treatments-xml/data/1A/32/36/1A323678620F223E09D9FA40FC9D6DC1.xml
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<document id="702776A591AD146DEF1DE09097FC786A" ID-CLB-Dataset="8243" ID-DOI="10.11646/zootaxa.4923.1.1" ID-GBIF-Dataset="0a027439-c601-4533-bd03-f0e5d4442801" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496757" ID-ZooBank="41D057A7-F855-47C2-A598-8000CFD83A5B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612332986149" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2021" docId="1A323678620F223E09D9FA40FC9D6DC1" docLanguage="en" docName="zootaxa.4923.1.1.pdf" docOrigin="Zootaxa 4923 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lepanus cameroni Gunter &amp; Weir 2021, new species" docType="treatment" docVersion="11" lastPageNumber="35" masterDocId="E60B4E00622E221C094EFFBDFFC26960" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="34" updateTime="1698926692322" updateUser="ExternalLinkService">
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<mods:title id="146547DEEDC529B62B9C2DBC2F7AC3AC">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): review of the L. ustulatus, L. storeyi, and L. nitidus species groups and description of eight new species</mods:title>
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<mods:namePart id="678F6E4B943BA3BFB7A5C53B368E5FE8">Gunter, Nicole L.</mods:namePart>
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<mods:namePart id="E1E3B47749517C4E3D667FFC53D9AC1E">Weir, Thomas A.</mods:namePart>
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<treatment id="1A323678620F223E09D9FA40FC9D6DC1" ID-DOI="http://doi.org/10.5281/zenodo.4537941" ID-GBIF-Taxon="177324112" ID-Zenodo-Dep="4537941" LSID="urn:lsid:plazi:treatment:1A323678620F223E09D9FA40FC9D6DC1" httpUri="http://treatment.plazi.org/id/1A323678620F223E09D9FA40FC9D6DC1" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
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<taxonomicName id="559BFCED620F223D09D9FA40FDFD6F77" ID-CoL="7WWVN" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2021" box="[151,575,1533,1559]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="cameroni" status="sp. nov.">
<emphasis id="A0EF5B7C620F223D09D9FA40FEBB6F77" bold="true" box="[151,377,1533,1559]" italics="true" pageId="33" pageNumber="34">Lepanus cameroni</emphasis>
Gunter &amp; Weir
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,
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(
<figureCitation id="0AA09BEB620F223D09D1F99CFEDE6F5C" box="[159,284,1569,1596]" captionStart="FIGURE 2" captionStartId="28.[151,250,1798,1822]" captionTargetBox="[255,1320,187,1768]" captionTargetId="figure-26@28.[255,1332,181,1774]" captionTargetPageId="28" captionText="FIGURE 2. Some diagnostic characters of the Lepanus ustulatus species group. A, Dorsal view of male protibia of Lepanus ustulatus; B, dorsal view of female protibia of Lepanus ustulatus; C, dorsal view of male protibia of Lepanus globulus; D, male metatibia of Lepanus ustulatus; E, female metatibia of Lepanus ustulatus; F, male metatibia of Lepanus lemannae; G, lateral view of male protibia of Lepanus cameroni; H, lateral view of male protibia of Lepanus globulus; I, lateral view of male protibia of Lepanus cardwellensis; J, head of Lepanus cameroni showing eye canthus not dividing the eye; K, posterolateral view of pronotum of Lepanus globulus showing corners distinct; L, head of Lepanus globulus showing eye canthus almost dividing the eye; M, posterolateral view of pronotum of Lepanus cardwellensis showing corners rounded." figureDoi="http://doi.org/10.5281/zenodo.4496761" httpUri="https://zenodo.org/record/4496761/files/figure.png" pageId="33" pageNumber="34">Figs. 2G, J</figureCitation>
;
<figureCitation id="0AA09BEB620F223D0867F99CFE8C6F5B" box="[297,334,1569,1595]" captionStart="FIGURE 3" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[185,1384,196,1902]" captionTargetId="figure-19@29.[183,1404,181,1919]" captionTargetPageId="29" captionText="FIGURE 3. Dorsal habitus of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4496763" httpUri="https://zenodo.org/record/4496763/files/figure.png" pageId="33" pageNumber="34">3A</figureCitation>
;
<figureCitation id="0AA09BEB620F223D0814F99CFEBD6F5B" box="[346,383,1569,1595]" captionStart="FIGURE 4" captionStartId="30.[151,250,1263,1287]" captionTargetBox="[157,1422,197,1228]" captionTargetId="figure-26@30.[151,1436,181,1239]" captionTargetPageId="30" captionText="FIGURE 4. Pygidial configuration of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496765" httpUri="https://zenodo.org/record/4496765/files/figure.png" pageId="33" pageNumber="34">4A</figureCitation>
;
<figureCitation id="0AA09BEB620F223D08C5F99CFE6D6F5C" box="[395,431,1569,1596]" captionStart="FIGURE 5" captionStartId="31.[151,250,1943,1967]" captionTargetBox="[217,1358,189,1913]" captionTargetId="figure-19@31.[213,1373,181,1919]" captionTargetPageId="31" captionText="FIGURE 5. Left, dorsal and right view of aedeagi of species within the Lepanus ustulatus species group. A, Lepanus cameroni; B, Lepanus cardwellensis; C, Lepanus globulus; D, Lepanus lemannae; E, Lepanus ustulatus. All specimens in comparative scale. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4496767" httpUri="https://zenodo.org/record/4496767/files/figure.png" pageId="33" pageNumber="34">5A</figureCitation>
;
<figureCitation id="0AA09BEB620F223D08F2F99CFE1C6F5C" box="[444,478,1569,1596]" captionStart="FIGURE 6" captionStartId="32.[151,250,999,1023]" captionTargetBox="[154,1436,187,969]" captionTargetId="figure-26@32.[151,1436,181,975]" captionTargetPageId="32" captionText="FIGURE 6. A, Map of known records of Lepanus ustulatus; B, predicted distribution of Lepanus ustulatus; C, map of known records of Lepanus cameroni." figureDoi="http://doi.org/10.5281/zenodo.4496769" httpUri="https://zenodo.org/record/4496769/files/figure.png" pageId="33" pageNumber="34">6C</figureCitation>
)
</paragraph>
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<emphasis id="A0EF5B7C620F223D09D9F9D4FEE16FE3" bold="true" box="[151,291,1641,1667]" pageId="33" pageNumber="34">Type series.</emphasis>
<materialsCitation id="22F38D33620F223D0864F9D4FA8C6FC7" ID-GBIF-Occurrence="3034608809" collectingDate="1992-10-24" collectionCode="ANIC" collectorName="T. Weir &amp; P. Zborowski " latitude="-12.45" location="West Claudie River" longLatPrecision="776" longitude="143.14" pageId="33" pageNumber="34" specimenCount="1" typeStatus="holotype">
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<typeStatus id="4D2039CC620F223D0864F9D4FE596FE3" box="[298,411,1641,1667]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
: ♂
</emphasis>
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QLD
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,
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,
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closed forest” / “
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Database 25 023716” / “
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8, det
<collectorName id="3F6EE2B8620F223D0D5AF930FB406FC7" box="[1044,1154,1677,1703]" pageId="33" pageNumber="34">T.A. Weir</collectorName>
2018” (in
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)
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.
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<typeStatus id="4D2039CC620F223D0C17F930FF176FAB" pageId="33" pageNumber="34" type="paratype">Paratypes</typeStatus>
(2x ♂, 3x
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):
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ENE Mount Tozer
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,
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,
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,
<collectorName id="3F6EE2B8620F223D0DC4F90CFAD16FAB" box="[1162,1299,1713,1739]" pageId="33" pageNumber="34">J.C. Cardale</collectorName>
(25-058143
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♂ in
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);
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,
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<collectorName id="3F6EE2B8620F223D0DF0F968FAA66F8F" box="[1214,1380,1749,1775]" pageId="33" pageNumber="34">G.B. Monteith</collectorName>
(25- 060831
<specimenCount id="849D4CE7620F223D09BFF944FE4B6E74" box="[241,393,1785,1812]" pageId="33" pageNumber="34" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620F223D08E7F944FE2F6E73" box="[425,493,1785,1811]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="34">ANIC</collectionCode>
also as COL1463)
</materialsCitation>
;
<materialsCitation id="22F38D33620F223D0B84F944FCA16E57" ID-GBIF-Occurrence="3034608988" collectingDate="1971-06-01" collectingDateMax="1986-06-30" collectingDateMin="1971-06-01" collectionCode="ANIC, QM" collectorName="T. A. Weir &amp; A. Calder" latitude="-12.75" location="Claudie River" longLatPrecision="21" longitude="143.21666" pageId="33" pageNumber="34" specimenCount="3" specimenCount-female="2" stateProvince="York" typeStatus="paratype">
<locationDeviation id="83969E4C620F223D0B84F944FC116E74" box="[714,979,1785,1812]" location="Mount Tozer" pageId="33" pageNumber="34">
<quantity id="55632A8B620F223D0B84F944FCC06E73" box="[714,770,1785,1811]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="33" pageNumber="34" unit="km" value="6.0">6 km</quantity>
ENE Mount Tozer
</locationDeviation>
[
<geoCoordinate id="F7AFE1A9620F223D0AAEF944FBAA6E73" box="[992,1128,1785,1811]" degrees="12" direction="south" minutes="44" orientation="latitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="-12.733334">12°4400”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620F223D0D3DF944FACE6E74" box="[1139,1292,1785,1812]" degrees="143" direction="east" minutes="16" orientation="longitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="143.26666">143°1600”E</geoCoordinate>
],
<date id="E625A1AE620F223D0C53F944FA5A6E73" box="[1309,1432,1785,1811]" pageId="33" pageNumber="34" value="1986-06-30">
<collectingDate id="F6615846620F223D0C53F944FA5A6E73" box="[1309,1432,1785,1811]" pageId="33" pageNumber="34" value="1986-06-30">30.vi.1986</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620F223D09D9F8A0FECB6E57" box="[151,265,1821,1847]" pageId="33" pageNumber="34">T.A. Weir</collectorName>
&amp;
<collectorName id="3F6EE2B8620F223D087AF8A0FE646E57" box="[308,422,1821,1847]" pageId="33" pageNumber="34">A. Calder</collectorName>
(25-058144
<specimenCount id="849D4CE7620F223D0B71F8A0FD206E58" box="[575,738,1821,1848]" pageId="33" pageNumber="34" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620F223D0A43F8A0FC946E57" box="[781,854,1821,1847]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="34">ANIC</collectionCode>
);
</materialsCitation>
<materialsCitation id="22F38D33620F223D0A20F8A3FBEB6E3B" ID-GBIF-Occurrence="3034608397" collectingDate="1971-06-01" collectingDateMax="1986-06-30" collectingDateMin="1971-06-01" collectionCode="ANIC, QM" collectorName="G. B. Monteith" latitude="-12.75" location="Claudie River" longLatPrecision="21" longitude="143.21666" pageId="33" pageNumber="34" specimenCount="3" specimenCount-female="2" stateProvince="York" typeStatus="paratype">
Iron Range, Cape
<collectingRegion id="505F498C620F223D0D08F8A0FBBF6E57" box="[1094,1149,1821,1847]" country="United Kingdom" name="York" pageId="33" pageNumber="34">York</collectingRegion>
Peninsula, [
<geoCoordinate id="F7AFE1A9620F223D0C41F8A0FA556E57" box="[1295,1431,1821,1847]" degrees="12" direction="south" minutes="44" orientation="latitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="-12.733334">12°4400”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620F223D09D9F8FCFEF26E3C" box="[151,304,1857,1884]" degrees="143" direction="east" minutes="17" orientation="longitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="143.28334">143°1700”E</geoCoordinate>
],
<date id="E625A1AE620F223D0809F8FCFE0D6E3B" box="[327,463,1857,1883]" pageId="33" pageNumber="34" value="1971-06-01" valueMax="1971-06-09" valueMin="1971-06-01">
<collectingDate id="F6615846620F223D0809F8FCFE0D6E3B" box="[327,463,1857,1883]" pageId="33" pageNumber="34" value="1971-06-01" valueMax="1971-06-09" valueMin="1971-06-01">19.vi.1971</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620F223D0890F8FCFD476E3B" box="[478,645,1857,1883]" pageId="33" pageNumber="34">G.B. Monteith</collectorName>
(25-058468
<specimenCount id="849D4CE7620F223D0A53F8FCFC626E3B" box="[797,928,1857,1883]" pageId="33" pageNumber="34" type="generic">1 specimen</specimenCount>
♂ in
<collectionCode id="F48A1FAB620F223D0AA2F8FCFBDC6E3B" box="[1004,1054,1857,1883]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="33" pageNumber="34" type="Museum">QM</collectionCode>
);
</materialsCitation>
<materialsCitation id="22F38D33620F223D0D7BF8FCFB1A6E1F" ID-GBIF-Occurrence="3034608430" collectingDate="1971-06-01" collectingDateMax="1986-06-30" collectingDateMin="1971-06-01" collectionCode="ANIC, QM" collectorName="J. F. Lawrence" latitude="-12.75" location="Claudie River" longLatPrecision="21" longitude="143.21666" pageId="33" pageNumber="34" specimenCount="3" specimenCount-female="2" stateProvince="York" typeStatus="paratype">
<location id="9744D1B5620F223D0D7BF8FCFB176E3B" LSID="urn:lsid:plazi:treatment:1A323678620F223E09D9FA40FC9D6DC1:9744D1B5620F223D0D7BF8FCFB176E3B" box="[1077,1237,1857,1883]" latitude="-12.75" longLatPrecision="21" longitude="143.21666" name="Claudie River" pageId="33" pageNumber="34" stateProvince="York">Claudie River</location>
near Iron Range [
<geoCoordinate id="F7AFE1A9620F223D09D0F8D8FEEA6EE0" box="[158,296,1893,1920]" degrees="12" direction="south" minutes="45" orientation="latitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="-12.75">12°4500”S</geoCoordinate>
,
<geoCoordinate id="F7AFE1A9620F223D087DF8D8FE096EE0" box="[307,459,1893,1920]" degrees="143" direction="east" minutes="13" orientation="longitude" pageId="33" pageNumber="34" precision="15" seconds="00" value="143.21666">143°1300”E</geoCoordinate>
],
<date id="E625A1AE620F223D0890F8D8FD486EE0" box="[478,650,1893,1920]" pageId="33" pageNumber="34" value="1978-07-19" valueMax="1978-07-25" valueMin="1978-07-19">
<collectingDate id="F6615846620F223D0890F8D8FD486EE0" box="[478,650,1893,1920]" pageId="33" pageNumber="34" value="1978-07-19" valueMax="1978-07-25" valueMin="1978-07-19">1925.vii.1978</collectingDate>
</date>
,
<collectorName id="3F6EE2B8620F223D0BD8F8DBFCF66EE0" box="[662,820,1894,1920]" pageId="33" pageNumber="34">J.F. Lawrence</collectorName>
, (25-057894
<specimenCount id="849D4CE7620F223D0A87F8D8FBA66EE0" box="[969,1124,1893,1920]" pageId="33" pageNumber="34" type="female">1 specimen ♀</specimenCount>
in
<collectionCode id="F48A1FAB620F223D0DC8F8D8FB126E1F" box="[1158,1232,1893,1919]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="33" pageNumber="34">ANIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620F223E0989F834FB846AE1" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="9224876E620F223D0989F834FBAA6E8C" blockId="33.[151,1437,1641,2028]" pageId="33" pageNumber="34">
<emphasis id="A0EF5B7C620F223D0989F834FE9B6EC3" bold="true" box="[199,345,1929,1955]" pageId="33" pageNumber="34">Description.</emphasis>
Head and thorax dark brown to black, elytra predominately light brown to dark brown with humeri and apical edges reddish brown. Antennal clubs with apical antennomere (3rd) white, basal 2 antennomeres (1st, 2nd) light grey. Dorsal surfaces nitid, with extremely fine setae (often rubbed off).
</paragraph>
<paragraph id="9224876E620C223E0989FF2AFB3B69D1" blockId="34.[151,1437,150,1185]" box="[199,1273,150,177]" pageId="34" pageNumber="35">
Total length:
<quantity id="55632A8B620C223E0812FF2BFE2F69D1" box="[348,493,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="34" pageNumber="35" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">4.5 4.7 mm</quantity>
.
<typeStatus id="4D2039CC620C223E08B6FF2AFDA269D1" box="[504,608,151,177]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
measurements: Total length
<quantity id="55632A8B620C223E0AECFF2AFC3969D1" box="[930,1019,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="34" pageNumber="35" unit="mm" value="4.6">4.6 mm</quantity>
; elytral width
<quantity id="55632A8B620C223E0DD2FF2AFB3769D1" box="[1180,1269,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="34" pageNumber="35" unit="mm" value="2.9">2.9 mm</quantity>
.
</paragraph>
<paragraph id="9224876E620C223E0989FF06FBD0687D" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">
<emphasis id="A0EF5B7C620C223E0989FF06FECA69B5" box="[199,264,187,213]" italics="true" pageId="34" pageNumber="35">Male.</emphasis>
Head: Width to length ratio 37: 25. Surface smooth and nitid, very finely punctate but clypeal area with a few very fine reticulations. Dorsal part of eyes wide, separated by an interocular space approximately 1011 times eye width (53: 5); eye canthus not dividing eye, leaving a gap of about 4 facets.
</paragraph>
<paragraph id="9224876E620C223E0989FE9AFEAF68E9" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">Prothorax: Pronotum with posterolateral angles distinct, lateral margins bordered. Pronotal surface smooth, nitid, very finely punctate. Hypomeral surface very finely reticulate, hypomeral stria absent. Pronotum width to length ratio 63: 40.</paragraph>
<paragraph id="9224876E620C223E0989FE2EFCD36895" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">Elytra: Surface smooth, nitid, striae superficial, impunctate. Stria 6 extending to the elytral base, and stria 7 not extending to the elytral base. 8th stria with a few punctures where it curves inwards posteriorly. Ratio of length of the elytra along suture to maximum elytral width 70: 77.</paragraph>
<paragraph id="9224876E620C223E0989FE42FEAC6BE5" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">Legs: Protibia with a distinct, roughly crenulate ridge on the underside. Protibia with 2 teeth on the outer edge, front edge truncate, apical digit broad with a blunt point, apical spur absent. Mesotibia with a brush of setae apically on the inner side. Metatibia with inner edge nearly straight and finely crenulate. Basal metatarsomere with slight lobe on inner edge.</paragraph>
<paragraph id="9224876E620C223E0989FD32FAAB6BAD" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">Abdomen: Pygidium with a transverse sinuate fold across the base that is curved and not impressed medially. Pygidial disc with a large flat, reticulate central area that occupies more than half the disc. Ventrite 6 punctate.</paragraph>
<paragraph id="9224876E620C223E0989FD6AFA9F6A59" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">Pterothorax: Mesometaventral suture slightly arched. Medial lobe of metaventrite smooth, nitid, virtually impunctate, broadly margined beside mesocoxae. Lateral lobe of metaventrite finely punctate, setose. Mesoventrite smooth. nitid, and notched anteriorly. Mesepimeron very finely reticulate. Metanepisternum finely reticulate.</paragraph>
<paragraph id="9224876E620C223E0989FCFEFB846AE1" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">
<emphasis id="A0EF5B7C620C223E0989FCFEFEE16A3D" box="[199,291,835,861]" italics="true" pageId="34" pageNumber="35">Female.</emphasis>
Protibia with front edge truncate and apical spur arising directly from front edge. Apical digit absent. Metatibia with inner edge not crenulate. Mesotibia without a brush of setae apically.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620C223E0989FC36FD596AA9" pageId="34" pageNumber="35" type="etymology">
<paragraph id="9224876E620C223E0989FC36FD596AA9" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">
<emphasis id="A0EF5B7C620C223E0989FC36FE8C6AC5" bold="true" box="[199,334,907,933]" pageId="34" pageNumber="35">Etymology.</emphasis>
This species is named for Stephen Cameron, Purdue University, for his mentorship and contribution to many of our Australian beetle projects.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620C223E0989FC6FFD756D71" pageId="34" pageNumber="35" type="distribution">
<paragraph id="9224876E620C223E0989FC6FFD756D71" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">
<emphasis id="A0EF5B7C620C223E0989FC6FFEA36A8C" bold="true" box="[199,353,978,1004]" pageId="34" pageNumber="35">Distribution.</emphasis>
Only known from rainforests of the Iron Range in the Cape
<collectingRegion id="505F498C620C223E0AB9FC6EFBEC6A8D" box="[1015,1070,979,1005]" country="United Kingdom" name="York" pageId="34" pageNumber="35">York</collectingRegion>
Peninsula (CYP) bioregion (
<figureCitation id="0AA09BEB620C223E0C21FC6EFF7B6D72" captionStart="FIGURE 6" captionStartId="32.[151,250,999,1023]" captionTargetBox="[154,1436,187,969]" captionTargetId="figure-26@32.[151,1436,181,975]" captionTargetPageId="32" captionText="FIGURE 6. A, Map of known records of Lepanus ustulatus; B, predicted distribution of Lepanus ustulatus; C, map of known records of Lepanus cameroni." figureDoi="http://doi.org/10.5281/zenodo.4496769" httpUri="https://zenodo.org/record/4496769/files/figure.png" pageId="34" pageNumber="35">Fig. 6C</figureCitation>
) and likely represents a short-range endemic.
</paragraph>
</subSubSection>
<subSubSection id="DA81D4E5620C223E0989FBA7FC9D6DC1" pageId="34" pageNumber="35" type="discussion">
<paragraph id="9224876E620C223E0989FBA7FC9D6DC1" blockId="34.[151,1437,150,1185]" pageId="34" pageNumber="35">
<emphasis id="A0EF5B7C620C223E0989FBA7FE8D6D54" bold="true" box="[199,335,1050,1076]" pageId="34" pageNumber="35">Comments.</emphasis>
Only known from six specimens collected in either dung traps or pan traps. This species has been referred to under the informal name of “
<taxonomicName id="559BFCED620C223E0B11FB82FCC06D39" authority="CYP" authorityName="CYP" box="[607,770,1087,1113]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="A0EF5B7C620C223E0B11FB82FD7C6D38" box="[607,702,1087,1112]" italics="true" pageId="34" pageNumber="35">Lepanus</emphasis>
CYP
</taxonomicName>
8”. The sequenced specimen in
<bibRefCitation id="F60AFA9F620C223E0D2DFB82FA866D39" author="Gunter, N. L. &amp; Weir, T. A." box="[1123,1348,1087,1113]" pageId="34" pageNumber="35" pagination="41 - 80" refId="ref76546" refString="Gunter, N. L. &amp; Weir, T. A. (2019 a) Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): key to species groups and description of 14 new species from the L. pygmaeus species group. Zootaxa, 4564 (1), 41 - 80. https: // doi. org / 10.11646 / zootaxa. 4564.1.2" type="journal article" year="2019">
Gunter
<emphasis id="A0EF5B7C620C223E0DF6FB82FB306D39" box="[1208,1266,1087,1113]" italics="true" pageId="34" pageNumber="35">et al.</emphasis>
(2019)
</bibRefCitation>
is from Gordons Creek and is designated as a
<typeStatus id="4D2039CC620C223E0B1FFBDEFD736D1D" box="[593,689,1123,1149]" pageId="34" pageNumber="35" type="paratype">paratype</typeStatus>
here (ANIC 25-053831 also as COL1463) and was recovered as sister to
<taxonomicName id="559BFCED620C223E09B8FB3AFEB66DC1" authorityName="DET" authorityYear="1971" baseAuthorityName="Macleay" baseAuthorityYear="1887" box="[246,372,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="globulus">
<emphasis id="A0EF5B7C620C223E09B8FB3AFEB66DC1" box="[246,372,1159,1185]" italics="true" pageId="34" pageNumber="35">L. globulus</emphasis>
</taxonomicName>
within the
<taxonomicName id="559BFCED620C223E08BDFB3AFDB76DC1" baseAuthorityName="Lansberge" baseAuthorityYear="1874" box="[499,629,1159,1185]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="ustulatus">
<emphasis id="A0EF5B7C620C223E08BDFB3AFDB76DC1" box="[499,629,1159,1185]" italics="true" pageId="34" pageNumber="35">L. ustulatus</emphasis>
</taxonomicName>
species group clade.
</paragraph>
</subSubSection>
</treatment>
</document>