379 lines
50 KiB
XML
379 lines
50 KiB
XML
<document id="E66D31A1A4C061CACF244CDBCF468A47" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E30E8C3CA2EDF941FF1BFA3D" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zephronia laotica Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="209" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="205" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="E61129D2D40AE10AC71CE61725AF7492">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="591A10998781E1084215F6DB15E6F02F">Wesener, Thomas</mods:namePart>
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<mods:title id="1FEAC2B1E5A9EBC2732D931608C2FE15">Zootaxa</mods:title>
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<mods:part id="262F3A46146D79265A5CFB8703D14B99">
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<mods:date id="1A52DB5A2D2F26C611C2A50BB2C66880">2019</mods:date>
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<treatment id="03EB5506E30E8C3CA2EDF941FF1BFA3D" ID-DOI="http://doi.org/10.5281/zenodo.5934516" ID-GBIF-Taxon="154474553" ID-Zenodo-Dep="5934516" LSID="urn:lsid:plazi:treatment:03EB5506E30E8C3CA2EDF941FF1BFA3D" httpUri="http://treatment.plazi.org/id/03EB5506E30E8C3CA2EDF941FF1BFA3D" lastPageId="8" lastPageNumber="209" pageId="4" pageNumber="205">
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<subSubSection id="C358B79BE30E8C30A2EDF941FDEFF954" box="[151,518,1678,1704]" pageId="4" pageNumber="205" type="nomenclature">
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<paragraph id="8BFDE410E30E8C30A2EDF941FDEFF954" blockId="4.[151,518,1678,1738]" box="[151,518,1678,1704]" pageId="4" pageNumber="205">
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<heading id="D0B5537CE30E8C30A2EDF941FDEFF954" bold="true" box="[151,518,1678,1704]" fontSize="11" level="1" pageId="4" pageNumber="205" reason="1">
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<emphasis id="B9363802E30E8C30A2EDF941FDEFF954" bold="true" box="[151,518,1678,1704]" pageId="4" pageNumber="205">
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<taxonomicName id="4C429F93E30E8C30A2EDF941FE87F954" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,366,1678,1704]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
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<emphasis id="B9363802E30E8C30A2EDF941FE87F954" bold="true" box="[151,366,1678,1704]" italics="true" pageId="4" pageNumber="205">Zephronia laotica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E30E8C30A30CF941FDEFF954" box="[374,518,1678,1704]" pageId="4" pageNumber="205" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C358B79BE30E8C30A2EDF97EFECAF936" box="[151,291,1713,1738]" pageId="4" pageNumber="205" type="description">
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<paragraph id="8BFDE410E30E8C30A2EDF97EFECAF936" blockId="4.[151,518,1678,1738]" box="[151,291,1713,1738]" pageId="4" pageNumber="205">
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<figureCitation id="1379F895E30E8C30A2EDF97EFEF6F936" box="[151,287,1713,1738]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="5.[151,250,1870,1892]" captionStartId-1="6.[151,250,1933,1955]" captionStartId-2="7.[151,250,1943,1965]" captionTargetBox-0="[267,1320,445,1848]" captionTargetBox-1="[186,1403,553,1923]" captionTargetBox-2="[168,1419,193,1925]" captionTargetId-0="figure@5.[267,1320,445,1849]" captionTargetId-1="figure@6.[186,1403,553,1923]" captionTargetId-2="figure@7.[168,1419,193,1925]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 2. Distribution map and habitus photograph of Zephronia species, Z. laotica n. sp." captionText-1="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." captionText-2="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri-0="https://zenodo.org/record/2601076/files/figure.png" httpUri-1="https://zenodo.org/record/2601078/files/figure.png" httpUri-2="https://zenodo.org/record/2601080/files/figure.png" pageId="4" pageNumber="205">Figures 2–4</figureCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358B79BE30E8C30A2EDF936FCFCF8A6" pageId="4" pageNumber="205" type="materials_examined">
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<paragraph id="8BFDE410E30E8C30A2EDF936FCFCF8A6" blockId="4.[151,1437,1785,2026]" pageId="4" pageNumber="205">
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<emphasis id="B9363802E30E8C30A2EDF936FE90F8EE" bold="true" box="[151,377,1785,1810]" pageId="4" pageNumber="205">Material examined</emphasis>
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: Type specimens. 1 M
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<typeStatus id="54F95AB2E30E8C30A0FBF936FD0AF8EE" box="[641,739,1785,1810]" pageId="4" pageNumber="205" type="holotype">holotype</typeStatus>
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(ZFMK MYR3502) from
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<collectingCountry id="F355A480E30E8C30A66AF935FBA0F8ED" box="[1040,1097,1786,1809]" name="Laos" pageId="4" pageNumber="205">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E30E8C30A62FF936FAAEF8EE" box="[1109,1351,1785,1810]" country="Laos" name="Champasak" pageId="4" pageNumber="205">Champasak Province</collectingRegion>
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, east of Mekong, Garden of Erawan Riverside Hotel (
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<geoCoordinate id="EE7682D7E30E8C30A0E0F8D1FCD8F8CA" box="[666,817,1821,1846]" direction="north" orientation="latitude" pageId="4" pageNumber="205" precision="1" value="15.1075">N 15°6'27.0"</geoCoordinate>
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,
