281/zenodo.208740" ID-GBIF-Dataset="219d1ad7-0bc2-4982-b619-951254c1291b" ID-ISSN="1175-5326" ID-Zenodo-Dep="208740" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460551926673" checkinUser="plazi" docAuthor="Van, Barry W. M., Guinot, Danièle, Corral, José Carmelo & Artal, Pedro" docDate="2012" docId="E552DD5FFF8A0B66A7D8FCC3226D3B01" docLanguage="en" docName="zt03534p052.pdf" docOrigin="Zootaxa 3534" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Graptocarcinus urbasaensis Van, Guinot, Corral & Artal, 2012, n. sp." docType="treatment" docVersion="7" lastPageNumber="50" masterDocId="196BA527FF8D0B6CA74FFFF5236A3815" masterDocTitle="Graptocarcininae n. subfam., an extinct subfamily of Dynomenidae Ortmann, 1892 (Crustacea, Brachyura, Podotremata)" masterLastPageNumber="52" masterPageNumber="40" pageNumber="47" updateTime="1698318472179" updateUser="plazi">
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<mods:mods id="04D34620E7953FCBCD0546FF81DE371C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E8973E1F933C92D23CCABFEC864454F0">
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<mods:title id="D453D9EC6AE59EFBC4DEAB105C1CA8EA">Graptocarcininae n. subfam., an extinct subfamily of Dynomenidae Ortmann, 1892 (Crustacea, Brachyura, Podotremata)</mods:title>
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</mods:titleInfo>
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<mods:name id="C4CB215A67141593EAE4BC80CBCE296E" type="personal">
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<mods:roleTerm id="80725FB890F7EAAE9A31E17C7F79FD53">Author</mods:roleTerm>
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<mods:namePart id="D44753B2F92DFD30B7967D6324D38497">Van, Barry W. M.</mods:namePart>
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<mods:name id="8C99514E9F57849820661903261AA297" type="personal">
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<mods:roleTerm id="B70D2A924E60A264BB95D822AC38B52F">Author</mods:roleTerm>
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<mods:namePart id="BEBCC9A91594CDC9707151E911D0F3DA">Guinot, Danièle</mods:namePart>
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</mods:name>
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<mods:name id="F4AE15166CCEBDEC0455BEA646896C77" type="personal">
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<mods:roleTerm id="94082C08D32E8D58F97C12F2980FD238">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35F0748F44AF725D01C94C40031117D9">Corral, José Carmelo</mods:namePart>
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</mods:name>
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<mods:name id="9547D571F295390DB27322DEF2FE1058" type="personal">
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<mods:roleTerm id="24813141BFC926FA8515D291C946C62D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="858FE07C31947632BAF5D16C5BCD58EC">Artal, Pedro</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="734A9714FFE57C54AC2AE26BA1DB9670">text</mods:typeOfResource>
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<mods:title id="AF8841B177418C55E8C717C6B878E4DB">Zootaxa</mods:title>
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<mods:part id="0E5F57B9DCC7FD586E439D8A274DCA5C">
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<mods:date id="01D6828E3DBFE912597F10BD0ECF29AC">2012</mods:date>
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<mods:detail id="3A6BE7063F510C869DC101D1905D2F0E" type="volume">
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<mods:number id="EDE99A70670746C85B0697F271D27626">3534</mods:number>
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<mods:start id="40662BEECCEDF1903C24FFE82E4728E0">40</mods:start>
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<mods:classification id="29B831C54BF206982EE4DED06CEEE442">journal article</mods:classification>
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<mods:identifier id="47CBEF91184DEC6FECFE490466533552" type="DOI">10.5281/zenodo.208740</mods:identifier>
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<mods:identifier id="C54707C22CD593DD73E55FC571662FBA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E6CEDB61AB38A5085E7E406AB83DFCDE" type="Zenodo-Dep">208740</mods:identifier>
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<treatment id="E552DD5FFF8A0B66A7D8FCC3226D3B01" ID-DOI="http://doi.org/10.5281/zenodo.6173704" ID-GBIF-Taxon="119530298" ID-Zenodo-Dep="6173704" LSID="urn:lsid:plazi:treatment:E552DD5FFF8A0B66A7D8FCC3226D3B01" httpUri="http://treatment.plazi.org/id/E552DD5FFF8A0B66A7D8FCC3226D3B01" lastPageId="10" lastPageNumber="50" pageId="7" pageNumber="47">
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<subSubSection id="25E13FC2FF8A0B6BA7D8FCC3227B3B64" pageId="7" pageNumber="47" type="nomenclature">
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<paragraph id="6D446C49FF8A0B6BA7D8FCC3215E3B45" blockId="7.[151,564,822,881]" box="[151,564,822,848]" pageId="7" pageNumber="47">
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<heading id="360CDB25FF8A0B6BA7D8FCC3215E3B45" bold="true" box="[151,564,822,848]" fontSize="11" level="1" pageId="7" pageNumber="47" reason="1">
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<emphasis id="5F8FB05BFF8A0B6BA7D8FCC3215E3B45" bold="true" box="[151,564,822,848]" pageId="7" pageNumber="47">
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||
<taxonomicName id="AAFB17CAFF8A0B6BA7D8FCC322803B45" ID-CoL="eca454e8-a54d-4328-aa21-4e1da7c1f187" box="[151,490,822,848]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
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<emphasis id="5F8FB05BFF8A0B6BA7D8FCC322803B45" bold="true" box="[151,490,822,848]" italics="true" pageId="7" pageNumber="47">Graptocarcinus urbasaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="44BC0D20FF8A0B6BA6BDFCC3215E3B45" box="[498,564,822,848]" pageId="7" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="6D446C49FF8A0B6BA7D8FCAC227B3B64" blockId="7.[151,564,822,881]" box="[151,273,857,881]" pageId="7" pageNumber="47">
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(
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<figureCitation id="F5C070CCFF8A0B6BA7D0FCAC23853B64" box="[159,239,857,881]" captionStart="FIGURE 2" captionStartId="5.[151,250,1905,1928]" captionTargetBox="[151,1435,320,1884]" captionTargetId="figure@5.[151,1436,319,1884]" captionTargetPageId="5" captionText="FIGURE 2. Graptocarcinus urbasaensis n. sp., MGSB 78.337 a – c (holotype, female), Santonian, Contrasta, Álava, Spain. A, dorsal view; B, right lateral view; C, ventral view; D, frontal view; E, posterior view; F, left cheliped. a 1, a 2, a 3, a 4, a 5, a 6, abdominal somites 1, 2, 3, 4, 5, 6; d, depression; u, uropod; t, telson. Scale bars: 10 mm. Specimen whitened with ammonium chloride sublimate prior to photography." httpUri="https://zenodo.org/record/208742/files/figure.png" pageId="7" pageNumber="47">Figs. 2</figureCitation>
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,
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<figureCitation id="F5C070CCFF8A0B6BA7B5FCAC22623B64" box="[250,264,857,881]" captionStart="FIGURE 3" captionStartId="8.[151,250,1874,1897]" captionTargetBox="[151,1435,320,1869]" captionTargetId="figure@8.[151,1436,315,1881]" captionTargetPageId="8" captionText="FIGURE 3. Graptocarcinus urbasaensis n. sp., MCNA 8562 (paratype, male), Santonian, Egibil quarry, Olazti-Olazagutía, Navarra, Spain. A, dorsal view; B, frontal view; C, ventral view; D, posterior view; E, detail of abdomen and pereiopod coxae; F, detail of posterior view. a 1, a 2, a 3, a 4, a 5, a 6, abdominal somites 1, 2, 3, 4, 5, 6; ab, articulating bulge; cx 1, cx 2, cx 3, cx 4, cx 5, P 1 – P 5 coxae; cmxp 3, mxp 3 coxa; d, depression; gr, granules; mP 5, P 5 merus; pr, prominence; pt, pterygostome; u, uropod; t, telson. Scale bars: 10 mm. Specimen whitened with ammonium chloride sublimate prior to photography." httpUri="https://zenodo.org/record/208743/files/figure.png" pageId="7" pageNumber="47">3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="25E13FC2FF8A0B6BA7D8FC5727963BAC" box="[151,1276,929,954]" pageId="7" pageNumber="47" type="etymology">
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<paragraph id="6D446C49FF8A0B6BA7D8FC5727963BAC" blockId="7.[151,1436,929,2033]" box="[151,1276,929,954]" pageId="7" pageNumber="47">
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<emphasis id="5F8FB05BFF8A0B6BA7D8FC5722743BAF" bold="true" box="[151,286,930,954]" pageId="7" pageNumber="47">Etymology.</emphasis>
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From Urbasa, the Spanish name of the mountain range where the fossils were collected.
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</paragraph>
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</subSubSection>
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<subSubSection id="25E13FC2FF8A0B6BA788FC3021643C78" pageId="7" pageNumber="47" type="materials_examined">
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<paragraph id="6D446C49FF8A0B6BA788FC3021643C78" blockId="7.[151,1436,929,2033]" pageId="7" pageNumber="47">
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<emphasis id="5F8FB05BFF8A0B6BA788FC3022D83BCB" bold="true" box="[199,434,965,990]" pageId="7" pageNumber="47">Material examined.</emphasis>
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<typeStatus id="B240D2EBFF8A0B6BA6F3FC3021433BC8" box="[444,553,965,989]" pageId="7" pageNumber="47" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="0BEAF48CFF8A0B6BA57BFC3021E93BC8" box="[564,643,965,989]" pageId="7" pageNumber="47">MGSB</collectionCode>
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78.337a–c (female), carapace with abdomen, left-hand cheliped, and pereiopods, Santonian, Contrasta, Álava province,
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<collectingCountry id="15EC2CD9FF8A0B6BA59DFC1C207F3C14" box="[722,789,1001,1025]" name="Spain" pageId="7" pageNumber="47">Spain</collectingCountry>
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.
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<typeStatus id="B240D2EBFF8A0B6BA46FFC1F20FB3C17" box="[800,913,1002,1026]" pageId="7" pageNumber="47" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="0BEAF48CFF8A0B6BA4D3FC1C209A3C14" box="[924,1008,1001,1025]" httpUri="http://grbio.org/cool/c5sb-vm81" name="Museo de Ciencias naturals de Alava" pageId="7" pageNumber="47">MCNA</collectionCode>
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8562 (male), well preserved carapace with abdomen, left-hand cheliped, and pereiopods, Santonian, Egibil quarry (“Cantera de Margas”), Olazti- Olazagutía, Navarra Province,
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<collectingCountry id="15EC2CD9FF8A0B6BA6BBFBC4215D3C5C" box="[500,567,1073,1097]" name="Spain" pageId="7" pageNumber="47">Spain</collectingCountry>
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.
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<collectionCode id="0BEAF48CFF8A0B6BA50CFBC421F83C5C" box="[579,658,1073,1097]" pageId="7" pageNumber="47">MGSB</collectionCode>
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28.148a–b, carapace and isolated cheliped, Maastrichtian, Puerto de Urbasa, Navarra Province,
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<collectingCountry id="15EC2CD9FF8A0B6BA688FBA021603C78" box="[455,522,1109,1133]" name="Spain" pageId="7" pageNumber="47">Spain</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="25E13FC2FF8A0B66A788FB8C226D3B01" lastPageId="10" lastPageNumber="50" pageId="7" pageNumber="47" type="description">
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<paragraph id="6D446C49FF8A0B6BA788FB8C272A3E08" blockId="7.[151,1436,929,2033]" pageId="7" pageNumber="47">
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<emphasis id="5F8FB05BFF8A0B6BA788FB8C223F3C87" bold="true" box="[199,341,1145,1170]" pageId="7" pageNumber="47">Description.</emphasis>
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Carapace relatively large (maximum carapace width
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<quantity id="AA03C1ACFF8A0B6BA4E7FB8C20983C84" box="[936,1010,1145,1169]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="7" pageNumber="47" unit="mm" value="34.0">34mm</quantity>
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), wider than long, width 1.25x length, widest slightly posterior to midlength. Dorsal surface convex in longitudinal section, gently tumid in transverse section, with strongly inclined sides; surface ornamented with large, blunt granules, cuticle with numerous minute setal pits. Cervical groove indistinct, only defined in medial third, broadly V-shaped. Branchial groove obsolete, branchiocardiac grooves curved, crescent-shaped, diverging anteriorly. Anterolateral margins long, evenly curved, well defined, sharp, without teeth, set with granules; corner of lateral margins produced as sharp flange; posterolateral margin weakly defined, blunt, straight in dorsal view; posterior margin relatively narrow, straight in dorsal view, markedly concave
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<emphasis id="5F8FB05BFF8A0B6BA621FA8322DF3D98" box="[366,437,1398,1421]" italics="true" pageId="7" pageNumber="47">per se</emphasis>
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, distinctly thickened. Front weakly projecting, strongly downturned, broadly triangular, continuous with supraorbital margin; supraorbital margin oblique, arched, continuous above orbits, thickened, rimmed, without notches; infraorbital margin short, without notches. Orbits oblique, elongated, well defined, outer orbital corner rounded. Subhepatic region, pterygostome flattened, grooved, surface non-ornamented; cervical groove ventrally present as shallow groove. Pleural line absent; branchiostegite low, inclined.
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</paragraph>
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<paragraph id="6D446C49FF8A0B64A788F9DC27CC3909" blockId="7.[151,1436,929,2033]" lastBlockId="8.[151,1436,151,284]" lastPageId="8" lastPageNumber="48" pageId="7" pageNumber="47">Mxp3 operculiform, coxae elongated, twisted; endopod: basis separated from ischium by complete, distinct suture, ischium smooth, widened distally, inner margin arched; exopod: incompletely preserved, rather wide, straight. Abdomen entirely covering thoracic sternum (i.e. in width, length) in both sexes, thus in contact with pereiopod and mxp3 coxae; sexual dimorphism distinct, female abdomen much wider than in male (31% of maximum carapace width, versus 23% in male); all somites separate, somites 2–6 subequal in length, width; somites with median portion vaulted, lateral portions produced as anteriorly directed flanges. First somite subdorsal, conspicuously short, strongly curved, matching curvature of concave posterior carapace margin; somite 6 with salient dorsal uropods along majority of somite length. Telson elongate, posterior corners rounded, posterior part with subvertical margins with weak depressions, anteriorly broadly triangular with straight sides, rounded tip. Coxal abdominal holding mechanism present in both sexes at P1–P4 coxae; P1 coxa with granules, P2–P4 coxae with distinct granular prominence, overhanging lateral portions of abdominal somites. P1 strongly developed; basis, ischium separated; merus thick, rounded triangular in section, lower margin arched; carpus short, granular; chelipeds robust; propodus, carpus proximally with distinctive articulating bulge; palm tumid, coarsely granular, with rounded upper, lower margins; fingers short, stout, dactylus with furrow parallel to arched upper margin, both fingers with single crushing tooth, distally spoon-tipped. P2–P4 (partially preserved) robust; coxae large, anteriorly with distinctive coxo-sternal sliding arch; merus long, oval in section; carpus short; elements of P1–P4 granular. P5 (partially preserved) conspicuously reduced, carried subdorsally in oblique position, smooth.</paragraph>
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<caption id="39843CC1FF850B64A7D8F8A7264A3FF6" httpUri="https://zenodo.org/record/208743/files/figure.png" pageId="8" pageNumber="48" targetBox="[151,1435,320,1869]" targetPageId="8">
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<paragraph id="6D446C49FF850B64A7D8F8A7264A3FF6" blockId="8.[151,1436,1874,2019]" pageId="8" pageNumber="48">
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<emphasis id="5F8FB05BFF850B64A7D8F8A7227F3F7D" bold="true" box="[151,277,1874,1897]" pageId="8" pageNumber="48">FIGURE 3.</emphasis>
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<taxonomicName id="AAFB17CAFF850B64A652F8A621543F7C" box="[285,574,1875,1897]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
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<emphasis id="5F8FB05BFF850B64A652F8A621543F7C" box="[285,574,1875,1897]" italics="true" pageId="8" pageNumber="48">Graptocarcinus urbasaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="44BC0D20FF850B64A508F8A621E83F7C" box="[583,642,1875,1897]" pageId="8" pageNumber="48" rank="species">
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<emphasis id="5F8FB05BFF850B64A508F8A621163F7D" bold="true" box="[583,636,1875,1896]" pageId="8" pageNumber="48">n. sp</emphasis>
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.
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</taxonomicNameLabel>
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, MCNA 8562 (paratype, male), Santonian, Egibil quarry, Olazti-Olazagutía, Navarra, Spain. A, dorsal view; B, frontal view; C, ventral view; D, posterior view; E, detail of abdomen and pereiopod coxae; F, detail of posterior view.
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<emphasis id="5F8FB05BFF850B64A6E6F86521E33FB0" bold="true" box="[425,649,1936,1957]" pageId="8" pageNumber="48">a1, a2, a3, a4, a5, a6,</emphasis>
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abdominal somites 1, 2, 3, 4, 5, 6;
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<emphasis id="5F8FB05BFF850B64A4BCF865277E3FB0" bold="true" box="[1011,1044,1936,1957]" pageId="8" pageNumber="48">ab,</emphasis>
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articulating bulge;
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<emphasis id="5F8FB05BFF850B64A393F86523AB3FD1" bold="true" pageId="8" pageNumber="48">cx1, cx2, cx3, cx4, cx5,</emphasis>
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P1–P5 coxae;
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<emphasis id="5F8FB05BFF850B64A62CF85A22DB3FD1" bold="true" box="[355,433,1967,1988]" pageId="8" pageNumber="48">cmxp3,</emphasis>
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mxp3 coxa;
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<emphasis id="5F8FB05BFF850B64A572F85A213B3FD1" bold="true" box="[573,593,1967,1988]" pageId="8" pageNumber="48">d,</emphasis>
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depression;
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<emphasis id="5F8FB05BFF850B64A597F85A219E3FD1" bold="true" box="[728,756,1967,1988]" pageId="8" pageNumber="48">gr,</emphasis>
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granules;
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<emphasis id="5F8FB05BFF850B64A42BF85A20F63FD1" bold="true" box="[868,924,1967,1988]" pageId="8" pageNumber="48">mP5,</emphasis>
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P5 merus;
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<emphasis id="5F8FB05BFF850B64A357F85A275F3FD1" bold="true" box="[1048,1077,1967,1988]" pageId="8" pageNumber="48">pr,</emphasis>
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prominence;
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<emphasis id="5F8FB05BFF850B64A388F85A27893FD1" bold="true" box="[1223,1251,1967,1988]" pageId="8" pageNumber="48">pt,</emphasis>
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pterygostome;
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<emphasis id="5F8FB05BFF850B64A2C8F85A26F63FD1" bold="true" box="[1415,1436,1967,1988]" pageId="8" pageNumber="48">u,</emphasis>
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uropod;
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<emphasis id="5F8FB05BFF850B64A7A3F83B23903FF6" bold="true" box="[236,250,1998,2019]" pageId="8" pageNumber="48">t,</emphasis>
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telson. Scale bars: 10mm. Specimen whitened with ammonium chloride sublimate prior to photography.
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</paragraph>
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</caption>
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<caption id="39843CC1FF840B65A7D8F90626EB3F73" httpUri="https://zenodo.org/record/208744/files/figure.png" pageId="9" pageNumber="49" targetBox="[151,1435,193,1758]" targetPageId="9">
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<paragraph id="6D446C49FF840B65A7D8F90626EB3F73" blockId="9.[151,1436,1779,1894]" pageId="9" pageNumber="49">
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<emphasis id="5F8FB05BFF840B65A7D8F90622793F1C" bold="true" box="[151,275,1779,1802]" pageId="9" pageNumber="49">FIGURE 4.</emphasis>
|
||
<taxonomicName id="AAFB17CAFF840B65A655F90121AC3F1F" authority="Alcock, 1899" authorityName="Alcock" authorityYear="1899" box="[282,710,1780,1802]" class="Malacostraca" family="Dynomenidae" genus="Acanthodromia" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="49" phylum="Arthropoda" rank="species" species="margarita">
|
||
<emphasis id="5F8FB05BFF840B65A655F90121493F1F" box="[282,547,1780,1802]" italics="true" pageId="9" pageNumber="49">Acanthodromia margarita</emphasis>
|
||
(Alcock, 1899)
|
||
</taxonomicName>
|
||
, MNHN-B28812 (male and female), Balicasag Island, Panglao, Bohol, Visayas, Philippines. A, dorsal view of male, with markedly reduced, obliquely oriented P5; B, posterior view of male; C, ventral view of female, abdomen removed, showing thoracic sternum; D, ventral view of female, E, ventral view of male.
|
||
<emphasis id="5F8FB05BFF840B65A233F8C423B63F70" bold="true" pageId="9" pageNumber="49">a1, a2, a3,</emphasis>
|
||
abdominal somites 1, 2, 3;
|
||
<emphasis id="5F8FB05BFF840B65A6BBF8A5218E3F70" bold="true" box="[500,740,1872,1893]" pageId="9" pageNumber="49">cx1, cx2, cx3, cx4, cx5,</emphasis>
|
||
P1–P5 coxae;
|
||
<emphasis id="5F8FB05BFF840B65A434F8A520F73F70" bold="true" box="[891,925,1872,1893]" pageId="9" pageNumber="49">P5,</emphasis>
|
||
pereiopod 5;
|
||
<emphasis id="5F8FB05BFF840B65A366F8A5272D3F70" bold="true" box="[1065,1095,1872,1893]" pageId="9" pageNumber="49">pr,</emphasis>
|
||
prominence. Scale bars: 5mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6D446C49FF840B65A788F87821B63FF8" pageId="9" pageNumber="49">
|
||
<table id="1FFB9EE9FF84F493A7D8F87826F63FF8" box="[151,1436,1933,2029]" gridcols="3" gridrows="3" pageId="9" pageNumber="49">
|
||
<tr id="D3CB6E0BFF84F493A7D8F87826F63FB0" box="[151,1436,1933,1957]" gridrow="0" pageId="9" pageNumber="49">
|
||
<th id="901A0777FF84F493A7D8F87821733FB0" box="[151,537,1933,1957]" gridcol="0" gridrow="0" pageId="9" pageNumber="49">
|
||
<emphasis id="5F8FB05BFF840B65A788F87822513FB0" bold="true" box="[199,315,1933,1957]" pageId="9" pageNumber="49">Remarks.</emphasis>
|
||
Other species of
|
||
</th>
|
||
<th id="901A0777FF84F493A56FF878208D3FB0" box="[544,999,1933,1957]" gridcol="1" gridrow="0" pageId="9" pageNumber="49">
|
||
<taxonomicName id="AAFB17CAFF840B65A56FF87B21BA3FB0" box="[544,720,1934,1957]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5F8FB05BFF840B65A56FF87B21BA3FB0" box="[544,720,1934,1957]" italics="true" pageId="9" pageNumber="49">Graptocarcinus</emphasis>
|
||
</taxonomicName>
|
||
are only known from
|
||
</th>
|
||
<th id="901A0777FF84F493A4BEF87826F63FB0" box="[1009,1436,1933,1957]" gridcol="2" gridrow="0" pageId="9" pageNumber="49">carapace material. The carapace of</th>
|
||
</tr>
|
||
<tr id="D3CB6E0BFF84F493A7D8F84426F63FDC" box="[151,1436,1969,1993]" gridrow="1" pageId="9" pageNumber="49">
|
||
<th id="901A0777FF84F493A7D8F84421733FDC" box="[151,537,1969,1993]" gridcol="0" gridrow="1" pageId="9" pageNumber="49">
|
||
<taxonomicName id="AAFB17CAFF840B65A7D8F847222D3FDC" box="[151,327,1970,1993]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5F8FB05BFF840B65A7D8F847222D3FDC" box="[151,327,1970,1993]" italics="true" pageId="9" pageNumber="49">Graptocarcinus</emphasis>
|
||
</taxonomicName>
|
||
spp. also exhibits
|
||
</th>
|
||
<td id="901A0777FF84F493A56FF844208D3FDC" box="[544,999,1969,1993]" gridcol="1" gridrow="1" pageId="9" pageNumber="49">only a few distinctive characters, which</td>
|
||
<td id="901A0777FF84F493A4BEF84426F63FDC" box="[1009,1436,1969,1993]" gridcol="2" gridrow="1" pageId="9" pageNumber="49">makes comparison with other species</td>
|
||
</tr>
|
||
<tr id="D3CB6E0BFF84F493A7D8F82026F63FF8" box="[151,1436,2005,2029]" gridrow="2" pageId="9" pageNumber="49" rowspan-2="1">
|
||
<th id="901A0777FF84F493A7D8F82021733FF8" box="[151,537,2005,2029]" gridcol="0" gridrow="2" pageId="9" pageNumber="49">difficult. Nevertheless, differences</th>
|
||
<td id="901A0777FF84F493A56FF820208D3FF8" box="[544,999,2005,2029]" gridcol="1" gridrow="2" pageId="9" pageNumber="49">can be observed.</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="6D446C49FF870B66A788FF6D209338E2" blockId="10.[151,1436,151,788]" pageId="10" pageNumber="50">
|
||
<taxonomicName id="AAFB17CAFF870B66A788FF6D216E38A5" box="[199,516,152,176]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
|
||
<emphasis id="5F8FB05BFF870B66A788FF6D216E38A5" box="[199,516,152,176]" italics="true" pageId="10" pageNumber="50">Graptocarcinus urbasaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44BC0D20FF870B66A544FF6D212338A5" box="[523,585,152,176]" pageId="10" pageNumber="50" rank="species">
|
||
<emphasis id="5F8FB05BFF870B66A544FF6D212338A5" bold="true" box="[523,585,152,176]" pageId="10" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<emphasis id="5F8FB05BFF870B66A591FF6D205C38A5" box="[734,822,152,176]" italics="true" pageId="10" pageNumber="50">G. m u i r i</emphasis>
|
||
by the absence of an anterior mesogastric process and frontal sulcus (present in
|
||
<emphasis id="5F8FB05BFF870B66A6FEFF48216238C1" box="[433,520,189,212]" italics="true" pageId="10" pageNumber="50">G. m u i r i</emphasis>
|
||
); the supraorbital margin being more vaulted than
|
||
<emphasis id="5F8FB05BFF870B66A30EFF4827F238C1" box="[1089,1176,189,212]" italics="true" pageId="10" pageNumber="50">G. m u i r i</emphasis>
|
||
; and having the corner of the lateral carapace margins produced as a sharp flange (blunt in
|
||
<emphasis id="5F8FB05BFF870B66A4C2FF15208F38ED" box="[909,997,224,248]" italics="true" pageId="10" pageNumber="50">G. m u i r i</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6D446C49FF870B66A788FEF022EF39DA" blockId="10.[151,1436,151,788]" pageId="10" pageNumber="50">
|
||
<taxonomicName id="AAFB17CAFF870B66A788FEF022043909" box="[199,366,260,284]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
|
||
<emphasis id="5F8FB05BFF870B66A788FEF022043909" box="[199,366,260,284]" italics="true" pageId="10" pageNumber="50">G. urbasaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44BC0D20FF870B66A637FEF122D33909" box="[376,441,260,284]" pageId="10" pageNumber="50" rank="species">
|
||
<emphasis id="5F8FB05BFF870B66A637FEF122D33909" bold="true" box="[376,441,260,284]" pageId="10" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="AAFB17CAFF870B66A519FEF021A43909" box="[598,718,261,284]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A519FEF021A43909" box="[598,718,261,284]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of an anterior mesogastric process and frontal sulcus (present in
|
||
<taxonomicName id="AAFB17CAFF870B66A62CFEDD22B23955" box="[355,472,296,320]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A62CFEDD22B23955" box="[355,472,296,320]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
); the supraorbital margin being more vaulted than in
|
||
<taxonomicName id="AAFB17CAFF870B66A37BFEDD27C23955" box="[1076,1192,296,320]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A37BFEDD27C23955" box="[1076,1192,296,320]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
; having the corner of the lateral carapace margins produced as a sharp flange (blunt in
|
||
<taxonomicName id="AAFB17CAFF870B66A435FEB820853971" box="[890,1007,333,356]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A435FEB820853971" box="[890,1007,333,356]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
); the carapace being wider relative to its length; having sparser granules on the dorsal carapace surface (more granular in
|
||
<taxonomicName id="AAFB17CAFF870B66A31AFE8527A1399D" box="[1109,1227,368,392]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A31AFE8527A1399D" box="[1109,1227,368,392]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
); the anterolateral carapace margin not being strongly granular as in
|
||
<taxonomicName id="AAFB17CAFF870B66A59AFE60202639B9" box="[725,844,405,428]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A59AFE60202639B9" box="[725,844,405,428]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
; and the posterolateral carapace margin being blunt (sharp in
|
||
<taxonomicName id="AAFB17CAFF870B66A64FFE4D221F39C5" box="[256,373,440,464]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A64FFE4D221F39C5" box="[256,373,440,464]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6D446C49FF870B66A788FE2820173A29" blockId="10.[151,1436,151,788]" pageId="10" pageNumber="50">
|
||
<taxonomicName id="AAFB17CAFF870B66A788FE28220439E1" box="[199,366,476,500]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
|
||
<emphasis id="5F8FB05BFF870B66A788FE28220439E1" box="[199,366,476,500]" italics="true" pageId="10" pageNumber="50">G. urbasaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44BC0D20FF870B66A639FE2922DD39E1" box="[374,439,476,500]" pageId="10" pageNumber="50" rank="species">
|
||
<emphasis id="5F8FB05BFF870B66A639FE2922DD39E1" bold="true" box="[374,439,476,500]" pageId="10" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="AAFB17CAFF870B66A51DFE2821A339E1" box="[594,713,476,500]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="bellonii">
|
||
<emphasis id="5F8FB05BFF870B66A51DFE2821A339E1" box="[594,713,476,500]" italics="true" pageId="10" pageNumber="50">G. bellonii</emphasis>
|
||
</taxonomicName>
|
||
by absence of the anterior mesogastric process / frontal sulcus (present in
|
||
<taxonomicName id="AAFB17CAFF870B66A65AFDF522E33A0D" box="[277,393,512,536]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="bellonii">
|
||
<emphasis id="5F8FB05BFF870B66A65AFDF522E33A0D" box="[277,393,512,536]" italics="true" pageId="10" pageNumber="50">G. bellonii</emphasis>
|
||
</taxonomicName>
|
||
); having much larger orbits relative to the carapace than
|
||
<taxonomicName id="AAFB17CAFF870B66A343FDF527EA3A0D" box="[1036,1152,512,536]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="bellonii">
|
||
<emphasis id="5F8FB05BFF870B66A343FDF527EA3A0D" box="[1036,1152,512,536]" italics="true" pageId="10" pageNumber="50">G. bellonii</emphasis>
|
||
</taxonomicName>
|
||
; and the corner of lateral carapace margins produced as a sharp flange (blunt in
|
||
<emphasis id="5F8FB05BFF870B66A5B6FDD020033A29" box="[761,873,548,572]" italics="true" pageId="10" pageNumber="50">G. b e l l o n i i</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6D446C49FF870B66A788FDB220AD3A91" blockId="10.[151,1436,151,788]" pageId="10" pageNumber="50">
|
||
The claw figured by
|
||
<bibRefCitation id="096A11B8FF870B66A6FFFDBD21033A4A" author="Jagt" box="[432,617,583,608]" pageId="10" pageNumber="50" refString="Jagt, J. W. M., Fraaije, R. H. B., Van Bakel, B. W. M. & Artal, P. (2010) Necrocarcinus ornatissimus Forir, 1887, and Prehepatus werneri Fraaye & Collins, 1987 (upper Maastrichtian, the Netherlands) revisited, with notes on other Cretaceous dynomenid crabs (Decapoda, Brachyura). In: Castro, P., Davie, P. J. F., Ng, P. K. L. & Richer de Forges, B. (Eds.), Studies on Brachyura: a homage to Daniele Guinot. Crustaceana Monographs, 11, 173 - 195." type="journal article" year="2010">
|
||
Jagt
|
||
<emphasis id="5F8FB05BFF870B66A6ABFDBC217E3A75" box="[484,532,584,608]" italics="true" pageId="10" pageNumber="50">et al</emphasis>
|
||
. (2010
|
||
</bibRefCitation>
|
||
: fig.4e–h) as
|
||
<taxonomicName id="AAFB17CAFF870B66A5B1FDBD20C53A4A" box="[766,943,584,607]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="5F8FB05BFF870B66A5B1FDBD20C53A4A" box="[766,943,584,607]" italics="true" pageId="10" pageNumber="50">Graptocarcinus</emphasis>
|
||
</taxonomicName>
|
||
sp. differs from the claws of
|
||
<taxonomicName id="AAFB17CAFF870B66A3B8FDBD26F63A75" box="[1271,1436,584,608]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="urbasaensis" status="sp. nov.">
|
||
<emphasis id="5F8FB05BFF870B66A3B8FDBD26F63A75" box="[1271,1436,584,608]" italics="true" pageId="10" pageNumber="50">G. urbasaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="44BC0D20FF870B66A7D8FD9923BF3A91" box="[151,213,620,644]" pageId="10" pageNumber="50" rank="species">
|
||
<emphasis id="5F8FB05BFF870B66A7D8FD9923BF3A91" bold="true" box="[151,213,620,644]" pageId="10" pageNumber="50">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="AAFB17CAFF870B66A643FD9822EA3A91" box="[268,384,621,644]" class="Malacostraca" family="Dynomenidae" genus="Graptocarcinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="texanus">
|
||
<emphasis id="5F8FB05BFF870B66A643FD9822EA3A91" box="[268,384,621,644]" italics="true" pageId="10" pageNumber="50">G. texanus</emphasis>
|
||
</taxonomicName>
|
||
, the fingers being more spatulate and without teeth.
|
||
</paragraph>
|
||
<paragraph id="6D446C49FF870B66A788FD7A226D3B01" blockId="10.[151,1436,151,788]" pageId="10" pageNumber="50">
|
||
<bibRefCitation id="096A11B8FF870B66A788FD7A215A3AB2" author="Amezarri" box="[199,560,655,680]" pageId="10" pageNumber="50" refString="Amezarri Grandal, E., Galan Fernandez, R., Saiz de Omenaca, J. & Via Boada, L. (1977) Nota previa sobre un nuevo yacimiento de Braquiuros Cretacicos, en Ribamontan al Mar (Santander). Breviora Geologica Asturica, 4, 49 - 50." type="journal article" year="1977">
|
||
Amézarri Grandal
|
||
<emphasis id="5F8FB05BFF870B66A6EDFD6422B23ABD" box="[418,472,656,680]" italics="true" pageId="10" pageNumber="50">et al</emphasis>
|
||
. (1977
|
||
</bibRefCitation>
|
||
: 49, 50) illustrated the dorsal and ventral side of a well preserved female specimen with pereiopods, abdomen and mxp3 that corresponds to the new species described here, but they failed to formally describe and name it. This publication is rare, never referred to, and the whereabouts of the specimen is unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |