treatments-xml/data/31/33/B0/3133B07BFF821055966EA0545FF1F9ED.xml

<document id="14D0417105EFCC0A110A36F49F267DE6" ID-CLB-Dataset="55540" ID-DOI="10.5281/zenodo.5741445" ID-GBIF-Dataset="00b3a6f6-2b7f-4b65-a3ba-58775fa446ff" ID-ISSN="1536-9307" ID-Zenodo-Dep="5741445" ID-ZooBank="15B1EA02-BFD2-43CB-80FD-B8B12BA6C0BC" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638264778786" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Stockmann, Mark &amp; Šťáhlavský, František" docDate="2020" docId="3133B07BFF821055966EA0545FF1F9ED" docLanguage="en" docName="Euscorpius.298.1-40.pdf" docOrigin="Euscorpius 298" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Compsobuthus ullrichi Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský 2020, sp. n." docType="treatment" docUuid="37EEF1EA-5294-4BAB-B92D-9F568946A2F4" docUuidSource="ZooBank" docVersion="9" lastPageNumber="38" masterDocId="CD0AC803FFA010729761A4635C51FFDF" masterDocTitle="Notes on Compsobuthus: redescription of C. arabicus Levy et al., 1973 from Arabia, and description of two new species from North Africa (Scorpiones: Buthidae)" masterLastPageNumber="40" masterPageNumber="1" pageNumber="33" updateTime="1699471349907" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="AEE37F4E922100728854A312414AF265">Notes on Compsobuthus: redescription of C. arabicus Levy et al., 1973 from Arabia, and description of two new species from North Africa (Scorpiones: Buthidae)</mods:title>
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<mods:namePart id="10F8F45CAD89BBCDEDBFD350C69049BB">Kovařík, František</mods:namePart>
<mods:affiliation id="020340E6FCF20C12DFFD36BDC349D50D">P. O. Box 27, CZ- 145 01 Praha 45, Czech Republic &amp; Department of Zoology, Charles University, ViničnÁ 7, CZ- 128 44 Praha 2, Czech Republic</mods:affiliation>
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<mods:namePart id="3244D6F4C7B38D6A52A25ED68C95D756">Lowe, Graeme</mods:namePart>
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<treatment id="3133B07BFF821055966EA0545FF1F9ED" ID-DOI="http://doi.org/10.5281/zenodo.6546405" ID-GBIF-Taxon="195304348" ID-Zenodo-Dep="6546405" LSID="urn:lsid:zoobank.org:act:37EEF1EA-5294-4BAB-B92D-9F568946A2F4" httpUri="http://treatment.plazi.org/id/3133B07BFF821055966EA0545FF1F9ED" lastPageId="39" lastPageNumber="38" pageId="34" pageNumber="33">
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(
<figureCitation id="21A11DE8FF8210509788A03A5DF3FBAE" box="[233,418,1113,1137]" captionStart-0="Figures 160–167" captionStart-1="Figures 168–171" captionStart-2="Figures 172–179" captionStart-3="Figures 180–189" captionStart-4="Figures 190–209" captionStartId-0="32.[115,192,1926,1948]" captionStartId-1="33.[115,192,1986,2008]" captionStartId-2="35.[113,190,1958,1980]" captionStartId-3="36.[117,194,1601,1623]" captionStartId-4="37.[117,194,1900,1922]" captionTargetBox-0="[127,1468,243,1890]" captionTargetBox-1="[146,1454,231,1976]" captionTargetBox-2="[117,1468,265,1932]" captionTargetBox-3="[113,1474,234,1577]" captionTargetBox-4="[113,1473,240,1878]" captionTargetId-0="figure-15@32.[113,1474,238,1905]" captionTargetId-1="figure-9@33.[113,1474,223,1980]" captionTargetId-2="figure-8@35.[113,1474,238,1938]" captionTargetId-3="figure-15@36.[113,1474,234,1590]" captionTargetId-4="figure-114@37.[113,1474,234,1886]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="35" captionTargetPageId-3="36" captionTargetPageId-4="37" captionText-0="Figures 160–167: Hemispermatophores. Figures 160–163. Compsobuthus turieli sp. n., male holotype, whole hemispermatophore (160), capsule in convex compressed (161), anterior (162), and posterior (163) views. Figures 164–167. C. ullrichi sp. n., male holotype, whole hemispermatophore (164), capsule in posterior (165), convex compressed (166), and anterior (167) views. Scale bars: 1 mm (160, 164), 200 μm (161–163, 165–167)." captionText-1="Figures 168–171: Figures 168–169. Compsobuthus ullrichi sp. n., male holotype in dorsal (168) and ventral (169) views. Figures 170–171. C. levyi, male holotype in dorsal (170) and ventral (171) views. Scale bar: 10 mm." captionText-2="Figures 172–179: Figures 172, 174–176. Compsobuthus levyi, male holotype, telson lateral (172), metasoma and telson lateral (174), dorsal (175), and ventral (176) views. Figures 173, 177–179. C. ullrichi sp. n., male holotype, telson lateral (173), metasoma and telson lateral (177), dorsal (178), and ventral (179) views. Scale bar: 10 mm (174–179)." captionText-3="Figures 180–189: Figures 180–181, 184–187. Compsobuthus ullrichi sp. n., male holotype, carapace and tergites I–IV (180), posterior coxosternal area and sternites (181), and left legs I–IV, retrolateral aspects (184–187). Figures 182–183, 188–189. C. levyi, male holotype, carapace and tergites I–III (182), posterior coxosternal area and sternites (183), and left legs III–IV, retrolateral aspects (188–189)." captionText-4="Figures 190–209: Pedipalps. Figures 190–200. Compsobuthus ullrichi sp. n., male holotype, chela dorsal (190), external (191) and ventrointernal (192), patella dorsal (193), external (194) and ventral (195), trochanter and femur dorsointernal (196), dorsal (197), and ventral (198) views, and movable (199) and fixed (200) fingers dentition. Figures 201–209. C. levyi, male holotype, chela dorsal (201), external (202) and ventrointernal (203), patella dorsal (204), external (205) and ventral (206), trochanter and femur dorsal (207) views, and movable (208) and fixed (209) finger dentition. Trichobothrial pattern is indicated in Figures 190–194, 196–197, 201–205, 207 (white circles)." figureDoi-0="http://doi.org/10.5281/zenodo.5741503" figureDoi-1="http://doi.org/10.5281/zenodo.5741505" figureDoi-2="http://doi.org/10.5281/zenodo.5741507" figureDoi-3="http://doi.org/10.5281/zenodo.5741509" figureDoi-4="http://doi.org/10.5281/zenodo.5741511" httpUri-0="https://zenodo.org/record/5741503/files/figure.png" httpUri-1="https://zenodo.org/record/5741505/files/figure.png" httpUri-2="https://zenodo.org/record/5741507/files/figure.png" httpUri-3="https://zenodo.org/record/5741509/files/figure.png" httpUri-4="https://zenodo.org/record/5741511/files/figure.png" pageId="34" pageNumber="33">Figures 164–209</figureCitation>
, 
<figureCitation id="21A11DE8FF82105096CDA03A5E5CFBAE" box="[428,525,1113,1137]" captionStart="Figures 210–224" captionStartId="38.[116,193,1874,1896]" captionTargetBox="[113,1474,234,1856]" captionTargetId="figure-15@38.[113,1474,234,1862]" captionText="Figures 210–224: Mitotic metaphases (210, 213, 216, 219, 222), male postpachytenes (211, 214, 217, 220, 223), and ideograms (212, 215, 218, 221, 224) (y axis: % of the diploid chromosome length, dark grey marks: chromosomes in multivalent association) of Compsobuthus species from Arabia and North Africa. Figure 210. Female of C. acutecarinatus (2n=22). Figures 211–212. Male of C. acutecarinatus (2n=22, 11II). Figures 213–215. Male of C. acutecarinatus (2n=22, 9II+CIV). Figures 216–218. Male of C. arabicus (2n=22, 9II+CIV). Figures 219–221. Male of C. maindroni (2n=22, 11II). Figures 222–224. Male of C. ullrichi sp. n. (2n=22, 9II+CIV).Arrows show chromosomes in multivalent association during postpachytene (214, 217, 223). Scale bar: 10 μm" figureDoi="http://doi.org/10.5281/zenodo.6544482" httpUri="https://zenodo.org/record/6544482/files/figure.png" pageId="34" pageNumber="33">222–224</figureCitation>
, 
<tableCitation id="F41834D6FF8210509576A03A5EDDFBAE" box="[535,652,1113,1137]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="34.[116,172,986,1007]" captionStartId-1="39.[117,173,697,718]" captionTargetId-0="graphics-458@34.[119,1471,235,973]" captionTargetId-1="graphics-629@39.[118,1476,236,680]" captionTargetPageId-0="34" captionTargetPageId-1="39" captionText-0="Table 3. Comparative measurements of adults of Compsobuthus ullrichi sp. n., C. levyi and C. kabateki. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 4. The diploid number, the postpachytene configuration, the percentage of the total chromosome length of the diploid set (% TCL) and the origin of Compsobuthus species cytogenetically investigated. Specimens karyotyped: C. acutecarinatus (Simon, 1882): Oman, Dhalkut beach, 16.6989°N 53.2214°E, 1♂; E of Aydam, 17.8887°N 53.06640°E, 1♂; W of Qairoon Hairitti,17.2564°N 54.0220°E, 2♂; WadiAsh Shuwaymiyyah, 17.9293°N 55.5510°E, 1♂1♀. C. arabicus Levy, Amitai &amp; Shulov, 1973: Oman, Al Wasil, 22.4914°N 58.7151°E, 2♂; Jabal Bani Jabir, 22.8132°N 59.0589°E, 1♂; E of Aydam, 17.8887°N 53.06640°E, 1♂; W of Wahiba, 22.0299°N 58.1849°E, 1♂. C. maindroni (Kraepelin, 1900): Oman, Khatm Ha‘ir Asfar, 24.7750°N 55.9146°E, 2♂; Say Plateau, 23.1185°N 57.6311°E, 2♂. C. ullrichi sp. n., 1♂ holotype." httpUri-0="http://table.plazi.org/id/EDE551E5FF8210509715A7B95EC7FBD2" httpUri-1="http://table.plazi.org/id/EDE551E5FF8710559714A6DA5F1EFCA3" pageId="34" pageNumber="33" tableUuid-0="EDE551E5FF8210509715A7B95EC7FBD2" tableUuid-1="EDE551E5FF8710559714A6DA5F1EFCA3">Tables 3–4</tableCitation>
)
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<emphasis id="8BEEDD7FFF821050971DA01A5D63FB6E" bold="true" pageId="34" pageNumber="33">http://zoobank.org/urn:lsid:zoobank.org:act:7EEF1EA-5294</emphasis>
-4BAB-B92D-9F568946A2F4
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<materialsCitation id="09F20B30FF82105096FBA15A5EF0FAAE" ID-GBIF-Occurrence="3766759327" collectingDate="2019-02" collectionCode="FKCP" collectorName="A. Ullrich" country="Egypt" latitude="28.483334" location="DahAb" longLatPrecision="1232" longitude="34.5" municipality="DahAb" pageId="34" pageNumber="33" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<paragraph id="B925016DFF8210509713A1FA5EDFFA0E" blockId="34.[114,772,1433,1489]" pageId="34" pageNumber="33">