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<geoCoordinate id="EE7682D7E30E8C30A147F8D1FC05F8CA" box="[829,1004,1821,1846]" direction="east" orientation="longitude" pageId="4" pageNumber="205" precision="1" value="105.82064">E 105°49'14.3"</geoCoordinate>
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), leg. Siegfried Huber,
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<date id="FFFCC2D0E30E8C30A689F8D2FA71F8CA" box="[1267,1432,1821,1846]" pageId="4" pageNumber="205" value="2012-11-10" valueMax="2012-11-15" valueMin="2012-11-10">10–15.xi.2012</date>
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. 1 F
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<typeStatus id="54F95AB2E30E8C30A2B8F88DFECAF8A6" box="[194,291,1858,1882]" pageId="4" pageNumber="205" type="paratype">paratype</typeStatus>
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(ZFMK MYR8098), same data as
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<typeStatus id="54F95AB2E30E8C30A0D6F88EFCFBF8A6" box="[684,786,1857,1882]" pageId="4" pageNumber="205" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE30E8C31A2BDF8AAFE7AFE74" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205" type="diagnosis">
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<paragraph id="8BFDE410E30E8C31A2BDF8AAFE7AFE74" blockId="4.[151,1437,1785,2026]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="206" pageId="4" pageNumber="205">
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<emphasis id="B9363802E30E8C30A2BDF8AAFEA8F882" bold="true" box="[199,321,1893,1918]" pageId="4" pageNumber="205">Diagnosis.</emphasis>
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<taxonomicName id="4C429F93E30E8C30A333F8AAFDFAF881" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[329,531,1893,1917]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
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<emphasis id="B9363802E30E8C30A333F8AAFDFAF881" box="[329,531,1893,1917]" italics="true" pageId="4" pageNumber="205">Zephronia laotica</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E30E8C30A060F8AAFDB3F882" bold="true" box="[538,602,1893,1918]" pageId="4" pageNumber="205">
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<taxonomicNameLabel id="A2058579E30E8C30A060F8AAFDB3F882" box="[538,602,1893,1918]" pageId="4" pageNumber="205" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to the
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<taxonomicName id="4C429F93E30E8C30A173F8AAFC94F881" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[777,893,1893,1917]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B9363802E30E8C30A173F8AAFC94F881" box="[777,893,1893,1917]" italics="true" pageId="4" pageNumber="205">Zephronia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E30E8C30A1FFF8A9FBF8F882" box="[901,1041,1893,1918]" pageId="4" pageNumber="205" sensu="stricto">sensu stricto</taxonomicNameLabel>
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group based on the position of the organ of Tömösváry, located next to the aberrant ocelli, not inside the antennal groove (Semeyuk
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<emphasis id="B9363802E30E8C30A774F845FAABF85D" box="[1294,1346,1929,1953]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
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. 2018).
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<taxonomicName id="4C429F93E30E8C30A2EDF862FEE2F839" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,267,1965,1989]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B9363802E30E8C30A2EDF862FEE2F839" box="[151,267,1965,1989]" italics="true" pageId="4" pageNumber="205">Zephronia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E30E8C30A36EF861FE48F83A" box="[276,417,1965,1990]" pageId="4" pageNumber="205" sensu="stricto">sensu stricto</taxonomicNameLabel>
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encompasses:
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<taxonomicName id="4C429F93E30E8C30A034F861FCD5F83A" authority="Gray, 1832" authorityName="Gray" authorityYear="1832" box="[590,828,1965,1990]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="ovalis">
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<emphasis id="B9363802E30E8C30A034F861FD59F839" box="[590,688,1965,1989]" italics="true" pageId="4" pageNumber="205">Z. ovalis</emphasis>
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<bibRefCitation id="EFD399E1E30E8C30A0C0F861FCD5F83A" author="Gray, G." box="[698,828,1966,1990]" pageId="4" pageNumber="205" pagination="124 - 142" refId="ref24522" refString="Gray, G. (1832) The Myriapods (Myriapoda, Fab.). in Griffith E & Pidgeon E, The Class Insecta arranged by the Baron Cuvier, 1, 124 - 142, pl. 1 - 2." type="journal article" year="1832">Gray, 1832</bibRefCitation>
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</taxonomicName>
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;
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<taxonomicName id="4C429F93E30E8C30A130F861FACAF83A" authority="Semenyuk et al., 2018" authorityName="Semenyuk" authorityYear="2018" box="[842,1315,1965,1990]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="konkakinhensis">
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<emphasis id="B9363802E30E8C30A130F861FBFCF839" box="[842,1045,1965,1989]" italics="true" pageId="4" pageNumber="205">Z. konkakinhensis</emphasis>
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<bibRefCitation id="EFD399E1E30E8C30A664F862FACAF83A" author="Semenyuk, I. & Golovatch, S. I. & Wesener, T." box="[1054,1315,1965,1990]" pageId="4" pageNumber="205" pagination="535 - 550" refId="ref25362" refString="Semenyuk, I., Golovatch, S. I. & Wesener, T. (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida: Zephroniidae). Zootaxa, 4459 (3), 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3.7" type="journal article" year="2018">
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Semenyuk
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<emphasis id="B9363802E30E8C30A6E7F861FB39F839" box="[1181,1232,1965,1989]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
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., 2018
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</bibRefCitation>
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||
</taxonomicName>
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;
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<taxonomicName id="4C429F93E30E8C30A74BF861FE63F816" authority="Semenyuk et al. 2018" authorityName="Semenyuk" authorityYear="2018" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="montis">
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<emphasis id="B9363802E30E8C30A74BF861FA75F839" box="[1329,1436,1966,1989]" italics="true" pageId="4" pageNumber="205">Z. montis</emphasis>
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||
<bibRefCitation id="EFD399E1E30E8C30A2EDF81EFE63F816" author="Semenyuk, I. & Golovatch, S. I. & Wesener, T." box="[151,394,2001,2026]" pageId="4" pageNumber="205" pagination="535 - 550" refId="ref25362" refString="Semenyuk, I., Golovatch, S. I. & Wesener, T. (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida: Zephroniidae). Zootaxa, 4459 (3), 535 - 550. https: // doi. org / 10.11646 / zootaxa. 4459.3.7" type="journal article" year="2018">
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||
Semenyuk
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<emphasis id="B9363802E30E8C30A36EF81DFEACF815" box="[276,325,2001,2025]" italics="true" pageId="4" pageNumber="205">et al</emphasis>
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||
. 2018
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</bibRefCitation>
|
||
</taxonomicName>
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from
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<collectingCountry id="F355A480E30E8C30A3B4F81EFDD9F816" box="[462,560,2001,2026]" name="Vietnam" pageId="4" pageNumber="205">Vietnam</collectingCountry>
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||
;
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||
<taxonomicName id="4C429F93E30E8C30A046F81DFCA4F816" authority="Hirst, 1907" authorityName="Hirst" authorityYear="1907" box="[572,845,2001,2026]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="205" phylum="Arthropoda" rank="species" species="siamensis">
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||
<emphasis id="B9363802E30E8C30A046F81DFD2FF815" box="[572,710,2002,2025]" italics="true" pageId="4" pageNumber="205">Z. siamensis</emphasis>
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||
Hirst, 1907
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</taxonomicName>
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||
, the only species of the genus known from
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<collectingCountry id="F355A480E30E8C30A74BF81EFA71F816" box="[1329,1432,2001,2026]" name="Thailand" pageId="4" pageNumber="205">Thailand</collectingCountry>
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||
; and
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<taxonomicName id="4C429F93E30F8C31A2BDFF56FE1EFF4C" authority="Attems, 1953" authorityName="Attems" authorityYear="1953" box="[199,503,151,176]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="dawydoffi">
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||
<emphasis id="B9363802E30F8C31A2BDFF56FEBDFF4C" box="[199,340,152,176]" italics="true" pageId="5" pageNumber="206">Z. dawydoffi</emphasis>
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||
<bibRefCitation id="EFD399E1E30F8C31A326FF57FE1EFF4C" author="Attems, C." box="[348,503,151,176]" pageId="5" pageNumber="206" pagination="133 - 230" refId="ref23903" refString="Attems, C. (1953) Myriopoden von Indochina, Expedition von Dr. C. Dawydoff (1938 - 1939). Memoires du Museum national d'histoire naturelle, N. S., serie A, 5 (3), 133 - 230." type="journal article" year="1953">Attems, 1953</bibRefCitation>
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</taxonomicName>
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, the only giant pill-millipede species known from
|
||
<collectingCountry id="F355A480E30F8C31A648FF58FB40FF4C" box="[1074,1193,151,176]" name="Cambodia" pageId="5" pageNumber="206">Cambodia</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C429F93E30F8C31A6CEFF57FA97FF4C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1204,1406,152,176]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
|
||
<emphasis id="B9363802E30F8C31A6CEFF57FA97FF4C" box="[1204,1406,152,176]" italics="true" pageId="5" pageNumber="206">Zephronia laotica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E30F8C31A7FFFF58FF51FF29" bold="true" pageId="5" pageNumber="206">
|
||
<taxonomicNameLabel id="A2058579E30F8C31A7FFFF58FF51FF29" pageId="5" pageNumber="206" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E30F8C31A32EFF72FDF6FF28" box="[340,543,188,212]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="konkakinhensis">
|
||
<emphasis id="B9363802E30F8C31A32EFF72FDF6FF28" box="[340,543,188,212]" italics="true" pageId="5" pageNumber="206">Z. konkakinhensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C429F93E30F8C31A021FF72FD2FFF28" box="[603,710,189,212]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="montis">
|
||
<emphasis id="B9363802E30F8C31A021FF72FD2FFF28" box="[603,710,189,212]" italics="true" pageId="5" pageNumber="206">Z. montis</emphasis>
|
||
</taxonomicName>
|
||
in the walking legs that lack strong teeth on the femur, and
|
||
<taxonomicName id="4C429F93E30F8C31A7FFFF72FF33FF04" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="B9363802E30F8C31A7FFFF72FF33FF04" italics="true" pageId="5" pageNumber="206">Z. ovalis</emphasis>
|
||
</taxonomicName>
|
||
which has weak teeth at the mesal margin of the femur (
|
||
<figureCitation id="1379F895E30F8C31A113FF10FC2BFF04" box="[873,962,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="5" pageNumber="206">Fig. 3B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E30F8C31A1A3FF2FFB4AFF04" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[985,1187,224,248]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
|
||
<emphasis id="B9363802E30F8C31A1A3FF2FFB4AFF04" box="[985,1187,224,248]" italics="true" pageId="5" pageNumber="206">Zephronia laotica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E30F8C31A6D6FF10FB05FF04" bold="true" box="[1196,1260,223,248]" pageId="5" pageNumber="206">
|
||
<taxonomicNameLabel id="A2058579E30F8C31A6D6FF10FB05FF04" box="[1196,1260,223,248]" pageId="5" pageNumber="206" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E30F8C31A7FFFF2EFF33FEE0" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="B9363802E30F8C31A7FFFF2EFF33FEE0" italics="true" pageId="5" pageNumber="206">Z. ovalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C429F93E30F8C31A29DFECAFE98FEE0" authorityName="Hirst" authorityYear="1907" box="[231,369,261,284]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="siamensis">
|
||
<emphasis id="B9363802E30F8C31A29DFECAFE98FEE0" box="[231,369,261,284]" italics="true" pageId="5" pageNumber="206">Z. siamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C429F93E30F8C31A3DCFECAFDD8FEE0" authorityName="Attems" authorityYear="1953" box="[422,561,260,284]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="dawydoffi">
|
||
<emphasis id="B9363802E30F8C31A3DCFECAFDD8FEE0" box="[422,561,260,284]" italics="true" pageId="5" pageNumber="206">Z. dawydoffi</emphasis>
|
||
</taxonomicName>
|
||
also in the differently shaped telopoditomeres 3 and 4 of the anterior telopods, the elongation of telopoditomere 3 and the small size of telopoditomere 4 are unique (
|
||
<figureCitation id="1379F895E30F8C31A635FEE8FB7CFEBC" box="[1103,1173,295,320]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1965]" captionTargetBox="[168,1419,193,1925]" captionTargetId="figure@7.[168,1419,193,1925]" captionTargetPageId="7" captionText="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="5" pageNumber="206">Figs 4</figureCitation>
|
||
B–D), a different bright green colour of the tergites (
|
||
<figureCitation id="1379F895E30F8C31A3ABFE83FDC3FE98" box="[465,554,332,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1870,1892]" captionTargetBox="[267,1320,445,1848]" captionTargetId="figure@5.[267,1320,445,1849]" captionTargetPageId="5" captionText="FIGURE 2. Distribution map and habitus photograph of Zephronia species, Z. laotica n. sp." httpUri="https://zenodo.org/record/2601076/files/figure.png" pageId="5" pageNumber="206">Fig. 2A</figureCitation>
|
||
), leg 3 lacking an apical spine, and the presence of two locking carinae on each side of the anal shield.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3DB498E30F8C31A2EDF881FBA9F898" box="[151,1088,1870,1893]" httpUri="https://zenodo.org/record/2601076/files/figure.png" pageId="5" pageNumber="206" startId="5.[151,250,1870,1892]" targetBox="[267,1320,445,1848]" targetPageId="5">
|
||
<paragraph id="8BFDE410E30F8C31A2EDF881FBA9F898" blockId="5.[151,1088,1870,1893]" box="[151,1088,1870,1893]" pageId="5" pageNumber="206">
|
||
<emphasis id="B9363802E30F8C31A2EDF881FEFBF898" bold="true" box="[151,274,1870,1892]" pageId="5" pageNumber="206">FIGURE 2.</emphasis>
|
||
Distribution map and habitus photograph of
|
||
<taxonomicName id="4C429F93E30F8C31A0A1F881FCAAF898" authorityName="Wesener" authorityYear="2019" box="[731,835,1870,1892]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E30F8C31A0A1F881FCAAF898" box="[731,835,1870,1892]" italics="true" pageId="5" pageNumber="206">Zephronia</emphasis>
|
||
</taxonomicName>
|
||
species,
|
||
<taxonomicName id="4C429F93E30F8C31A1E5F89FFBE9F898" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[927,1024,1870,1893]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="206" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
|
||
<emphasis id="B9363802E30F8C31A1E5F89FFBE9F898" box="[927,1024,1870,1893]" italics="true" pageId="5" pageNumber="206">Z. laotica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2058579E30F8C31A67DF881FBA9F898" box="[1031,1088,1870,1892]" pageId="5" pageNumber="206" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C358B79BE30F8C3CA2BDF85EFEB4FA84" lastPageId="8" lastPageNumber="209" pageId="5" pageNumber="206" type="description">
|
||
<paragraph id="8BFDE410E30F8C32A2BDF85EFC72FF04" blockId="5.[151,1436,1937,2034]" lastBlockId="6.[151,1437,151,501]" lastPageId="6" lastPageNumber="207" pageId="5" pageNumber="206">
|
||
<emphasis id="B9363802E30F8C31A2BDF85EFEB8F856" bold="true" box="[199,337,1937,1962]" pageId="5" pageNumber="206">Description</emphasis>
|
||
. Based on
|
||
<typeStatus id="54F95AB2E30F8C31A3B4F85EFDD9F856" box="[462,560,1937,1962]" pageId="5" pageNumber="206" type="holotype">holotype</typeStatus>
|
||
male and
|
||
<typeStatus id="54F95AB2E30F8C31A0DEF85DFCEDF856" box="[676,772,1938,1962]" pageId="5" pageNumber="206" type="paratype">paratype</typeStatus>
|
||
female.
|
||
<emphasis id="B9363802E30F8C31A2BDF87AFE9FF832" bold="true" box="[199,374,1973,1998]" pageId="5" pageNumber="206">Measurements</emphasis>
|
||
:
|
||
<typeStatus id="54F95AB2E30F8C31A3FEF87AFE05F832" box="[388,492,1973,1998]" pageId="5" pageNumber="206" type="holotype">Holotype</typeStatus>
|
||
male. Body length =
|
||
<quantity id="4CBA49F5E30F8C31A0A4F87AFCADF831" box="[734,836,1973,1998]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" pageId="5" pageNumber="206" unit="mm" value="15.6">15.6 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E30F8C31A6FAF87AFB3EF831" box="[1152,1239,1973,1998]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="5" pageNumber="206" unit="mm" value="6.7">6.7 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E30F8C31A703F879FF2AF80D" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="206" unit="mm" value="7.2">7.2 mm</quantity>
|
||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E30F8C31A0E6F815FD1DF80D" box="[668,756,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="5" pageNumber="206" unit="mm" value="4.2">4.2 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E30F8C31A1E3F815FC06F80D" box="[921,1007,2010,2034]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="206" unit="mm" value="4.3">4.3 mm</quantity>
|
||
(= highest). Female: body length = ca
|
||
<quantity id="4CBA49F5E30C8C32A2EDFF57FF16FF53" box="[151,255,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="6" pageNumber="207" unit="mm" value="14.1">14.1 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E30C8C32A03FFF58FD7FFF4C" box="[581,662,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="6" pageNumber="207" unit="mm" value="6.1">6.1mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E30C8C32A13AFF58FC7EFF53" box="[832,919,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="6" pageNumber="207" unit="mm" value="6.9">6.9 mm</quantity>
|
||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E30C8C32A703FF57FF2CFF28" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="6" pageNumber="207" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E30C8C32A31DFF73FE54FF28" box="[359,445,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="207" unit="mm" value="4.2">4.2 mm</quantity>
|
||
(= highest).
|
||
<emphasis id="B9363802E30C8C32A031FF73FD25FF29" bold="true" box="[587,716,188,213]" pageId="6" pageNumber="207">Coloration</emphasis>
|
||
: in preserved specimens, head and collum green. Tergites bright green, posterior half light brown. Antennae and legs green (
|
||
<figureCitation id="1379F895E30C8C32A149FF10FC67FF04" box="[819,910,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1870,1892]" captionTargetBox="[267,1320,445,1848]" captionTargetId="figure@5.[267,1320,445,1849]" captionTargetPageId="5" captionText="FIGURE 2. Distribution map and habitus photograph of Zephronia species, Z. laotica n. sp." httpUri="https://zenodo.org/record/2601076/files/figure.png" pageId="6" pageNumber="207">Fig. 2A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E30C8C32A2BDFECBFE52FE08" blockId="6.[151,1437,151,501]" pageId="6" pageNumber="207">
|
||
<emphasis id="B9363802E30C8C32A2BDFECBFEEEFEE1" bold="true" box="[199,263,260,285]" pageId="6" pageNumber="207">Head</emphasis>
|
||
: wide and short. Eyes with>75 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 3. Sizes of antennomeres 1=2=3=4=5<<6. Antennomere 6 densely pubescent, with a few sensilla basiconica surrounding apical disc. Shape sexually dimorphic: in female cylindrical, in male laterally flattened, axe-shaped. Male apical disc with 61/65 apical sensory cones, that of female with 32/34 apical cones. Organ of Tömösváry located at a projected brim close to the ocelli at margin of antennal groove.
|
||
<emphasis id="B9363802E30C8C32A283FE78FE2EFE2C" bold="true" box="[249,455,439,464]" pageId="6" pageNumber="207">Gnathochilarium</emphasis>
|
||
: structure typical of the order. Sensory cones of palpi all located in single field.
|
||
<emphasis id="B9363802E30C8C32A2EDFE13FEFCFE09" bold="true" box="[151,277,476,501]" pageId="6" pageNumber="207">Mandibles</emphasis>
|
||
not examined.
|
||
</paragraph>
|
||
<caption id="DF3DB498E30C8C32A2EDF842FAA0F81C" httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="6" pageNumber="207" startId="6.[151,250,1933,1955]" targetBox="[186,1403,553,1923]" targetPageId="6">
|
||
<paragraph id="8BFDE410E30C8C32A2EDF842FAA0F81C" blockId="6.[151,1436,1933,2016]" pageId="6" pageNumber="207">
|
||
<emphasis id="B9363802E30C8C32A2EDF842FEE5F85F" bold="true" box="[151,268,1933,1955]" pageId="6" pageNumber="207">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E30C8C32A363F842FE27F85F" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[281,462,1933,1955]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
|
||
<emphasis id="B9363802E30C8C32A363F842FE27F85F" box="[281,462,1933,1955]" italics="true" pageId="6" pageNumber="207">Zephronia laotica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E30C8C32A3AEF842FDE4F85F" bold="true" box="[468,525,1933,1955]" pageId="6" pageNumber="207">
|
||
<taxonomicNameLabel id="A2058579E30C8C32A3AEF842FDE4F85F" box="[468,525,1933,1955]" pageId="6" pageNumber="207" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9363802E30C8C32A060F842FDA0F85F" bold="true" box="[538,585,1933,1955]" pageId="6" pageNumber="207">A, B</emphasis>
|
||
: male holotype (ZFMK MYR3502);
|
||
<emphasis id="B9363802E30C8C32A1BEF842FC1CF85F" bold="true" box="[964,1013,1933,1955]" pageId="6" pageNumber="207">C, D</emphasis>
|
||
: female paratype (ZFMK MYR8098);
|
||
<emphasis id="B9363802E30C8C32A7F9F842FA7CF85F" bold="true" box="[1411,1429,1933,1955]" pageId="6" pageNumber="207">A</emphasis>
|
||
: coxa of first left leg with first stigmatic plate;
|
||
<emphasis id="B9363802E30C8C32A017F863FD97F83E" bold="true" box="[621,638,1964,1986]" pageId="6" pageNumber="207">B</emphasis>
|
||
: 9th right leg;
|
||
<emphasis id="B9363802E30C8C32A16CF863FCD8F83E" bold="true" box="[790,817,1964,1986]" pageId="6" pageNumber="207">C:</emphasis>
|
||
coxa and prefemur of second left leg with vulva;
|
||
<emphasis id="B9363802E30C8C32A754F863FAA9F83E" bold="true" box="[1326,1344,1964,1986]" pageId="6" pageNumber="207">D</emphasis>
|
||
: subanal plate. Abbreviations:
|
||
<emphasis id="B9363802E30C8C32A30BF805FE78F81C" bold="true" box="[369,401,1994,2016]" pageId="6" pageNumber="207">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E30C8C32A392F805FE15F81C" bold="true" box="[488,508,1994,2016]" pageId="6" pageNumber="207">O</emphasis>
|
||
= operculum of vulva;
|
||
<emphasis id="B9363802E30C8C32A091F805FCF8F81C" bold="true" box="[747,785,1994,2016]" pageId="6" pageNumber="207">Pre</emphasis>
|
||
= prefemur;
|
||
<emphasis id="B9363802E30C8C32A1EFF805FC21F81C" bold="true" box="[917,968,1994,2016]" pageId="6" pageNumber="207">St-Pl</emphasis>
|
||
= stigmatic plate. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3DB498E30D8C33A2EDF858FA8CF816" httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="7" pageNumber="208" startId="7.[151,250,1943,1965]" targetBox="[168,1419,193,1925]" targetPageId="7">
|
||
<paragraph id="8BFDE410E30D8C33A2EDF858FA8CF816" blockId="7.[151,1436,1943,2026]" pageId="7" pageNumber="208">
|
||
<emphasis id="B9363802E30D8C33A2EDF858FEE6F851" bold="true" box="[151,271,1943,1965]" pageId="7" pageNumber="208">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E30D8C33A367F858FE3DF851" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[285,468,1943,1965]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="laotica" status="sp. nov.">
|
||
<emphasis id="B9363802E30D8C33A367F858FE3DF851" box="[285,468,1943,1965]" italics="true" pageId="7" pageNumber="208">Zephronia laotica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E30D8C33A3A7F858FDF1F851" bold="true" box="[477,536,1943,1965]" pageId="7" pageNumber="208">
|
||
<taxonomicNameLabel id="A2058579E30D8C33A3A7F858FDF1F851" box="[477,536,1943,1965]" pageId="7" pageNumber="208" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype (ZFMK MYR3502);
|
||
<emphasis id="B9363802E30D8C33A1E7F858FC46F851" bold="true" box="[925,943,1943,1965]" pageId="7" pageNumber="208">A</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E30D8C33A720F858FA82F851" bold="true" box="[1370,1387,1943,1965]" pageId="7" pageNumber="208">B</emphasis>
|
||
: left anterior telopod, anterior view;
|
||
<emphasis id="B9363802E30D8C33A3A3F87AFE1DF837" bold="true" box="[473,500,1973,1995]" pageId="7" pageNumber="208">C:</emphasis>
|
||
left anterior telopod, posterior view;
|
||
<emphasis id="B9363802E30D8C33A10BF87AFC6AF837" bold="true" box="[881,899,1973,1995]" pageId="7" pageNumber="208">D</emphasis>
|
||
: left anterior telopod, lateral view;
|
||
<emphasis id="B9363802E30D8C33A697F87AFB17F837" bold="true" box="[1261,1278,1973,1995]" pageId="7" pageNumber="208">E</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="B9363802E30D8C33A3F9F81BFE7BF816" bold="true" box="[387,402,2004,2026]" pageId="7" pageNumber="208">F</emphasis>
|
||
: right posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E30D8C33A1B2F81BFBF1F816" bold="true" box="[968,1048,2004,2026]" pageId="7" pageNumber="208">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E3028C3CA2BDFF58FAA1FD78" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9363802E3028C3CA2BDFF58FE6FFF4C" bold="true" box="[199,390,151,176]" pageId="8" pageNumber="209">Stigmatic plates</emphasis>
|
||
: first stigmatic plate triangular, apex broadly rounded, slightly curved towards coxa 1 (
|
||
<figureCitation id="1379F895E3028C3CA715FF58FF55FF28" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Fig. 3A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3028C3CA2AAFF73FE90FF29" bold="true" box="[208,377,188,213]" pageId="8" pageNumber="209">Laterotergites</emphasis>
|
||
: first laterotergite tip weakly extended, pointed process. Second laterotergite with even shorter process, well-rounded apex.
|
||
<emphasis id="B9363802E3028C3CA39FFF10FDD4FF04" bold="true" box="[485,573,223,248]" pageId="8" pageNumber="209">Collum</emphasis>
|
||
: glabrous, except for margins which have a few isolated and long setae.
|
||
<emphasis id="B9363802E3028C3CA2EDFECBFEBDFEE1" bold="true" box="[151,340,260,285]" pageId="8" pageNumber="209">Thoracic shield</emphasis>
|
||
: surface as in tergites. Shallow grooves filled with numerous long setae, no keels.
|
||
<emphasis id="B9363802E3028C3CA74FFECBFA7DFEE1" bold="true" box="[1333,1428,260,285]" pageId="8" pageNumber="209">Tergites</emphasis>
|
||
: surface shiny, glabrous, weakly coriaceous. Tips of paratergites of midbody tergites straight (
|
||
<figureCitation id="1379F895E3028C3CA750FEE8FA66FEBC" box="[1322,1423,295,320]" captionStart="FIGURE 2" captionStartId="5.[151,250,1870,1892]" captionTargetBox="[267,1320,445,1848]" captionTargetId="figure@5.[267,1320,445,1849]" captionTargetPageId="5" captionText="FIGURE 2. Distribution map and habitus photograph of Zephronia species, Z. laotica n. sp." httpUri="https://zenodo.org/record/2601076/files/figure.png" pageId="8" pageNumber="209">Fig. 2A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3028C3CA2EDFE83FEC7FE99" bold="true" box="[151,302,332,357]" pageId="8" pageNumber="209">Endotergum</emphasis>
|
||
: inner section with few longer setae. Middle area with a single row of dense round, shallow, cuticular impressions; distance between impressions slightly smaller than their diameter. Apically, 1–2 dense rows of short marginal bristles, the longest just protruding above tergal margin. Bristles not smooth, but with numerous small spicules.
|
||
<emphasis id="B9363802E3028C3CA37AFE78FE6EFE2C" bold="true" box="[256,391,439,464]" pageId="8" pageNumber="209">Anal shield</emphasis>
|
||
: slightly sexually dimorphic, in female large and well-rounded, in male more rectangular, in both sexes glabrous. Surface similar to that of tergites. Underside with two locking carinae; first short, as long as those of tergites, second as long as first, but much weaker.
|
||
<emphasis id="B9363802E3028C3CA146FE30FC9DFDE4" bold="true" box="[828,884,511,536]" pageId="8" pageNumber="209">Legs</emphasis>
|
||
: leg-pair 1 with 1 or 2 ventral spines, leg-pair 2 with 4, leg-pair 3 with 6. First three leg-pairs without an apical spine. Leg-pairs 4–21 with 7–9 ventral spines and 3–5 apical spines. In leg-pair 9, femur 1.5 times, tarsus 3.5 times longer than wide (
|
||
<figureCitation id="1379F895E3028C3CA643FD88FB79FD9C" box="[1081,1168,583,608]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Fig. 3B</figureCitation>
|
||
). All podomeres setose. Coxa with a large, well-rounded process. Coxal process absent on leg-pair 1 (
|
||
<figureCitation id="1379F895E3028C3CA181FDA3FBBAFD78" box="[1019,1107,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Fig. 3A</figureCitation>
|
||
) and 2 (
|
||
<figureCitation id="1379F895E3028C3CA6D4FDA3FAE0FD78" box="[1198,1289,620,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Figs 3C</figureCitation>
|
||
,
|
||
<figureCitation id="1379F895E3028C3CA76FFDA3FAD5FD78" box="[1301,1340,620,644]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1965]" captionTargetBox="[168,1419,193,1925]" captionTargetId="figure@7.[168,1419,193,1925]" captionTargetPageId="7" captionText="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="8" pageNumber="209">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3028C3CA2BDFD40FE67FD0C" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9363802E3028C3CA2BDFD40FE1EFD54" bold="true" box="[199,503,655,680]" pageId="8" pageNumber="209">Female sexual characters</emphasis>
|
||
: vulva small, covering 1/2 of coxa, located at mesal margin, extending mesally to basal third of prefemur length (
|
||
<figureCitation id="1379F895E3028C3CA385FD7BFDB3FD30" box="[511,602,692,717]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Fig. 3C</figureCitation>
|
||
). Operculum rounded.
|
||
<emphasis id="B9363802E3028C3CA119FD7BFBE0FD31" bold="true" box="[867,1033,692,717]" pageId="8" pageNumber="209">Subanal plate</emphasis>
|
||
: large, tall, triangular, with a wellrounded tip (
|
||
<figureCitation id="1379F895E3028C3CA35CFD18FE68FD0C" box="[294,385,727,752]" captionStart="FIGURE 3" captionStartId="6.[151,250,1933,1955]" captionTargetBox="[186,1403,553,1923]" captionTargetId="figure@6.[186,1403,553,1923]" captionTargetPageId="6" captionText="FIGURE 3. Zephronia laotica n. sp., A, B: male holotype (ZFMK MYR3502); C, D: female paratype (ZFMK MYR8098); A: coxa of first left leg with first stigmatic plate; B: 9th right leg; C: coxa and prefemur of second left leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601078/files/figure.png" pageId="8" pageNumber="209">Fig. 3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3028C3CA2BDFD33FEB4FA84" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9363802E3028C3CA2BDFD33FE36FCE9" bold="true" box="[199,479,764,789]" pageId="8" pageNumber="209">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, triangular, sclerotized plate (
|
||
<figureCitation id="1379F895E3028C3CA749FD33FA79FCE8" box="[1331,1424,764,789]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1965]" captionTargetBox="[168,1419,193,1925]" captionTargetId="figure@7.[168,1419,193,1925]" captionTargetPageId="7" captionText="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="8" pageNumber="209">Fig. 4A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3028C3CA2EDFCD0FE86FCC4" bold="true" box="[151,367,799,824]" pageId="8" pageNumber="209">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3028C3CA3FBFCD0FE24FCC4" box="[385,461,799,824]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1965]" captionTargetBox="[168,1419,193,1925]" captionTargetId="figure@7.[168,1419,193,1925]" captionTargetPageId="7" captionText="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="8" pageNumber="209">Figs 4</figureCitation>
|
||
B–D): first telopoditomere rectangular, slightly longer than wide. Telopoditomere 2 large, as long as telopoditomeres 3 and 4 combined. Process of telopoditomere 2 located posteriorly, but partly visible laterally in anterior view. Process of telopoditomere 2 wide, broader than telopoditomeres 3 and 4. Process of telopoditomere 2 wide, visible mesally in anterior view. Process of telopoditomere 2 tapering apically, wellrounded, protruding as high as basal part of telopoditomere 4. Telopoditomere 3 slender, 1.5 longer than wide, twice as long as telopoditomere 4. Telopoditomere 4 very short, in posterior aspect without any teeth, but with two prominent spines. All podomeres covered with long sparse setae, except for central part of telopoditomere 1 and posterior surfaces of 3 and 4.
|
||
<emphasis id="B9363802E3028C3CA398FBD3FD55FBC9" bold="true" box="[482,700,1052,1077]" pageId="8" pageNumber="209">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3028C3CA0B0FBD3FCAAFBC8" box="[714,835,1052,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1943,1965]" captionTargetBox="[168,1419,193,1925]" captionTargetId="figure@7.[168,1419,193,1925]" captionTargetPageId="7" captionText="FIGURE 4. Zephronia laotica n. sp., male holotype (ZFMK MYR3502); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601080/files/figure.png" pageId="8" pageNumber="209">Figs 4E, F</figureCitation>
|
||
): first telopoditomere stout and narrow, twice as wide as long. Immovable finger (process of telopoditomere 2) as long as movable finger and consisting of telopoditomeres 3 and 4. Immovable finger wide, 1.9X longer than wide, weakly tapering apically, not curved. Margin towards movable finger genus-characteristic: with two large, triangular, membranous lobes, a spine, and several rows of circular sclerotized spots. Telopoditomere 3 elongated, typical of the genus: twice as long as telopoditomere 4. Margin near immovable finger with a large membranous lobe and a single long and slender spine, posterior surface with a row of 11 crenulate and sclerotized teeth. Telopoditomere 4 slender, only 2/3 as wide as telopoditomere 3,
|
||
<date id="FFFCC2D0E3028C3CA3F2FAD7FE42FACC" box="[392,427,1304,1328]" pageId="8" pageNumber="209">2X</date>
|
||
longer than wide, slightly tapering apically. Telopoditomere 4 with at inner margin two long spines and a single membranous lobe. Entire telopod with few setae, mostly at margins at on anterior side of telopoditomere 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3028C3CA2BDFA4BFF1BFA3D" pageId="8" pageNumber="209" type="etymology">
|
||
<paragraph id="8BFDE410E3028C3CA2BDFA4BFF1BFA3D" blockId="8.[151,1437,151,1473]" pageId="8" pageNumber="209">
|
||
<emphasis id="B9363802E3028C3CA2BDFA4BFE48FA61" bold="true" box="[199,417,1412,1437]" pageId="8" pageNumber="209">Derivatio nominis</emphasis>
|
||
: laotica, noun in apposition, after
|
||
<collectingCountry id="F355A480E3028C3CA140FA4AFC9AFA60" box="[826,883,1413,1436]" name="Laos" pageId="8" pageNumber="209">Laos</collectingCountry>
|
||
, as this is the first
|
||
<taxonomicName id="4C429F93E3028C3CA626FA4BFB39FA60" authorityName="Wesener" authorityYear="2019" box="[1116,1232,1412,1436]" class="Diplopoda" family="Zephronidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="209" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9363802E3028C3CA626FA4BFB39FA60" box="[1116,1232,1412,1436]" italics="true" pageId="8" pageNumber="209">Zephronia</emphasis>
|
||
</taxonomicName>
|
||
known from this country.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |