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<mods:identifier id="985F58972EC46644D29EB471525DCBC6" type="Pensoft-Pub">1314-2003-240-1</mods:identifier>
<mods:identifier id="70D4C70003BBA92586E958830DB55738" type="Pensoft-UUID">B699D9DE2B435B1093DE3C38C703D430</mods:identifier>
</mods:mods>
<treatment id="2DF816101B8D61477591A7D3C5505AB7" LSID="urn:lsid:plazi:treatment:2DF816101B8D61477591A7D3C5505AB7" httpUri="http://treatment.plazi.org/id/2DF816101B8D61477591A7D3C5505AB7" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="5C33D14BAA1B6D5DA4352163C0E90C82" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="C1CB028BE40423FB1267AB5628EEAEFB" pageId="0" pageNumber="1">
<taxonomicName id="5FBA02ACCCC7ED72743A1E3DD237D08E" ID-CoL="9CL73" authority="Benth., J. Bot. (Hooker) 4: 416. 1842." authorityName="Benth., J. Bot. (Hooker) 4: 416. 1842." class="Magnoliopsida" family="Fabaceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Leucaena Benth., J. Bot. (Hooker) 4: 416. 1842.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="E5B975956B056231129FF56387ACB52A" pageId="0" pageNumber="1" type="description">
<paragraph id="6523A609C91E0630800B48D9AB7FEC7D" pageId="0" pageNumber="1">
<figureCitation id="0B43D0E9DA08A260CEFA0C2EE0B8CAE7" captionStart="Figure 144" captionStartId="F151" captionText="Figure 144. Habit and growth forms of genera of the Dichrostachys clade A herbaceous to weakly woody thicket of shrubby Neptunia plena (L.) Benth. in seasonally inundated ruderal habitat, Bolivia (Saerkinen 2159) B, C medium-sized to large trees of Lemurodendron capuronii Villiers &amp; P. Guinet, north-western Madagascar (Koenen 429) D small tree of Leucaena matudae (Zarate) C. E. Hughes in seasonally dry tropical forest, Guerrero, Mexico (Hughes 2153) E shrub of Kanaloa kahoolawensis Lorence &amp; K. R. Wood cultivated, National Tropical Botanical Garden, Kaua'i, Hawaii F small treelet of Desmanthus fruticosus Rose, Baja California, Mexico (Hughes 1532) G prostrate woody suffrutex of Desmanthus acuminatus Benth., Potosi, Bolivia (Hughes 2314) H shrub of Mimozyganthus carinatus (Griseb.) Burkart, Santa Cruz, Bolivia (Hughes 2476) I shrub of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart, Mendoza, Argentina J shrublet of Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet, Puebla, Mexico (Hughes 1784) K small treelet of Dichrostachys cinerea (L.) Wight &amp; Arn., Kruger National Park, South Africa L small treelet of Alantsilodendron pilosum Villiers, S. Madagascar (Luckow 4162) M treelet of Gagnebina pterocarpa (Lam.) Baill., Madagascar N shrub of Dichrostachys venosa Villiers, in dry spiny forest, southern Madagascar. Photo credits A, D-H, J CE Hughes B, C E Koenen I Estepa Patagonica K B Dupont L M Luckow M C Commenge naturetropicale N D Du Puy." figureDoi="10.3897/phytokeys.240.101716.figure144" httpUri="https://binary.pensoft.net/fig/1014816" pageId="0" pageNumber="1">Figs 144</figureCitation>
<figureCitation id="21AAF98BA6350914BF5C8929315C7105" captionStart="Figure 145" captionStartId="F152" captionText="Figure 145. Inflorescences and flowers of genera of the Dichrostachys clade A Lemurodendron capuronii Villiers &amp; P. Guinet (Koenen 435) B inflorescence of Neptunia plena (L.) Benth., basal flowers sterile with showy yellow petaloid staminodia (Wood 26650) C inflorescence of Leucaena esculenta (Sesse &amp; Mocino ex DC.) Benth. (Hughes 1326) D inflorescences and tergeminate leaves of Kanaloa kahoolawensis Lorence &amp; K. R. Wood E fascicles of capitate inflorescences in axils of coevally developing leaves, Schleinitzia fosbergii Nevling &amp; Niezgoda F inflorescences of Desmanthus bicornutus S. Watson showing basal sterile flowers with exserted white showy staminodia (Hughes 1526) G capitate inflorescences of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart H semi-spherical capitate inflorescences of Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet (Hughes 1784) I inflorescence of Alantsilodendron ramosum Villiers J spicate inflorescences of Gagnebina pterocarpa (Lam.) Baill., the basal flowers sterile with showy white staminodia K spicate inflorescence of Dichrostachys cinerea (L.) Wight &amp; Arn., basal flowers sterile with showy pink staminodia L compact spikes of Dichrostachys akataensis Villiers the basal flowers sterile with showy pale pink staminodia. Photo credits A E Koenen B, C, F, H, J, K CE Hughes D A Palomino E Lauren Gutierrez G D Cabral, Flora Mondocina I, L D Du Puy." figureDoi="10.3897/phytokeys.240.101716.figure145" httpUri="https://binary.pensoft.net/fig/1014817" pageId="0" pageNumber="1">, 145</figureCitation>
<figureCitation id="1DF3E6C9B2689FB3B0D63ABA2630652C" captionStart="Figure 146" captionStartId="F153" captionText="Figure 146. Fruits of genera of the Dichrostachys clade A leaves and pod of Lemurodendron capuronii Villiers &amp; P. Guinet, north-eastern Madagascar (Koenen 429) B Neptunia plena (L.) Benth. fruits dehiscing along the dorsal suture (Wood 26650) C ripe pods of Leucaena cruziana Britton &amp; Rose, Oaxaca, Mexico (Hughes 1300) D cluster of unripe pods on a single inflorescence of Leucaena leucocephala (Lam.) de Wit subsp. glabrata (Rose) S. Zarate, Honduras (Hughes 1557) E ripe pods of Desmanthus leptophyllus Kunth dehiscing along both sutures (Hughes 3098) F unripe fruit of Mezcala balsensis (J. L. Contr.) C. E. Hughes &amp; J. L. Contr., held erect, Guerrero, Mexico G ripe fruits of Mezcala balsensis dehiscing from the apex along both sutures H, I unripe and ripe fruits of Kanaloa kahoolawensis Lorence &amp; K. R. Wood, cultivated at the Olinda rare plant propagation facility, Maui, Hawaii, USA J ripe fruits of Schleinitzia insularum (Guilemin) Burkart, the valves tardily splitting along both sutures but indehiscent K winged fruits of Gagnebina pterocarpa (Lam.) Baill., Madagascar L unripe, lomentiform fruits of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart which break up into one- or few-seeded fragments when ripe M ripe fruits of Mimozyganthus carinatus (Griseb.) Burkart, S Bolivia (Hughes 2476) N ripe fruits, elastically dehisced from the apex, Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet, Puebla, Mexico (Hughes 1784) O unripe coiled fruits of Dichrostachys cinerea (L.) Wight &amp; Arn. P ripe fruits dehiscing elastically from the apex, Dichrostachys venosa Villiers, southern Madagascar Q ripe fruits of Alantsilodendron pilosum Villiers, Madagascar. Photo credits A E Koenen B-E, M, N C Hughes F, G J Luis Contreras H, I A Palomino J G McCormack, Cook Islands Biodiversity K, P, Q D Du Puy L I Specogno, Flora Mendocina O Atamari (CC-BY-SA- 3.0)." figureDoi="10.3897/phytokeys.240.101716.figure146" httpUri="https://binary.pensoft.net/fig/1014818" pageId="0" pageNumber="1">, 146</figureCitation>
<figureCitation id="E88A521CF51435C1D6765CCD09777CD6" captionStart="Figure 149" captionStartId="F156" captionText="Figure 149. Distribution of Leucaena based on quality-controlled digitised herbarium records. Note that the pantropically cultivated and widely introduced species Leucaena leucocephala (Lam.) de Wit, whose true native distribution remains unknown (Hughes 1998 a), is not included in the map. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure149" httpUri="https://binary.pensoft.net/fig/1014821" pageId="0" pageNumber="1">, 149</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7E571EB918CADE58F2B094FA127BA293" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="931E77A65F862EC949DB656B6CD94C59" pageId="0" pageNumber="1">
<taxonomicName id="D471406BCF579C44C9B512D2ABCC75FD" class="Magnoliopsida" family="Fabaceae" genus="Ryncholeucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ryncholeucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Ryncholeucaena</taxonomicName>
Britton &amp; Rose, Fl. N. Amer. 23: 130. 1928. Type: 
<taxonomicName id="6B03CDFFFEE2B3F223A88F304E898403" class="Magnoliopsida" family="Fabaceae" genus="Ryncholeucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ryncholeucaena greggii" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="greggii">Ryncholeucaena greggii</taxonomicName>
(S. Watson) Britton &amp; Rose [≡ 
<taxonomicName id="E652652843FEF30F74D32AECEB068815" class="Magnoliopsida" family="Fabaceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena greggii" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="greggii">Leucaena greggii</taxonomicName>
S. Watson]
</paragraph>
<paragraph id="A5513B48F71DC6608E7C5A1CD4C47904" pageId="0" pageNumber="1">
<taxonomicName id="01372F6C0537411C4CD57DBE93416F81" class="Magnoliopsida" family="Fabaceae" genus="Caudoleucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caudoleucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Caudoleucaena</taxonomicName>
Britton &amp; Rose, Fl. N. Amer. 23: 130. 1928. Type: 
<taxonomicName id="EC8BD370C70024D9C6BA0DAD634B5F17" class="Magnoliopsida" family="Fabaceae" genus="Caudoleucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Caudoleucaena retusa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="retusa">Caudoleucaena retusa</taxonomicName>
(Benth.) Britton &amp; Rose [≡ 
<taxonomicName id="D24EC012873189810B3EEB3FE48B7520" class="Magnoliopsida" family="Fabaceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena retusa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="retusa">Leucaena retusa</taxonomicName>
Benth.]
</paragraph>
</subSubSection>
<subSubSection id="88F873F3C08441A9C310DE3470BCB8E0" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="4F1FFEAAC2F6FC48BBDB74A1BE11B80A" pageId="0" pageNumber="1">
<typeStatus id="07785EB043786F6B7AF31B5B74330F7B">Type</typeStatus>
.
</paragraph>
<paragraph id="454FB9564366E134074DEDB2CEC0B846" pageId="0" pageNumber="1">
<taxonomicName id="6E829E1E9FFE7340384F9FC39DF1B753" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena diversifolia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="diversifolia">
<emphasis id="BD79B9A80EE4343B5D97C218CDB08FA1" italics="true" pageId="0" pageNumber="1">Leucaena diversifolia</emphasis>
</taxonomicName>
(Schlecht.) Benth. [≡ 
<taxonomicName id="E2CACE40227B5EB63A257B799158815B" class="Magnoliopsida" family="Caesalpiniaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia diversifolia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="diversifolia">
<emphasis id="677E946FDC767F5ED66A1072AA291904" italics="true" pageId="0" pageNumber="1">Acacia diversifolia</emphasis>
</taxonomicName>
Schlecht.]
</paragraph>
</subSubSection>
<subSubSection id="88B2849D9B3523EC15D44BB4EAEF8010" pageId="0" pageNumber="1" type="description">
<paragraph id="134F40E461F4222CC21F24E99C2F902D" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="BFFF702DA704C5DC20A08438DCB74E24" pageId="0" pageNumber="1">
Unarmed small or medium-sized trees, 2-15 (20) m (Fig. 
<figureCitation id="CCD9F4C02F150002116AD3848852155D" captionStart="Figure 144" captionStartId="F151" captionText="Figure 144. Habit and growth forms of genera of the Dichrostachys clade A herbaceous to weakly woody thicket of shrubby Neptunia plena (L.) Benth. in seasonally inundated ruderal habitat, Bolivia (Saerkinen 2159) B, C medium-sized to large trees of Lemurodendron capuronii Villiers &amp; P. Guinet, north-western Madagascar (Koenen 429) D small tree of Leucaena matudae (Zarate) C. E. Hughes in seasonally dry tropical forest, Guerrero, Mexico (Hughes 2153) E shrub of Kanaloa kahoolawensis Lorence &amp; K. R. Wood cultivated, National Tropical Botanical Garden, Kaua'i, Hawaii F small treelet of Desmanthus fruticosus Rose, Baja California, Mexico (Hughes 1532) G prostrate woody suffrutex of Desmanthus acuminatus Benth., Potosi, Bolivia (Hughes 2314) H shrub of Mimozyganthus carinatus (Griseb.) Burkart, Santa Cruz, Bolivia (Hughes 2476) I shrub of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart, Mendoza, Argentina J shrublet of Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet, Puebla, Mexico (Hughes 1784) K small treelet of Dichrostachys cinerea (L.) Wight &amp; Arn., Kruger National Park, South Africa L small treelet of Alantsilodendron pilosum Villiers, S. Madagascar (Luckow 4162) M treelet of Gagnebina pterocarpa (Lam.) Baill., Madagascar N shrub of Dichrostachys venosa Villiers, in dry spiny forest, southern Madagascar. Photo credits A, D-H, J CE Hughes B, C E Koenen I Estepa Patagonica K B Dupont L M Luckow M C Commenge naturetropicale N D Du Puy." figureDoi="10.3897/phytokeys.240.101716.figure144" httpUri="https://binary.pensoft.net/fig/1014816" pageId="0" pageNumber="1">144D</figureCitation>
), brachyblasts absent. 
<emphasis id="EAA37580BAD1A73F055BD7F2A0DDC4E0" bold="true" pageId="0" pageNumber="1">Stipules</emphasis>
persistent or caducous, ovate or subulate. 
<emphasis id="237301F2E171266309F6C2DCB87F3902" bold="true" pageId="0" pageNumber="1">Leaves</emphasis>
bipinnate, extrafloral nectaries between proximal pair of pinnae, sometimes between other pinnae pairs and on pinnular rachis, to 75 on a single leaf (
<taxonomicName id="CF479F706BB6AC7A247414F266C409C3" lsidName="L. trichandra" pageId="0" pageNumber="1" rank="species" species="trichandra">
<emphasis id="03C86828DC9904FB756845477B174E58" italics="true" pageId="0" pageNumber="1">L. trichandra</emphasis>
</taxonomicName>
(Zucc.) Urban); pinnae 1-20 (60) pairs, opposite; leaflets 3-65 (85) pairs per pinna, variable in size and shape (linear, oblong, elliptic). 
<emphasis id="F1943A129E58538CB34946AE0AE39257" bold="true" pageId="0" pageNumber="1">Inflorescences</emphasis>
capitate, peduncle with a distal, or sub-distal involucel of united bracts, 1 (4) in leaf axils or on 1-2-branched terminal panicles exserted beyond the leaves (Fig. 
<figureCitation id="EDDDE074AD2726A6FCA94D740CA6FAFA" captionStart="Figure 145" captionStartId="F152" captionText="Figure 145. Inflorescences and flowers of genera of the Dichrostachys clade A Lemurodendron capuronii Villiers &amp; P. Guinet (Koenen 435) B inflorescence of Neptunia plena (L.) Benth., basal flowers sterile with showy yellow petaloid staminodia (Wood 26650) C inflorescence of Leucaena esculenta (Sesse &amp; Mocino ex DC.) Benth. (Hughes 1326) D inflorescences and tergeminate leaves of Kanaloa kahoolawensis Lorence &amp; K. R. Wood E fascicles of capitate inflorescences in axils of coevally developing leaves, Schleinitzia fosbergii Nevling &amp; Niezgoda F inflorescences of Desmanthus bicornutus S. Watson showing basal sterile flowers with exserted white showy staminodia (Hughes 1526) G capitate inflorescences of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart H semi-spherical capitate inflorescences of Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet (Hughes 1784) I inflorescence of Alantsilodendron ramosum Villiers J spicate inflorescences of Gagnebina pterocarpa (Lam.) Baill., the basal flowers sterile with showy white staminodia K spicate inflorescence of Dichrostachys cinerea (L.) Wight &amp; Arn., basal flowers sterile with showy pink staminodia L compact spikes of Dichrostachys akataensis Villiers the basal flowers sterile with showy pale pink staminodia. Photo credits A E Koenen B, C, F, H, J, K CE Hughes D A Palomino E Lauren Gutierrez G D Cabral, Flora Mondocina I, L D Du Puy." figureDoi="10.3897/phytokeys.240.101716.figure145" httpUri="https://binary.pensoft.net/fig/1014817" pageId="0" pageNumber="1">145C</figureCitation>
), composed mainly of hermaphrodite flowers, sterile flowers with staminodia absent; bracteoles subtending each flower bud persistent, peltate, sometimes exserted in bud, round, lanceolate or caudate. 
<emphasis id="A3C10E7D54B92AA39F71E5E06CCD1306" bold="true" pageId="0" pageNumber="1">Flowers</emphasis>
with 5 sepals, valvate in bud, calyx obconic, tubular or campanulate or united along mid-portion; petals 5, valvate, usually free, white or pale green; stamens 10, filaments usually white, but occasionally pink, yellow or reddish, anthers usually pilose, occasionally glabrous, usually eglandular, but sometimes the connective with a small rounded or hooded apiculum; pollen highly variable among species, occurring in monads, acalymmate polyads, or calymmate tetrahedral tetrads, tricolporate or pantoporate, exine smooth, psilate or punctate; ovary sessile or sub-sessile, glabrous or hairy, stigma narrowly funnelform. 
<emphasis id="A63490A36CC9D8F7F5529D06DE3C980B" bold="true" pageId="0" pageNumber="1">Fruits</emphasis>
linear, flattened, (5) 10-20 (26)-seeded, valves chartaceous or coriaceous, inertly dehiscent along one or usually both sutures (Fig. 
<figureCitation id="F26DA7A165AE12A5FF0E30BB71D91C62" captionStart="Figure 146" captionStartId="F153" captionText="Figure 146. Fruits of genera of the Dichrostachys clade A leaves and pod of Lemurodendron capuronii Villiers &amp; P. Guinet, north-eastern Madagascar (Koenen 429) B Neptunia plena (L.) Benth. fruits dehiscing along the dorsal suture (Wood 26650) C ripe pods of Leucaena cruziana Britton &amp; Rose, Oaxaca, Mexico (Hughes 1300) D cluster of unripe pods on a single inflorescence of Leucaena leucocephala (Lam.) de Wit subsp. glabrata (Rose) S. Zarate, Honduras (Hughes 1557) E ripe pods of Desmanthus leptophyllus Kunth dehiscing along both sutures (Hughes 3098) F unripe fruit of Mezcala balsensis (J. L. Contr.) C. E. Hughes &amp; J. L. Contr., held erect, Guerrero, Mexico G ripe fruits of Mezcala balsensis dehiscing from the apex along both sutures H, I unripe and ripe fruits of Kanaloa kahoolawensis Lorence &amp; K. R. Wood, cultivated at the Olinda rare plant propagation facility, Maui, Hawaii, USA J ripe fruits of Schleinitzia insularum (Guilemin) Burkart, the valves tardily splitting along both sutures but indehiscent K winged fruits of Gagnebina pterocarpa (Lam.) Baill., Madagascar L unripe, lomentiform fruits of Prosopidastrum globosum (Gillies ex Hook. &amp; Arn.) Burkart which break up into one- or few-seeded fragments when ripe M ripe fruits of Mimozyganthus carinatus (Griseb.) Burkart, S Bolivia (Hughes 2476) N ripe fruits, elastically dehisced from the apex, Calliandropsis nervosa (Britton &amp; Rose) H. M. Hern. &amp; P. Guinet, Puebla, Mexico (Hughes 1784) O unripe coiled fruits of Dichrostachys cinerea (L.) Wight &amp; Arn. P ripe fruits dehiscing elastically from the apex, Dichrostachys venosa Villiers, southern Madagascar Q ripe fruits of Alantsilodendron pilosum Villiers, Madagascar. Photo credits A E Koenen B-E, M, N C Hughes F, G J Luis Contreras H, I A Palomino J G McCormack, Cook Islands Biodiversity K, P, Q D Du Puy L I Specogno, Flora Mendocina O Atamari (CC-BY-SA- 3.0)." figureDoi="10.3897/phytokeys.240.101716.figure146" httpUri="https://binary.pensoft.net/fig/1014818" pageId="0" pageNumber="1">146C, D</figureCitation>
). 
<emphasis id="6193DF1DCDD2B9C12844FB59E84D2F8E" bold="true" pageId="0" pageNumber="1">Seeds</emphasis>
transverse or occasionally oblique in pod, ovate to rhomboidal, pleurogram U-shaped.
</paragraph>
</subSubSection>
<subSubSection id="62CFF03F1AF0DF0EAB88C96A99C6B954" pageId="0" pageNumber="1" type="chromosome number">
<paragraph id="1F5AC1A929D27904E9119EAF98175DDD" pageId="0" pageNumber="1">Chromosome number.</paragraph>
<paragraph id="F587A063C48E4D9941795E5020C3797F" pageId="0" pageNumber="1">
2 
<emphasis id="2ADBDFADC47BB496ED734030CDD0C2EF" italics="true" pageId="0" pageNumber="1">n</emphasis>
= 52, 56, 104, 112 (
<bibRefCitation id="D0B227C8124F1C51DA627BAC358AEAFA" DOI="https://doi.org/10.1111/j.1095-8339.2000.tb00550.x" author="Cardoso, MB" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="549 - 556" refId="B160" refString="Cardoso, MB, Schifino-Wittmann, MT, Bodanese-Zanettini, MH, 2000. Taxonomic and evolutionary implications of intraspecific variability in chromosome numbers of species of Leucaena Benth. (Leguminosae). Botanical Journal of the Linnean Society 134: 549 - 556, DOI: https://doi.org/10.1111/j.1095-8339.2000.tb00550.x" title="Taxonomic and evolutionary implications of intraspecific variability in chromosome numbers of species of Leucaena Benth. (Leguminosae)." url="https://doi.org/10.1111/j.1095-8339.2000.tb00550.x" volume="134" year="2000">Cardoso et al. 2000</bibRefCitation>
); whole genus paleopolyploid; five documented neo-allopolyploid species (
<bibRefCitation id="1A09DE26DC32C719CB379642D27CA114" DOI="https://doi.org/10.3732/ajb.1100260" author="Govindarajulu, R" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="2064 - 2076" refId="B318" refString="Govindarajulu, R, Hughes, CE, Alexander, PJ, Bailey, CD, 2011a. The complex evolutionary dynamics of ancient and recent polyploidy in Leucaena (Leguminosae; Mimosoideae). American Journal of Botany 98 (12): 2064 - 2076, DOI: https://doi.org/10.3732/ajb.1100260" title="The complex evolutionary dynamics of ancient and recent polyploidy in Leucaena (Leguminosae; Mimosoideae)." url="https://doi.org/10.3732/ajb.1100260" volume="98" year="2011 a">Govindarajulu et al. 2011a</bibRefCitation>
); one relatively common named sterile triploid, 
<taxonomicName id="CC683B9ED6AD0BAADC1052E61E463C4D" isHybrid="true" lsidName="L. mixtec" pageId="0" pageNumber="1" rank="species" species="mixtec">
L. 
<normalizedToken id="FA07B61C08ECDC20365F7BF3CBE681B7" originalValue="×">x</normalizedToken>
mixtec
</taxonomicName>
C.E. Hughes &amp; S.A. Harris (
<bibRefCitation id="E4134B32D7B662B98AE0598D86C748D2" DOI="https://doi.org/10.1007/BF00986251" author="Hughes, CE" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="1" pagination="177 - 197" refId="B384" refString="Hughes, CE, Harris, SA, 1994. The characterisation and identification of a naturally occurring hybrid in the genus Leucaena (Leguminosae: Mimosoideae). Plant Systematics and Evolution 192: 177 - 197, DOI: https://doi.org/10.1007/BF00986251" title="The characterisation and identification of a naturally occurring hybrid in the genus Leucaena (Leguminosae: Mimosoideae)." url="https://doi.org/10.1007/BF00986251" volume="192" year="1994">Hughes and Harris 1994</bibRefCitation>
) and one named homoploid hybrid, 
<taxonomicName id="6ECEBA34187B93205A3D47A4E1451FA3" isHybrid="true" lsidName="L. spontanea" pageId="0" pageNumber="1" rank="species" species="spontanea">
L. 
<normalizedToken id="429FA3131CBE5B3E1EE26FACEA2213E5" originalValue="×">x</normalizedToken>
spontanea
</taxonomicName>
C.E. Hughes &amp; S.A. Harris, at the tetraploid level (
<bibRefCitation id="F37DFFB7DC70D0C58B4780DECA2DD8DD" DOI="https://doi.org/10.1007/BF00985221" author="Hughes, CE" journalOrPublisher="Plant Systematics and Evolution" pageId="0" pageNumber="1" pagination="53 - 77" refId="B385" refString="Hughes, CE, Harris, SA, 1998. A second spontaneous hybrid in the genus Leucaena (Leguminosae, Mimosoideae). Plant Systematics and Evolution 212: 53 - 77, DOI: https://doi.org/10.1007/BF00985221" title="A second spontaneous hybrid in the genus Leucaena (Leguminosae, Mimosoideae)." url="https://doi.org/10.1007/BF00985221" volume="212" year="1998">Hughes and Harris 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="73D1390679D016D785DB55EA4CD86B92" pageId="0" pageNumber="1" type="distribution">
<paragraph id="89722CA2DE080996980B230BA0867703" pageId="0" pageNumber="1">Included species and geographic distribution.</paragraph>
<paragraph id="75385432BD7333AC5FE38856A908E415" pageId="0" pageNumber="1">
Twenty-four species plus two named hybrids (
<bibRefCitation id="C19BE37D5991BAE8F1DA93C834100BD5" DOI="https://doi.org/10.2307/25027876" author="Hughes, CE" journalOrPublisher="Systematic Botany Monographs" pageId="0" pageNumber="1" pagination="1 - 244" refId="B381" refString="Hughes, CE, 1998a. Monograph of Leucaena (Leguminosae - Mimosoideae). Systematic Botany Monographs 55: 1 - 244, DOI: https://doi.org/10.2307/25027876" title="Monograph of Leucaena (Leguminosae - Mimosoideae)." url="https://doi.org/10.2307/25027876" volume="55" year="1998 a">Hughes 1998a</bibRefCitation>
; 
<bibRefCitation id="5BDD2544910000AC85D7B19BB117DCF2" DOI="https://doi.org/10.3732/ajb.1100259" author="Govindarajulu, R" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="2049 - 2063" refId="B319" refString="Govindarajulu, R, Hughes, CE, Bailey, CD, 2011b. Phylogenetic and population genetic assessment of diploid Leucaena reveal cryptic species diversity and patterns of allopatric speciation. American Journal of Botany 98 (12): 2049 - 2063, DOI: https://doi.org/10.3732/ajb.1100259" title="Phylogenetic and population genetic assessment of diploid Leucaena reveal cryptic species diversity and patterns of allopatric speciation." url="https://doi.org/10.3732/ajb.1100259" volume="98" year="2011 b">Govindarajulu et al. 2011b</bibRefCitation>
). Mexico 11 endemic species with two extending to the USA (Texas and New Mexico) and four species to Central America; Central America five endemic species (Guatemala to Panama); one species in South America; one species pantropically introduced (Fig. 
<figureCitation id="430749976AFCE0263704FAAD311BFD2E" captionStart="Figure 149" captionStartId="F156" captionText="Figure 149. Distribution of Leucaena based on quality-controlled digitised herbarium records. Note that the pantropically cultivated and widely introduced species Leucaena leucocephala (Lam.) de Wit, whose true native distribution remains unknown (Hughes 1998 a), is not included in the map. See Suppl. material 1 for the source of occurrence data." figureDoi="10.3897/phytokeys.240.101716.figure149" httpUri="https://binary.pensoft.net/fig/1014821" pageId="0" pageNumber="1">149</figureCitation>
).
</paragraph>
<caption id="8236DC9B41930F596C72D5FD09663E6E" doi="10.3897/phytokeys.240.101716.figure149" httpUri="https://binary.pensoft.net/fig/1014821" pageId="0" pageNumber="1" start="Figure 149" startId="F156">
<paragraph id="6A9D1E52076858BB290701A7ECDCEA4E" pageId="0" pageNumber="1">
<emphasis id="8E59F08D2133F5ABAB6111FCB41123B9" bold="true" pageId="0" pageNumber="1">Figure 149.</emphasis>
Distribution of 
<taxonomicName id="95F752F73A3ED9804ADE8C67E28877DB" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="A254626A6A56924BFE9F6F65F841B3DF" italics="true" pageId="0" pageNumber="1">Leucaena</emphasis>
</taxonomicName>
based on quality-controlled digitised herbarium records. Note that the pantropically cultivated and widely introduced species 
<taxonomicName id="F0116090EC7ECE9AD3149AC6AFFA5DC5" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena leucocephala" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="leucocephala">
<emphasis id="C3ED1773B017C6B2C9AE22523A2B7B5D" italics="true" pageId="0" pageNumber="1">Leucaena leucocephala</emphasis>
</taxonomicName>
(Lam.) de Wit, whose true native distribution remains unknown (
<bibRefCitation id="21877FDD4AF04BF6C4B854E9201161A0" DOI="https://doi.org/10.2307/25027876" author="Hughes, CE" journalOrPublisher="Systematic Botany Monographs" pageId="0" pageNumber="1" pagination="1 - 244" refId="B381" refString="Hughes, CE, 1998a. Monograph of Leucaena (Leguminosae - Mimosoideae). Systematic Botany Monographs 55: 1 - 244, DOI: https://doi.org/10.2307/25027876" title="Monograph of Leucaena (Leguminosae - Mimosoideae)." url="https://doi.org/10.2307/25027876" volume="55" year="1998 a">Hughes 1998a</bibRefCitation>
), is not included in the map. See Suppl. material 1 for the source of occurrence data.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="AAEBD6D23CAA90324BB7CCEE2AB313D9" pageId="0" pageNumber="1" type="ecology">
<paragraph id="3D97A93AE8A6ABCE7967297EA87263CB" pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph id="737483F72116AAE21E36EFC9351C54E9" pageId="0" pageNumber="1">
Seasonally dry tropical forest and semi-arid thorn scrub; two species extending to warm temperate scrublands and one species to rainforests. Seed dispersal passive. All species deciduous. Nodulating, symbiosome present. Bee-pollinated (
<taxonomicName id="587030F1B4D7FC2D6881D4472CD947EB" class="Insecta" family="Apidae" genus="Xylocopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Xylocopa" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthopoda" rank="genus">
<emphasis id="BE001E6AFD540E3187853C90231460E1" italics="true" pageId="0" pageNumber="1">Xylocopa</emphasis>
</taxonomicName>
and other smaller bees).
</paragraph>
</subSubSection>
<subSubSection id="030C06692B34F8A275FD42D1F20E8D12" pageId="0" pageNumber="1" type="etymology">
<paragraph id="44C7F4D22CE670D4E771CF1FFE836B71" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="12F75C4A16C7E00FD49E4E4D1B77783C" pageId="0" pageNumber="1">
From 
<taxonomicName id="9A2472A4E28A713F85B89E1EE0859544" genus="Leucaino" lsidName="Leucaino" pageId="0" pageNumber="1" rank="genus">
<emphasis id="82DF0F13708F46FBE58955A2F55C64DA" italics="true" pageId="0" pageNumber="1">Leucaino</emphasis>
</taxonomicName>
(Latin = becoming white), probably referring to the predominantly white stamen filaments.
</paragraph>
</subSubSection>
<subSubSection id="548A6972E9F8513E35F5814EFFCC1408" pageId="0" pageNumber="1" type="human uses">
<paragraph id="554A1174EDF9328F0596E9EA08CF68B4" pageId="0" pageNumber="1">Human uses.</paragraph>
<paragraph id="80C8F8491D430F26B18E5BDD5EC69543" pageId="0" pageNumber="1">
Unripe seeds of 13 species are used as 
<taxonomicName id="40F4F6D8C5BD6EDF3682A98344ED8E59" class="Magnoliopsida" family="Fabaceae" genus="Hoffmannseggia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Hoffmannseggia minor" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="minor">Hoffmannseggia minor</taxonomicName>
foods in Mexico and are widely cultivated and incipiently domesticated in backyards in south-central Mexico (
<bibRefCitation id="99778788059DD9AF2012BE942C12D08C" author="Hughes, CE" journalOrPublisher="University of Oxford, United Kingdom" pageId="0" pageNumber="1" refId="B382" refString="Hughes, CE, 1998b. Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37. University of Oxford, United Kingdom" title="Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37." year="1998 b">Hughes 1998b</bibRefCitation>
), spawning spontaneous interspecific hybrids (
<bibRefCitation id="422A36D57C0FA08357AAE0BD0AA3B6E2" DOI="https://doi.org/10.1073/pnas.0702193104" author="Hughes, CE" journalOrPublisher="Proceedings of the National Academy of Sciences of the United States of America" pageId="0" pageNumber="1" pagination="14389 - 14394" refId="B387" refString="Hughes, CE, Govindarajulu, R, Robertson, A, Filer, DL, Harris, SA, Bailey, CD, 2007. Serendipitous backyard hybridization and the origin of crops. Proceedings of the National Academy of Sciences of the United States of America 104 (36): 14389 - 14394, DOI: https://doi.org/10.1073/pnas.0702193104" title="Serendipitous backyard hybridization and the origin of crops." url="https://doi.org/10.1073/pnas.0702193104" volume="104" year="2007">Hughes et al. 2007</bibRefCitation>
). One species, 
<taxonomicName id="95431B3DCE50CD6BADFC5F23F0141716" lsidName="L. leucocephala" pageId="0" pageNumber="1" rank="species" species="leucocephala">
<emphasis id="9FFCB48535A13C118DC9511D6C47B26D" italics="true" pageId="0" pageNumber="1">L. leucocephala</emphasis>
</taxonomicName>
(Lam.) de Wit, cultivated pantropically, naturalised and weedy in many places. Used in tropical agroforestry especially for livestock fodder (dubbed the alfalfa of the tropics), green manure, poles, firewood and soil conservation (
<bibRefCitation id="F4AE10782D8FB0D63795EC710264FF9A" author="Hughes, CE" journalOrPublisher="University of Oxford, United Kingdom" pageId="0" pageNumber="1" refId="B382" refString="Hughes, CE, 1998b. Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37. University of Oxford, United Kingdom" title="Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37." year="1998 b">Hughes 1998b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1E9DA35FFA9C8708149A5008ED0F2A3C" pageId="0" pageNumber="1" type="notes">
<paragraph id="E0AA802B106BFF3C38B4912A5B62DCFB" pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph id="8CB371D8F4B3C19083853DF3F158BAC7" pageId="0" pageNumber="1">
The 
<taxonomicName id="A6E49C93749FCF0C3748B050045E6D91" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Leucaena</taxonomicName>
subclade (
<taxonomicName id="BE4F6929ADFBA7CA800EBE366A80549B" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="B824EAEA33083FFDDD61DF0D12C5FC5C" italics="true" pageId="0" pageNumber="1">Leucaena</emphasis>
</taxonomicName>
, 
<taxonomicName id="AB687F3E918C21C07AEDBB6FBD25C6A2" class="Magnoliopsida" family="Fabaceae" genus="Schleinitzia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Schleinitzia" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="7F75602F007174F0BD3517F56D4F374C" italics="true" pageId="0" pageNumber="1">Schleinitzia</emphasis>
</taxonomicName>
, 
<taxonomicName id="03F38CB6318CC3A823A67A24B6172DBB" class="Magnoliopsida" family="Leguminosae" genus="Mezcala" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Mezcala" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="9316B045A60B1EFB0FE9F1234C08EC55" italics="true" pageId="0" pageNumber="1">Mezcala</emphasis>
</taxonomicName>
, 
<taxonomicName id="D0113F1262D4B297458C277C2771A47D" class="Magnoliopsida" family="Leguminosae" genus="Kanaloa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Kanaloa" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="F7B6C66AC7C8E189E6BBE6DE9132618E" italics="true" pageId="0" pageNumber="1">Kanaloa</emphasis>
</taxonomicName>
plus 
<taxonomicName id="75372077653FF8F8CEBFFE925DAA7230" class="Magnoliopsida" family="Leguminosae" genus="Desmanthus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Desmanthus" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="7F04E0DDFF56342E032A20BF3E5C01A8" italics="true" pageId="0" pageNumber="1">Desmanthus</emphasis>
</taxonomicName>
) is equivalent to the informal 
<taxonomicName id="DAA6BFBB5DB30CC65C0CB739C117E1F9" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Leucaena</taxonomicName>
group of 
<bibRefCitation id="4501C32A3BC01EB344A58CE62570CF49" author="Hughes, CE" editor="Klitgaard, B" journalOrPublisher="Royal Botanic Gardens, Kew, United Kingdom" pageId="0" pageNumber="1" pagination="221 - 238" refId="B386" refString="Hughes, CE, Bailey, CD, Krosnick, S, Luckow, MA, 2003. Relationships among genera of the informal Dichrostachys and Leucaena groups (Mimosoideae) inferred from nuclear ribosomal ITS sequences. In: Klitgaard, B, Bruneau, A, Eds., Advances in Legume Systematics, Part 10, Higher level systematics. Royal Botanic Gardens, Kew, United Kingdom: 221 - 238" title="Relationships among genera of the informal Dichrostachys and Leucaena groups (Mimosoideae) inferred from nuclear ribosomal ITS sequences." volumeTitle="Advances in Legume Systematics, Part 10, Higher level systematics." year="2003">Hughes et al. (2003)</bibRefCitation>
. 
<taxonomicName id="CB0348142197D93583D9759E657F4BF9" class="Magnoliopsida" family="Asteraceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena" order="Fabales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis id="428A5B638647135F57BEDF8565028686" italics="true" pageId="0" pageNumber="1">Leucaena</emphasis>
</taxonomicName>
can be distinguished from the other genera within this subclade by its hairy anthers which are present in all but three species and which are unique within 
<taxonomicName id="4AD7C3918B6C9AF60E96BC87D4ECEB9D" family="Asteraceae" kingdom="Plantae" lsidName="" order="Fabales" pageId="0" pageNumber="1" rank="tribe" tribe="Mimoseae">Mimoseae</taxonomicName>
. The genus is the focus of various genome sequencing initiatives (
<bibRefCitation id="5CB8CBED35902D834AFA1ED09A3EC423" DOI="https://doi.org/10.1038/srep16958" author="Dugas, DV" journalOrPublisher="Scientific Reports" pageId="0" pageNumber="1" pagination="1 - 13" refId="B241" refString="Dugas, DV, Hernandez, D, Koenen, EJ, Schwarz, E, Straub, S, Hughes, CE, Jansen, RK, Nageswara-Rao, M, Staats, M, Trujillo, JT, Hajrah, NH, Alharbi, NS, Al-Malki, AL, Sabir, JSM, Bailey, CD, 2015. Mimosoid legume plastome evolution: IR expansion, tandem repeat expansions and accelerated rate of evolution in clpP. Scientific Reports 5 (1): 1 - 13, DOI: https://doi.org/10.1038/srep16958" title="Mimosoid legume plastome evolution: IR expansion, tandem repeat expansions and accelerated rate of evolution in clpP." url="https://doi.org/10.1038/srep16958" volume="5" year="2015">Dugas et al. 2015</bibRefCitation>
; 
<bibRefCitation id="A103E29A61B2DE8C78051174D64324C3" DOI="https://doi.org/10.1093/gbe/evy179" author="Kovar, L" journalOrPublisher="Genome Biology and Evolution" pageId="0" pageNumber="1" pagination="2501 - 2517" refId="B445" refString="Kovar, L, Nageswara-Rao, M, Ortega-Rodriguez, S, Dugas, DV, Straub, S, Cronn, R, Strickler, SR, Hughes, CE, Hanley, KA, Rodriguez, DN, Langhorst, BW, Dimalanta, DT, Bailey, CD, 2018. PacBio-based mitochondrial genome assembly of Leucaena trichandra (Leguminosae) and an intrageneric assessment of mitochondrial RNA editing. Genome Biology and Evolution 10 (9): 2501 - 2517, DOI: https://doi.org/10.1093/gbe/evy179" title="PacBio-based mitochondrial genome assembly of Leucaena trichandra (Leguminosae) and an intrageneric assessment of mitochondrial RNA editing." url="https://doi.org/10.1093/gbe/evy179" volume="10" year="2018">Kovar et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="750C4E700124D90CBA883741DCBD17B9" pageId="0" pageNumber="1" type="taxonomic references">
<paragraph id="F36D8DD989902B920984F8FDBD1C19EE" pageId="0" pageNumber="1">Taxonomic references.</paragraph>
<paragraph id="FAEE1294A4591E24EC19296B631E80F4" pageId="0" pageNumber="1">
<bibRefCitation id="760314BFFC70896CB48730789E423273" DOI="https://doi.org/10.3732/ajb.1100259" author="Govindarajulu, R" journalOrPublisher="American Journal of Botany" pageId="0" pageNumber="1" pagination="2049 - 2063" refId="B319" refString="Govindarajulu, R, Hughes, CE, Bailey, CD, 2011b. Phylogenetic and population genetic assessment of diploid Leucaena reveal cryptic species diversity and patterns of allopatric speciation. American Journal of Botany 98 (12): 2049 - 2063, DOI: https://doi.org/10.3732/ajb.1100259" title="Phylogenetic and population genetic assessment of diploid Leucaena reveal cryptic species diversity and patterns of allopatric speciation." url="https://doi.org/10.3732/ajb.1100259" volume="98" year="2011 b">Govindarajulu et al. (2011b)</bibRefCitation>
; 
<bibRefCitation id="582DAB99334A9E820A87BD81800EBA41" DOI="https://doi.org/10.2307/25027876" author="Hughes, CE" journalOrPublisher="Systematic Botany Monographs" pageId="0" pageNumber="1" pagination="1 - 244" refId="B381" refString="Hughes, CE, 1998a. Monograph of Leucaena (Leguminosae - Mimosoideae). Systematic Botany Monographs 55: 1 - 244, DOI: https://doi.org/10.2307/25027876" title="Monograph of Leucaena (Leguminosae - Mimosoideae)." url="https://doi.org/10.2307/25027876" volume="55" year="1998 a">Hughes (1998a</bibRefCitation>
, 
<bibRefCitation id="8E03BC4D82B02B94E258F534F88C1C8C" author="Hughes, CE" journalOrPublisher="University of Oxford, United Kingdom" pageId="0" pageNumber="1" refId="B382" refString="Hughes, CE, 1998b. Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37. University of Oxford, United Kingdom" title="Leucaena. A Genetic Resources Handbook. Tropical Forest Paper 37." year="1998 b">b</bibRefCitation>
), with illustrations; 
<bibRefCitation id="47BF397CCE68FDCA9CFC5DCF69855990" DOI="https://doi.org/10.1073/pnas.0702193104" author="Hughes, CE" journalOrPublisher="Proceedings of the National Academy of Sciences of the United States of America" pageId="0" pageNumber="1" pagination="14389 - 14394" refId="B387" refString="Hughes, CE, Govindarajulu, R, Robertson, A, Filer, DL, Harris, SA, Bailey, CD, 2007. Serendipitous backyard hybridization and the origin of crops. Proceedings of the National Academy of Sciences of the United States of America 104 (36): 14389 - 14394, DOI: https://doi.org/10.1073/pnas.0702193104" title="Serendipitous backyard hybridization and the origin of crops." url="https://doi.org/10.1073/pnas.0702193104" volume="104" year="2007">Hughes et al. (2007)</bibRefCitation>
; 
<bibRefCitation id="598220528DE1DD285573A2BFA26BF8BA" author="Zarate, S" journalOrPublisher="Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Botanica" pageId="0" pageNumber="1" pagination="83 - 162" refId="B928" refString="Zarate, S, 1994. Revision del genero Leucaena Benth. en Mexico. Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Botanica 65: 83 - 162" title="Revision del genero Leucaena Benth. en Mexico." volume="65" year="1994">
<normalizedToken id="507B65E58EF8899AD0E2652B4152A25E" originalValue="Zárate">Zarate</normalizedToken>
(1994)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>