ETYMOLOGY. It is a pleasure to name this species after Alex Ullrich (
<collectingCountry id="C18D41FDFF82105097ADA1DA5D64FA0E" box="[204,309,1465,1489]" name="Germany" pageId="34" pageNumber="33">Germany</collectingCountry>
), collector of the male 
<typeStatus id="6621BFCFFF8210509548A1DA5EDBFA0E" box="[553,650,1465,1489]" pageId="34" pageNumber="33" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="F18052E6FF8210509713A19A597AFB70" pageId="34" pageNumber="33" type="diagnosis">
<paragraph id="B925016DFF8210509713A19A597AFB70" blockId="34.[114,773,1529,2033]" lastBlockId="34.[817,1475,1079,1199]" pageId="34" pageNumber="33">
DIAGNOSIS (
<specimenCount id="AF9CCAE4FF8210509797A19A5D5AF9CE" box="[246,267,1529,1553]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
). Total length 
<quantity id="7E62AC88FF82105096D2A19A5E54F9CE" box="[435,517,1529,1553]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="34" pageNumber="33" unit="mm" value="28.0">28 mm</quantity>
. Male pedipalp fingers with dentate margins proximally undulate, chela L/W ratio: 4.75. Base color uniform yellow to yellowish brown with dark spot on fifth metasomal segment. Carapace and tergites finely granular with some larger granules. Anterior margin of carapace bearing 8 symmetrically distributed spinules. Pedipalp femur L/ Carapace L ratio: 
<specimenCount id="AF9CCAE4FF8210509565A2D95E49F90E" box="[516,536,1722,1745]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
0.95. Movable finger of pedipalp chela with 10–11 rows of granules, all rows with external and internal accessory denticles (‘ 
<emphasis id="8BEEDD7FFF8210509536A2995EFBF8CE" box="[599,682,1786,1809]" italics="true" pageId="34" pageNumber="33">werneri</emphasis>
’ group; Levy &amp; Amitai, 1980). Manus of pedipalp chela shorter than fixed finger. Pedipalp chela L/movable finger L ratio: 
<specimenCount id="AF9CCAE4FF82105095D2A3595E96F88E" box="[691,711,1850,1873]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
1.47. Metasoma I–II with 10 carinae, III–IV with 8 carinae. All metasomal segments longer than wide; metasoma L/W ratios 
<specimenCount id="AF9CCAE4FF8210509713A3F95CD8F86E" box="[114,137,1946,1969]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
: III 1.94, IV 2.25, V 2.80. Metasoma V W/D ratio 
<specimenCount id="AF9CCAE4FF82105095FEA3F95EE7F86E" box="[671,694,1946,1969]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
: 0.945. Ventral intercarinal surfaces of metasoma with macrosetae. Pectine teeth: 
<specimenCount id="AF9CCAE4FF8210509674A3B95D78F82E" box="[277,297,2010,2033]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
22–23. Pectine L/ Metasoma V W ratio: 
<specimenCount id="AF9CCAE4FF8210509450A05B5F14FB90" box="[817,837,1080,1103]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
2.31. Sternites smooth medially and finely granulated marginally; metasoma granulated. Sternites VI–VII with 4 crenulate carinae. Telson rather bulbous, aculeus shorter than vesicle. Subaculear tubercle weakly developed.
</paragraph>
</subSubSection>
<subSubSection id="F18052E6FF8210559450A0BB5DF6F9CD" lastPageId="39" lastPageNumber="38" pageId="34" pageNumber="33" type="description">
<paragraph id="B925016DFF8210509450A0BB5964FA90" blockId="34.[817,1475,1239,2031]" pageId="34" pageNumber="33">
DESCRIPTION (
<specimenCount id="AF9CCAE4FF82105094BAA0BB5FBEFB2F" box="[987,1007,1240,1264]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
<typeStatus id="6621BFCFFF8210509367A0BB582BFB2F" box="[1030,1146,1240,1264]" pageId="34" pageNumber="33" type="holotype">HOLOTYPE</typeStatus>
). Total length 
<quantity id="7E62AC88FF8210509221A0BB59EEFB30" box="[1344,1471,1240,1264]" metricMagnitude="-2" metricUnit="m" metricValue="2.8129999999999997" pageId="34" pageNumber="33" unit="mm" value="28.13">28.13 mm</quantity>
. Female unknown. The habitus is shown in 
<figureCitation id="21A11DE8FF821050927FA09459EEFAD0" box="[1310,1471,1271,1295]" captionStart="Figures 168–171" captionStartId="33.[115,192,1986,2008]" captionTargetBox="[146,1454,231,1976]" captionTargetId="figure-9@33.[113,1474,223,1980]" captionTargetPageId="33" captionText="Figures 168–171: Figures 168–169. Compsobuthus ullrichi sp. n., male holotype in dorsal (168) and ventral (169) views. Figures 170–171. C. levyi, male holotype in dorsal (170) and ventral (171) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741505" httpUri="https://zenodo.org/record/5741505/files/figure.png" pageId="34" pageNumber="33">Figs. 168–169</figureCitation>
. Trichobothriotaxy of pedipalps is shown in 
<figureCitation id="21A11DE8FF821050927EA17459EEFAEF" box="[1311,1471,1303,1328]" captionStart="Figures 190–209" captionStartId="37.[117,194,1900,1922]" captionTargetBox="[113,1473,240,1878]" captionTargetId="figure-114@37.[113,1474,234,1886]" captionTargetPageId="37" captionText="Figures 190–209: Pedipalps. Figures 190–200. Compsobuthus ullrichi sp. n., male holotype, chela dorsal (190), external (191) and ventrointernal (192), patella dorsal (193), external (194) and ventral (195), trochanter and femur dorsointernal (196), dorsal (197), and ventral (198) views, and movable (199) and fixed (200) fingers dentition. Figures 201–209. C. levyi, male holotype, chela dorsal (201), external (202) and ventrointernal (203), patella dorsal (204), external (205) and ventral (206), trochanter and femur dorsal (207) views, and movable (208) and fixed (209) finger dentition. Trichobothrial pattern is indicated in Figures 190–194, 196–197, 201–205, 207 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5741511" httpUri="https://zenodo.org/record/5741511/files/figure.png" pageId="34" pageNumber="33">Figs. 190–197</figureCitation>
. Male with pedipalp fingers proximally undulate.
</paragraph>
<paragraph id="B925016DFF8210509450A1345FEAFA70" blockId="34.[817,1475,1239,2031]" pageId="34" pageNumber="33">
<emphasis id="8BEEDD7FFF8210509450A1345FFDFAB0" bold="true" box="[817,940,1367,1391]" pageId="34" pageNumber="33">Coloration</emphasis>
(
<figureCitation id="21A11DE8FF82105094D8A1345807FAB0" box="[953,1110,1367,1391]" captionStart="Figures 168–171" captionStartId="33.[115,192,1986,2008]" captionTargetBox="[146,1454,231,1976]" captionTargetId="figure-9@33.[113,1474,223,1980]" captionTargetPageId="33" captionText="Figures 168–171: Figures 168–169. Compsobuthus ullrichi sp. n., male holotype in dorsal (168) and ventral (169) views. Figures 170–171. C. levyi, male holotype in dorsal (170) and ventral (171) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5741505" httpUri="https://zenodo.org/record/5741505/files/figure.png" pageId="34" pageNumber="33">Figs. 168–169</figureCitation>
). The base color is uniform yellow to yellowish brown, with fuscosity on the anterior half of metasoma V.
</paragraph>
<paragraph id="B925016DFF8210569450A1D45D4DF8F2" blockId="34.[817,1475,1239,2031]" lastBlockId="36.[115,773,1717,2030]" lastPageId="36" lastPageNumber="35" pageId="34" pageNumber="33">
<emphasis id="8BEEDD7FFF8210509450A1D4580BFA0F" bold="true" box="[817,1114,1463,1488]" pageId="34" pageNumber="33">Carapace and mesosoma</emphasis>
(
<figureCitation id="21A11DE8FF8210509311A1D45942FA0F" box="[1136,1299,1463,1488]" captionStart="Figures 180–189" captionStartId="36.[117,194,1601,1623]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-15@36.[113,1474,234,1590]" captionTargetPageId="36" captionText="Figures 180–189: Figures 180–181, 184–187. Compsobuthus ullrichi sp. n., male holotype, carapace and tergites I–IV (180), posterior coxosternal area and sternites (181), and left legs I–IV, retrolateral aspects (184–187). Figures 182–183, 188–189. C. levyi, male holotype, carapace and tergites I–III (182), posterior coxosternal area and sternites (183), and left legs III–IV, retrolateral aspects (188–189)." figureDoi="http://doi.org/10.5281/zenodo.5741509" httpUri="https://zenodo.org/record/5741509/files/figure.png" pageId="34" pageNumber="33">Figs. 180–181</figureCitation>
). The carapace is covered by granules of different sizes. The carinae are moderately to strongly developed and granular. The anterior margin of the carapace is weakly concave medially, and bears 8 symmetrically distributed spinules (macrosetae). The tergites are partially, coarsely granulated. Tergites I–VI are tricarinate, with strong, denticulate median and lateral carinae. Each carina terminates in a spiniform process that in the lateral carinae extends well past the posterior margin of the tergite. Tergite VII is pentacarinate, with lateral pairs of carinae strong, serratocrenulate, median pairs moderate, crenulate; median carina is weak and confined to the anterior half of the segment. Pectinal tooth count: 
<specimenCount id="AF9CCAE4FF8210509240A35B5964F890" box="[1313,1333,1848,1871]" pageId="34" pageNumber="33" type="male">♂</specimenCount>
22–23. The pectine marginal tips extend to the posterior margin of sternite IV. The pectines have 3 marginal lamellae and 7–9 middle lamellae. The lamellae bear numerous dark setae, and each fulcrum bears 2–3 dark setae. Sternites are smooth medially and finely granulated marginally. The posterior areas of sternites lack a broad glabrous patch. Sternites VI–VII bear 4 crenulate carinae which are more strongly granulated on VII. The other sternites bear one pair of carinae on the medial side of the spiracles.
</paragraph>
<caption id="EDE551E5FF8310519710A3C55EAAF829" ID-DOI="http://doi.org/10.5281/zenodo.5741507" ID-Zenodo-Dep="5741507" httpUri="https://zenodo.org/record/5741507/files/figure.png" pageId="35" pageNumber="34" startId="35.[113,190,1958,1980]" targetBox="[117,1468,265,1932]" targetPageId="35">
<paragraph id="B925016DFF8310519710A3C55EAAF829" blockId="35.[113,1472,1958,2038]" pageId="35" pageNumber="34">
<emphasis id="8BEEDD7FFF8310519710A3C55D48F864" bold="true" box="[113,281,1958,1980]" pageId="35" pageNumber="34">Figures 172–179</emphasis>
: 
<emphasis id="8BEEDD7FFF8310519647A3C55DCCF864" bold="true" box="[294,413,1958,1980]" pageId="35" pageNumber="34">Figures 172</emphasis>
, 
<emphasis id="8BEEDD7FFF83105196CBA3C55DAFF864" bold="true" box="[426,510,1958,1980]" pageId="35" pageNumber="34">174–176</emphasis>
. 
<taxonomicName id="7E9A7AEEFF831051956BA3C55E9DF864" authorityName="KovarIk" authorityYear="2012" box="[522,716,1958,1979]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF831051956BA3C55E9DF864" box="[522,716,1958,1979]" italics="true" pageId="35" pageNumber="34">Compsobuthus levyi</emphasis>
</taxonomicName>
, male holotype, telson lateral (172), metasoma and telson lateral (174), dorsal (175), and ventral (176) views. 
<emphasis id="8BEEDD7FFF83105196FCA3A05E42F807" bold="true" box="[413,531,1987,2009]" pageId="35" pageNumber="34">Figures 173</emphasis>
, 
<emphasis id="8BEEDD7FFF831051957FA3A05E23F807" bold="true" box="[542,626,1987,2008]" pageId="35" pageNumber="34">177–179</emphasis>
. 
<taxonomicName id="7E9A7AEEFF831051951CA3A05EB1F807" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[637,736,1987,2008]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="34" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF831051951CA3A05EB1F807" box="[637,736,1987,2008]" italics="true" pageId="35" pageNumber="34">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8310519584A3A05F48F807" box="[741,793,1987,2008]" pageId="35" pageNumber="34" rank="species">
<emphasis id="8BEEDD7FFF8310519584A3A05EADF807" bold="true" box="[741,764,1987,2008]" pageId="35" pageNumber="34">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF8310519466A3A05F45F807" bold="true" box="[775,788,1987,2008]" pageId="35" pageNumber="34">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, telson lateral (173), metasoma and telson lateral (177), dorsal (178), and ventral (179) views. Scale bar: 10 mm (174–179).
</paragraph>
</caption>
<caption id="EDE551E5FF8410569714A2225908F94E" ID-DOI="http://doi.org/10.5281/zenodo.5741509" ID-Zenodo-Dep="5741509" httpUri="https://zenodo.org/record/5741509/files/figure.png" pageId="36" pageNumber="35" startId="36.[117,194,1601,1623]" targetBox="[113,1474,234,1577]" targetPageId="36">
<paragraph id="B925016DFF8410569714A2225908F94E" blockId="36.[116,1475,1601,1681]" pageId="36" pageNumber="35">
<emphasis id="8BEEDD7FFF8410569714A2225D4EF989" bold="true" box="[117,287,1601,1623]" pageId="36" pageNumber="35">Figures 180–189</emphasis>
: 
<emphasis id="8BEEDD7FFF841056964EA2225D8BF989" bold="true" box="[303,474,1601,1623]" pageId="36" pageNumber="35">Figures 180–181</emphasis>
, 
<emphasis id="8BEEDD7FFF8410569688A2225E6CF989" bold="true" box="[489,573,1601,1622]" pageId="36" pageNumber="35">184–187</emphasis>
. 
<taxonomicName id="7E9A7AEEFF841056952CA2225F7DF989" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[589,812,1601,1622]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF841056952CA2225F7DF989" box="[589,812,1601,1622]" italics="true" pageId="36" pageNumber="35">Compsobuthus ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8410569454A2225F3FF989" box="[821,878,1601,1622]" pageId="36" pageNumber="35" rank="species">
<emphasis id="8BEEDD7FFF8410569454A2225F1DF989" bold="true" box="[821,844,1601,1622]" pageId="36" pageNumber="35">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF841056943AA2225F39F989" bold="true" box="[859,872,1601,1622]" pageId="36" pageNumber="35">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, carapace and tergites I–IV (180), posterior coxosternal area and sternites (181), and left legs I–IV, retrolateral aspects (184–187). 
<emphasis id="8BEEDD7FFF84105694A2A23D583AF9AC" bold="true" box="[963,1131,1630,1652]" pageId="36" pageNumber="35">Figures 182–183</emphasis>
, 
<emphasis id="8BEEDD7FFF8410569318A23D589CF9AC" bold="true" box="[1145,1229,1630,1651]" pageId="36" pageNumber="35">188–189</emphasis>
. 
<taxonomicName id="7E9A7AEEFF84105693BBA23D5974F9AC" authorityName="KovarIk" authorityYear="2012" box="[1242,1317,1630,1651]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="35" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF84105693BBA23D5974F9AC" box="[1242,1317,1630,1651]" italics="true" pageId="36" pageNumber="35">C. levyi</emphasis>
</taxonomicName>
, male holotype, carapace and tergites I–III (182), posterior coxosternal area and sternites (183), and left legs III–IV, retrolateral aspects (188–189).
</paragraph>
</caption>
<paragraph id="B925016DFF8410569712A35558D8F8D2" blockId="36.[115,773,1717,2030]" lastBlockId="36.[818,1476,1717,2030]" pageId="36" pageNumber="35">
<emphasis id="8BEEDD7FFF8410569712A3555D27F891" bold="true" box="[115,374,1846,1870]" pageId="36" pageNumber="35">Metasoma and telson</emphasis>
(
<figureCitation id="21A11DE8FF84105696EFA3565E21F891" box="[398,624,1845,1870]" captionStart="Figures 172–179" captionStartId="35.[113,190,1958,1980]" captionTargetBox="[117,1468,265,1932]" captionTargetId="figure-8@35.[113,1474,238,1938]" captionTargetPageId="35" captionText="Figures 172–179: Figures 172, 174–176. Compsobuthus levyi, male holotype, telson lateral (172), metasoma and telson lateral (174), dorsal (175), and ventral (176) views. Figures 173, 177–179. C. ullrichi sp. n., male holotype, telson lateral (173), metasoma and telson lateral (177), dorsal (178), and ventral (179) views. Scale bar: 10 mm (174–179)." figureDoi="http://doi.org/10.5281/zenodo.5741507" httpUri="https://zenodo.org/record/5741507/files/figure.png" pageId="36" pageNumber="35">Figs. 173, 177–179</figureCitation>
). Metasomal segments I–II with 10 carinae, III–IV with 8 carinae, and V with 5 carinae. Median lateral carinae of metasoma III reduced and indicated only by several granules posteriorly. All segments sparsely setose and granulate. Accessory rows of granules are present on dorsal surfaces of segments as well as on the ventral surface of segment V. The telson is rather bulbous, with the aculeus a little shorter than the vesicle. A subaculear tubercle is indicated.
</paragraph>
<paragraph id="B925016DFF8410569453A37659E1F831" blockId="36.[818,1476,1717,2030]" pageId="36" pageNumber="35">
<emphasis id="8BEEDD7FFF8410569453A3765FCEF8F2" bold="true" box="[818,927,1813,1837]" pageId="36" pageNumber="35">Pedipalps</emphasis>
(
<figureCitation id="21A11DE8FF84105694D1A376581AF8F1" box="[944,1099,1813,1838]" captionStart="Figures 190–209" captionStartId="37.[117,194,1900,1922]" captionTargetBox="[113,1473,240,1878]" captionTargetId="figure-114@37.[113,1474,234,1886]" captionTargetPageId="37" captionText="Figures 190–209: Pedipalps. Figures 190–200. Compsobuthus ullrichi sp. n., male holotype, chela dorsal (190), external (191) and ventrointernal (192), patella dorsal (193), external (194) and ventral (195), trochanter and femur dorsointernal (196), dorsal (197), and ventral (198) views, and movable (199) and fixed (200) fingers dentition. Figures 201–209. C. levyi, male holotype, chela dorsal (201), external (202) and ventrointernal (203), patella dorsal (204), external (205) and ventral (206), trochanter and femur dorsal (207) views, and movable (208) and fixed (209) finger dentition. Trichobothrial pattern is indicated in Figures 190–194, 196–197, 201–205, 207 (white circles)." figureDoi="http://doi.org/10.5281/zenodo.5741511" httpUri="https://zenodo.org/record/5741511/files/figure.png" pageId="36" pageNumber="35">Figs. 190–200</figureCitation>
). The pedipalps are smooth, partly finely granulated and sparsely hirsute. The femur bears 5 carinae, the patella 7 granular carinae, the chela 5 rather smooth carinae. The movable and fixed fingers bear 10–11 rows of granules, all with external and internal accessory granules. Pedipalp chela L/W ratio: 
<specimenCount id="AF9CCAE4FF8410569326A3D5580AF812" box="[1095,1115,1974,1997]" pageId="36" pageNumber="35" type="male">♂</specimenCount>
4.75. Manus of chela shorter than fixed finger. Pedipalp chela L/movable finger L ratio: 
<specimenCount id="AF9CCAE4FF8410569200A3B55924F832" box="[1377,1397,2006,2029]" pageId="36" pageNumber="35" type="male">♂</specimenCount>
1.47.
</paragraph>
<caption id="EDE551E5FF8510579714A30F596AF82A" ID-DOI="http://doi.org/10.5281/zenodo.5741511" ID-Zenodo-Dep="5741511" httpUri="https://zenodo.org/record/5741511/files/figure.png" pageId="37" pageNumber="36" startId="37.[117,194,1900,1922]" targetBox="[113,1473,240,1878]" targetPageId="37">
<paragraph id="B925016DFF8510579714A30F596AF82A" blockId="37.[117,1476,1900,2037]" pageId="37" pageNumber="36">
<emphasis id="8BEEDD7FFF8510579714A30F5D73F85E" bold="true" box="[117,290,1900,1922]" pageId="37" pageNumber="36">Figures 190–209</emphasis>
: Pedipalps. 
<emphasis id="8BEEDD7FFF85105796C5A30F5E01F85E" bold="true" box="[420,592,1900,1922]" pageId="37" pageNumber="36">Figures 190–200</emphasis>
. 
<taxonomicName id="7E9A7AEEFF8510579503A30F5F15F85E" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[610,836,1900,1921]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8510579503A30F5F15F85E" box="[610,836,1900,1921]" italics="true" pageId="37" pageNumber="36">Compsobuthus ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF851057942EA30F5FDBF85D" box="[847,906,1900,1922]" pageId="37" pageNumber="36" rank="species">
<emphasis id="8BEEDD7FFF851057942EA30F5F37F85E" bold="true" box="[847,870,1900,1921]" pageId="37" pageNumber="36">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF8510579419A30F5FD4F85E" bold="true" box="[888,901,1900,1921]" pageId="37" pageNumber="36">n</emphasis>
.
</taxonomicNameLabel>
, male holotype, chela dorsal (190), external (191) and ventrointernal (192), patella dorsal (193), external (194) and ventral (195), trochanter and femur dorsointernal (196), dorsal (197), and ventral (198) views, and movable (199) and fixed (200) fingers dentition. 
<emphasis id="8BEEDD7FFF851057958AA3C55FC0F864" bold="true" box="[747,913,1958,1980]" pageId="37" pageNumber="36">Figures 201–209</emphasis>
. 
<taxonomicName id="7E9A7AEEFF85105794FCA3C55FB4F864" authorityName="KovarIk" authorityYear="2012" box="[925,997,1958,1979]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="36" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF85105794FCA3C55FB4F864" box="[925,997,1958,1979]" italics="true" pageId="37" pageNumber="36">C. levyi</emphasis>
</taxonomicName>
, male holotype, chela dorsal (201), external (202) and ventrointernal (203), patella dorsal (204), external (205) and ventral (206), trochanter and femur dorsal (207) views, and movable (208) and fixed (209) finger dentition. Trichobothrial pattern is indicated in Figures 190–194, 196–197, 201–205, 207 (white circles).
</paragraph>
</caption>
<caption id="EDE551E5FF8610549715A3315EAFF828" ID-DOI="http://doi.org/10.5281/zenodo.6544482" ID-Zenodo-Dep="6544482" httpUri="https://zenodo.org/record/6544482/files/figure.png" pageId="38" pageNumber="37" startId="38.[116,193,1874,1896]" subCaptionStartIDs="38.[413,481,1931,1953] 38.[904,981,1931,1953] 38.[116,193,1960,1982] 38.[738,815,1960,1982] 38.[1301,1378,1960,1982] 38.[467,544,1989,2011]" subCaptionStarts="Figure 210 &amp; Figures 211 &amp; Figures 213 &amp; Figures 216 &amp; Figures 219 &amp; Figures 222" targetBox="[113,1474,234,1856]" targetPageId="38">
<paragraph id="B925016DFF8610549715A3315EAFF828" blockId="38.[116,1475,1874,2040]" pageId="38" pageNumber="37">
<emphasis id="8BEEDD7FFF8610549715A3315D4BF8B8" bold="true" box="[116,282,1874,1896]" pageId="38" pageNumber="37">Figures 210–224</emphasis>
: Mitotic metaphases (210, 213, 216, 219, 222), male postpachytenes (211, 214, 217, 220, 223), and ideograms (212, 215, 218, 221, 224) (y axis: % of the diploid chromosome length, dark grey marks: chromosomes in multivalent association) of 
<taxonomicName id="7E9A7AEEFF8610549389A30C5993F85A" box="[1256,1474,1903,1925]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8BEEDD7FFF8610549389A30C5926F85B" box="[1256,1399,1903,1924]" italics="true" pageId="38" pageNumber="37">Compsobuthus</emphasis>
species
</taxonomicName>
from Arabia and North Africa. 
<emphasis id="8BEEDD7FFF86105496FCA3E85E5AF87F" bold="true" box="[413,523,1931,1953]" pageId="38" pageNumber="37">Figure 210</emphasis>
. Female of 
<taxonomicName id="7E9A7AEEFF861054951FA3EF5F78F87E" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[638,809,1932,1953]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF861054951FA3EF5F78F87E" box="[638,809,1932,1953]" italics="true" pageId="38" pageNumber="37">C. acutecarinatus</emphasis>
</taxonomicName>
(2n=22). 
<emphasis id="8BEEDD7FFF86105494E9A3E8587CF87F" bold="true" box="[904,1069,1931,1953]" pageId="38" pageNumber="37">Figures 211–212</emphasis>
. Male of 
<taxonomicName id="7E9A7AEEFF86105493EAA3EF5967F87E" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[1163,1334,1932,1953]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF86105493EAA3EF5967F87E" box="[1163,1334,1932,1953]" italics="true" pageId="38" pageNumber="37">C. acutecarinatus</emphasis>
</taxonomicName>
(2n=22, 11II). 
<emphasis id="8BEEDD7FFF8610549715A3CB5D4AF862" bold="true" box="[116,283,1960,1982]" pageId="38" pageNumber="37">Figures 213–215</emphasis>
. Male of 
<taxonomicName id="7E9A7AEEFF8610549619A3CA5E75F861" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[376,548,1961,1982]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF8610549619A3CA5E75F861" box="[376,548,1961,1982]" italics="true" pageId="38" pageNumber="37">C. acutecarinatus</emphasis>
</taxonomicName>
(2n=22, 9II+CIV). 
<emphasis id="8BEEDD7FFF8610549583A3CB5FD8F862" bold="true" box="[738,905,1960,1982]" pageId="38" pageNumber="37">Figures 216–218</emphasis>
. Male of 
<taxonomicName id="7E9A7AEEFF8610549487A3CA5807F862" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" box="[998,1110,1960,1982]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="8BEEDD7FFF8610549487A3CA5807F862" box="[998,1110,1960,1982]" italics="true" pageId="38" pageNumber="37">C. arabicus</emphasis>
</taxonomicName>
(2n=22, 9II+CIV). 
<emphasis id="8BEEDD7FFF8610549274A3CB59EDF862" bold="true" box="[1301,1468,1960,1982]" pageId="38" pageNumber="37">Figures 219–221</emphasis>
. Male of 
<taxonomicName id="7E9A7AEEFF86105497A5A3A65D13F805" baseAuthorityName="Kraepelin" baseAuthorityYear="1901" box="[196,322,1989,2010]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="37" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="8BEEDD7FFF86105497A5A3A65D13F805" box="[196,322,1989,2010]" italics="true" pageId="38" pageNumber="37">C. maindroni</emphasis>
</taxonomicName>
(2n=22, 11II). 
<emphasis id="8BEEDD7FFF86105496B2A3A65E28F805" bold="true" box="[467,633,1989,2011]" pageId="38" pageNumber="37">Figures 222–224</emphasis>
. Male of 
<taxonomicName id="7E9A7AEEFF86105495B4A3A65F66F805" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[725,823,1989,2010]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="38" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
C. 
<emphasis id="8BEEDD7FFF8610549591A3A65F66F805" box="[752,823,1989,2010]" italics="true" pageId="38" pageNumber="37">ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF861054945DA3A65F20F804" box="[828,881,1989,2011]" pageId="38" pageNumber="37" rank="species">
<emphasis id="8BEEDD7FFF861054945DA3A65F02F805" bold="true" box="[828,851,1989,2010]" pageId="38" pageNumber="37">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF861054943FA3A65F3AF805" bold="true" box="[862,875,1989,2010]" pageId="38" pageNumber="37">n</emphasis>
.
</taxonomicNameLabel>
(2n=22, 9II+CIV). Arrows show chromosomes in multivalent association during postpachytene (214, 217, 223). Scale bar: 10 μm
</paragraph>
</caption>
<paragraph id="B925016DFF861054959FA3815F53F828" blockId="38.[116,1475,1874,2040]" box="[766,770,2018,2039]" pageId="38" pageNumber="37">.</paragraph>
<paragraph id="B925016DFF871055971CA49A597EFD7A" pageId="39" pageNumber="38">
<table id="CB9AF3CDFF87EF8D971CA49A59F7FD7A" box="[125,1446,249,677]" gridcols="6" gridrows="10" pageId="39" pageNumber="38">
<tr id="07AA032FFF87EF8D971CA49A59F7FEC6" box="[125,1446,249,281]" gridrow="0" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA49A5D07FEC6" box="[125,342,249,281]" gridcol="0" gridrow="0" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF871055971CA49A5C8AFEC7" bold="true" box="[125,219,249,280]" pageId="39" pageNumber="38">Species</emphasis>
</th>
<th id="447B6A53FF87EF8D96F8A49A5DE8FEC6" box="[409,441,249,281]" gridcol="1" gridrow="0" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF87105596F8A4995DE8FEC6" bold="true" box="[409,441,250,281]" pageId="39" pageNumber="38">2n</emphasis>
</th>
<th id="447B6A53FF87EF8D96B9A49A5F2FFEC6" box="[472,894,249,281]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF87105596B9A4995F5CFEC7" bold="true" box="[472,781,249,281]" pageId="39" pageNumber="38">Postpachytene % TCL</emphasis>
</th>
<th id="447B6A53FF87EF8D9374A49A59F7FEC6" box="[1045,1446,249,281]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559374A49A5940FEC7" bold="true" box="[1045,1297,249,280]" pageId="39" pageNumber="38">Country Reference</emphasis>
</th>
</tr>
<tr id="07AA032FFF87EF8D971CA54459F7FE9A" box="[125,1446,295,325]" gridrow="1" pageId="39" pageNumber="38" rowspan-0="1" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td id="447B6A53FF87EF8D96B9A5445ED9FE9A" box="[472,648,295,325]" gridcol="2" gridrow="1" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF87105596B9A5445ED9FE9A" bold="true" box="[472,648,295,325]" pageId="39" pageNumber="38">configuration</emphasis>
</td>
</tr>
<tr id="07AA032FFF87EF8D971CA53259F7FEAF" box="[125,1446,337,368]" gridrow="2" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA5325D07FEAF" box="[125,342,337,368]" gridcol="0" gridrow="2" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA5325D07FEAF" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[125,342,337,368]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF871055971CA5325D07FEAF" box="[125,342,337,368]" italics="true" pageId="39" pageNumber="38">C. acutecarinatus</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A5325DE8FEAF" box="[409,441,337,368]" gridcol="1" gridrow="2" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A5325ED9FEAF" box="[472,648,337,368]" gridcol="2" gridrow="2" pageId="39" pageNumber="38">11II</td>
<td id="447B6A53FF87EF8D95C8A5325F2FFEAF" box="[681,894,337,368]" gridcol="3" gridrow="2" pageId="39" pageNumber="38">2x 6.23 – 2x 3.46</td>
<td id="447B6A53FF87EF8D9374A532583CFEAF" box="[1045,1133,337,368]" gridcol="4" gridrow="2" pageId="39" pageNumber="38">Oman</td>
<td id="447B6A53FF87EF8D93EEA53259F7FEAF" box="[1167,1446,337,368]" gridcol="5" gridrow="2" pageId="39" pageNumber="38">present study</td>
</tr>
<tr id="07AA032FFF87EF8D971CA51E59F7FE43" box="[125,1446,381,412]" gridrow="3" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA51E5D07FE43" box="[125,342,381,412]" gridcol="0" gridrow="3" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA51E5D07FE43" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[125,342,381,412]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF871055971CA51E5D07FE43" box="[125,342,381,412]" italics="true" pageId="39" pageNumber="38">C. acutecarinatus</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A51E5DE8FE43" box="[409,441,381,412]" gridcol="1" gridrow="3" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A51E5ED9FE43" box="[472,648,381,412]" gridcol="2" gridrow="3" pageId="39" pageNumber="38">9II+CIV</td>
<td id="447B6A53FF87EF8D95C8A51E5F2FFE43" box="[681,894,381,412]" gridcol="3" gridrow="3" pageId="39" pageNumber="38">2x 6.08 – 2x 3.28</td>
<td id="447B6A53FF87EF8D9374A51E583CFE43" box="[1045,1133,381,412]" gridcol="4" gridrow="3" pageId="39" pageNumber="38">Oman</td>
<td id="447B6A53FF87EF8D93EEA51E59F7FE43" box="[1167,1446,381,412]" gridcol="5" gridrow="3" pageId="39" pageNumber="38">present study</td>
</tr>
<tr id="07AA032FFF87EF8D971CA5CA59F7FE17" box="[125,1446,425,456]" gridrow="4" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA5CA5D07FE17" box="[125,342,425,456]" gridcol="0" gridrow="4" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA5CA5D5AFE17" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" box="[125,267,425,456]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="8BEEDD7FFF871055971CA5CA5D5AFE17" box="[125,267,425,456]" italics="true" pageId="39" pageNumber="38">C. arabicus</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A5CA5DE8FE17" box="[409,441,425,456]" gridcol="1" gridrow="4" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A5CA5ED9FE17" box="[472,648,425,456]" gridcol="2" gridrow="4" pageId="39" pageNumber="38">9II+CIV</td>
<td id="447B6A53FF87EF8D95C8A5CA5F2FFE17" box="[681,894,425,456]" gridcol="3" gridrow="4" pageId="39" pageNumber="38">2x 6.58 – 2.55</td>
<td id="447B6A53FF87EF8D9374A5CA583CFE17" box="[1045,1133,425,456]" gridcol="4" gridrow="4" pageId="39" pageNumber="38">Oman</td>
<td id="447B6A53FF87EF8D93EEA5CA59F7FE17" box="[1167,1446,425,456]" gridcol="5" gridrow="4" pageId="39" pageNumber="38">present study</td>
</tr>
<tr id="07AA032FFF87EF8D971CA5B659F7FE2A" box="[125,1446,469,501]" gridrow="5" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA5B65D07FE2A" box="[125,342,469,501]" gridcol="0" gridrow="5" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA5B65D61FE2B" box="[125,304,469,500]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="eritreaensis">
<emphasis id="8BEEDD7FFF871055971CA5B65D61FE2B" box="[125,304,469,500]" italics="true" pageId="39" pageNumber="38">C. eritreaensis</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A5B65DE8FE2A" box="[409,441,469,501]" gridcol="1" gridrow="5" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A5B65ED9FE2A" box="[472,648,469,501]" gridcol="2" gridrow="5" pageId="39" pageNumber="38">11II</td>
<td id="447B6A53FF87EF8D95C8A5B65F2FFE2A" box="[681,894,469,501]" gridcol="3" gridrow="5" pageId="39" pageNumber="38">2x 6.70 – 2x 2.81</td>
<td id="447B6A53FF87EF8D9374A5B6583CFE2A" box="[1045,1133,469,501]" gridcol="4" gridrow="5" pageId="39" pageNumber="38">Eritrea</td>
<td id="447B6A53FF87EF8D93EEA5B659F7FE2A" box="[1167,1446,469,501]" gridcol="5" gridrow="5" pageId="39" pageNumber="38">
<bibRefCitation id="DD0B7C9CFF87105593EEA5B659DEFE2B" author="Kovarik" box="[1167,1423,469,501]" etAl="et al." firstAuthor="Kovarik" pageId="39" pageNumber="38" pagination="1 - 21" refId="ref15356" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2016 a. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part VI. Compsobuthus Vachon, 1949 (Buthidae) with description of C. eritreaensis sp. n. Euscorpius, 226: 1 - 21." type="journal article" year="2016" yearSuffix="a">Kovařík et al., 2016a</bibRefCitation>
</td>
</tr>
<tr id="07AA032FFF87EF8D971CA66259F7FDFE" box="[125,1446,513,545]" gridrow="6" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA6625D07FDFE" box="[125,342,513,545]" gridcol="0" gridrow="6" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA6615D71FDFF" baseAuthorityName="Kraepelin" baseAuthorityYear="1901" box="[125,288,513,544]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="8BEEDD7FFF871055971CA6615D71FDFF" box="[125,288,513,544]" italics="true" pageId="39" pageNumber="38">C. maindroni</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A6625DE8FDFE" box="[409,441,513,545]" gridcol="1" gridrow="6" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A6625ED9FDFE" box="[472,648,513,545]" gridcol="2" gridrow="6" pageId="39" pageNumber="38">11II</td>
<td id="447B6A53FF87EF8D95C8A6625F2FFDFE" box="[681,894,513,545]" gridcol="3" gridrow="6" pageId="39" pageNumber="38">2x 6.94 – 2x 3.12</td>
<td id="447B6A53FF87EF8D9374A662583CFDFE" box="[1045,1133,513,545]" gridcol="4" gridrow="6" pageId="39" pageNumber="38">Oman</td>
<td id="447B6A53FF87EF8D93EEA66259F7FDFE" box="[1167,1446,513,545]" gridcol="5" gridrow="6" pageId="39" pageNumber="38">present study</td>
</tr>
<tr id="07AA032FFF87EF8D971CA64E59F7FD92" box="[125,1446,557,589]" gridrow="7" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA64E5D07FD92" box="[125,342,557,589]" gridcol="0" gridrow="7" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA64D5D68FD93" authorityName="StAhlavsky et al." authorityYear="2014" baseAuthorityName="Birula" baseAuthorityYear="1905" box="[125,313,557,588]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="matthiesseni">
<emphasis id="8BEEDD7FFF871055971CA64D5D68FD93" box="[125,313,557,588]" italics="true" pageId="39" pageNumber="38">C. matthiesseni</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A64E5DE8FD92" box="[409,441,557,589]" gridcol="1" gridrow="7" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A64E5ED9FD92" box="[472,648,557,589]" gridcol="2" gridrow="7" pageId="39" pageNumber="38">11II</td>
<td id="447B6A53FF87EF8D95C8A64E5F2FFD92" box="[681,894,557,589]" gridcol="3" gridrow="7" pageId="39" pageNumber="38">6.28 – 3.12</td>
<td id="447B6A53FF87EF8D9374A64E583CFD92" box="[1045,1133,557,589]" gridcol="4" gridrow="7" pageId="39" pageNumber="38">Turkey</td>
<td id="447B6A53FF87EF8D93EEA64E59F7FD92" box="[1167,1446,557,589]" gridcol="5" gridrow="7" pageId="39" pageNumber="38">
<bibRefCitation id="DD0B7C9CFF87105593EEA64E59F7FD92" author="Stahlavsky" box="[1167,1446,557,589]" etAl="et al." firstAuthor="Stahlavsky" pageId="39" pageNumber="38" pagination="355 - 358" refId="ref16198" refString="STAHLAVSKY, F., H. KOC &amp; E. A. YAGMUR. 2014. The first record of karyotypes in Leiurus abdullahbayrami and Compsobuthus matthiesseni (Scorpiones: Buthidae) from Turkey. Northwestern Journal of Zoology, 10 (2): 355 - 358." type="journal article" year="2014">Šťáhlavský et al., 2014</bibRefCitation>
</td>
</tr>
<tr id="07AA032FFF87EF8D971CA63A59F7FDA6" box="[125,1446,601,633]" gridrow="8" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA63A5D07FDA6" box="[125,342,601,633]" gridcol="0" gridrow="8" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA6395D68FDA7" authorityName="StAhlavsky et al." authorityYear="2014" baseAuthorityName="Birula" baseAuthorityYear="1905" box="[125,313,601,632]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="matthiesseni">
<emphasis id="8BEEDD7FFF871055971CA6395D68FDA7" box="[125,313,601,632]" italics="true" pageId="39" pageNumber="38">C. matthiesseni</emphasis>
</taxonomicName>
</th>
<td id="447B6A53FF87EF8D96F8A63A5DE8FDA6" box="[409,441,601,633]" gridcol="1" gridrow="8" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A63A5ED9FDA6" box="[472,648,601,633]" gridcol="2" gridrow="8" pageId="39" pageNumber="38">7II+CVIII</td>
<td id="447B6A53FF87EF8D95C8A63A5F2FFDA6" box="[681,894,601,633]" gridcol="3" gridrow="8" pageId="39" pageNumber="38">6.68 – 3.15</td>
<td id="447B6A53FF87EF8D9374A63A583CFDA6" box="[1045,1133,601,633]" gridcol="4" gridrow="8" pageId="39" pageNumber="38">Turkey</td>
<td id="447B6A53FF87EF8D93EEA63A59F7FDA6" box="[1167,1446,601,633]" gridcol="5" gridrow="8" pageId="39" pageNumber="38">
<bibRefCitation id="DD0B7C9CFF87105593EEA63A59F7FDA6" author="Stahlavsky" box="[1167,1446,601,633]" etAl="et al." firstAuthor="Stahlavsky" pageId="39" pageNumber="38" pagination="355 - 358" refId="ref16198" refString="STAHLAVSKY, F., H. KOC &amp; E. A. YAGMUR. 2014. The first record of karyotypes in Leiurus abdullahbayrami and Compsobuthus matthiesseni (Scorpiones: Buthidae) from Turkey. Northwestern Journal of Zoology, 10 (2): 355 - 358." type="journal article" year="2014">Šťáhlavský et al., 2014</bibRefCitation>
</td>
</tr>
<tr id="07AA032FFF87EF8D971CA6E659F7FD7A" box="[125,1446,645,677]" gridrow="9" pageId="39" pageNumber="38">
<th id="447B6A53FF87EF8D971CA6E65D07FD7A" box="[125,342,645,677]" gridcol="0" gridrow="9" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF871055971CA6E55CABFD7A" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[125,250,646,677]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF871055971CA6E55CABFD7A" box="[125,250,646,677]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<emphasis id="8BEEDD7FFF8710559666A6E55D1AFD7A" bold="true" box="[263,331,646,677]" pageId="39" pageNumber="38">
<taxonomicNameLabel id="90DD6004FF8710559666A6E55D1AFD7A" box="[263,331,646,677]" pageId="39" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
<td id="447B6A53FF87EF8D96F8A6E65DE8FD7A" box="[409,441,645,677]" gridcol="1" gridrow="9" pageId="39" pageNumber="38">22</td>
<td id="447B6A53FF87EF8D96B9A6E65ED9FD7A" box="[472,648,645,677]" gridcol="2" gridrow="9" pageId="39" pageNumber="38">9II+CIV</td>
<td id="447B6A53FF87EF8D95C8A6E6583CFD7A" box="[681,1133,645,677]" colspan="2" colspanRight="1" gridcol="3" gridrow="9" pageId="39" pageNumber="38">7.66, 6.92, 2x 5.76 – 2x 3.10 Egypt</td>
<td id="447B6A53FF87EF8D93EEA6E659F7FD7A" box="[1167,1446,645,677]" gridcol="5" gridrow="9" pageId="39" pageNumber="38">present study</td>
</tr>
</table>
</paragraph>
<caption id="EDE551E5FF8710559714A6DA5F1EFCA3" ID-Table-UUID="EDE551E5FF8710559714A6DA5F1EFCA3" httpUri="http://table.plazi.org/id/EDE551E5FF8710559714A6DA5F1EFCA3" pageId="39" pageNumber="38" startId="39.[117,173,697,718]" targetBox="[125,1446,249,677]" targetIsTable="true" targetPageId="39">
<paragraph id="B925016DFF8710559714A6DA5EB4FD33" blockId="39.[117,1475,697,892]" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559714A6DA5CEEFD11" bold="true" box="[117,191,697,718]" pageId="39" pageNumber="38">Table 4</emphasis>
. The diploid number, the postpachytene configuration, the percentage of the total chromosome length of the diploid set (% TCL) and the origin of 
<taxonomicName id="7E9A7AEEFF8710559790A6B55D9AFD33" box="[241,459,726,748]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8BEEDD7FFF8710559790A6B55DD1FD34" box="[241,384,726,747]" italics="true" pageId="39" pageNumber="38">Compsobuthus</emphasis>
species
</taxonomicName>
cytogenetically investigated.
</paragraph>
<paragraph id="B925016DFF8710559714A6905F1EFCA3" blockId="39.[117,1475,697,892]" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559714A6905D01FCD7" box="[117,336,755,776]" italics="true" pageId="39" pageNumber="38">Specimens karyotyped</emphasis>
: 
<taxonomicName id="7E9A7AEEFF8710559603A6905EE3FCD6" authority="(Simon, 1882)" baseAuthorityName="Simon" baseAuthorityYear="1882" box="[354,690,755,777]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="acutecarinatus">
<emphasis id="8BEEDD7FFF8710559603A6905E42FCD7" box="[354,531,755,776]" italics="true" pageId="39" pageNumber="38">C. acutecarinatus</emphasis>
(Simon, 1882)
</taxonomicName>
: Oman, Dhalkut beach, 16.6989°N 53.2214°E, 1♂; E of Aydam, 17.8887°N 53.06640°E, 1♂; W of Qairoon Hairitti, 17.2564°N 54.0220°E, 2♂; Wadi Ash Shuwaymiyyah, 17.9293°N 55.5510°E, 1♂1♀. 
<taxonomicName id="7E9A7AEEFF871055927BA7735D0DFC9D" authority="Levy, Amitai &amp; Shulov, 1973" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="8BEEDD7FFF871055927BA77359D8FCFA" box="[1306,1417,784,805]" italics="true" pageId="39" pageNumber="38">C. arabicus</emphasis>
Levy, Amitai &amp; Shulov, 1973
</taxonomicName>
: Oman, Al Wasil, 22.4914°N 58.7151°E, 2♂; Jabal Bani Jabir, 22.8132°N 59.0589°E, 1♂; E of Aydam, 17.8887°N 53.06640°E, 1♂; W of Wahiba, 22.0299°N 58.1849°E, 1♂. 
<taxonomicName id="7E9A7AEEFF87105595CDA7295F8CFC80" authority="(Kraepelin, 1900)" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[684,989,841,863]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="maindroni">
<emphasis id="8BEEDD7FFF87105595CDA7295F7AFC81" box="[684,811,841,863]" italics="true" pageId="39" pageNumber="38">C. maindroni</emphasis>
(Kraepelin, 1900)
</taxonomicName>
: Oman, Khatm Ha‘ir Asfar, 24.7750°N 55.9146°E, 2♂; Say Plateau, 23.1185°N 57.6311°E, 2♂. 
<taxonomicName id="7E9A7AEEFF8710559544A7055ED9FCA4" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[549,648,870,891]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559544A7055ED9FCA4" box="[549,648,870,891]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<emphasis id="8BEEDD7FFF87105595EFA7055E95FCA4" bold="true" box="[654,708,870,891]" pageId="39" pageNumber="38">
<taxonomicNameLabel id="90DD6004FF87105595EFA7055E95FCA4" box="[654,708,870,891]" pageId="39" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 1♂ holotype.
</paragraph>
</caption>
<paragraph id="B925016DFF8710559710A7F95D67FB6D" blockId="39.[113,772,922,1554]" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559710A7F95CF7FC6D" bold="true" box="[113,166,922,946]" pageId="39" pageNumber="38">Legs</emphasis>
(
<figureCitation id="21A11DE8FF87105597DEA7F95D35FC6D" box="[191,356,922,946]" captionStart="Figures 180–189" captionStartId="36.[117,194,1601,1623]" captionTargetBox="[113,1474,234,1577]" captionTargetId="figure-15@36.[113,1474,234,1590]" captionTargetPageId="36" captionText="Figures 180–189: Figures 180–181, 184–187. Compsobuthus ullrichi sp. n., male holotype, carapace and tergites I–IV (180), posterior coxosternal area and sternites (181), and left legs I–IV, retrolateral aspects (184–187). Figures 182–183, 188–189. C. levyi, male holotype, carapace and tergites I–III (182), posterior coxosternal area and sternites (183), and left legs III–IV, retrolateral aspects (188–189)." figureDoi="http://doi.org/10.5281/zenodo.5741509" httpUri="https://zenodo.org/record/5741509/files/figure.png" pageId="39" pageNumber="38">Figs. 184–187</figureCitation>
). Legs III–IV bear moderate tibial spurs. Retrolateral and prolateral pedal spurs are present on all legs. The tarsomeres bear two rows of macrosetae on the ventral surface and several macrosetae on the other surfaces. Bristlecombs are absent. The femur bears 4 carinae, the patella 4–6 six carinae. The femur and patella bear only solitary macrosetae and are granulated except for external lateral surfaces which are smooth. Tarsal ungues moderately elongated, curved.
</paragraph>
<paragraph id="B925016DFF8710559710A0D95C93FA2D" blockId="39.[113,772,922,1554]" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559710A0D95D0DFB0D" bold="true" box="[113,348,1210,1234]" pageId="39" pageNumber="38">Hemispermatophore</emphasis>
(
<figureCitation id="21A11DE8FF871055960BA0D95E56FB0D" box="[362,519,1210,1234]" captionStart="Figures 160–167" captionStartId="32.[115,192,1926,1948]" captionTargetBox="[127,1468,243,1890]" captionTargetId="figure-15@32.[113,1474,238,1905]" captionTargetPageId="32" captionText="Figures 160–167: Hemispermatophores. Figures 160–163. Compsobuthus turieli sp. n., male holotype, whole hemispermatophore (160), capsule in convex compressed (161), anterior (162), and posterior (163) views. Figures 164–167. C. ullrichi sp. n., male holotype, whole hemispermatophore (164), capsule in posterior (165), convex compressed (166), and anterior (167) views. Scale bars: 1 mm (160, 164), 200 μm (161–163, 165–167)." figureDoi="http://doi.org/10.5281/zenodo.5741503" httpUri="https://zenodo.org/record/5741503/files/figure.png" pageId="39" pageNumber="38">Figs. 164–167</figureCitation>
). Flagelliform, elongate and slender, trunk ca. 6 times length of capsule region. Flagellum separated from external lobe, pars recta ca. 50% of trunk length, broad with anterior lamina, pars reflecta ca. 75% of trunk length, narrow, hyaline. Capsule region with 4 lobes at base of flagellum: posterior lobe longest, triangular, median lobe shortest, apically truncate, anterior lobe acuminate with long thin terminus. Basal lobe strong with broad base and sharp, falcate hook. Left and right hemispermatophores were similar.
</paragraph>
<paragraph id="B925016DFF8710559710A1995DF6F9CD" blockId="39.[113,772,922,1554]" box="[113,423,1530,1554]" pageId="39" pageNumber="38">
<emphasis id="8BEEDD7FFF8710559710A1995D49F9CD" bold="true" box="[113,280,1530,1554]" pageId="39" pageNumber="38">Measurements</emphasis>
. See 
<tableCitation id="F41834D6FF8710559633A1995DF5F9CD" box="[338,420,1530,1554]" captionStart="Table 3" captionStartId="34.[116,172,986,1007]" captionTargetId="graphics-458@34.[119,1471,235,973]" captionTargetPageId="34" captionText="Table 3. Comparative measurements of adults of Compsobuthus ullrichi sp. n., C. levyi and C. kabateki. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/EDE551E5FF8210509715A7B95EC7FBD2" pageId="39" pageNumber="38" tableUuid="EDE551E5FF8210509715A7B95EC7FBD2">Table 3</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F18052E6FF8710559710A2595FF1F9ED" pageId="39" pageNumber="38" type="discussion">
<paragraph id="B925016DFF8710559710A2595FF1F9ED" blockId="39.[113,772,1594,2034]" lastBlockId="39.[816,1475,922,1586]" pageId="39" pageNumber="38">
AFFINITIES. The described characters distinguish 
<taxonomicName id="7E9A7AEEFF87105595F0A2595F52F98D" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[657,771,1594,1618]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF87105595F0A2595F52F98D" box="[657,771,1594,1618]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8710559710A2395CE2F9AE" box="[113,179,1626,1650]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF8710559710A2395CDBF9AD" bold="true" box="[113,138,1626,1650]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF87105597FCA2395CFDF9AD" bold="true" box="[157,172,1626,1650]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
from all other known species of the genus. Among 
<taxonomicName id="7E9A7AEEFF8710559710A2195D3FF94D" box="[113,366,1658,1682]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="8BEEDD7FFF8710559710A2195D43F94D" box="[113,274,1658,1682]" italics="true" pageId="39" pageNumber="38">Compsobuthus</emphasis>
species
</taxonomicName>
known from the region of Middle East and 
<collectingCountry id="C18D41FDFF87105597B6A2F95D4AF96D" box="[215,283,1690,1714]" name="Egypt" pageId="39" pageNumber="38">Egypt</collectingCountry>
, 
<taxonomicName id="7E9A7AEEFF8710559646A2F95DC9F96D" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[295,408,1690,1714]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559646A2F95DC9F96D" box="[295,408,1690,1714]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF87105596C0A2F95D8EF96E" box="[417,479,1690,1714]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF87105596C0A2F95DEBF96D" bold="true" box="[417,442,1690,1714]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF87105596A8A2F95D89F96D" bold="true" box="[457,472,1690,1714]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
is most similar to 
<taxonomicName id="7E9A7AEEFF87105595CEA2F95D41F90D" authority="KovarIk, 2012" authorityName="KovarIk" authorityYear="2012" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF87105595CEA2F95F52F96D" box="[687,771,1690,1714]" italics="true" pageId="39" pageNumber="38">C. levyi</emphasis>
<bibRefCitation id="DD0B7C9CFF8710559710A2D95D41F90D" author="KovarIk" box="[113,272,1722,1748]" firstAuthor="KovarIk" pageId="39" pageNumber="38" pagination="1 - 10" refId="ref15157" refString="KOVARIK, F. 2012. Three new species of Compsobuthus Vachon, 1949 from Yemen, Jordan, Israel, and Somaliland (Scorpiones: Buthidae). Euscorpius, 150: 1 - 10." type="journal article" year="2012">KovařÍk, 2012</bibRefCitation>
</taxonomicName>
, from 
<collectingCountry id="C18D41FDFF8710559634A2D95DCCF90D" box="[341,413,1722,1746]" name="Jordan" pageId="39" pageNumber="38">Jordan</collectingCountry>
(type locality: near Qasr Burqu). The two species can be separated by several characters: males of 
<taxonomicName id="7E9A7AEEFF87105597EFA2995C8EF8CD" authorityName="KovarIk" authorityYear="2012" box="[142,223,1786,1810]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF87105597EFA2995C8EF8CD" box="[142,223,1786,1810]" italics="true" pageId="39" pageNumber="38">C. levyi</emphasis>
</taxonomicName>
have longer and narrower pedipalp and metasomal segments (see 
<tableCitation id="F41834D6FF8710559672A3795D35F8ED" box="[275,356,1818,1842]" captionStart="Table 1" captionStartId="6.[113,169,998,1019]" captionTargetId="graphics-45@6.[115,1476,237,975]" captionTargetPageId="6" captionText="Table 1. Comparative measurements of adults of Compsobuthus arabicus and C. turieli sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." httpUri="http://table.plazi.org/id/EDE551E5FFA610749710A7855E70FBC6" pageId="39" pageNumber="38" tableUuid="EDE551E5FFA610749710A7855E70FBC6">Table 1</tableCitation>
and 
<figureCitation id="21A11DE8FF87105596FCA3795E6EF8ED" box="[413,575,1818,1842]" captionStart="Figures 172–179" captionStartId="35.[113,190,1958,1980]" captionTargetBox="[117,1468,265,1932]" captionTargetId="figure-8@35.[113,1474,238,1938]" captionTargetPageId="35" captionText="Figures 172–179: Figures 172, 174–176. Compsobuthus levyi, male holotype, telson lateral (172), metasoma and telson lateral (174), dorsal (175), and ventral (176) views. Figures 173, 177–179. C. ullrichi sp. n., male holotype, telson lateral (173), metasoma and telson lateral (177), dorsal (178), and ventral (179) views. Scale bar: 10 mm (174–179)." figureDoi="http://doi.org/10.5281/zenodo.5741507" httpUri="https://zenodo.org/record/5741507/files/figure.png" pageId="39" pageNumber="38">Figs. 174–179</figureCitation>
). Pedipalp patella L/W ratio is 
<quantity id="7E62AC88FF871055979EA3595D1DF88D" box="[255,332,1850,1874]" metricMagnitude="-2" metricUnit="m" metricValue="7.7724" pageId="39" pageNumber="38" unit="in" value="3.06">3.06 in</quantity>
male 
<typeStatus id="6621BFCFFF87105596F0A3595DA1F88D" box="[401,496,1850,1874]" pageId="39" pageNumber="38" type="holotype">holotype</typeStatus>
<taxonomicName id="7E9A7AEEFF8710559698A3595E38F88D" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[505,617,1850,1874]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="8BEEDD7FFF8710559698A3595E38F88D" box="[505,617,1850,1874]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
, 
<quantity id="7E62AC88FF8710559518A3595F52F88D" box="[633,771,1850,1874]" metricMagnitude="-2" metricUnit="m" metricValue="8.2677" metricValueMax="8.2804" metricValueMin="8.255" pageId="39" pageNumber="38" unit="in" value="3.255" valueMax="3.26" valueMin="3.25">3.25–3.26 in</quantity>
males of 
<taxonomicName id="7E9A7AEEFF87105597BCA3395D62F8AD" authorityName="KovarIk" authorityYear="2012" box="[221,307,1882,1906]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF87105597BCA3395D62F8AD" box="[221,307,1882,1906]" italics="true" pageId="39" pageNumber="38">C. levyi</emphasis>
</taxonomicName>
; metasoma I L/W ratio is 
<quantity id="7E62AC88FF8710559506A3395E95F8AD" box="[615,708,1882,1906]" metricMagnitude="-2" metricUnit="m" metricValue="3.49758" pageId="39" pageNumber="38" unit="in" value="1.377">1.377 in</quantity>
male 
<taxonomicName id="7E9A7AEEFF8710559710A3195CB4F84D" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[113,229,1914,1938]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="8BEEDD7FFF8710559710A3195CB4F84D" box="[113,229,1914,1938]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
, 
<quantity id="7E62AC88FF8710559796A3195DD5F84D" box="[247,388,1914,1938]" metricMagnitude="-2" metricUnit="m" metricValue="3.8735" metricValueMax="3.9116" metricValueMin="3.8354" pageId="39" pageNumber="38" unit="in" value="1.525" valueMax="1.54" valueMin="1.51">1.51–1.54 in</quantity>
males of 
<taxonomicName id="7E9A7AEEFF871055969BA3195E01F84D" authorityName="KovarIk" authorityYear="2012" box="[506,592,1914,1938]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="levyi">
<emphasis id="8BEEDD7FFF871055969BA3195E01F84D" box="[506,592,1914,1938]" italics="true" pageId="39" pageNumber="38">C. levyi</emphasis>
</taxonomicName>
; and metasoma IV L/W ratio is 
<quantity id="7E62AC88FF8710559643A3F95D3FF86D" box="[290,366,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="5.6896" pageId="39" pageNumber="38" unit="in" value="2.24">2.24 in</quantity>
male 
<taxonomicName id="7E9A7AEEFF87105596D2A3F95E72F86D" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[435,547,1946,1970]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi">
<emphasis id="8BEEDD7FFF87105596D2A3F95E72F86D" box="[435,547,1946,1970]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
, 
<quantity id="7E62AC88FF8710559553A3F95EEDF86D" box="[562,700,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="6.2738" metricValueMax="6.2992" metricValueMin="6.2484" pageId="39" pageNumber="38" unit="in" value="2.4699999999999998" valueMax="2.48" valueMin="2.46">2.46–2.48 in</quantity>
males of 
<emphasis id="8BEEDD7FFF87105597F2A3D95CA2F80D" box="[147,243,1978,2002]" italics="true" pageId="39" pageNumber="38">
<taxonomicName id="7E9A7AEEFF87105597F2A3D95CBEF80D" authorityName="KovarIk" authorityYear="2012" box="[147,239,1978,2002]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="levyi">C. levyi</taxonomicName>
.
</emphasis>
In this region are found five other species of 
<taxonomicName id="7E9A7AEEFF8710559710A3B95D43F82D" authorityName="Vachon" authorityYear="1949" box="[113,274,2010,2034]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="8BEEDD7FFF8710559710A3B95D43F82D" box="[113,274,2010,2034]" italics="true" pageId="39" pageNumber="38">Compsobuthus</emphasis>
</taxonomicName>
belonging to the ‘ 
<emphasis id="8BEEDD7FFF8710559683A3B85E66F82D" box="[482,567,2011,2034]" italics="true" pageId="39" pageNumber="38">werneri</emphasis>
’ group of Levy &amp; Amitai (1980). From these, 
<taxonomicName id="7E9A7AEEFF8710559306A7F95889FC6D" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[1127,1240,922,946]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559306A7F95889FC6D" box="[1127,1240,922,946]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8710559380A7F95971FC6E" box="[1249,1312,922,946]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF8710559380A7F958ABFC6D" bold="true" box="[1249,1274,922,946]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF871055926BA7F95948FC6D" bold="true" box="[1290,1305,922,946]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
can be clearly differentiated by metasomal segment I being narrower than that of other species: 
<taxonomicName id="7E9A7AEEFF8710559343A7B95F37FBCD" authority="Levy, Amitai &amp; Shulov, 1973" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="carmelitis">
<emphasis id="8BEEDD7FFF8710559343A7B958E0FC2D" box="[1058,1201,986,1010]" italics="true" pageId="39" pageNumber="38">C. carmelitis</emphasis>
Levy, Amitai &amp; Shulov, 1973
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF87105594D1A7995FBEFBCD" box="[944,1007,1018,1042]" name="Israel" pageId="39" pageNumber="38">Israel</collectingCountry>
, and probably also 
<collectingCountry id="C18D41FDFF87105593B1A7995949FBCD" box="[1232,1304,1018,1042]" name="Jordan" pageId="39" pageNumber="38">Jordan</collectingCountry>
and 
<collectingCountry id="C18D41FDFF8710559235A79959C1FBCD" box="[1364,1424,1018,1042]" name="Syria" pageId="39" pageNumber="38">Syria</collectingCountry>
, has metasoma I of males wider than long, or as wide as long; 
<taxonomicName id="7E9A7AEEFF87105592C8A07958CDFB8D" authority="Lourenco et al., 2009" authorityName="Lourenco" authorityYear="2009" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="egyptiensis">
<emphasis id="8BEEDD7FFF87105592C8A0795FF8FB8D" italics="true" pageId="39" pageNumber="38">C. egyptiensis</emphasis>
Lourenco et al., 2009
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF8710559381A0595970FB8D" box="[1248,1313,1082,1106]" name="Egypt" pageId="39" pageNumber="38">Egypt</collectingCountry>
(type locality: NW of Siwa) has metasoma I L/W ratio 
<quantity id="7E62AC88FF8710559397A0395923FBAD" box="[1270,1394,1114,1138]" metricMagnitude="-2" metricUnit="m" metricValue="2.8829" metricValueMax="2.9718" metricValueMin="2.794" pageId="39" pageNumber="38" unit="in" value="1.135" valueMax="1.17" valueMin="1.1">1.1–1.17 in</quantity>
males; and 
<taxonomicName id="7E9A7AEEFF8710559407A01958CAFB4D" authority="KovarIk, 2003" authorityName="KovarIk" authorityYear="2003" box="[870,1179,1146,1172]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="kabateki">
<emphasis id="8BEEDD7FFF8710559407A0195FB8FB4D" box="[870,1001,1146,1170]" italics="true" pageId="39" pageNumber="38">C. kabateki</emphasis>
<bibRefCitation id="DD0B7C9CFF8710559496A01958CAFB4D" author="KovarIk" box="[1015,1179,1146,1172]" firstAuthor="KovarIk" pageId="39" pageNumber="38" pagination="87 - 112" refId="ref15081" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">KovařÍk, 2003</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF871055938DA019597CFB4D" box="[1260,1325,1146,1170]" name="Egypt" pageId="39" pageNumber="38">Egypt</collectingCountry>
(
<collectingRegion id="7B5ECF8FFF8710559222A01959D8FB4D" box="[1347,1417,1146,1170]" country="Egypt" name="Al Uqsur" pageId="39" pageNumber="38">Luxor</collectingRegion>
) has metasoma I L/W ratio 
<quantity id="7E62AC88FF871055934BA0F95826FB6D" box="[1066,1143,1178,1202]" metricMagnitude="-2" metricUnit="m" metricValue="2.7178" pageId="39" pageNumber="38" unit="in" value="1.07">1.07 in</quantity>
males. In contrast, 
<taxonomicName id="7E9A7AEEFF8710559230A0F95993FB6D" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[1361,1474,1178,1202]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559230A0F95993FB6D" box="[1361,1474,1178,1202]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8710559451A0D95F29FB0E" box="[816,888,1210,1234]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF8710559451A0D95F18FB0D" bold="true" box="[816,841,1210,1234]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF8710559403A0D95F20FB0D" bold="true" box="[866,881,1210,1234]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
has metasoma I L/W ratio 1.37. 
<taxonomicName id="7E9A7AEEFF8710559240A0D958BEFB2D" authority="Levy, Amitai &amp; Shulov, 1973" authorityName="Levy, Amitai &amp; Shulov" authorityYear="1973" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="longipalpis">
<emphasis id="8BEEDD7FFF8710559240A0D95FFAFB2D" italics="true" pageId="39" pageNumber="38">Compsobuthus longipalpis</emphasis>
Levy, Amitai &amp; Shulov, 1973
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF8710559251A0B95925FB2D" box="[1328,1396,1242,1266]" name="Egypt" pageId="39" pageNumber="38">Egypt</collectingCountry>
, 
<collectingCountry id="C18D41FDFF871055921EA0B959EFFB2D" box="[1407,1470,1242,1266]" name="Israel" pageId="39" pageNumber="38">Israel</collectingCountry>
, 
<collectingCountry id="C18D41FDFF8710559451A0995F2AFACD" box="[816,891,1274,1298]" name="Jordan" pageId="39" pageNumber="38">Jordan</collectingCountry>
, 
<collectingCountry id="C18D41FDFF87105594E6A0995846FACD" box="[903,1047,1274,1298]" name="Saudi Arabia" pageId="39" pageNumber="38">Saudi Arabia</collectingCountry>
, and 
<collectingCountry id="C18D41FDFF871055932EA09958DAFACD" box="[1103,1163,1274,1298]" name="Syria" pageId="39" pageNumber="38">Syria</collectingCountry>
, differs in having 14 rows of granules on the pedipalp movable finger and a total length of 
<quantity id="7E62AC88FF8710559431A1595F97FA8E" box="[848,966,1338,1362]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="39" pageNumber="38" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 mm</quantity>
(in 
<taxonomicName id="7E9A7AEEFF8710559497A1595839FA8D" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[1014,1128,1338,1362]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559497A1595839FA8D" box="[1014,1128,1338,1362]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF8710559310A15958E0FA8E" box="[1137,1201,1338,1362]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF8710559310A15958DBFA8D" bold="true" box="[1137,1162,1338,1362]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF87105593FBA15958F8FA8D" bold="true" box="[1178,1193,1338,1362]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
there are 10–11 rows of granules on the pedipalp movable finger and total length is &lt;
<quantity id="7E62AC88FF871055942AA1195FF1FA4D" box="[843,928,1402,1426]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="38" unit="mm" value="30.0">30 mm</quantity>
). 
<taxonomicName id="7E9A7AEEFF87105594D7A1195962FA4D" authority="Vachon, 1949" authorityName="Vachon" authorityYear="1949" box="[950,1331,1402,1426]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="schmiedeknechti">
<emphasis id="8BEEDD7FFF87105594D7A11958DFFA4D" box="[950,1166,1402,1426]" italics="true" pageId="39" pageNumber="38">C. schmiedeknechti</emphasis>
Vachon, 1949
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF871055921EA11959EFFA4D" box="[1407,1470,1402,1426]" name="Israel" pageId="39" pageNumber="38">Israel</collectingCountry>
, 
<collectingCountry id="C18D41FDFF8710559451A1F95F2AFA6D" box="[816,891,1434,1458]" name="Jordan" pageId="39" pageNumber="38">Jordan</collectingCountry>
, 
<collectingCountry id="C18D41FDFF87105594E9A1F95FBBFA6D" box="[904,1002,1434,1458]" name="Lebanon" pageId="39" pageNumber="38">Lebanon</collectingCountry>
, 
<collectingCountry id="C18D41FDFF8710559497A1F95863FA6D" box="[1014,1074,1434,1458]" name="Syria" pageId="39" pageNumber="38">Syria</collectingCountry>
, and 
<collectingCountry id="C18D41FDFF871055930CA1F958EBFA6D" box="[1133,1210,1434,1458]" name="Turkey" pageId="39" pageNumber="38">Turkey</collectingCountry>
differs in having 15–18 pectinal teeth in males; 
<taxonomicName id="7E9A7AEEFF871055932CA1D959D1FA0D" authority="KovarIk, 2003" authorityName="KovarIk" authorityYear="2003" box="[1101,1408,1466,1492]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="kabateki">
<emphasis id="8BEEDD7FFF871055932CA1D9589EFA0D" box="[1101,1231,1466,1490]" italics="true" pageId="39" pageNumber="38">C. kabateki</emphasis>
<bibRefCitation id="DD0B7C9CFF87105593BCA1D959D1FA0D" author="KovarIk" box="[1245,1408,1466,1492]" firstAuthor="KovarIk" pageId="39" pageNumber="38" pagination="87 - 112" refId="ref15081" refString="KOVARIK, F. 2003. Eight new species of Compsobuthus Vachon, 1949 from Africa and Asia (Scorpiones: Buthidae). Serket, 8 (3): 87 - 112." type="journal article" year="2003">KovařÍk, 2003</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="C18D41FDFF8710559451A1B95F20FA2D" box="[816,881,1498,1522]" name="Egypt" pageId="39" pageNumber="38">Egypt</collectingCountry>
(
<collectingRegion id="7B5ECF8FFF871055941EA1B95F94FA2D" box="[895,965,1498,1522]" country="Egypt" name="Al Uqsur" pageId="39" pageNumber="38">Luxor</collectingRegion>
) differs in having 19 pectinal teeth in males (in 
<taxonomicName id="7E9A7AEEFF8710559451A1995FF3F9CD" authority="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský, 2020" authorityName="Kovařík &amp; Lowe &amp; Stockmann &amp; Šťáhlavský" authorityYear="2020" box="[816,930,1530,1554]" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="39" pageNumber="38" phylum="Arthropoda" rank="species" species="ullrichi" status="sp. nov.">
<emphasis id="8BEEDD7FFF8710559451A1995FF3F9CD" box="[816,930,1530,1554]" italics="true" pageId="39" pageNumber="38">C. ullrichi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DD6004FF87105594CAA1995FBAF9CE" box="[939,1003,1530,1554]" pageId="39" pageNumber="38" rank="species">
<emphasis id="8BEEDD7FFF87105594CAA1995F95F9CD" bold="true" box="[939,964,1530,1554]" pageId="39" pageNumber="38">sp</emphasis>
. 
<emphasis id="8BEEDD7FFF87105594B5A1995FB2F9CD" bold="true" box="[980,995,1530,1554]" pageId="39" pageNumber="38">n</emphasis>
.
</taxonomicNameLabel>
there are 22–23 pectinal teeth in the male 
<typeStatus id="6621BFCFFF8710559451A2795FC5F9ED" box="[816,916,1562,1586]" pageId="39" pageNumber="38" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
</treatment>
</document>