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<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e62332" ID-GBIF-Dataset="c6cd0951-6d64-4b2f-b920-bb7ba439eb39" ID-GBIF-Taxon="189469055" ID-Pensoft-Pub="1864-8312-79-509" ID-Pensoft-UUID="F1742697BB0C564B9BA5036415F063D4" ID-ZooBank="951E28C6EAC0494EBC255DEF1991583B" LSID="urn:lsid:zoobank.org:act:8B7D21DD-872B-4C09-8776-7B7FAD1CBE57" ModsDocID="1864-8312-79-509" checkinTime="1633953041321" checkinUser="pensoft" docAuthor="Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W. &amp; Harms, Danilo" docDate="2021" docId="7247F8607839543890D1AA25132F08D1" docLanguage="en" docName="ArthropodSystPhyl 79: 509-533" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 79" docPubDate="2021-10-11" docSource="http://dx.doi.org/10.3897/asp.79.e62332" docTitle="Missulena davidi Greenberg &amp; Huey &amp; Framenau &amp; Harms 2021, sp. nov." docType="treatment" docUuid="8B7D21DD-872B-4C09-8776-7B7FAD1CBE57" docUuidSource="ZooBank" docVersion="3" id="F1742697BB0C564B9BA5036415F063D4" lastPageNumber="509" masterDocId="F1742697BB0C564B9BA5036415F063D4" masterDocTitle="Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome" masterLastPageNumber="533" masterPageNumber="509" pageId="0" pageNumber="509" updateTime="1635284731334" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Greenberg, Marleen R.</mods:namePart>
<mods:affiliation>Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; Fachbereich Biologie, Universitaet Hamburg, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">marleen.greenberg@outlook.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huey, Joel A.</mods:namePart>
<mods:affiliation>Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South Street, Murdoch 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia &amp; Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Harms, Danilo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
<mods:affiliation>Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-10-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>79</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>509</mods:start>
<mods:end>533</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.79.e62332</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e62332</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-79-509</mods:identifier>
<mods:identifier type="ZooBank">951E28C6EAC0494EBC255DEF1991583B</mods:identifier>
<mods:identifier type="Pensoft-UUID">F1742697BB0C564B9BA5036415F063D4</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="509" type="3. taxonomy">
<treatment ID-GBIF-Taxon="189469055" LSID="urn:lsid:zoobank.org:act:8B7D21DD-872B-4C09-8776-7B7FAD1CBE57" httpUri="http://treatment.plazi.org/id/7247F8607839543890D1AA25132F08D1" lastPageNumber="509" pageId="0" pageNumber="509">
<subSubSection pageId="0" pageNumber="509" type="nomenclature">
<paragraph pageId="0" pageNumber="509">
3.1.
<taxonomicName LSID="http://zoobank.org/8B7D21DD-872B-4C09-8776-7B7FAD1CBE57" authority="Greenberg &amp; Huey &amp; Framenau &amp; Harms, 2021" authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi" status="sp. nov.">Missulena davidi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="509">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="description">
<paragraph pageId="0" pageNumber="509">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">, 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">, 5</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Missulena davidi sp. nov. Variability of spermatheca in females: A allotype specimen WAMT 107393, clade I; B specimen WAMT 119995, clade IV; C specimen WAMT 126272, clade II; D specimen WAMT 122226, clade III. Systematic drawings based on allotype WAMT 107393: E spermatheca; F pattern of cheliceral teeth in the cheliceral groove. Scale bars: A-E 0.5 mm; F 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure6" httpUri="https://binary.pensoft.net/fig/597112" pageId="0" pageNumber="509">, 6</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of the Missulena davidi sp. nov. in the Pilbara in Western Australia, marked with colour according to clades of the phylogenetic tree (Fig. 2)." figureDoi="10.3897/asp.79.e62332.figure7" httpUri="https://binary.pensoft.net/fig/597113" pageId="0" pageNumber="509">, 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="type material">
<paragraph pageId="0" pageNumber="509">Type material.</paragraph>
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Holotype</emphasis>
: AUSTRALIA - Western Australia • ♂; Juna Downs Station, 113 km NW of Newman;
<geoCoordinate degrees="22" direction="south" minutes="41.23" orientation="latitude" precision="9" value="-22.687166">22°41.23S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="53.55" orientation="longitude" precision="9" value="118.8925">118°53.55E</geoCoordinate>
; 10 May 2011; C. Cole and P. Runham leg.; pit trap; WAM T119725 •
<emphasis bold="true" pageId="0" pageNumber="509">Allotype</emphasis>
: AUSTRALIA - Western Australia • ♀; Hope Downs, 73.7 km NW of Newman;
<geoCoordinate degrees="20" direction="south" minutes="59.42" orientation="latitude" precision="9" value="-20.990334">20°59.42S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="7.3" orientation="longitude" precision="92" value="119.121666">119°7.3E</geoCoordinate>
; 27 June 2010; G. Humphreys and P. Runham leg.; dug from burrow; WAM T107393 •
<emphasis bold="true" pageId="0" pageNumber="509">Paratypes</emphasis>
: AUSTRALIA - Western Australia • 1♀; same data as holotype;
<geoCoordinate degrees="22" direction="south" minutes="41.18" orientation="latitude" precision="9" value="-22.686333">22°41.18S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="53.58" orientation="longitude" precision="9" value="118.893">118°53.58E</geoCoordinate>
; dug from burrow; WAM T119711 • 1♀; Juna Downs Station, 114 km NW of Newman;
<geoCoordinate degrees="22" direction="south" minutes="43.55" orientation="latitude" precision="9" value="-22.725834">22°43.55S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="50.98" orientation="longitude" precision="9" value="118.84967">118°50.98E</geoCoordinate>
; 12 May 2011; P. Runham leg.; dug from burrow; WAM T119718 • 1♂; Juna Downs Station, 116 km NW of Newman;
<geoCoordinate degrees="22" direction="south" minutes="38.73" orientation="latitude" precision="9" value="-22.6455">22°38.73S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="54.17" orientation="longitude" precision="9" value="118.90283">118°54.17E</geoCoordinate>
; 11 May 2011; C. Cole and P. Runham leg.; pit trap; WAM T119726 • 1♂; same data as for preceding; WAM T119727 • 1♂; same data as for preceding; 10 May 2011; WAM T119728 • 1♂; same data as for preceding; WAM T119729 • 1♂; same data as for preceding; 117 km NW of Newman;
<geoCoordinate degrees="22" direction="south" minutes="37.73" orientation="latitude" precision="9" value="-22.628834">22°37.73S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="54.1" orientation="longitude" precision="92" value="118.901665">118°54.1E</geoCoordinate>
; WAM T119731 • 1♂; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="36.63" orientation="latitude" precision="9" value="-22.6105">22°36.63S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="56.38" orientation="longitude" precision="9" value="118.93967">118°56.38E</geoCoordinate>
; G. Humphreys and J. Tatler leg.; WAM T119733.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="materials_examined">
<paragraph pageId="0" pageNumber="509">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="509">
<materialsCitation ID-GBIF-Occurrence="3394036311" accessionNumber="T46798" collectingDate="2002-07-23" country="AUSTRALIA" latitude="-24.883333" location="Gascoyne Road" longLatPrecision="1249" longitude="113.65" municipality="Carnarvon" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<collectingCountry name="Australia">AUSTRALIA</collectingCountry>
-
<collectingRegion country="Australia" name="Western Australia">Western Australia</collectingRegion>
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Carnarvon</collectingMunicipality>
, 99
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:37D03DED3F82E15906DFD3E55B37B075" country="AUSTRALIA" latitude="-24.883333" longLatPrecision="1249" longitude="113.65" municipality="Carnarvon" name="Gascoyne Road" stateProvince="Western Australia">Gascoyne Road</location>
;
<geoCoordinate degrees="24" direction="south" minutes="53" orientation="latitude" precision="925" value="-24.883333">24°53S</geoCoordinate>
<geoCoordinate degrees="113" direction="east" minutes="39" orientation="longitude" precision="925" value="113.65">113°39E</geoCoordinate>
;
<collectingDate value="2002-07-23">23 July 2002</collectingDate>
; residents leg.; by hand; WAM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T46798">T46798</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036351" accessionNumber="T84005" collectingDate="2006-09-06" collectorName="S. Thompson" country="Australia" latitude="-22.335" location="Fortescue Metals Group" longLatPrecision="92" longitude="119.40383" municipality="Cloudbreak Mining Lease" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount type="male">1♂</specimenCount>
;
<collectingMunicipality>Cloudbreak Mining Lease</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:19A72F6C5297FE2B633BEFEAEEB3B8F3" country="Australia" latitude="-22.335" longLatPrecision="92" longitude="119.40383" municipality="Cloudbreak Mining Lease" name="Fortescue Metals Group" stateProvince="Western Australia">Fortescue Metals Group</location>
(site 25);
<geoCoordinate degrees="22" direction="south" minutes="20.1" orientation="latitude" precision="92" value="-22.335">22°20.1S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="24.23" orientation="longitude" precision="9" value="119.40383">119°24.23E</geoCoordinate>
;
<collectingDate value="2006-09-06">6 Sept. 2006</collectingDate>
;
<collectorName>S. Thompson</collectorName>
leg.; WAM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T84005">T84005</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036322" accessionNumber="T95397" collectingDate="2009-02-06" collectorName="P. Bolton, C. Weston" country="Australia" latitude="-23.375" location="Jimblebar" longLatPrecision="92" longitude="120.20966" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:9EB12D2C3A879B44B80B294C7F894954" country="Australia" latitude="-23.375" longLatPrecision="92" longitude="120.20966" name="Jimblebar" stateProvince="Western Australia">Jimblebar</location>
minesite,
<locationDeviation location="Newman">
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.5" unit="km" value="35.0">35 km</quantity>
E of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="22.5" orientation="latitude" precision="92" value="-23.375">23°22.5S</geoCoordinate>
<geoCoordinate degrees="120" direction="east" minutes="12.58" orientation="longitude" precision="9" value="120.20966">120°12.58E</geoCoordinate>
;
<collectingDate value="2009-02-06">6 Feb. 2009</collectingDate>
;
<collectorName>P. Bolton</collectorName>
and
<collectorName>C. Weston</collectorName>
leg.; active search; WAM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T95397">T95397</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036328" accessionNumber="T97637" collectingDate="2009-04-19" collectingMethod="dry pitfall trap" collectorName="N. Dight, L. Quinn" country="Australia" county="Murray Hills" latitude="-22.127832" location="Ecologia" longLatPrecision="12" longitude="118.515335" municipality="Mulga Downs Station" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<collectingCounty>Murray Hills</collectingCounty>
,
<collectingMunicipality>Mulga Downs Station</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:229588EBBFF55CC26C41CF7981772856" country="Australia" county="Murray Hills" latitude="-22.127832" longLatPrecision="12" longitude="118.515335" municipality="Mulga Downs Station" name="Ecologia" stateProvince="Western Australia">Ecologia</location>
project 1142;
<geoCoordinate degrees="22" direction="south" minutes="07.67" orientation="latitude" precision="9" value="-22.127832">22°07.67S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="30.92" orientation="longitude" precision="9" value="118.515335">118°30.92E</geoCoordinate>
;
<collectingDate value="2009-04-19">19 Apr. 2009</collectingDate>
;
<collectorName>N. Dight</collectorName>
and
<collectorName>L. Quinn</collectorName>
leg.;
<collectingMethod>dry pitfall trap</collectingMethod>
; WAM
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/T97637">T97637</accessionNumber>
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036313" collectingDate="2010-04-09" collectingMethod="pitfall" collectorName="J. Clark" country="Australia" latitude="-23.428833" location="Davidson Creek" longLatPrecision="84" longitude="120.44666" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:57E5DCC34BAF9F0387D516FFED8DCFD1" country="Australia" latitude="-23.428833" longLatPrecision="84" longitude="120.44666" name="Davidson Creek" stateProvince="Western Australia">Davidson Creek</location>
, ca.
<locationDeviation location="Newman">
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.5" unit="km" value="75.0">75 km</quantity>
E of Newman
</locationDeviation>
, vert site 6;
<geoCoordinate degrees="23" direction="south" minutes="25.73" orientation="latitude" precision="9" value="-23.428833">23°25.73S</geoCoordinate>
<geoCoordinate degrees="120" direction="east" minutes="26.8" orientation="longitude" precision="92" value="120.44666">120°26.8E</geoCoordinate>
;
<collectingDate value="2010-04-09">9 Apr. 2010</collectingDate>
;
<collectorName>J. Clark</collectorName>
leg.; dry
<collectingMethod>pitfall</collectingMethod>
; WAM T102165
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036323" collectingDate="2011-04-16" collectorName="R. Teale, M. Greenham" country="Australia" latitude="-23.0855" location="South Parmelia" longLatPrecision="12" longitude="119.318" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:61EA03B943472ABA76CC8357D10B28D2" country="Australia" latitude="-23.0855" longLatPrecision="12" longitude="119.318" name="South Parmelia" stateProvince="Western Australia">South Parmelia</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="4" metricUnit="m" metricValue="5.2" unit="km" value="52.0">52 km</quantity>
NW of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="5.13" orientation="latitude" precision="9" value="-23.0855">23°5.13S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="19.08" orientation="longitude" precision="9" value="119.318">119°19.08E</geoCoordinate>
;
<collectingDate value="2011-04-16">16 Apr. 2011</collectingDate>
;
<collectorName>R. Teale</collectorName>
and
<collectorName>M. Greenham</collectorName>
leg.; dug from burrow; WAM T113591
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036342" collectingDate="2011-04-14" collectorName="R. Teale, M. Greenham" country="Australia" latitude="-23.002832" location="Southern Flank" longLatPrecision="12" longitude="119.1395" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount type="male">1♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:E60158AF0E0656E11735FEA935B0139D" country="Australia" latitude="-23.002832" longLatPrecision="12" longitude="119.1395" name="Southern Flank" stateProvince="Western Australia">Southern Flank</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.2" unit="km" value="72.0">72 km</quantity>
NW of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="0.17" orientation="latitude" precision="9" value="-23.002832">23°0.17S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="8.37" orientation="longitude" precision="9" value="119.1395">119°8.37E</geoCoordinate>
;
<collectingDate value="2011-04-14">14 Apr. 2011</collectingDate>
;
<collectorName>R. Teale</collectorName>
and
<collectorName>M. Greenham</collectorName>
leg.; dug from burrow; WAM T113596
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036330" collectingDate="2011-04-14" collectorName="R. Teale, M. Greenham" country="Australia" latitude="-23.003" location="Southern Flank" longLatPrecision="12" longitude="119.13917" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount type="male">1♂</specimenCount>
; same locality;
<geoCoordinate degrees="23" direction="south" minutes="0.18" orientation="latitude" precision="9" value="-23.003">23°0.18S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="8.35" orientation="longitude" precision="9" value="119.13917">119°8.35E</geoCoordinate>
,
<collectingDate value="2011-04-14">14 Apr. 2011</collectingDate>
;
<collectorName>R. Teale</collectorName>
and
<collectorName>M. Greenham</collectorName>
leg.; dug from burrow; WAM T113598
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036312" collectingDate="2011-05-26" collectorName="M. Greenham, R. Teale" country="Australia" latitude="-22.6565" location="Newman" longLatPrecision="12" longitude="118.91817" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.138" unit="km" value="113.8">113.8 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:7457DFB6B81AF6AD7ABE656C65D740ED" country="Australia" latitude="-22.6565" longLatPrecision="12" longitude="118.91817" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="39.39" orientation="latitude" precision="9" value="-22.6565">22°39.39S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="55.09" orientation="longitude" precision="9" value="118.91817">118°55.09E</geoCoordinate>
;
<collectingDate value="2011-05-26">26 May 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>R. Teale</collectorName>
leg.; dug from burrow; WAM T113626
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036338" collectingDate="2011-05-30" collectorName="M. Greenham, R. Teale" country="Australia" latitude="-22.633667" location="Newman" longLatPrecision="12" longitude="118.869835" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.191" unit="km" value="119.1">119.1 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:E4EE66143230B26F3C1EECFB2E553B54" country="Australia" latitude="-22.633667" longLatPrecision="12" longitude="118.869835" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="38.02" orientation="latitude" precision="9" value="-22.633667">22°38.02S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="52.19" orientation="longitude" precision="9" value="118.869835">118°52.19E</geoCoordinate>
;
<collectingDate value="2011-05-30">30 May 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>R. Teale</collectorName>
leg.; dug from burrow; WAM T113660
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036353" collectingDate="2011-05-31" collectorName="M. Greenham, R. Teale" country="Australia" latitude="-22.627666" location="Newman" longLatPrecision="12" longitude="118.896" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.176" unit="km" value="117.6">117.6 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:1428A2E998699C8F343CE6C698E59D39" country="Australia" latitude="-22.627666" longLatPrecision="12" longitude="118.896" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="37.66" orientation="latitude" precision="9" value="-22.627666">22°37.66S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="53.76" orientation="longitude" precision="9" value="118.896">118°53.76E</geoCoordinate>
;
<collectingDate value="2011-05-31">31 May 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>R. Teale</collectorName>
leg.; dug from burrow; WAM T113665
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036337" collectingDate="2011-06-30" collectorName="C. Cole, N. Watson" country="Australia" latitude="-23.093834" location="Mudlark" longLatPrecision="12" longitude="118.7195" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:F4F45F19B841EA61CE2336CE4F87F289" country="Australia" latitude="-23.093834" longLatPrecision="12" longitude="118.7195" name="Mudlark" stateProvince="Western Australia">Mudlark</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.0699999999999998" unit="km" value="107.0">107 km</quantity>
W of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="5.63" orientation="latitude" precision="9" value="-23.093834">23°5.63S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="43.17" orientation="longitude" precision="9" value="118.7195">118°43.17E</geoCoordinate>
;
<collectingDate value="2011-06-30">30 June 2011</collectingDate>
;
<collectorName>C. Cole</collectorName>
and
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T116746
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036304" collectingDate="2011-06-30" collectorName="M. Greenham, J. Cairnes" country="Australia" latitude="-23.086666" location="Mudlark" longLatPrecision="92" longitude="118.68633" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:EEC340CA1ECCDBD14A151AA289FCB6F8" country="Australia" latitude="-23.086666" longLatPrecision="92" longitude="118.68633" name="Mudlark" stateProvince="Western Australia">Mudlark</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.1099999999999999" unit="km" value="111.0">111 km</quantity>
WNW of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="5.2" orientation="latitude" precision="92" value="-23.086666">23°5.2S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="41.18" orientation="longitude" precision="9" value="118.68633">118°41.18E</geoCoordinate>
;
<collectingDate value="2011-06-30">30 June 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>J. Cairnes</collectorName>
leg.; dug from burrow; WAM T116751
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036317" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. Greenham &amp; J. Cairnes" country="Australia" latitude="-23.087" location="Mudlark" longLatPrecision="12" longitude="118.686165" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="23" direction="south" minutes="5.22" orientation="latitude" precision="9" value="-23.087">23°5.22S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="41.17" orientation="longitude" precision="9" value="118.686165">118°41.17E</geoCoordinate>
; WAM T116755
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036335" collectingDate="2011-07-01" collectorName="M. Greenham, J. Cairnes" country="Australia" latitude="-23.038" location="Mudlark" longLatPrecision="12" longitude="118.68283" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:3107498B64ED983E1C3591FDA4203222" country="Australia" latitude="-23.038" longLatPrecision="12" longitude="118.68283" name="Mudlark" stateProvince="Western Australia">Mudlark</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.13" unit="km" value="113.0">113 km</quantity>
W of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="2.28" orientation="latitude" precision="9" value="-23.038">23°2.28S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="40.97" orientation="longitude" precision="9" value="118.68283">118°40.97E</geoCoordinate>
;
<collectingDate value="2011-07-01">1 July 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>J. Cairnes</collectorName>
leg.; dug from burrow; WAM T116776
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036343" collectingDate="2011-07-03" collectorName="C. Cole, N. Watson" country="Australia" latitude="-23.09" location="Mudlark" longLatPrecision="92" longitude="118.811165" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:CDB0509F66739956B5EDE29548DB831D" country="Australia" latitude="-23.09" longLatPrecision="92" longitude="118.811165" name="Mudlark" stateProvince="Western Australia">Mudlark</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.02" unit="km" value="102.0">102 km</quantity>
W of Newman
</locationDeviation>
;
<geoCoordinate degrees="23" direction="south" minutes="5.4" orientation="latitude" precision="92" value="-23.09">23°5.4S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="48.67" orientation="longitude" precision="9" value="118.811165">118°48.67E</geoCoordinate>
;
<collectingDate value="2011-07-03">3 July 2011</collectingDate>
;
<collectorName>C. Cole</collectorName>
and
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T116839
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036336" collectingDate="2011-07-06" collectorName="M. Greenham, J. Cairnes" country="Australia" latitude="-23.079666" location="Newman" longLatPrecision="12" longitude="118.858" municipality="Mudlark" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Mudlark</collectingMunicipality>
,
<quantity metricMagnitude="4" metricUnit="m" metricValue="9.4" unit="km" value="94.0">94 km</quantity>
W. of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:9ECB9E5E36CE745835A9F3EAB7E5A5A5" country="Australia" latitude="-23.079666" longLatPrecision="12" longitude="118.858" municipality="Mudlark" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="23" direction="south" minutes="4.78" orientation="latitude" precision="9" value="-23.079666">23°4.78S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="51.48" orientation="longitude" precision="9" value="118.858">118°51.48E</geoCoordinate>
;
<collectingDate value="2011-07-06">6 July 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>J. Cairnes</collectorName>
leg.; dug from burrow; WAM T116866
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036334" collectingDate="2011-07-26" collectorName="C. Cole, N. Watson" country="Australia" latitude="-23.0795" location="Newman" longLatPrecision="12" longitude="118.857834" municipality="Mudlark" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same locality;
<geoCoordinate degrees="23" direction="south" minutes="4.77" orientation="latitude" precision="9" value="-23.0795">23°4.77S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="51.47" orientation="longitude" precision="9" value="118.857834">118°51.47E</geoCoordinate>
;
<collectingDate value="2011-07-26">26 July 2011</collectingDate>
;
<collectorName>C. Cole</collectorName>
and
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T116868
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036327" collectingDate="2011-07-26" collectorName="D. Kamien, M. Greenham, Z. Hamilton" country="Australia" latitude="-22.675" location="Newman" longLatPrecision="92" longitude="119.34917" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.42" unit="km" value="84.2">84.2 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:95CF78F856FE66270B668EC37D6A092D" country="Australia" latitude="-22.675" longLatPrecision="92" longitude="119.34917" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="40.5" orientation="latitude" precision="92" value="-22.675">22°40.5S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="20.95" orientation="longitude" precision="9" value="119.34917">119°20.95E</geoCoordinate>
;
<collectingDate value="2011-07-26">26 July 2011</collectingDate>
;
<collectorName>D. Kamien</collectorName>
,
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; dug from burrow; WAM T116873
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036354" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.674667" location="Newman" longLatPrecision="85" longitude="119.348335" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="40.48" orientation="latitude" precision="9" value="-22.674667">22°40.48S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="20.9" orientation="longitude" precision="92" value="119.348335">119°20.9E</geoCoordinate>
; WAM T116874
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036339" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.640833" location="89.3 km NW of Newman" longLatPrecision="12" longitude="119.3205" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:C1B82D3C9609973E80C4E146148E09E7" country="Australia" latitude="-22.640833" longLatPrecision="12" longitude="119.3205" name="89.3 km NW of Newman" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.93" unit="km" value="89.3">89.3 km</quantity>
NW of Newman
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="38.45" orientation="latitude" precision="9" value="-22.640833">22°38.45S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="19.23" orientation="longitude" precision="9" value="119.3205">119°19.23E</geoCoordinate>
; WAM T116875
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036348" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.640833" location="Newman" longLatPrecision="12" longitude="119.320335" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.93" unit="km" value="89.3">89.3 km</quantity>
NW. of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:A640B2D98F934D110AD7A36E89E23CD7" country="Australia" latitude="-22.640833" longLatPrecision="12" longitude="119.320335" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="38.45" orientation="latitude" precision="9" value="-22.640833">22°38.45S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="19.22" orientation="longitude" precision="9" value="119.320335">119°19.22E</geoCoordinate>
; WAM T116881
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036349" collectingDate="2012-04-16" country="Australia" latitude="-22.227167" location="Cowra" longLatPrecision="12" longitude="119.013664" municipality="Mulga Downs Station" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Mulga Downs Station</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:5367650FD3F7A4565494F82C9E1AB876" country="Australia" latitude="-22.227167" longLatPrecision="12" longitude="119.013664" municipality="Mulga Downs Station" name="Cowra" stateProvince="Western Australia">Cowra</location>
, site 994-13;
<geoCoordinate degrees="22" direction="south" minutes="13.63" orientation="latitude" precision="9" value="-22.227167">22°13.63S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="0.82" orientation="longitude" precision="9" value="119.013664">119°0.82E</geoCoordinate>
;
<collectingDate value="2012-04-16">16 Apr. 2012</collectingDate>
; WAM T118328
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036306" collectingDate="2011-07-27" collectorName="D. Kamien, M. Greenham, Z. Hamilton" country="Australia" latitude="-22.668333" location="Newman" longLatPrecision="92" longitude="119.374664" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.48" unit="km" value="84.8">84.8 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:B024BFD986827652703C7CFFDD26B1E7" country="Australia" latitude="-22.668333" longLatPrecision="92" longitude="119.374664" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="40.1" orientation="latitude" precision="92" value="-22.668333">22°40.1S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="22.48" orientation="longitude" precision="9" value="119.374664">119°22.48E</geoCoordinate>
;
<collectingDate value="2011-07-27">27 July 2011</collectingDate>
;
<collectorName>D. Kamien</collectorName>
,
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; dug from burrow; WAM T119975
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036319" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.571667" location="99.1 km NW of Newman" longLatPrecision="92" longitude="119.285835" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:F3765455F1C65D2DFCA9BEB16842F224" country="Australia" latitude="-22.571667" longLatPrecision="92" longitude="119.285835" name="99.1 km NW of Newman" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="9.91" unit="km" value="99.1">99.1 km</quantity>
NW of Newman
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="34.3" orientation="latitude" precision="92" value="-22.571667">22°34.3S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="17.15" orientation="longitude" precision="9" value="119.285835">119°17.15E</geoCoordinate>
; WAM T119979
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036347" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.571667" location="99.1 km NW of Newman" longLatPrecision="92" longitude="119.28616" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="34.3" orientation="latitude" precision="92" value="-22.571667">22°34.3S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="17.17" orientation="longitude" precision="9" value="119.28616">119°17.17E</geoCoordinate>
; WAM T119980
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036333" collectingDate="2011-07-28" collectorName="D. Kamien, M. Greenham, Z. Hamilton" country="Australia" latitude="-22.512" location="Newman" longLatPrecision="12" longitude="119.25916" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.053" unit="km" value="105.3">105.3 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:15033C481B7E6E07801CB2A5F2FEBEAA" country="Australia" latitude="-22.512" longLatPrecision="12" longitude="119.25916" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="30.72" orientation="latitude" precision="9" value="-22.512">22°30.72S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="15.55" orientation="longitude" precision="9" value="119.25916">119°15.55E</geoCoordinate>
;
<collectingDate value="2011-07-28">28 July 2011</collectingDate>
;
<collectorName>D. Kamien</collectorName>
,
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; dug from burrow; WAM T119984
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036350" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.512" location="Newman" longLatPrecision="12" longitude="119.258835" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="30.72" orientation="latitude" precision="9" value="-22.512">22°30.72S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="15.53" orientation="longitude" precision="9" value="119.258835">119°15.53E</geoCoordinate>
; WAM T119993
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036315" collectingDate="2011-07-29" collectorName="D. Kamien, M. Greenham, Z. Hamilton" country="Australia" latitude="-22.700333" location="Newman" longLatPrecision="12" longitude="119.378" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.12" unit="km" value="81.2">81.2 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:34F9F92A282695FBBF7447C8DF4B05BE" country="Australia" latitude="-22.700333" longLatPrecision="12" longitude="119.378" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="42.02" orientation="latitude" precision="9" value="-22.700333">22°42.02S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="22.68" orientation="longitude" precision="9" value="119.378">119°22.68E</geoCoordinate>
;
<collectingDate value="2011-07-29">29 July 2011</collectingDate>
;
<collectorName>D. Kamien</collectorName>
,
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; dug from burrow; WAM T119995
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036308" collectingDate="2011-07-31" collectorName="D. Kamien, M. Greenham, Z. Hamilton" country="Australia" latitude="-22.653334" location="Newman" longLatPrecision="92" longitude="119.413666" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.52" unit="km" value="85.2">85.2 km</quantity>
NW. of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:701B071F8A41A68D1437958631582F85" country="Australia" latitude="-22.653334" longLatPrecision="92" longitude="119.413666" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="39.2" orientation="latitude" precision="92" value="-22.653334">22°39.2S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="24.82" orientation="longitude" precision="9" value="119.413666">119°24.82E</geoCoordinate>
;
<collectingDate value="2011-07-31">31 July 2011</collectingDate>
;
<collectorName>D. Kamien</collectorName>
,
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; dug from burrow; WAM T120018
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036324" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.653334" location="Newman" longLatPrecision="92" longitude="119.41383" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="39.2" orientation="latitude" precision="92" value="-22.653334">22°39.2S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="24.83" orientation="longitude" precision="9" value="119.41383">119°24.83E</geoCoordinate>
; WAM T120019
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036325" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="D. Kamien &amp; M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.653334" location="18.9 km NE of Tom Price" longLatPrecision="92" longitude="119.413666" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount type="male">1♂</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:3E72EF78D07BA69C00995C0CC926C80C" country="Australia" latitude="-22.653334" longLatPrecision="92" longitude="119.413666" name="18.9 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.89" unit="km" value="18.9">18.9 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="39.2" orientation="latitude" precision="92" value="-22.653334">22°39.2S</geoCoordinate>
<geoCoordinate degrees="119" direction="east" minutes="24.82" orientation="longitude" precision="9" value="119.413666">119°24.82E</geoCoordinate>
; WAM T120081
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036307" collectingDate="2012-02-20" collectorName="C. Cole" country="Australia" latitude="-22.328333" location="Koodaideri Corridor West" longLatPrecision="92" longitude="118.61017" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:8AD0AD8D9B05F4900A24CB6BC4C61DBF" country="Australia" latitude="-22.328333" longLatPrecision="92" longitude="118.61017" name="Koodaideri Corridor West" stateProvince="Western Australia">Koodaideri Corridor West</location>
,
<locationDeviation location="Tom Price">
<quantity metricMagnitude="4" metricUnit="m" metricValue="9.37" unit="km" value="93.7">93.7 km</quantity>
NE of Tom Price
</locationDeviation>
;
<geoCoordinate degrees="22" direction="south" minutes="19.7" orientation="latitude" precision="92" value="-22.328333">22°19.7S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="36.61" orientation="longitude" precision="9" value="118.61017">118°36.61E</geoCoordinate>
;
<collectingDate value="2012-02-20">20 Feb. 2012</collectingDate>
;
<collectorName>C. Cole</collectorName>
leg.; burrow search; WAM T122209
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036329" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.265333" location="89.4 km NE of Tom Price" longLatPrecision="85" longitude="118.52167" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:9D01D0991BDA9ED88A1B91B620ED43BA" country="Australia" latitude="-22.265333" longLatPrecision="85" longitude="118.52167" name="89.4 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.94" unit="km" value="89.4">89.4 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="15.92" orientation="latitude" precision="9" value="-22.265333">22°15.92S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="31.3" orientation="longitude" precision="92" value="118.52167">118°31.3E</geoCoordinate>
; WAM T122217
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036352" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.135334" location="71.7 km NE of Tom Price" longLatPrecision="12" longitude="118.13617" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:AC12DC6A20037A47EDB91FDAF472C386" country="Australia" latitude="-22.135334" longLatPrecision="12" longitude="118.13617" name="71.7 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.17" unit="km" value="71.7">71.7 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="8.12" orientation="latitude" precision="9" value="-22.135334">22°8.12S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="8.17" orientation="longitude" precision="9" value="118.13617">118°8.17E</geoCoordinate>
; WAM T122224
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036346" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.155167" location="70.1 km NE of Tom Price" longLatPrecision="12" longitude="118.1345" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:4A1FC69A6C158504A829E6ED277F7303" country="Australia" latitude="-22.155167" longLatPrecision="12" longitude="118.1345" name="70.1 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.01" unit="km" value="70.1">70.1 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="9.31" orientation="latitude" precision="9" value="-22.155167">22°9.31S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="8.07" orientation="longitude" precision="9" value="118.1345">118°8.07E</geoCoordinate>
; WAM T122226
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036302" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.156166" location="70.1 km NE of Tom Price" longLatPrecision="12" longitude="118.1345" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="9.37" orientation="latitude" precision="9" value="-22.156166">22°9.37S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="8.07" orientation="longitude" precision="9" value="118.1345">118°8.07E</geoCoordinate>
; leaf litter rake; WAM T122235
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036331" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.2325" location="82.1 km NE of Tom Price" longLatPrecision="12" longitude="118.414665" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:C711432FB4FB0A34912171F4B61327D8" country="Australia" latitude="-22.2325" longLatPrecision="12" longitude="118.414665" name="82.1 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="8.209999999999999" unit="km" value="82.1">82.1 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="13.95" orientation="latitude" precision="9" value="-22.2325">22°13.95S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="24.88" orientation="longitude" precision="9" value="118.414665">118°24.88E</geoCoordinate>
; burrow search; WAM T122252
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036326" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="C. Cole" country="Australia" latitude="-22.028" location="77.3 km NE of Tom Price" longLatPrecision="12" longitude="118.00367" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:4920D7CF0749A975FA8A6C9942FAECD3" country="Australia" latitude="-22.028" longLatPrecision="12" longitude="118.00367" name="77.3 km NE of Tom Price" stateProvince="Western Australia">
<quantity metricMagnitude="4" metricUnit="m" metricValue="7.7299999999999995" unit="km" value="77.3">77.3 km</quantity>
NE of Tom Price
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="1.68" orientation="latitude" precision="9" value="-22.028">22°1.68S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="0.22" orientation="longitude" precision="9" value="118.00367">118°0.22E</geoCoordinate>
; WAM T122254
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036318" collectingDate="2012-03-29" collectorName="C. Cole, N. Watson" country="Australia" latitude="-22.892" location="Newman" longLatPrecision="12" longitude="118.76483" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.116" unit="km" value="111.6">111.6 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:47E75014279CACD872AEC759FEA26370" country="Australia" latitude="-22.892" longLatPrecision="12" longitude="118.76483" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="53.52" orientation="latitude" precision="9" value="-22.892">22°53.52S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="45.89" orientation="longitude" precision="9" value="118.76483">118°45.89E</geoCoordinate>
;
<collectingDate value="2012-03-29">29 Mar. 2012</collectingDate>
;
<collectorName>C. Cole</collectorName>
and
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T122822
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036340" collectingDate="2012-03-31" collectorName="C. Cole, N. Watson" country="Australia" latitude="-22.908667" location="Newman" longLatPrecision="12" longitude="118.7175" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.154" unit="km" value="115.4">115.4 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:385C02FFF8BC5373B623BE78EE29220B" country="Australia" latitude="-22.908667" longLatPrecision="12" longitude="118.7175" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="54.52" orientation="latitude" precision="9" value="-22.908667">22°54.52S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="43.05" orientation="longitude" precision="9" value="118.7175">118°43.05E</geoCoordinate>
;
<collectingDate value="2012-03-31">31 Mar. 2012</collectingDate>
;
<collectorName>C. Cole</collectorName>
and
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T122843
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036344" collectingDate="2012-04-01" collectorName="N. Watson, P. Brooshooft" country="Australia" latitude="-22.880833" location="Newman" longLatPrecision="12" longitude="118.687" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.186" unit="km" value="118.6">118.6 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:EACD70FA32766587029A08AD5AB76A6E" country="Australia" latitude="-22.880833" longLatPrecision="12" longitude="118.687" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="52.85" orientation="latitude" precision="9" value="-22.880833">22°52.85S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="41.22" orientation="longitude" precision="9" value="118.687">118°41.22E</geoCoordinate>
;
<collectingDate value="2012-04-01">1 Apr. 2012</collectingDate>
;
<collectorName>N. Watson</collectorName>
and
<collectorName>P. Brooshooft</collectorName>
leg.; dug from burrow; WAM T122865
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036314" collectingDate="2012-04-01" collectorName="N. Watson" country="Australia" latitude="-22.8655" location="Newman" longLatPrecision="12" longitude="118.64117" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.24" unit="km" value="124.0">124 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:5E4B773860683E8422263C06616F9F99" country="Australia" latitude="-22.8655" longLatPrecision="12" longitude="118.64117" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="51.93" orientation="latitude" precision="9" value="-22.8655">22°51.93S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="38.47" orientation="longitude" precision="9" value="118.64117">118°38.47E</geoCoordinate>
;
<collectingDate value="2012-04-01">1 Apr. 2012</collectingDate>
;
<collectorName>N. Watson</collectorName>
leg.; dug from burrow; WAM T122872
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036303" collectingDate="2012-04-27" collectingMethod="pitfall" collectorName="E. S. Volschenk" country="Australia" latitude="-23.2885" location="Paraburdoo" longLatPrecision="85" longitude="118.285" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount type="male">1♂</specimenCount>
;
<quantity metricMagnitude="4" metricUnit="m" metricValue="6.35" unit="km" value="63.5">63.5 km</quantity>
ESE of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:582C2F9A727B8BF473CF81545ECB2C5D" country="Australia" latitude="-23.2885" longLatPrecision="85" longitude="118.285" name="Paraburdoo" stateProvince="Western Australia">Paraburdoo</location>
, site 1000-tur01;
<geoCoordinate degrees="23" direction="south" minutes="17.31" orientation="latitude" precision="9" value="-23.2885">23°17.31S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="17.1" orientation="longitude" precision="92" value="118.285">118°17.1E</geoCoordinate>
;
<collectingDate value="2012-04-27">27 Apr. 2012</collectingDate>
;
<collectorName>E.S. Volschenk</collectorName>
leg.; wet
<collectingMethod>pitfall</collectingMethod>
; WAM T125176
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036316" collectingDate="2012-03-28" collectorName="G. Humphreys, M. Greenham" country="Australia" latitude="-22.1285" location="Western Corridor" longLatPrecision="12" longitude="118.092834" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; Koodaideri
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:F95A78FEC78D8581EAA6AEC461143BA2" country="Australia" latitude="-22.1285" longLatPrecision="12" longitude="118.092834" name="Western Corridor" stateProvince="Western Australia">Western Corridor</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.175" unit="km" value="217.5">217.5 km</quantity>
NW of Newman
</locationDeviation>
;
<geoCoordinate degrees="22" direction="south" minutes="7.71" orientation="latitude" precision="9" value="-22.1285">22°7.71S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="5.57" orientation="longitude" precision="9" value="118.092834">118°5.57E</geoCoordinate>
;
<collectingDate value="2012-03-28">28 Mar. 2012</collectingDate>
;
<collectorName>G. Humphreys</collectorName>
and
<collectorName>M. Greenham</collectorName>
leg.; dug from burrow; WAM T125307
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036341" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="G. Humphreys &amp; M. Greenham" country="Australia" latitude="-22.1285" location="194.8 km NW of Newman" longLatPrecision="12" longitude="118.1285" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:4467194FCBD90BD797AE26E4187A24FA" country="Australia" latitude="-22.1285" longLatPrecision="12" longitude="118.1285" name="194.8 km NW of Newman" stateProvince="Western Australia">
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.948" unit="km" value="194.8">194.8 km</quantity>
NW of Newman
</location>
;
<geoCoordinate degrees="22" direction="south" minutes="7.71" orientation="latitude" precision="9" value="-22.1285">22°7.71S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="7.71" orientation="longitude" precision="9" value="118.1285">118°7.71E</geoCoordinate>
; WAM T125308
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036301" collectingDate="2012-03-29" collectorName="G. Humphreys, J. King" country="Australia" latitude="-22.135666" location="Western Corridor" longLatPrecision="12" longitude="118.107666" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; Koodaideri
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:CF919316AA3DB0AB0AEACADA5780261A" country="Australia" latitude="-22.135666" longLatPrecision="12" longitude="118.107666" name="Western Corridor" stateProvince="Western Australia">Western Corridor</location>
,
<locationDeviation location="Newman">
<quantity metricMagnitude="5" metricUnit="m" metricValue="2.1399999999999997" unit="km" value="214.0">214 km</quantity>
NW of Newman
</locationDeviation>
;
<geoCoordinate degrees="22" direction="south" minutes="8.14" orientation="latitude" precision="9" value="-22.135666">22°8.14S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="6.46" orientation="longitude" precision="9" value="118.107666">118°6.46E</geoCoordinate>
;
<collectingDate value="2012-03-29">29 Mar. 2012</collectingDate>
;
<collectorName>G. Humphreys</collectorName>
and
<collectorName>J. King</collectorName>
leg.; dug from burrow; WAM T125316
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036355" collectingDate="2011-11-19" collectorName="M. Greenham, Z. Hamilton" country="Australia" latitude="-22.605333" location="Newman" longLatPrecision="12" longitude="118.91917" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.182" unit="km" value="118.2">118.2 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:3396CC214014C823E0FC9074960E27B3" country="Australia" latitude="-22.605333" longLatPrecision="12" longitude="118.91917" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="36.32" orientation="latitude" precision="9" value="-22.605333">22°36.32S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="55.15" orientation="longitude" precision="9" value="118.91917">118°55.15E</geoCoordinate>
;
<collectingDate value="2011-11-19">19 Nov. 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; WAM T126257
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036345" collectingDate="2011-11-18" collectorName="M. Greenham, Z. Hamilton" country="Australia" latitude="-22.611166" location="Newman" longLatPrecision="12" longitude="118.907166" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.183" unit="km" value="118.3">118.3 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:E4C4A97882E8C57BAF071D768F6FAE87" country="Australia" latitude="-22.611166" longLatPrecision="12" longitude="118.907166" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="36.67" orientation="latitude" precision="9" value="-22.611166">22°36.67S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="54.43" orientation="longitude" precision="9" value="118.907166">118°54.43E</geoCoordinate>
;
<collectingDate value="2011-11-18">18 Nov. 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; WAM T126260
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036305" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.611166" location="Newman" longLatPrecision="12" longitude="118.907166" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding; WAM T126264
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036332" collectingDate="2011-11-21" collectorName="M. Greenham, Z. Hamilton" country="Australia" latitude="-22.615" location="Newman" longLatPrecision="21" longitude="118.954994" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<quantity metricMagnitude="5" metricUnit="m" metricValue="1.14" unit="km" value="114.0">114 km</quantity>
NW of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:A70DDB60C943C4DADE77AD93648FC7AA" country="Australia" latitude="-22.615" longLatPrecision="21" longitude="118.954994" name="Newman" stateProvince="Western Australia">Newman</location>
;
<geoCoordinate degrees="22" direction="south" minutes="36" orientation="latitude" precision="15" seconds="54" value="-22.615">22°3654″S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="57" orientation="longitude" precision="15" seconds="18" value="118.954994">118°5718″E</geoCoordinate>
;
<collectingDate value="2011-11-21">21 Nov. 2011</collectingDate>
;
<collectorName>M. Greenham</collectorName>
and
<collectorName>Z. Hamilton</collectorName>
leg.; WAM T126272
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036321" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. Greenham &amp; Z. Hamilton" country="Australia" latitude="-22.6145" location="Newman" longLatPrecision="85" longitude="118.955" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
; same data as for preceding;
<geoCoordinate degrees="22" direction="south" minutes="36.87" orientation="latitude" precision="9" value="-22.6145">22°36.87S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="57.3" orientation="longitude" precision="92" value="118.955">118°57.3E</geoCoordinate>
; WAM T126276
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036320" collectingDate="2015-03-15" collectorName="C. Stevenson, M. S. Harvey, M. Hillyer" country="Australia" latitude="-22.483833" location="Karijini National Park" longLatPrecision="12" longitude="118.1475" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:87CA23F2278A279755D2102B12C1EB7F" country="Australia" latitude="-22.483833" longLatPrecision="12" longitude="118.1475" name="Karijini National Park" stateProvince="Western Australia">Karijini National Park</location>
, ca.
<locationDeviation location="Hancock Gorge">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
SW of Hancock Gorge
</locationDeviation>
;
<geoCoordinate degrees="22" direction="south" minutes="29.03" orientation="latitude" precision="9" value="-22.483833">22°29.03S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="8.85" orientation="longitude" precision="9" value="118.1475">118°8.85E</geoCoordinate>
;
<collectingDate value="2015-03-15">15 Mar. 2015</collectingDate>
;
<collectorName>C. Stevenson</collectorName>
,
<collectorName>M.S. Harvey</collectorName>
and
<collectorName>M. Hillyer</collectorName>
leg.; WAM T135548
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036309" collectingDate="2015-03-17" collectorName="M. S. Harvey" country="Australia" latitude="-22.658" location="Dales Gorge" longLatPrecision="12" longitude="118.434166" municipality="Karijini National Park" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<collectingMunicipality>Karijini National Park</collectingMunicipality>
, ca.
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.5" unit="km" value="25.0">25 km</quantity>
SSW. of
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:C1C250236B1CC4E4A300534B477C9C61" country="Australia" latitude="-22.658" longLatPrecision="12" longitude="118.434166" municipality="Karijini National Park" name="Dales Gorge" stateProvince="Western Australia">Dales Gorge</location>
;
<geoCoordinate degrees="22" direction="south" minutes="39.48" orientation="latitude" precision="9" value="-22.658">22°39.48S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="26.05" orientation="longitude" precision="9" value="118.434166">118°26.05E</geoCoordinate>
;
<collectingDate value="2015-03-17">17 Mar. 2015</collectingDate>
;
<collectorName>M.S. Harvey</collectorName>
et al. leg.; WAM T135563
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3394036310" collectingDate="2015-03-15" collectorName="J. Huey" country="Australia" latitude="-22.568666" location="Karijini National Park" longLatPrecision="12" longitude="118.09967" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount type="female">1♀</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7247F8607839543890D1AA25132F08D1:330323F0093E38B97BD26266E475227D" country="Australia" latitude="-22.568666" longLatPrecision="12" longitude="118.09967" name="Karijini National Park" stateProvince="Western Australia">Karijini National Park</location>
, ca.
<locationDeviation location="Mt Bruce">
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
NW of Mt Bruce
</locationDeviation>
;
<geoCoordinate degrees="22" direction="south" minutes="34.12" orientation="latitude" precision="9" value="-22.568666">22°34.12S</geoCoordinate>
<geoCoordinate degrees="118" direction="east" minutes="5.98" orientation="longitude" precision="9" value="118.09967">118°5.98E</geoCoordinate>
;
<collectingDate value="2015-03-15">15 Mar. 2015</collectingDate>
;
<collectorName>J. Huey</collectorName>
et al. leg.; WAM T135591
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="diagnosis">
<paragraph pageId="0" pageNumber="509">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="509">
Males of
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. share the red colouration of chelicerae and pars cephalica with
<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
Harms and Harvey, 2013,
<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
</taxonomicName>
Walckenaer, 1805,
<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
O. Pickard-Cambridge, 1877,
<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov. that are morphologically most similar. They differ from
<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
by having strong, conical spines of the rastellum (thin and not conical in the former) and a longer carapace (&gt;3.00 mm;
<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
</taxonomicName>
up to 2.8 mm). They differ from
<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
</taxonomicName>
and
<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
by having spines on patellae III and IV only and not on all four legs (on patellae I and II 1 spine, respectively).
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. males have more cuspules on maxillae and labium than those of
<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov. (
<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov.: 5 at labium, 30 at maxillae;
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov.: 15-10 at labium, 35-100 at maxillae).
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. males differ from
<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
</taxonomicName>
sp. nov. by the ridge present in the cheliceral groove. Females of
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. have uniformly red chelicerae that they share with
<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
; however, the fourth leg of
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov. is the longest of all legs, whilst in
<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
the longest leg is the first. Additionally, there are no cuspules recorded on the labium or the maxillae in
<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
</taxonomicName>
females.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="description">
<paragraph pageId="0" pageNumber="509">Description.</paragraph>
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">MALE</emphasis>
(based on holotype; WAM T119725). Total length 9.8.
<emphasis bold="true" pageId="0" pageNumber="509">Colour</emphasis>
: pars cephalica and chelicerae reddish-orange (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3C</figureCitation>
); a slim, black ring surrounding the PME (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4E</figureCitation>
); pars thoracica brown with a light, metallic blue sheen (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3C</figureCitation>
); abdomen greyish with a strong, metallic blue sheen on the dorsal side, ventrally more brownish with a faint hint of purple (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3D, E</figureCitation>
); sternum orange, slightly fading into olive with 8 sigilla in similar colour (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4D</figureCitation>
); labium and maxillae orange with a dark olive spot on the base of labium (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3F</figureCitation>
); legs olive-yellowish fading into light brown ventrally, dorsally brown (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3A, B</figureCitation>
); spinnerets beige-coloured (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Carapace</emphasis>
: 3.86 long and 4.67 wide; clypeus 0.31; pars cephalica covers 2.45 of its length, is highly elevated and slightly granulated with very few setae (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3G</figureCitation>
); pars thoracica also granulated with bands of faint, radial fissures and with two notches close to the abdomen (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Eyes</emphasis>
: OQ 4 times wider than long; outer width of each eye pair AME 0.66, ALE 2.44, PME 1.48 and PLE 2.21; diameter of AME 0.19, ALE 0.22, PME 0.15, PLE 0.19; anterior eyes very slightly recurved; posterior eyes strongly recurved (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Chelicerae</emphasis>
: 2.04 long and 1.47 wide on the base; edges rounded and recurved with the widest point being 1.55 close to the chelicerae base (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3C</figureCitation>
); small, faint files along the outer margin of each chelicera; evenly spread setae along the inner margin and the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 7 (left 8) strong, conical spines (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4F</figureCitation>
); over 25 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth and a general cheliceral teeth area in between those 2 clear rows (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4A, J</figureCitation>
); prolateral row with approx. 9 teeth; retrolateral row with 4 teeth; intermediate area with 10 small teeth.
<emphasis bold="true" pageId="0" pageNumber="509">Maxillae</emphasis>
: 2.08 long and 1.44 wide; at least 80-100 weakly developed cuspules along entire anterior margin (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3F</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Labium</emphasis>
: 0.96 long and 0.86 wide on the base; conical; at least 40 weakly developed cuspules anteriorly (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3F</figureCitation>
); labiosternal junction visible (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4D</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Sternum</emphasis>
: 2.6 long and 2.38 wide; ovoid (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4D</figureCitation>
); setae of various length somewhat densely but disordered along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair smallest and hardly visible, second pair also very small and circular, third pair significantly larger than second (roughly 5 times bigger) in the shape of an elongated oval, and posterior pair biggest (roughly 1.5 times the size of the third pair) in the shape of a drop, all sigilla slightly depressed.
<emphasis bold="true" pageId="0" pageNumber="509">Abdomen</emphasis>
: 3.88 long and 3.4 wide; shape of a rounded trapezoid (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3D</figureCitation>
); 4 spinnerets, PLS 1.12 long, 0.48 wide; PMS 0.43 long, 0.23 wide.
<emphasis bold="true" pageId="0" pageNumber="509">Pedipalp</emphasis>
: length of trochanter 1.49, femur 3.68, patella 1.68, tibia 4.01, tarsus 0.72; all segments with setae, tibia ventrally covered with comparably long setae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4I</figureCitation>
); tibia rather thin and slightly recurved, 1.00 wide on the widest point from dorsal and prolateral view (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4G-I</figureCitation>
); bulb roughly pyriform (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4G-I</figureCitation>
), two strongly sclerotized sections connected by a velar median structure (
<normalizedToken originalValue="“haematodocha”">&quot;haematodocha&quot;</normalizedToken>
); embolus short with an intumescence in proximal region; tip of embolus triangular with a small lamella and a tooth best visible from prolateral view (paratype, Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3H, I</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Legs</emphasis>
: brown setae of various sizes on all sides of the legs and bent strongly towards the exterior with the exceptions of some long, dorsal setae on tibia I and IV; ventral preening comb on tarsi and metatarsi III and IV.
<emphasis bold="true" pageId="0" pageNumber="509">Leg spination</emphasis>
: leg I: tibia rv0, v5, pl0, d0; metatarsus rv3, v8, pl2, d0; tarsus rv3, v3, pl4, d0; leg II: tibia rv0, v8, pl0, d0; metatarsus rv0, v11, pl0, d0; tarsus rv3, v3, pl2, d0; leg III: tibia rv4, v8, pl0, d11; metatarsus rv4, v11, pl0, d11; tarsus rv5, v9, pl4, d5; leg IV: tibia rv0, v5, pl0, d0; metatarsus rv0, v16, pl0, d2; tarsus rv4, v10, pl3, d3; patella I with one spine prolateral close to the tibia and patella II with one spine ventrally also close to the tibia; patella III with ca. 23 spines prolateral to dorsal (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4B</figureCitation>
), 1 spine retrolateral; patella IV with one spine dorsal close to the tibia and approx. 12 small spines (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Missulena davidi sp. nov. Male holotype (WAMT 119725): A chelicerae with cheliceral groove, ventral view; B patella III, dorsal view; C patella IV, dorsal view; D sternum, ventral view; E eye region, dorsal view; F rastellum, frontal view; G right pedipalp, retrolateral view; H same, ventral view; I same, prolateral view; J pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm." figureDoi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509">4C</figureCitation>
) and ca. 18 prolateral, also very small.
<emphasis bold="true" pageId="0" pageNumber="509">Leg measurements</emphasis>
: Leg I: femur 3.72, patella 1.29, tibia 2.72, metatarsus 2.37, tarsus 1.37, total 11.47. Leg II: femur 3.31, patella 1.36, tibia 2.43, metatarsus 2.26, tarsus 1.38, total 10.74. Leg III: femur 2.72, patella 1.26, tibia 1.93, metatarsus 1.92, tarsus 1.35, total 9.18. Leg IV: femur 3.45, patella 1.47, tibia 2.76, metatarsus 2.41, tarsus 1.51, total 11.6. Formula: 4&gt;1&gt;2&gt;3.
</paragraph>
<caption doi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Figure 3.</emphasis>
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. Male holotype (WAMT119725):
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
habitus, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
same, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
carapace, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
abdomen, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
same, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
maxillae, labium, and chelicerae, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">G</emphasis>
carapace, lateral view. Male paratype (WAMT119727), left pedipalp,
<emphasis bold="true" pageId="0" pageNumber="509">H</emphasis>
embolus with embolar tooth, prolateral view;
<emphasis bold="true" pageId="0" pageNumber="509">I</emphasis>
same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100
<normalizedToken originalValue="µm">µm</normalizedToken>
; I 40
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</caption>
<caption doi="10.3897/asp.79.e62332.figure4" httpUri="https://binary.pensoft.net/fig/597110" pageId="0" pageNumber="509" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Figure 4.</emphasis>
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. Male holotype (WAMT119725):
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
chelicerae with cheliceral groove, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
patella III, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
patella IV, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
sternum, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
eye region, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
rastellum, frontal view;
<emphasis bold="true" pageId="0" pageNumber="509">G</emphasis>
right pedipalp, retrolateral view;
<emphasis bold="true" pageId="0" pageNumber="509">H</emphasis>
same, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">I</emphasis>
same, prolateral view;
<emphasis bold="true" pageId="0" pageNumber="509">J</emphasis>
pattern of cheliceral teeth in cheliceral groove. Scale bars: A, D-H 2.0 mm; B, C 0.5 mm; J 1.0 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">FEMALE</emphasis>
(based on allotype; WAM T107393). Total length 23.9.
<emphasis bold="true" pageId="0" pageNumber="509">Colour</emphasis>
: Carapace brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); chelicerae reddish-orangish with a darker spot (dark reddish-brown) on each chelicerae base (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); eye region light reddish colour similar to chelicerae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); abdomen greyish-brown with a faint, dorsal, metallic blue sheen (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5B</figureCitation>
); sternum light brown fading into a reddish-brown towards labium (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5C</figureCitation>
), sigilla darker brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5C</figureCitation>
); labium und maxillae reddish-brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5E</figureCitation>
); legs brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5F, G</figureCitation>
); spinnerets lighter brown (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5B</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Carapace</emphasis>
: 7.31 long and 9.88 wide; clypeus 0.74; pars cephalica covers 4.72 of its length, is highly elevated and smooth (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5D</figureCitation>
) with some setae going along the margin of the chelicerae as well as vertically in a line from the AME to fovea plus some random setae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); pars thoracica smooth surface with bands of faint, radial fissures (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Eyes</emphasis>
: OQ 4.9 times wider than long; width of each eye pair AME 0.91, APE 6.01, PME 3.59 and PLE 5.54; diameter of AME 0.33, ALE 0.36, PME 0.25, PLE 0.3; anterior eyes in one straight line; posterior eyes clearly recurved (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5H</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Chelicerae</emphasis>
: 5.65 long and 4.15 wide on the base; edges rounded and recurved with the widest point being 4.73 close to the chelicerae base (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); long setae along the inner margin increasing in amount towards rastellum; short and fewer setae along the outer margin and no setae in centre (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5A</figureCitation>
); rastellum present with 10-14 conical spines on each chelicera and long, densely disordered setae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5I</figureCitation>
); approx. 25 setae cover anterior base of fang; inner margin of cheliceral furrow with two main rows of cheliceral teeth and a small cheliceral teeth area in between (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Missulena davidi sp. nov. Variability of spermatheca in females: A allotype specimen WAMT 107393, clade I; B specimen WAMT 119995, clade IV; C specimen WAMT 126272, clade II; D specimen WAMT 122226, clade III. Systematic drawings based on allotype WAMT 107393: E spermatheca; F pattern of cheliceral teeth in the cheliceral groove. Scale bars: A-E 0.5 mm; F 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure6" httpUri="https://binary.pensoft.net/fig/597112" pageId="0" pageNumber="509">6F</figureCitation>
); prolateral row with 11 teeth, the first 3 teeth next to the
<normalizedToken originalValue="fangs">fang's</normalizedToken>
base are partially grown together; retrolateral row with 10 teeth; intermediate area with approx. 32 small teeth.
<emphasis bold="true" pageId="0" pageNumber="509">Maxillae</emphasis>
: 4.48 long and 3.67 wide; at least 150-170 strongly developed cuspules along entire anterior margin (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Labium</emphasis>
: 4 long and 2.27 wide on the base; conical; at least 60 cuspules (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5E</figureCitation>
); anterior pair of sigilla reach labiosternal junction; labiosternal junction clearly developed (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5C</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Sternum</emphasis>
: 6.32 long and 5.7 wide; oval (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5C</figureCitation>
); setae of various length somewhat densely but disordered along the margin and a similar amount of setae of various sizes spread evenly over the sternum; 4 pairs of sigilla, anterior pair small and hardly visible, second pair (anterior-posterior) smallest and divided in circles, third pair larger than second and roughly in the shape of an elongated oval, and posterior pair biggest (roughly 4 times the size of the third pair); all sigilla depressed, two anterior pairs just slightly, two posterior pairs strongly.
<emphasis bold="true" pageId="0" pageNumber="509">Abdomen</emphasis>
: 10.87 long and 9.12 wide; surface covered with horizontal wrinkles and dense setae (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5B</figureCitation>
); PLS 2.84 long and 1.5 wide; PMS 1.43 long and 0.6 wide.
<emphasis bold="true" pageId="0" pageNumber="509">Pedipalp</emphasis>
: Length of trochanter 1.5, femur 4.62, patella 1.79, tibia 3.23, tarsus 2.96; approx. 17 spines spread prolateral, retrolateral and ventral on tarsus.
<emphasis bold="true" pageId="0" pageNumber="509">Genitalia</emphasis>
: one pair of simple and rounded spermathecae, sperm ducts relatively short (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Missulena davidi sp. nov. Variability of spermatheca in females: A allotype specimen WAMT 107393, clade I; B specimen WAMT 119995, clade IV; C specimen WAMT 126272, clade II; D specimen WAMT 122226, clade III. Systematic drawings based on allotype WAMT 107393: E spermatheca; F pattern of cheliceral teeth in the cheliceral groove. Scale bars: A-E 0.5 mm; F 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure6" httpUri="https://binary.pensoft.net/fig/597112" pageId="0" pageNumber="509">6A-E</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Legs</emphasis>
: densely covered in brown setae of various sizes on all sides of the legs and bent towards the exterior with the exceptions of some long, dorsal setae on tibia, metatarsus and tarsus.
<emphasis bold="true" pageId="0" pageNumber="509">Leg spination</emphasis>
: leg I: tibia rv0, v0, pl0, d0; metatarsus rv1, v2, pl0, d0; tarsus rv6, v13, pl6, d0; leg II: tibia rv0, v0, pl0, d0; metatarsus rv1, v3, pl0, d0; tarsus rv6, v11, pl3, d0; leg III: tibia rv1, v0, pl2, d8; metatarsus rv0, v1, pl0, d18; tarsus rv7, v11, pl7, d6; leg IV: tibia rv0, v0, pl1, d5; metatarsus rv0, v3, pl4, d2; tarsus rv1, v15, pl9, d3; patellae I and II aspinose; patella III with ca. 28 spines prolateral to dorsal (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5F</figureCitation>
); patella IV with ca. 18 spines prolateral to dorsal (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Missulena davidi sp. nov. Female allotype (WAMT 107393): A carapace, dorsal view; B abdomen, dorsal view; C sternum, ventral view; D carapace, lateral view; E maxillae, labium and chelicerae, ventral view; F patella III, dorsal view; G patella IV, dorsal view; H eye region, dorsal view; I rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509">5G</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="509">Leg measurement</emphasis>
: leg I: femur 5.2, patella 2.16, tibia 3.4, metatarsus 2.5, tarsus 1.93, total 15.19. leg II: femur 5.32, patella 2.54, tibia 3.02, metatarsus 2.86, tarsus 2.04, total 15.78. leg III: femur 5.03, patella 2.64, tibia 2.39, metatarsus 3.08, tarsus 2.15, total 15.29. leg IV: femur 5.6, patella 2.61, tibia 3.66, metatarsus 3.31, tarsus 2.24, total 17.42. Formula: 4&gt;2&gt;3&gt;1.
</paragraph>
<caption doi="10.3897/asp.79.e62332.figure5" httpUri="https://binary.pensoft.net/fig/597111" pageId="0" pageNumber="509" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Figure 5.</emphasis>
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. Female allotype (WAMT107393):
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
carapace, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
abdomen, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
sternum, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
carapace, lateral view;
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
maxillae, labium and chelicerae, ventral view;
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
patella III, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">G</emphasis>
patella IV, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">H</emphasis>
eye region, dorsal view;
<emphasis bold="true" pageId="0" pageNumber="509">I</emphasis>
rastellum, frontal view. Scale bars: A, B 4.0 mm; C-I 2.0 mm.
</paragraph>
</caption>
<caption doi="10.3897/asp.79.e62332.figure6" httpUri="https://binary.pensoft.net/fig/597112" pageId="0" pageNumber="509" start="Figure 6" startId="F6">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Figure 6.</emphasis>
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. Variability of spermatheca in females:
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
allotype specimen WAMT107393, clade I;
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
specimen WAMT119995, clade IV;
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
specimen WAMT126272, clade II;
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
specimen WAMT122226, clade III. Systematic drawings based on allotype WAMT107393:
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
spermatheca;
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
pattern of cheliceral teeth in the cheliceral groove. Scale bars: A-E 0.5 mm; F 2.0 mm
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="etymology">
<paragraph pageId="0" pageNumber="509">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="509">
The specific epithet is a patronym in honour of the senior
<normalizedToken originalValue="authors">author's</normalizedToken>
husband, David A. Greenberg.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="distribution">
<paragraph pageId="0" pageNumber="509">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="509">
Pilbara region of Western Australia, excluding the northern Pilbara subregion, extending into the Little Sandy Desert region. The known linear range of this species is 295 km (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of the Missulena davidi sp. nov. in the Pilbara in Western Australia, marked with colour according to clades of the phylogenetic tree (Fig. 2)." figureDoi="10.3897/asp.79.e62332.figure7" httpUri="https://binary.pensoft.net/fig/597113" pageId="0" pageNumber="509">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="genetic structure">
<paragraph pageId="0" pageNumber="509">Genetic structure.</paragraph>
<paragraph pageId="0" pageNumber="509">
The species is highly structured across its range with four genetic clades that have sympatric distributions (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum Likelihood phylogeny of all Missulena specimens. The ultimate outgroup taxon Atrax has been removed from the figure for convenience. All bootstrap values below 80 have been removed. Genetic clades within Missulena davidi sp. nov. are colour-coded: clade I brown, clade II red, clade III blue, clade IV pink." figureDoi="10.3897/asp.79.e62332.figure2" httpUri="https://binary.pensoft.net/fig/597108" pageId="0" pageNumber="509">2</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution records of the Missulena davidi sp. nov. in the Pilbara in Western Australia, marked with colour according to clades of the phylogenetic tree (Fig. 2)." figureDoi="10.3897/asp.79.e62332.figure7" httpUri="https://binary.pensoft.net/fig/597113" pageId="0" pageNumber="509">7</figureCitation>
).
</paragraph>
<caption doi="10.3897/asp.79.e62332.figure7" httpUri="https://binary.pensoft.net/fig/597113" pageId="0" pageNumber="509" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Figure 7.</emphasis>
Distribution records of the
<taxonomicName authorityName="Greenberg &amp; Huey &amp; Framenau &amp; Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena davidi" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">Missulena davidi</emphasis>
</taxonomicName>
sp. nov. in the Pilbara in Western Australia, marked with colour according to clades of the phylogenetic tree (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Maximum Likelihood phylogeny of all Missulena specimens. The ultimate outgroup taxon Atrax has been removed from the figure for convenience. All bootstrap values below 80 have been removed. Genetic clades within Missulena davidi sp. nov. are colour-coded: clade I brown, clade II red, clade III blue, clade IV pink." figureDoi="10.3897/asp.79.e62332.figure2" httpUri="https://binary.pensoft.net/fig/597108" pageId="0" pageNumber="509">2</figureCitation>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="remarks">
<paragraph pageId="0" pageNumber="509">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="509">
This species had been labelled
<normalizedToken originalValue="“MYG045”">&quot;MYG045&quot;</normalizedToken>
in previous barcoding studies (
<bibRefCitation DOI="https://doi.org/10.1071/IS13058" author="Castalanelli, MA" journalOrPublisher="Invertebrate Systematics" pageId="0" pageNumber="509" pagination="375 - 385" refId="B7" refString="Castalanelli, MA, Teale, R, Rix, MG, Kennington, WJ, Harvey, MS, 2014. Barcoding of mygalomorph spiders (Araneae: Mygalomorphae) in the Pilbara bioregion of Western Australia reveals a highly diverse biota. Invertebrate Systematics 28: 375 - 385, DOI: https://doi.org/10.1071/IS13058" title="Barcoding of mygalomorph spiders (Araneae: Mygalomorphae) in the Pilbara bioregion of Western Australia reveals a highly diverse biota." url="https://doi.org/10.1071/IS13058" volume="28" year="2014">Castalanelli et al. 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="509" type="variability">
<paragraph pageId="0" pageNumber="509">Variability.</paragraph>
<paragraph pageId="0" pageNumber="509">
Assessment of 19 characters in male
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov. and 18 characters in females (Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Summary of scored characters in M. davidi sp. nov., M. manningensis sp. nov. and M. iugum sp. nov .. M = Median, n = sample size." httpUri="http://table.plazi.org/id/2CC31655A1189263F478D67DBB848526" pageId="0" pageNumber="509" tableUuid="2CC31655A1189263F478D67DBB848526">1</tableCitation>
) highlights substantial variation in many characters within and between sampling localities, in both the male and female specimens examined.
</paragraph>
<caption ID-Table-UUID="2CC31655A1189263F478D67DBB848526" httpUri="http://table.plazi.org/id/2CC31655A1189263F478D67DBB848526" pageId="0" pageNumber="509" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="509">
<emphasis bold="true" pageId="0" pageNumber="509">Table 1.</emphasis>
Summary of scored characters in
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
</taxonomicName>
sp. nov.. M = Median, n = sample size.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="509">
<table pageId="0" pageNumber="509">
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="3">
<emphasis bold="true" pageId="0" pageNumber="509">Characters</emphasis>
</td>
<td colspan="4" pageId="0" pageNumber="509" rowspan="1">
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="509">males (n = 12)</emphasis>
</td>
<td colspan="5" pageId="0" pageNumber="509" rowspan="1">
<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
</taxonomicName>
-
<emphasis bold="true" pageId="0" pageNumber="509">females (n = 26)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="3">
<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="509">(n = 1)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="3">
<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="509">(n = 2)</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="3" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">Clades</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="509">All clades combined</emphasis>
</td>
<td colspan="4" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">Clades</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="509">All clades combined</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">I (n = 4)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">II (n = 4)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">III (n = 4)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">I (n = 10)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">II (n = 7)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">III (n = 7)</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="509">IV (n = 5)</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Carapace length [mm]</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.86-4.2</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.95-4.74</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.58-4.11</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.58-4.74 M: 4.03</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5.19-8.55</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5.12-8.54</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6.75-8.59</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6.21-8.76</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5.12-8.97 M: 7.91</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.6</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">3.68-3.87</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Carapace width [mm]</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.67-4.96</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.56-5.55</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.23-4.88</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.23-5.55 M: 4.77</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6.28-10.7</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6.71-10.44</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8.39-12</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8.14-10.85</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6.28-12 M: 10.28</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.61</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4.57-4.98</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Carapace height [mm]</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.73-2.54</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.92-2.24</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.75-2.31</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.73-2.54 M: 2.16</td>
<td colspan="5" pageId="0" pageNumber="509" rowspan="1">Not measured</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.96</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">1.67-1.69</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Pars cephalica ratio to carapace (in %)</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">63.4-64.5</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">63.2-67.3</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">62.0-64.8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">62-67% M: 64%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">62.6-70.3</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">65.6-74.6</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">61.2-71.9</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">62.5-73.4</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">61-75% M: 67%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">63%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">60-62%</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">PLE ratio to ALE (in %)</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">90.5-98.2</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">89.7-95.4</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">92.4-96.0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">90-98% M: 94%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">84.3-93.0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">84.6-90.4</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">86.1-94.3</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">85.5-94.7</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">84-95% M: 90%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">95%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">87-90%</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">PME ratio to ALE (in %)</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">60.7-64.5</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">59.6-62.9</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">59.0-64.1</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">59-64% M: 62%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">57.0-67.8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">55.7-60.6</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">52.2-65.0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">56.9-65.9</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">52-68% M: 60%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">64%</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">65-69%</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Cuspules right maxillae</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">50-80</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">55-60</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">50-90</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">50-90 M: 60</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">75-200</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">130-180</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">120-230</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">130 -200</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">75-230 M: 170</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">70-100</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Cuspules left maxillae</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">45-70</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">55-80</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">35-70</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">35-80 M: 60</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">80-190</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">110-160</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">120-220</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">90-150</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">80-220 M: 150</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">70-85</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Cuspules labium</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">25-30</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">15-25</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">20-30</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">15-40 M: 25</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">35-65</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">30-55</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">35-100</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">25-65</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">25-100 M: 48</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">30-35</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Ridge in the Cheliceral groove</td>
<td colspan="4" pageId="0" pageNumber="509" rowspan="1">Not present</td>
<td colspan="5" pageId="0" pageNumber="509" rowspan="1">Not present</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Not present</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Present</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Right rastellum</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5-8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-9</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5-8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-9 M: 6</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8-15</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">11-17</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">7-18</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5-12</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5-18 M: 12</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Left rastellum</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">5-10</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">6-9</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-10 M: 8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">7-19</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">11-17</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-16</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">7-15</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">4-19 M: 12</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">8-9</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Spination 1st patella</td>
<td colspan="3" pageId="0" pageNumber="509" rowspan="4">Not individually measured</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0-1; pl: 1-7; d: 0</td>
<td colspan="4" pageId="0" pageNumber="509" rowspan="4">Not individually measured</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0; pl: 0; d: 0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 1; pl: 8; d: 0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 3-9; pl: 9-12; d: 0</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Spination 2nd patella</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 1; pl: 0; d: 0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0; pl: 0; d: 0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 1; pl: 1; d: 0</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 2; pl: 0; d: 0</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Spination 3rd patella</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 1-2; v: 0; pl/d: 23-25</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0; pl/d: 26-28</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 1; v: 0; pl/d: 26</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 4; v: 2-3; pl/d: 27</td>
</tr>
<tr pageId="0" pageNumber="509">
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">Supination 4th patella</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0-1; v: 0; pl: 16-18; d: 1 + ~12 (small)</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0; pl/d: 17-26</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 0; pl: 14 (small); d: 15 (small)</td>
<td colspan="1" pageId="0" pageNumber="509" rowspan="1">rl: 0; v: 3 (or just thickened setae); pl: 6 (small); d: 7 (small)</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="509">
In both sexes there is substantial variation in body colouration. In males, the chelicera and pars cephalica ranged from a bright red (specimen WAM T119729; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13A</figureCitation>
) to a red with a shade of orange (male: T120081; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13B</figureCitation>
) whereas abdominal colour in dorsal varied between green, purple and blue metallic tones to no metallic sheen at all (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13G-H</figureCitation>
). Colouration of the pars thoracica ranged from light brown (T119733) to nearly black (T119731) and the orange to olive transition on the sternum and coxa was also highly variable (compare T84005; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13F</figureCitation>
with T113596; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13E</figureCitation>
). In females, cheliceral colouration ranged between dark red (T116874; Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14A</figureCitation>
) to a light orange (T119979 or T116776; Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14B</figureCitation>
). The abdomen in some specimens had a metallic sheen and the sternum varied between uniformly orange (T116776; Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14F</figureCitation>
) to a full transition from orange to dark red (T119711; Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14G</figureCitation>
). Colour of legs and pars thoracica ranged between light brown (T116776) to dark brown (T125308). The colour variations still hold true if considering possible artefacts of preservation, i.e. varying trapping liquids, ethanol concentration during storage, and time of preservation.
</paragraph>
<paragraph pageId="0" pageNumber="509">
Variability was also high in rastellum spination and cuspule counts. Some male specimens had about twice as many rastellum spines (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13L, M</figureCitation>
) and/or cuspules on both maxillae and labium than others. Female variability was even higher with some specimens having up to four times more maxillary cuspules (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14H, I</figureCitation>
) or nearly five times more spines on the rastellum (17; Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14N, O</figureCitation>
) than others. The spination of the patellae was also variable which included spinal counts, size, and positioning of the individual spines. While male specimens showed variation on the first, third and fourth patella, females only had spines on the third and fourth patella (see Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Summary of scored characters in M. davidi sp. nov., M. manningensis sp. nov. and M. iugum sp. nov .. M = Median, n = sample size." httpUri="http://table.plazi.org/id/2CC31655A1189263F478D67DBB848526" pageId="0" pageNumber="509" tableUuid="2CC31655A1189263F478D67DBB848526">1</tableCitation>
for details). The shape of the sternum in females varied considerably between ovoid and round, and the shape, size and position of the sternal sigilla differed substantially (Fig.
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14F, G</figureCitation>
). The second pair of sigilla was subdivided into two depressions in some female specimens but not in others. Variation in body size was also substantial (Fig.
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Missulena davidi sp. nov., male and female variability in carapace length and width in mm; n = 24 females, 12 males." figureDoi="10.3897/asp.79.e62332.figure15" httpUri="https://binary.pensoft.net/fig/597121" pageId="0" pageNumber="509">15</figureCitation>
) and the carapace length ranged from 3.58 to 4.74 in males and 5.12 to 8.97 in females.
</paragraph>
<paragraph pageId="0" pageNumber="509">
Eye ratio of the PLE and PME pair width in relation to the ALE pair width varied less than other characters. While still showing variation in males and females its range was less extreme (within 8% in males and 16% in females) than in other characters (Figs
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Variability in Missulena davidi sp. nov. males: A carapace, dorsal view (T 119729, clade III); B carapace, dorsal view (T 120081, clade II); C carapace, dorsal view (T 119727, clade II); D carapace, dorsal view (T 119733, clade III); E sternum, ventral view (T 113596, clade I); F sternum, ventral view (T 84005, clade I); G abdomen, dorsal view (T 119727, clade II); H abdomen, dorsal view (T 125176, clade III); I eye region, dorsal view (T 120081, clade II); J eye region, dorsal view (T 84005, clade I); K eye region, dorsal view (T 119729, clade III); L rastellum, anterior view (T 84005, clade I); M rastellum, anterior view (T 119726, clade II). Scale bars: A-M 2.0 mm" figureDoi="10.3897/asp.79.e62332.figure13" httpUri="https://binary.pensoft.net/fig/597119" pageId="0" pageNumber="509">13I-K</figureCitation>
,
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Variability in Missulena davidi sp. nov. females: A carapace, dorsal view (T 116874, clade IV); B carapace, dorsal view (T 116776, clade I); C carapace, dorsal view (T 113591, clade III); D carapace, dorsal view (T 122226, clade III); E carapace, dorsal view (T 126272, clade II); F sternum, ventral view (T 116776, clade I); G sternum, ventral view (T 119711, clade I); H left maxilla, ventral view (T 119979, clade IV); I right maxillae, ventral view (T 116868, clade I); J eye region, dorsal view (T 116839, clade I); K eye region, dorsal view (T 119979, clade IV); L eye region, dorsal view (T 122865, clade I); M eye region, dorsal view (T 126264, clade III); N rastellum, anterior view (T 125316, clade III); O rastellum, anterior view (T 102165, clade I). Scale bars: A-G 4.0 mm; H-O 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure14" httpUri="https://binary.pensoft.net/fig/597120" pageId="0" pageNumber="509">14J-M</figureCitation>
; see boxplot Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Missulena davidi sp. nov., male and female variability in PLE and PME width in relation to the ALE width shown in %; n = 25 females, 12 males." figureDoi="10.3897/asp.79.e62332.figure16" httpUri="https://binary.pensoft.net/fig/597122" pageId="0" pageNumber="509">16</figureCitation>
). Male bulb structure also varied little (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Missulena davidi sp. nov. Male holotype (WAMT 119725): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E same, ventral view; F maxillae, labium, and chelicerae, ventral view; G carapace, lateral view. Male paratype (WAMT 119727), left pedipalp, H embolus with embolar tooth, prolateral view; I same, retrolateral view. Scale bars: A, B 4.0 mm; C-G 2.0 mm; H 100 µm; I 40 µm." figureDoi="10.3897/asp.79.e62332.figure3" httpUri="https://binary.pensoft.net/fig/597109" pageId="0" pageNumber="509">3 H, I</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="509">There was no correlation between geographical distance of samples and morphological divergence. For example, two females of the clade III (WAM T126260 and T126264) from one locality showed substantial differences in the number of rastellum spines (9 vs 17 respectively), which was also notable for four male specimens of the clades II and III (T119726-T119729) from the same location (4 to 10 spines). Similar results were inferred for cuspule count in these males (55 to 80 cuspules on the maxillae).</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>