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<mods:title id="3DFB726104D15C4E454E8A721650B8DE">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<taxonomicName id="52F9BCF330C352FDEF32F944730307C1" ID-CoL="7XMPH" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" box="[549,1039,1763,1790]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="curvicornis">
Onthophagus curvicornis
<bibRefCitation id="F168BA8130C352FDEE76F943730307C1" author="Latreille P. A." box="[865,1039,1764,1790]" pageId="153" pageNumber="154" pagination="197 - 293" refId="ref127349" refString="Latreille P. A. 1809. Insectes de l'Amerique equinoxiale, recueillis pendant le voyage de MM. de Humboldt et Bonpland [livraison 4: pp. 197 - 293]. In: Humboldt A. &amp; Bonpland A. 1805 - 1810. Recueil d'observations de zoologie et d'anatomie comparee, faites dans l'ocean Atlantique, dans l'interieur du nouveau continent et dans la mer du Sud pendant les annees 1799, 1800, 1801, 1802 et 1803. Premier volume. Schoell and Dufour, Paris. [See comments for the previous entry. Evenhuis (1997 a) associated the publication of plate 23, containing the Onthophagus curvicornis figure, with livraisons &quot; 5 - 6, &quot; not 4 as the corresponding text, but he did so apparently tentatively. This is immaterial to the year of publication of the plate, however, as all three of these livraisons have been dated to 1809 (Evenhuis 1997 a; Bousquet 2016); the month and day of publication are unknown for all.]" type="journal article" year="1809">Latreille, 1809</bibRefCitation>
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<figureCitation id="0DC2DBF530C352FDEFF1F8A07441061D" box="[742,845,1799,1826]" captionStart="Fig" captionStartId="154.[189,232,1963,1989]" captionTargetBox="[286,1309,278,1880]" captionTargetId="figure-17@154.[274,1314,265,1913]" captionTargetPageId="154" captionText="Fig. 59. Species distribution. A. Onthophagus bidentatus Drapiez, 1819. B. Onthophagus curvicornis Latreille, 1809. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945627" httpUri="https://zenodo.org/record/13945627/files/figure.png" pageId="153" pageNumber="154">Fig. 59B</figureCitation>
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<treatmentCitation id="1458E16130C352FDEDAAF8EF75DF065C" author="Latreille P. A." box="[189,723,1864,1891]" page="220" pageId="153" pageNumber="154" year="1809">
<taxonomicName id="52F9BCF330C352FDEDAAF8EF75DF065C" ID-CoL="7XMPH" authority="Latreille, 1809: 220" authorityName="Latreille" authorityPageNumber="220" authorityYear="1809" box="[189,723,1864,1891]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="153" pageNumber="154" phylum="Arthropoda" rank="species" species="curvicornis">
Onthophagus curvicornis
<bibRefCitation id="F168BA8130C352FDECFBF8EF75DF065C" author="Latreille P. A." box="[492,723,1864,1891]" pageId="153" pageNumber="154" pagination="197 - 293" refId="ref127349" refString="Latreille P. A. 1809. Insectes de l'Amerique equinoxiale, recueillis pendant le voyage de MM. de Humboldt et Bonpland [livraison 4: pp. 197 - 293]. In: Humboldt A. &amp; Bonpland A. 1805 - 1810. Recueil d'observations de zoologie et d'anatomie comparee, faites dans l'ocean Atlantique, dans l'interieur du nouveau continent et dans la mer du Sud pendant les annees 1799, 1800, 1801, 1802 et 1803. Premier volume. Schoell and Dufour, Paris. [See comments for the previous entry. Evenhuis (1997 a) associated the publication of plate 23, containing the Onthophagus curvicornis figure, with livraisons &quot; 5 - 6, &quot; not 4 as the corresponding text, but he did so apparently tentatively. This is immaterial to the year of publication of the plate, however, as all three of these livraisons have been dated to 1809 (Evenhuis 1997 a; Bousquet 2016); the month and day of publication are unknown for all.]" type="journal article" year="1809">Latreille, 1809: 220</bibRefCitation>
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, pl. 23 fig. 2 (original description). Type locality:
<collectingCountry id="EDEE87E030C352FDE81AF8EF727F065D" box="[1293,1395,1864,1890]" name="Ecuador" pageId="153" pageNumber="154">Ecuador</collectingCountry>
:
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: San Francisco de Las Pampas,
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. Name-bearing type:
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(MZUF), designated by
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<bibRefCitation id="F168BA8130C352FDEC83F82875480696" author="Rossini M." box="[404,580,1935,1961]" pageId="153" pageNumber="154" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini (2021)</bibRefCitation>
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, not examined.
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</paragraph>
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<emphasis id="A78D1B6230C052FEEDAAF80C761006FA" bold="true" box="[189,284,1963,1989]" pageId="154" pageNumber="155">Fig. 59.</emphasis>
Species distribution.
<emphasis id="A78D1B6230C052FEEF37F80C753806FA" bold="true" box="[544,564,1963,1989]" pageId="154" pageNumber="155">A</emphasis>
.
<taxonomicName id="52F9BCF330C052FEEF53F80C731706FA" authority="Drapiez, 1819" authorityName="Drapiez" authorityYear="1819" box="[580,1051,1963,1989]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="A78D1B6230C052FEEF53F80C746906FA" box="[580,869,1963,1989]" italics="true" pageId="154" pageNumber="155">Onthophagus bidentatus</emphasis>
<bibRefCitation id="F168BA8130C052FEEE67F80C731706FA" author="Drapiez M." box="[880,1051,1963,1989]" pageId="154" pageNumber="155" pagination="130 - 138" refId="ref116912" refString="Drapiez M. 1819. Descriptions de quelques insectes nouveaux. Annales generales des Sciences physiques 1 (45 - 55): 130 - 138." type="journal article" year="1819">Drapiez, 1819</bibRefCitation>
</taxonomicName>
.
<emphasis id="A78D1B6230C052FEE93FF80C733606FA" bold="true" box="[1064,1082,1963,1989]" pageId="154" pageNumber="155">B</emphasis>
.
<taxonomicName id="52F9BCF330C052FEE95DF80C766106D7" authority="Latreille, 1809" authorityName="Latreille" authorityYear="1809" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="154" pageNumber="155" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="A78D1B6230C052FEE95DF80C727B06FA" box="[1098,1399,1963,1989]" italics="true" pageId="154" pageNumber="155">Onthophagus curvicornis</emphasis>
<bibRefCitation id="F168BA8130C052FEEDAAF869766106D7" author="Latreille P. A." box="[189,365,1998,2024]" pageId="154" pageNumber="155" pagination="197 - 293" refId="ref127349" refString="Latreille P. A. 1809. Insectes de l'Amerique equinoxiale, recueillis pendant le voyage de MM. de Humboldt et Bonpland [livraison 4: pp. 197 - 293]. In: Humboldt A. &amp; Bonpland A. 1805 - 1810. Recueil d'observations de zoologie et d'anatomie comparee, faites dans l'ocean Atlantique, dans l'interieur du nouveau continent et dans la mer du Sud pendant les annees 1799, 1800, 1801, 1802 et 1803. Premier volume. Schoell and Dufour, Paris. [See comments for the previous entry. Evenhuis (1997 a) associated the publication of plate 23, containing the Onthophagus curvicornis figure, with livraisons &quot; 5 - 6, &quot; not 4 as the corresponding text, but he did so apparently tentatively. This is immaterial to the year of publication of the plate, however, as all three of these livraisons have been dated to 1809 (Evenhuis 1997 a; Bousquet 2016); the month and day of publication are unknown for all.]" type="journal article" year="1809">Latreille, 1809</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C77030C152FFEDAAFEAE73880079" blockId="155.[189,1400,265,503]" pageId="155" pageNumber="156">
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<treatmentCitation id="1458E16130C152FFEDAAFEAE758A001B" author="Kirsch T." box="[189,646,265,292]" page="215" pageId="155" pageNumber="156" year="1866">
<taxonomicName id="52F9BCF330C152FFEDAAFEAE758A001B" ID-CoL="7XNLB" authority="Kirsch, 1866: 215" authorityName="Kirsch" authorityPageNumber="215" authorityYear="1866" box="[189,646,265,292]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="A78D1B6230C152FFEDAAFEAE76A7001C" box="[189,427,265,291]" italics="true" pageId="155" pageNumber="156">Onthophagus minax</emphasis>
<bibRefCitation id="F168BA8130C152FFECA5FEAE758A001B" author="Kirsch T." box="[434,646,265,292]" pageId="155" pageNumber="156" pagination="173 - 216" refId="ref125658" refString="Kirsch T. 1866. Beitrage zur Kaferfauna von Bogota. Berliner entomologische Zeitschrift 10: 173 - 216. https: // doi. org / 10.1002 / mmnd. 18660100113" type="journal article" year="1866">Kirsch, 1866: 215</bibRefCitation>
</taxonomicName>
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(original description). Type locality:
<collectingCountry id="EDEE87E030C152FFE957FEAE73B4001C" box="[1088,1208,265,291]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
:
<collectingRegion id="573D099230C152FFE9D4FEAE727F001C" box="[1219,1395,265,291]" country="Colombia" name="Cundinamarca" pageId="155" pageNumber="156">Cundinamarca</collectingRegion>
:
<collectingRegion id="573D099230C152FFEDFCFE8B76480079" box="[235,324,300,326]" country="Colombia" name="Bogota" pageId="155" pageNumber="156">Bogotá</collectingRegion>
. Name-bearing type:
<typeStatus id="4A4279D230C152FFEF28FE8B75A70079" box="[575,683,300,326]" pageId="155" pageNumber="156" type="lectotype">lectotype</typeStatus>
(SMTD), designated by
<treatmentCitation id="1458E16130C152FFEED8FE8B738D0079" author="Rossini M." box="[975,1153,300,326]" pageId="155" pageNumber="156" year="2021">
<bibRefCitation id="F168BA8130C152FFEED8FE8B738D0079" author="Rossini M." box="[975,1153,300,326]" pageId="155" pageNumber="156" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini (2021)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="9546C77030C152FFEDAAFEE8753F00C8" blockId="155.[189,1400,265,503]" pageId="155" pageNumber="156">
<treatmentCitationGroup id="B5E9E05E30C152FFEDAAFEE8753F00C8" pageId="155" pageNumber="156">
<treatmentCitation id="1458E16130C152FFEDAAFEE875D30056" author="d'Orbigny H." box="[189,735,335,362]" page="69" pageId="155" pageNumber="156" year="1902">
<taxonomicName id="52F9BCF330C152FFEDAAFEE875D30056" ID-CoL="7XNBX" authority="d'Orbigny, 1902: 69" authorityName="d'Orbigny" authorityPageNumber="69" authorityYear="1902" box="[189,735,335,362]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="laevatus">
<emphasis id="A78D1B6230C152FFEDAAFEE876C20056" box="[189,462,335,361]" italics="true" pageId="155" pageNumber="156">Onthophagus laevatus</emphasis>
<bibRefCitation id="F168BA8130C152FFECC9FEE875D30056" author="d'Orbigny H." box="[478,735,335,362]" pageId="155" pageNumber="156" pagination="1 - 324" refId="ref134097" refString="d'Orbigny H. 1902. Memoire sur les Onthophagides d'Afrique. Annales de la Societe entomologique de France 71: 1 - 324. Available from https: // www. biodiversitylibrary. org / page / 8251405 [accessed 16 Feb. 2024]." type="journal article" year="1902">dOrbigny, 1902: 69</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality: unknown. The lectotypes provenance label indicates “Delagoa”, i.e., Delagoa Bay, or, as the locality is modernly called,
<collectingRegion id="573D099230C152FFEC54FE317692008F" box="[323,414,406,432]" country="Mozambique" name="Maputo" pageId="155" pageNumber="156">Maputo</collectingRegion>
Bay, in
<collectingCountry id="EDEE87E030C152FFEF1EFE3175AB008F" box="[521,679,406,432]" name="Mozambique" pageId="155" pageNumber="156">Mozambique</collectingCountry>
, but this is certainly incorrect, as the species is exclusively South American (
<treatmentCitation id="1458E16130C152FFECD7FE1E756800EC" author="Rossini M." box="[448,612,441,467]" pageId="155" pageNumber="156" year="2021">
<bibRefCitation id="F168BA8130C152FFECD7FE1E756800EC" author="Rossini M." box="[448,612,441,467]" pageId="155" pageNumber="156" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini 2021</bibRefCitation>
</treatmentCitation>
). Name-bearing type:
<typeStatus id="4A4279D230C152FFEE61FE1E74EE00EC" box="[886,994,441,467]" pageId="155" pageNumber="156" type="lectotype">lectotype</typeStatus>
(MNHN), designated by
<treatmentCitation id="1458E16130C152FFE809FE1E763700C9" author="Rossini M." pageId="155" pageNumber="156" year="2021">
<bibRefCitation id="F168BA8130C152FFE809FE1E763700C9" author="Rossini M." pageId="155" pageNumber="156" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini (2021)</bibRefCitation>
</treatmentCitation>
, not examined by us.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C152FFEDAAFDB974A704F4" blockId="155.[189,1400,541,1765]" pageId="155" pageNumber="156">
<treatmentCitationGroup id="B5E9E05E30C152FFEDAAFDB974A704F4" pageId="155" pageNumber="156">
<taxonomicName id="52F9BCF330C152FFEDAAFDB976E60308" ID-CoL="7XMPH" authorityName="Latreille" authorityYear="1809" box="[189,490,542,568]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="A78D1B6230C152FFEDAAFDB976E60308" box="[189,490,542,568]" italics="true" pageId="155" pageNumber="156">Onthophagus curvicornis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C152FFEF1BFDB975F60307" author="Harold E." box="[524,762,541,568]" page="1028" pageId="155" pageNumber="156" year="1869">
<bibRefCitation id="F168BA8130C152FFEF1BFDB975F60307" author="Harold E." box="[524,762,541,568]" pageId="155" pageNumber="156" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1028</bibRefCitation>
</treatmentCitation>
(catalog, cited for Quito);
<treatmentCitation id="1458E16130C152FFE92BFDB973A90307" author="Harold E." box="[1084,1189,542,568]" page="29" pageId="155" pageNumber="156" year="1880">
<bibRefCitation id="F168BA8130C152FFE92BFDB973A90307" author="Harold E." box="[1084,1189,542,568]" pageId="155" pageNumber="156" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">1880: 29</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E16130C152FFEDFBFDE676F60364" author="Taschenberg E. L." box="[236,506,577,603]" page="184" pageId="155" pageNumber="156" year="1870">
<bibRefCitation id="F168BA8130C152FFEDFBFDE676F60364" author="Taschenberg E. L." box="[236,506,577,603]" pageId="155" pageNumber="156" pagination="177 - 199" refId="ref139821" refString="Taschenberg E. L. 1870. Neue Kafer aus Colombien und Ecuador. Zeitschrift fur die Gesammten Naturwissenschaft 35: 177 - 199." type="journal article" year="1870">Taschenberg 1870: 184</bibRefCitation>
</treatmentCitation>
(cited for
<collectingRegion id="573D099230C152FFEF66FDE675A60364" box="[625,682,577,603]" country="Ecuador" name="Loja" pageId="155" pageNumber="156">Loja</collectingRegion>
,
<collectingCountry id="EDEE87E030C152FFEFA3FDE674170364" box="[692,795,577,603]" name="Ecuador" pageId="155" pageNumber="156">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEE47FDE6734A0364" author="Bates H. W." box="[848,1094,577,603]" page="66" pageId="155" pageNumber="156" year="1886">
<bibRefCitation id="F168BA8130C152FFEE47FDE6734A0364" author="Bates H. W." box="[848,1094,577,603]" firstAuthor="Bates" pageId="155" pageNumber="156" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1886">Bates 18861890: 66</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E16130C152FFE80FFDE676420340" author="Campos F." page="57" pageId="155" pageNumber="156" year="1921">
<bibRefCitation id="F168BA8130C152FFE80FFDE676420340" author="Campos F." pageId="155" pageNumber="156" pagination="26 - 100" refId="ref113384" refString="Campos F. 1921. Estudios sobre la fauna entomologica del Ecuador. Revista del Colegio Nacional Vicente Rocafuerte 6: 26 - 100." type="journal article" year="1921">Campos 1921: 57</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030C152FFECB8FDC3751A0341" box="[431,534,612,638]" name="Ecuador" pageId="155" pageNumber="156">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEF5BFDC374BF0340" author="Gillet J. J. E. &amp; Boucomont A." box="[588,947,612,639]" page="205" pageId="155" pageNumber="156" year="1927">
<bibRefCitation id="F168BA8130C152FFEF5BFDC374BF0340" author="Gillet J. J. E. &amp; Boucomont A." box="[588,947,612,639]" pageId="155" pageNumber="156" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 205</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C152FFE97BFDC2727E0341" author="Boucomont A." box="[1132,1394,612,639]" page="308" pageId="155" pageNumber="156" year="1932">
<bibRefCitation id="F168BA8130C152FFE97BFDC2727E0341" author="Boucomont A." box="[1132,1394,612,639]" pageId="155" pageNumber="156" pagination="293 - 332" refId="ref112419" refString="Boucomont A. 1932. Synopsis des Onthophagus d'Amerique du Sud (Col. Scarab.). Annales de la Societe entomologique de France 101: 293 - 332. https: // doi. org / 10.1080 / 21686351.1932.12280217" type="journal article" year="1932">Boucomont 1932: 308</bibRefCitation>
</treatmentCitation>
, 323 (synopsis, key). —
<treatmentCitation id="1458E16130C152FFEF15FD20741F039E" author="Blackwelder R. E." box="[514,787,647,673]" page="211" pageId="155" pageNumber="156" year="1944">
<bibRefCitation id="F168BA8130C152FFEF15FD20741F039E" author="Blackwelder R. E." box="[514,787,647,673]" pageId="155" pageNumber="156" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C152FFEE9FFD207354039E" author="Guerin J." box="[904,1112,647,674]" page="262" pageId="155" pageNumber="156" year="1953">
<bibRefCitation id="F168BA8130C152FFEE9FFD207354039E" author="Guerin J." box="[904,1112,647,674]" pageId="155" pageNumber="156" refId="ref121980" refString="Guerin J. 1953. Coleopteros do Brasil. Faculdade de Filosofia, Ciencias e Letras da Universidade de Sao Paulo, Sao Paulo." type="book" year="1953">Guérin 1953: 262</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C152FFE9DCFD2F727A039D" author="Roze J. A." box="[1227,1398,647,674]" page="45" pageId="155" pageNumber="156" year="1955">
<bibRefCitation id="F168BA8130C152FFE9DCFD2F727A039D" author="Roze J. A." box="[1227,1398,647,674]" pageId="155" pageNumber="156" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 45</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C152FFEC9DFD0D750403FB" box="[394,520,682,708]" name="Venezuela" pageId="155" pageNumber="156">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEF2BFD0D748203FA" author="Vulcano M. A. &amp; Pereira F. S." box="[572,910,682,709]" page="565" pageId="155" pageNumber="156" year="1967">
<bibRefCitation id="F168BA8130C152FFEF2BFD0D748203FA" author="Vulcano M. A. &amp; Pereira F. S." box="[572,910,682,709]" pageId="155" pageNumber="156" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 565</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130C152FFE914FD0D73CE03FA" author="Blanco J." box="[1027,1218,682,709]" page="47" pageId="155" pageNumber="156" year="1988">
<bibRefCitation id="F168BA8130C152FFE914FD0D73CE03FA" author="Blanco J." box="[1027,1218,682,709]" pageId="155" pageNumber="156" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 47</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C152FFEDFEFD6976C903D8" author="Havranek D." box="[233,453,717,744]" page="61" pageId="155" pageNumber="156" year="1989">
<bibRefCitation id="F168BA8130C152FFEDFEFD6976C903D8" author="Havranek D." box="[233,453,717,744]" pageId="155" pageNumber="156" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C152FFEF39FD6975E703D7" author="Deloya C." box="[558,747,718,744]" page="22" pageId="155" pageNumber="156" year="1992">
<bibRefCitation id="F168BA8130C152FFEF39FD6975E703D7" author="Deloya C." box="[558,747,718,744]" pageId="155" pageNumber="156" pagination="1 - 11" refId="ref116640" refString="Deloya C. 1992. Necrophilous Scarabaeidae and Trogidae beetles of tropical deciduous forest in Tepexco, Puebla, Mexico. Acta Zoologica Mexicana (n. s.) 52: 1 - 11. https: // doi. org / 10.21829 / azm. 1992.49521952" type="journal article" year="1992">Deloya 1992: 22</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C152FFEE42FD69739103D8" author="Zunino M. &amp; Halffter G." box="[853,1181,717,744]" page="161" pageId="155" pageNumber="156" year="1997">
<bibRefCitation id="F168BA8130C152FFEE42FD69739103D8" author="Zunino M. &amp; Halffter G." box="[853,1181,717,744]" pageId="155" pageNumber="156" pagination="157 - 178" refId="ref142302" refString="Zunino M. &amp; Halffter G. 1997. Sobre Onthophagus Latreille, 1802 americanos (Coleoptera: Scarabaeidae: Scarabaeinae). Elytron 11: 157 - 178." type="journal article" year="1997">Zunino &amp; Halffter 1997: 161</bibRefCitation>
</treatmentCitation>
(classification). —
<treatmentCitation id="1458E16130C152FFEDFDFD5676C50234" author="Escobar F." box="[234,457,753,779]" page="209" pageId="155" pageNumber="156" year="2000">
<bibRefCitation id="F168BA8130C152FFEDFDFD5676C50234" author="Escobar F." box="[234,457,753,779]" pageId="155" pageNumber="156" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 209</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C152FFEF61FD5675E30234" box="[630,751,753,779]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEE3EFD5673460234" author="Vaz-de-Mello F. Z." box="[809,1098,753,779]" page="194" pageId="155" pageNumber="156" year="2000">
<bibRefCitation id="F168BA8130C152FFEE3EFD5673460234" author="Vaz-de-Mello F. Z." box="[809,1098,753,779]" pageId="155" pageNumber="156" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C152FFE9EFFD5672490234" box="[1272,1349,753,779]" name="Brazil" pageId="155" pageNumber="156">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEDFBFCB375090211" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[236,517,788,814]" page="139" pageId="155" pageNumber="156" year="2001">
<bibRefCitation id="F168BA8130C152FFEDFBFCB375090211" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[236,517,788,814]" pageId="155" pageNumber="156" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230C152FFEC5BFCB2768B0211" box="[332,391,788,814]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2001: 139
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C152FFEFA6FCB374260211" box="[689,810,788,814]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
);
<treatmentCitation id="1458E16130C152FFEE29FCB374E30210" author="Medina C. A. &amp; Escobar F. &amp; Kattan G. H." box="[830,1007,788,815]" page="182" pageId="155" pageNumber="156" year="2002">
<bibRefCitation id="F168BA8130C152FFEE29FCB374E30210" author="Medina C. A. &amp; Escobar F. &amp; Kattan G. H." box="[830,1007,788,815]" pageId="155" pageNumber="156" pagination="181 - 187" refId="ref130836" refString="Medina C. A., Escobar F. &amp; Kattan G. H. 2002. Diversity and habitat use of dung beetles in a restored Andean landscape. Biotropica 34: 181 - 187. https: // doi. org / 10.1111 / j. 1744 - 7429.2002. tb 00255. x" type="journal article" year="2002">2002: 182185</bibRefCitation>
</treatmentCitation>
(ecology). —
<treatmentCitation id="1458E16130C152FFE98CFCB3766D026E" author="Cancino E. R. &amp; Blanco J. M. C." page="120" pageId="155" pageNumber="156" year="2002">
<bibRefCitation id="F168BA8130C152FFE98CFCB3766D026E" author="Cancino E. R. &amp; Blanco J. M. C." pageId="155" pageNumber="156" refId="ref113409" refString="Cancino E. R. &amp; Blanco J. M. C. 2002. Artropodos Terrestres de los Estados de Tamaulipas y Nuevo Leon, Mexico (Vol. 4). Universidad Autonoma de Tamaulipas, Unidad Academica Multidisciplinaria Agronomia y Ciencias, Centro de Investigacion y Desarrollo Agropecuario, Forestal y de la Fauna. Universidad Autonoma de Tamaulipas, Victoria, Tamaulipas." type="book" year="2002">Cancino &amp; Blanco 2002: 120</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130C152FFECC5FC907599026E" author="Zunino M." box="[466,661,823,849]" page="71" pageId="155" pageNumber="156" year="2003">
<bibRefCitation id="F168BA8130C152FFECC5FC907599026E" author="Zunino M." box="[466,661,823,849]" pageId="155" pageNumber="156" pagination="66 - 74" refId="ref142252" refString="Zunino M. 2003 Tribu Onthophagini. In: Moron M. A. (ed.) Atlas de los Escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 66 - 74. Argania, Barcelona." type="book chapter" year="2003">Zunino 2003: 71</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E030C152FFEE07FC907463026E" box="[784,879,823,849]" name="Mexico" pageId="155" pageNumber="156">Mexico</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEEB1FC90737F026E" author="Noriega J. A." box="[934,1139,823,849]" page="40" pageId="155" pageNumber="156" year="2004">
<bibRefCitation id="F168BA8130C152FFEEB1FC90737F026E" author="Noriega-Alvarado J. A." box="[934,1139,823,849]" firstAuthor="Noriega" pageId="155" pageNumber="156" pagination="37 - 44" refId="ref132303" refString="Noriega-Alvarado J. A. 2004. Preliminary checklist of the scarab community (Coleoptera: Scarabaeidae) at CIEM, Tinigua National Park, Meta-Colombia. Field Studies of Fauna and Flora, La Macarena, Colombia 14: 37 - 44." type="journal article" year="2004">Noriega 2004: 40</bibRefCitation>
</treatmentCitation>
(checklist for Tinigua Park,
<collectingCountry id="EDEE87E030C152FFEC39FCFD76AB024B" box="[302,423,858,884]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFECCAFCFC746C024B" author="Ramirez G. J. C. &amp; Locarno P. L. C." box="[477,864,858,885]" page="61" pageId="155" pageNumber="156" year="2004">
<bibRefCitation id="F168BA8130C152FFECCAFCFC746C024B" author="Ramirez G. J. C. &amp; Locarno P. L. C." box="[477,864,858,885]" pageId="155" pageNumber="156" pagination="59 - 63" refId="ref136737" refString="Ramirez G. J. C. &amp; Locarno P. L. C. 2004. Escarabajos Scarabaeinae saprofagos (Coleoptera: Scarabaeidae) en un bosque muy humedo premontano de los Andes Occidentales Colombianos. Ecologia Aplicada 3: 59 - 63. https: // doi. org / 10.21704 / rea. v 3 i 1 - 2.271" type="journal article" year="2004">Ramírez &amp; Locarno 2004: 6162</bibRefCitation>
</treatmentCitation>
(ecology, list). —
<treatmentCitation id="1458E16130C152FFE92EFCFD73F1024A" author="Krajcik M." box="[1081,1277,858,885]" page="99" pageId="155" pageNumber="156" year="2006">
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<emphasis id="A78D1B6230C152FFE90FFCD9724802A8" box="[1048,1348,894,920]" italics="true" pageId="155" pageNumber="156">Onthophagus curvicornis</emphasis>
var.
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Pulido-Herrera
<emphasis id="A78D1B6230C152FFEF51FC05758C0284" box="[582,640,929,955]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2007: 307
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<bibRefCitation id="F168BA8130C152FFE816FC0676C702E1" author="Orozco J. &amp; Perez M." firstAuthor="Orozco" journalOrPublisher="Revista Brasileira de Entomologia" pageId="155" pageNumber="156" pagination="36 - 40" part="52" publicationUrl="https://doi.org/10.1590/S0085-56262008000100007" refId="ref134201" refString="Orozco J. &amp; Perez M. 2008. Escarabajos coprofagos (Coleoptera, Scarabaeoidea) del Parque Nacional Los Estoraques (Norte de Santander, Colombia). Revista Brasileira de Entomologia 52: 36 - 40. https: // doi. org / 10.1590 / S 0085 - 56262008000100007" title="Escarabajos coprofagos (Coleoptera, Scarabaeoidea) del Parque Nacional Los Estoraques (Norte de Santander, Colombia)" type="journal article" year="2008">Orozco &amp; Pérez 2008: 3839</bibRefCitation>
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<bibRefCitation id="F168BA8130C152FFEFFBFC63738202E1" author="Concha-Lozada C. M. &amp; Gallego M. C. &amp; Pardo-Locarno L. C." box="[748,1166,964,990]" pageId="155" pageNumber="156" pagination="43 - 55" refId="ref114591" refString="Concha-Lozada C. M., Gallego M. C. &amp; Pardo-Locarno L. C. 2010. Fragmentacion de ecosistemas montanos e impactos estructurales y poblacionales sobre la comunidad de escarabajos coprofagos (Coleoptera: Scarabaeinae) en el Alto Rio Cauca, Popayan, Colombia. Boletin Cientifico. Centro de Museos. Museo de Historia Natural 14: 43 - 55." type="journal article" year="2010">
Concha-Lozada
<emphasis id="A78D1B6230C152FFEEA6FC6274E202E1" box="[945,1006,964,990]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2010: 4344
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, 4749, 52 (list for Popayán,
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Carvajal
<emphasis id="A78D1B6230C152FFEF92FC4F75CE053E" box="[645,706,999,1025]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2011: 322323
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(checklist for
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<treatmentCitation id="1458E16130C152FFE9C1FC40761A051B" author="Deloya A. C. L." page="386" pageId="155" pageNumber="156" year="2011">
<bibRefCitation id="F168BA8130C152FFE9C1FC40761A051B" author="Deloya A. C. L." pageId="155" pageNumber="156" pagination="383 - 390" refId="ref116688" refString="Deloya A. C. L. 2011. Escarabajos coprofagos y necrofagos (Insecta: Coleoptera: Scarabaeidae). In: Aragon A., Hernandez F., Ortiz V. &amp; Mavil J. (eds) La Biodiversidad en Veracruz, Estudio de Estado. Diversidad de Especies. Comision Nacional para el Conocimiento y Uso de la Biodiversidad, Gobierno del Estado de Veracruz: 383 - 390. Universidad Veracruzana, Instituto de Ecologia, A. C. Mexico, Xalapa." type="book chapter" year="2011">Deloya 2011: 386</bibRefCitation>
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<bibRefCitation id="F168BA8130C152FFECD9FBAD75E4051A" author="Giraldo C. &amp; Escobar F. &amp; Chara J. D. &amp; Calle Z." box="[462,744,1034,1061]" pageId="155" pageNumber="156" pagination="115 - 122" refId="ref121491" refString="Giraldo C., Escobar F., Chara J. D. &amp; Calle Z. 2011. The adoption of silvopastoral systems promotes the recovery of ecological processes regulated by dung beetles in the Colombian Andes. Insect Conservation and Diversity 4: 115 - 122. https: // doi. org / 10.1111 / j. 1752 - 4598.2010.00112. x" type="journal article" year="2011">
Giraldo
<emphasis id="A78D1B6230C152FFEF27FBAC7560051B" box="[560,620,1034,1060]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2011: 118
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(comments). —
<treatmentCitation id="1458E16130C152FFEEA6FBAD721D051A" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[945,1297,1034,1061]" page="39" pageId="155" pageNumber="156" year="2012">
<bibRefCitation id="F168BA8130C152FFEEA6FBAD721D051A" author="Cultid-Medina C. A. &amp; Medina C. A. U. &amp; Martinez-Quintero B. G. &amp; Escobar A. F. V. &amp; Constantino L. M. C. &amp; Betancur N. J." box="[945,1297,1034,1061]" pageId="155" pageNumber="156" refId="ref114949" refString="Cultid-Medina C. A., Medina C. A. U., Martinez-Quintero B. G., Escobar A. F. V., Constantino L. M. C. &amp; Betancur N. J. 2012. Escarabajos Coprofagos (Scarabaeinae) del Eje Cafetero: Guia para el Estudio Ecologico. WCS Colombia, Cenicafe, Villa Maria, Caldas." type="book" year="2012">
Cultid-Medina
<emphasis id="A78D1B6230C152FFE971FBAC73AE051B" box="[1126,1186,1034,1060]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2012: 39
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<bibRefCitation id="F168BA8130C152FFEDFCFB8A76680578" author="Cultid-Medina C. A. &amp; Martinez-Quintero B. G. &amp; Escobar F. &amp; de Ulloa P. C." box="[235,356,1069,1096]" pageId="155" pageNumber="156" pagination="617 - 626" refId="ref115017" refString="Cultid-Medina C. A., Martinez-Quintero B. G., Escobar F. &amp; de Ulloa P. C. 2015. Movement and population size of two dung beetle species in an Andean agricultural landscape dominated by sun-grown coffee. Journal of Insect Conservation 19: 617 - 626. https: // doi. org / 10.1007 / s 10841 - 015 - 9784 - 3" type="journal article" year="2015">2015: 624</bibRefCitation>
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<bibRefCitation id="F168BA8130C152FFEF26FB8974530577" author="Delgado L. &amp; Mora-Aguilar E. F. &amp; Escobar-Hernandez F." box="[561,863,1070,1096]" pageId="155" pageNumber="156" pagination="319 - 332" refId="ref116588" refString="Delgado L., Mora-Aguilar E. F. &amp; Escobar-Hernandez F. 2012. Scarabaeoidea (Coleoptera) of the municipality of Xalapa, Veracruz, Mexico: Inventory and analysis. The Coleopterists Bulletin 66: 319 - 332. https: // doi. org / 10.1649 / 072.066.0405" type="journal article" year="2012">
Delgado
<emphasis id="A78D1B6230C152FFEFB7FB8975D70577" box="[672,731,1070,1096]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2012: 331
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<treatmentCitation id="1458E16130C152FFEEDBFB8973EB0577" author="Noriega J. A. &amp; Palacio M. J. &amp; Monroy-G. J. D. &amp; Valencia E." box="[972,1255,1070,1096]" page="43" pageId="155" pageNumber="156" year="2012">
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Noriega
<emphasis id="A78D1B6230C152FFE920FB89737D0577" box="[1079,1137,1070,1096]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2012: 43
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Ferrer-Paris
<emphasis id="A78D1B6230C152FFEFFEFBF5742B0554" box="[745,807,1105,1131]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2013: 99100
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, 109 (list). —
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, 199, 201, 203 (list, Appendix 8, 11, 13). —
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Molina
<emphasis id="A78D1B6230C152FFEE5EFB1C748805EB" box="[841,900,1210,1236]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2016: 128130
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<bibRefCitation id="F168BA8130C152FFECB7FB79740905C7" author="Alburqueque D. S. &amp; Vaz-de-Mello F. Z. &amp; Cherre A. U. &amp; Timana C. P." box="[416,773,1246,1272]" pageId="155" pageNumber="156" pagination="108 - 116" refId="ref109138" refString="Alburqueque D. S., Vaz-de-Mello F. Z., Cherre A. U. &amp; Timana C. P. 2017. Coleopteros (Coleoptera: Scarabaeidae) de los bosques de niebla, Ramos y Chin Chin, Ayabaca-Huancabamba, Piura-Peru. INDES Revista de Investigacion para el Desarrollo Sustentable 3: 108 - 116. https: // doi. org / 10.25127 / indes. 20153.138" type="journal article" year="2017">
Albuquerque
<emphasis id="A78D1B6230C152FFEF51FB79758E05C7" box="[582,642,1246,1272]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2017: 211
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). —
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, 2426 (biology). —
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Torres
<emphasis id="A78D1B6230C152FFEF3BFAA5756B0424" box="[556,615,1281,1307]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2017: 4849
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,
<collectingCountry id="EDEE87E030C152FFE919FAA6738B0424" box="[1038,1159,1281,1307]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFE9A9FAA676A20401" author="Villada-Bedoya S. &amp; Cultid-Medina C. A. &amp; Escobar F. &amp; Guevara R. &amp; Zurita G." page="197" pageId="155" pageNumber="156" year="2017">
<bibRefCitation id="F168BA8130C152FFE9A9FAA676A20401" author="Villada-Bedoya S. &amp; Cultid-Medina C. A. &amp; Escobar F. &amp; Guevara R. &amp; Zurita G." pageId="155" pageNumber="156" pagination="195 - 205" refId="ref141161" refString="Villada-Bedoya S., Cultid-Medina C. A., Escobar F., Guevara R. &amp; Zurita G. 2017. Edge effects on dung beetle assemblages in an Andean mosaic of forest and coffee plantations. Biotropica 49: 195 - 205. https: // doi. org / 10.1111 / btp. 12373" type="journal article" year="2017">
Villada-Bedoya
<emphasis id="A78D1B6230C152FFEDFDFA8276240401" box="[234,296,1316,1342]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2017: 197
</bibRefCitation>
</treatmentCitation>
, 200, 203 (ecology). —
<treatmentCitation id="1458E16130C152FFEFCBFA83731C0401" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[732,1040,1316,1342]" page="97" pageId="155" pageNumber="156" year="2018">
<bibRefCitation id="F168BA8130C152FFEFCBFA83731C0401" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[732,1040,1316,1342]" pageId="155" pageNumber="156" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B6230C152FFEE4AFA8274970401" box="[861,923,1316,1342]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2018: 97
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E030C152FFE968FA8373EA0401" box="[1151,1254,1316,1342]" name="Ecuador" pageId="155" pageNumber="156">Ecuador</collectingCountry>
);
<treatmentCitation id="1458E16130C152FFE9EAFA83727A0401" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[1277,1398,1316,1342]" httpUri="http://treatment.plazi.org/id/E9FA5E7AD68F84ED2BFA6F25746F5E61" page="186" pageId="155" pageNumber="156" year="2019">
<bibRefCitation id="F168BA8130C152FFE9EAFA83727A0401" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[1277,1398,1316,1342]" pageId="155" pageNumber="156" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 186</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E030C152FFEC8CFAE0750E045E" box="[411,514,1351,1377]" name="Ecuador" pageId="155" pageNumber="156">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEF57FAE0746B045E" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[576,871,1351,1377]" page="104" pageId="155" pageNumber="156" year="2018">
<bibRefCitation id="F168BA8130C152FFEF57FAE0746B045E" author="Giraldo C. &amp; Montoya S. &amp; Escobar F." box="[576,871,1351,1377]" pageId="155" pageNumber="156" refId="ref121556" refString="Giraldo C., Montoya S. &amp; Escobar F. 2018. Escarabajos del Estiercol en Paisajes Ganaderos de Colombia. Fundacion CIPAV, Cali, Colombia." type="book" year="2018">
Giraldo
<emphasis id="A78D1B6230C152FFEFB3FAEF75ED045E" box="[676,737,1351,1377]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2018: 104
</bibRefCitation>
</treatmentCitation>
, 111, 132 (guide). —
<treatmentCitation id="1458E16130C152FFE961FAE0727A045E" author="Giron Y. &amp; Calderon C. &amp; Noriega J." box="[1142,1398,1351,1377]" page="83" pageId="155" pageNumber="156" year="2018">
<bibRefCitation id="F168BA8130C152FFE961FAE0727A045E" author="Giron Y. &amp; Calderon C. &amp; Noriega J." box="[1142,1398,1351,1377]" pageId="155" pageNumber="156" pagination="83 - 98" refId="ref121585" refString="Giron Y., Calderon C. &amp; Noriega J. 2018. Estructura del ensamblaje de escarabajos coprofagos (Coleoptera: Scarabaeinae) lo largo de un gradiente altitudinal en el Refugio de vida silvestre Laquipampa, Ferrenafe, Lambayeque, Peru. Revista de Ciencia, Tecnologia y Humanidades 7: 83 - 98." type="journal article" year="2018">
Girón
<emphasis id="A78D1B6230C152FFE9D3FAEF720E045E" box="[1220,1282,1351,1377]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2018: 83
</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E16130C152FFECBEFACD75BC04BA" author="Rossini M. &amp; Vaz-de-Mello F. Z. &amp; Zunino M." box="[425,688,1386,1413]" page="9" pageId="155" pageNumber="156" year="2018">
<bibRefCitation id="F168BA8130C152FFECBEFACD75BC04BA" author="Rossini M. &amp; Vaz-de-Mello F. Z. &amp; Zunino M." box="[425,688,1386,1413]" pageId="155" pageNumber="156" pagination="1 - 21" refId="ref137851" refString="Rossini M., Vaz-de-Mello F. Z. &amp; Zunino M. 2018 b. Toward a comprehensive taxonomic revision of the &quot; hirculus &quot; group of American Onthophagus Latreille, 1802 (Coleoptera, Scarabaeidae, Scarabaeinae). European Journal of Taxonomy 432: 1 - 21. https: // doi. org / 10.5852 / ejt. 2018.432" type="journal article" year="2018">
Rossini
<emphasis id="A78D1B6230C152FFEF10FACC754D04BB" box="[519,577,1386,1412]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2018b: 9
</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="1458E16130C152FFEE4CFACC73D904BB" author="Martinez D. C. &amp; Morales I." box="[859,1237,1386,1413]" page="1689" pageId="155" pageNumber="156" year="2020">
<bibRefCitation id="F168BA8130C152FFEE4CFACC73D904BB" author="Martinez D. C. &amp; Morales I." box="[859,1237,1386,1413]" pageId="155" pageNumber="156" pagination="1679 - 1693" refId="ref130356" refString="Martinez D. C. &amp; Morales I. 2020. Annotated list of beetles (Insecta, Coleoptera) in an urban area of the Eastern Andes of Colombia. CheckList 16 (6): 1679 - 1693. https: // doi. org / 10.15560 / 16.6.1679" type="journal article" year="2020">Martínez &amp; Morales 2020: 1689</bibRefCitation>
</treatmentCitation>
(checklist for the Eastern Andes,
<collectingCountry id="EDEE87E030C152FFECDAFA29754A0497" box="[461,582,1422,1448]" name="Colombia" pageId="155" pageNumber="156">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEF69FA29748D0498" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[638,897,1421,1448]" page="136" pageId="155" pageNumber="156" year="2020">
<bibRefCitation id="F168BA8130C152FFEF69FA29748D0498" author="Nieto M. J. A. &amp; Giraldo E. C. &amp; Quevedo V. C. J. &amp; Chara J. &amp; Medina A. C." box="[638,897,1421,1448]" pageId="155" pageNumber="156" pagination="134 - 141" refId="ref132145" refString="Nieto M. J. A., Giraldo E. C., Quevedo V. C. J., Chara J. &amp; Medina A. C. 2020. Escarabajos estercoleros asociados a sistemas de ganaderia sostenible en diferentes regiones de Colombia. Biota Colombiana 21: 134 - 141. https: // doi. org / 10.21068 / c 2020. v 21 n 02 a 09" type="journal article" year="2020">
Nieto
<emphasis id="A78D1B6230C152FFEFDFFA29740F0497" box="[712,771,1422,1448]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2020: 136
</bibRefCitation>
</treatmentCitation>
(report). —
<treatmentCitation id="1458E16130C152FFE901FA29762404F4" author="Carrion-Paladines V. &amp; Fries A. &amp; Munoz A. &amp; Castillo E. &amp; Garcia-Ruiz R." page="9" pageId="155" pageNumber="156" year="2021">
<bibRefCitation id="F168BA8130C152FFE901FA29762404F4" author="Carrion-Paladines V. &amp; Fries A. &amp; Munoz A. &amp; Castillo E. &amp; Garcia-Ruiz R." pageId="155" pageNumber="156" pagination="1 - 22" refId="ref113769" refString="Carrion-Paladines V., Fries A., Munoz A., Castillo E. &amp; Garcia-Ruiz R. 2021. Effects of land-use change on the community structure of the dung beetle (Scarabaeinae) in an altered ecosystem in southern Ecuador. Insects 12: 1 - 22. https: // doi. org / 10.3390 / insects 12040306" type="journal article" year="2021">
Carrión-Paladines
<emphasis id="A78D1B6230C152FFE9E6FA2972200497" box="[1265,1324,1422,1448]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2021: 910
</bibRefCitation>
</treatmentCitation>
, 14, 1617 (ecology). —
<treatmentCitation id="1458E16130C152FFEF43FA16742604F4" author="Rossini M." box="[596,810,1457,1483]" httpUri="http://treatment.plazi.org/id/039387FECD67FFD7FF71B0F7EC9B8DDD" page="266" pageId="155" pageNumber="156" year="2021">
<bibRefCitation id="F168BA8130C152FFEF43FA16742604F4" author="Rossini M." box="[596,810,1457,1483]" pageId="155" pageNumber="156" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini 2021: 266</bibRefCitation>
</treatmentCitation>
(revision).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C152FFEDAAFA73744A0767" blockId="155.[189,1400,541,1765]" pageId="155" pageNumber="156">
<treatmentCitationGroup id="B5E9E05E30C152FFEDAAFA73744A0767" pageId="155" pageNumber="156">
<taxonomicName id="52F9BCF330C152FFEDAAFA7376A004D1" ID-CoL="7XNLB" authorityName="Kirsch" authorityYear="1866" box="[189,428,1492,1518]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="minax">
<emphasis id="A78D1B6230C152FFEDAAFA7376A004D1" box="[189,428,1492,1518]" italics="true" pageId="155" pageNumber="156">Onthophagus minax</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C152FFECDBFA7375B604D1" author="Harold E." box="[460,698,1492,1519]" page="1028" pageId="155" pageNumber="156" year="1869">
<bibRefCitation id="F168BA8130C152FFECDBFA7375B604D1" author="Harold E." box="[460,698,1492,1519]" pageId="155" pageNumber="156" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">Harold 1869d: 1028</bibRefCitation>
</treatmentCitation>
(cited for Bogotá, cited as synonym of
<taxonomicName id="52F9BCF330C152FFE981FA73723304D1" ID-CoL="7XMPH" authorityName="Latreille" authorityYear="1809" box="[1174,1343,1492,1518]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="A78D1B6230C152FFE981FA73723304D1" box="[1174,1343,1492,1518]" italics="true" pageId="155" pageNumber="156">O. curvicornis</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C152FFEDFBFA50756F072D" author="Gillet J. J. E. &amp; Boucomont A." box="[236,611,1527,1554]" page="205" pageId="155" pageNumber="156" year="1927">
<bibRefCitation id="F168BA8130C152FFEDFBFA50756F072D" author="Gillet J. J. E. &amp; Boucomont A." box="[236,611,1527,1554]" pageId="155" pageNumber="156" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 205</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C152FFEE30FA507330072E" author="Blackwelder R. E." box="[807,1084,1527,1553]" page="211" pageId="155" pageNumber="156" year="1944">
<bibRefCitation id="F168BA8130C152FFEE30FA507330072E" author="Blackwelder R. E." box="[807,1084,1527,1553]" pageId="155" pageNumber="156" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
</treatmentCitation>
(list, cited as synonym of
<taxonomicName id="52F9BCF330C152FFEDFAF9BC7699070B" ID-CoL="7XMPH" authorityName="Latreille" authorityYear="1809" box="[237,405,1563,1589]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="A78D1B6230C152FFEDFAF9BC7699070B" box="[237,405,1563,1589]" italics="true" pageId="155" pageNumber="156">O. curvicornis</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C152FFECC0F9BD748A070A" author="Pulido-Herrera L. A. &amp; Zunino M." box="[471,902,1562,1589]" page="101" pageId="155" pageNumber="156" year="2007">
<bibRefCitation id="F168BA8130C152FFECC0F9BD748A070A" author="Pulido-Herrera L. A. &amp; Zunino M." box="[471,902,1562,1589]" pageId="155" pageNumber="156" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 101</bibRefCitation>
</treatmentCitation>
(cited as synonym of
<taxonomicName id="52F9BCF330C152FFE982F9BC7233070B" ID-CoL="7XMPH" authorityName="Latreille" authorityYear="1809" box="[1173,1343,1563,1589]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="curvicornis">
<emphasis id="A78D1B6230C152FFE982F9BC7233070B" box="[1173,1343,1563,1589]" italics="true" pageId="155" pageNumber="156">O. curvicornis</emphasis>
</taxonomicName>
). —
<treatmentCitation id="1458E16130C152FFEDFBF999752D0768" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[236,545,1597,1624]" page="186" pageId="155" pageNumber="156" year="2019">
<bibRefCitation id="F168BA8130C152FFEDFBF999752D0768" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[236,545,1597,1624]" pageId="155" pageNumber="156" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230C152FFEC7EF99976AF0767" box="[361,419,1598,1624]" italics="true" pageId="155" pageNumber="156">et al.</emphasis>
2019: 186
</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E030C152FFEFC4F99974360767" box="[723,826,1598,1624]" name="Ecuador" pageId="155" pageNumber="156">Ecuador</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030C152FFEDAAF9C673B307FE" blockId="155.[189,1400,541,1765]" pageId="155" pageNumber="156">
<treatmentCitationGroup id="B5E9E05E30C152FFEDAAF9C673B307FE" pageId="155" pageNumber="156">
<taxonomicName id="52F9BCF330C152FFEDAAF9C676C40744" ID-CoL="7XNBX" authorityName="d'Orbigny" authorityYear="1902" box="[189,456,1633,1659]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="laevatus">
<emphasis id="A78D1B6230C152FFEDAAF9C676C40744" box="[189,456,1633,1659]" italics="true" pageId="155" pageNumber="156">Onthophagus laevatus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C152FFECFCF9C675E50744" author="d'Orbigny H." box="[491,745,1633,1660]" page="437" pageId="155" pageNumber="156" year="1905">
<bibRefCitation id="F168BA8130C152FFECFCF9C675E50744" author="d'Orbigny H." box="[491,745,1633,1660]" pageId="155" pageNumber="156" pagination="381 - 536" refId="ref134143" refString="d'Orbigny H. 1905. Onthophagides provenant du voyage de M. Alluaud dans l'Afrique orientale et descriptions d'autres especes africaines nouvelles. Annales de la Societe entomologique de France 74: 381 - 536. Available from https: // www. biodiversitylibrary. org / page / 32495308 [accessed 16 Feb. 2024]." type="journal article" year="1905">dOrbigny 1905: 437</bibRefCitation>
</treatmentCitation>
(comments);
<treatmentCitation id="1458E16130C152FFEE85F9C673460744" author="d'Orbigny H." box="[914,1098,1633,1659]" page="640" pageId="155" pageNumber="156" year="1913">
<bibRefCitation id="F168BA8130C152FFEE85F9C673460744" author="d'Orbigny H." box="[914,1098,1633,1659]" pageId="155" pageNumber="156" pagination="1 - 742" refId="ref134201" refString="d'Orbigny H. 1913. Synopsis des Onthophagides d'Afrique. Annales de la Societe entomologique de France 82: 1 - 742. Available from https: // www. biodiversitylibrary. org / page / 8239919 [accessed 16 Feb. 2024]." type="journal article" year="1913">1913: 174, 640</bibRefCitation>
</treatmentCitation>
(key, list). —
<treatmentCitation id="1458E16130C152FFE9E8F9C6766F07A0" author="Peringuey L." page="587" pageId="155" pageNumber="156" year="1908">
<bibRefCitation id="F168BA8130C152FFE9E8F9C6766F07A0" author="Peringuey L." pageId="155" pageNumber="156" pagination="547 - 752" refId="ref135677" refString="Peringuey L. 1908. Descriptive catalogue of the Coleoptera of South Africa. Additions and corrections. Transactions of the South African philosophical Society 13: 547 - 752." type="journal article" year="1908">Peringuey 1908: 587</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E16130C152FFEF43F92374C207A1" author="Gillet J. J. E. &amp; Boucomont A." box="[596,974,1668,1695]" page="177" pageId="155" pageNumber="156" year="1927">
<bibRefCitation id="F168BA8130C152FFEF43F92374C207A1" author="Gillet J. J. E. &amp; Boucomont A." box="[596,974,1668,1695]" pageId="155" pageNumber="156" refId="ref121284" refString="Gillet J. J. E. &amp; Boucomont A. 1927. Scarabaeidae: Coprinae, Termitotroginae (Partes 38 et 90). W. Junk, Berlin. [We are not sure about the publication date of this book. The title page gives the range &quot; 1911 - 1927 &quot;, which could be interpreted as indicating that the book was published in parts between those years. What we suspect, however, is that 1911 refers to the first part of the Coprinae catalogue, published by Gillet (1911 b) - i. e., our previous reference -, whereas the continuation of the Coprinae catalogue, now added by the Termitotroginae, was published at once in 1927 with the joint-authorship of Gillet and Boucomont.]" type="book" year="1927">Gillet &amp; Boucomont 1927: 177</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130C152FFE981F923727A07A1" author="Ferreira M. C." box="[1174,1398,1668,1694]" page="666" pageId="155" pageNumber="156" year="1972">
<bibRefCitation id="F168BA8130C152FFE981F923727A07A1" author="Ferreira M. C." box="[1174,1398,1668,1694]" pageId="155" pageNumber="156" pagination="5 - 1088" refId="ref119126" refString="Ferreira M. C. 1972. Os escarabideos de Africa (sul do Saara). I. Revista de Entomologia de Mocambique 11: 5 - 1088." type="journal article" year="1972">Ferreira 1972: 666</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E030C152FFEC49F90076F307FE" box="[350,511,1703,1729]" name="Mozambique" pageId="155" pageNumber="156">Mozambique</collectingCountry>
). —
<treatmentCitation id="1458E16130C152FFEF21F900746407FE" author="Daniel G. M. &amp; Genier F." box="[566,872,1703,1729]" page="28" pageId="155" pageNumber="156" year="2019">
<bibRefCitation id="F168BA8130C152FFEF21F900746407FE" author="Daniel G. M. &amp; Genier F." box="[566,872,1703,1729]" pageId="155" pageNumber="156" pagination="25 - 32" refId="ref116132" refString="Daniel G. M. &amp; &amp; Genier F. 2019. A preliminary checklist of the dung beetle species of Mozambique (Coleoptera: Scarabaeidae: Scarabaeinae). Catharsius 19: 25 - 32." type="journal article" year="2019">Daniel &amp; Génier 2019: 28</bibRefCitation>
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(checklist for
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).
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</paragraph>
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<taxonomicName id="52F9BCF330C152FFEDAAF96D75AD07DB" ID-CoL="7XMPH" authorityName="Latreille" authorityYear="1809" box="[189,673,1738,1765]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="155" pageNumber="156" phylum="Arthropoda" rank="species" species="curvicornis" subGenus="Onthophagus">
<emphasis id="A78D1B6230C152FFEDAAF96D765707DB" box="[189,347,1738,1764]" italics="true" pageId="155" pageNumber="156">Onthophagus</emphasis>
(
<emphasis id="A78D1B6230C152FFEC7BF96D750607DB" box="[364,522,1738,1764]" italics="true" pageId="155" pageNumber="156">Onthophagus</emphasis>
)
<emphasis id="A78D1B6230C152FFEF0CF96C75AD07DB" box="[539,673,1739,1764]" italics="true" pageId="155" pageNumber="156">curvicornis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130C152FFEFA9F96D74D107DA" author="Vaz-de-Mello F. Z." box="[702,989,1738,1765]" page="194" pageId="155" pageNumber="156" year="2000">
<bibRefCitation id="F168BA8130C152FFEFA9F96D74D107DA" author="Vaz-de-Mello F. Z." box="[702,989,1738,1765]" pageId="155" pageNumber="156" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030C152FFE990F96D73D807DB" box="[1159,1236,1738,1764]" name="Brazil" pageId="155" pageNumber="156">Brazil</collectingCountry>
).
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</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30C152F8EDAAF8AB76E20079" lastPageId="156" lastPageNumber="157" pageId="155" pageNumber="156" type="materials_examined">
<paragraph id="9546C77030C152FFEDAAF8AB76A60619" blockId="155.[189,426,1804,1830]" box="[189,426,1804,1830]" pageId="155" pageNumber="156">
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<emphasis id="A78D1B6230C152FFEDAAF8AB76A60619" bold="true" box="[189,426,1804,1830]" pageId="155" pageNumber="156">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030C152F8EDAAF89C76E20079" blockId="155.[189,1399,1849,1983]" lastBlockId="156.[189,1398,263,327]" lastPageId="156" lastPageNumber="157" pageId="155" pageNumber="156">
<materialsCitation id="2591CD2D30C152FFEDAAF89C74C3066A" ID-GBIF-Occurrence="4979307386" box="[189,975,1849,1877]" collectingDate="2006-08-02" collectionCode="CEMT" collectorName="El Manteco" country="Venezuela" location="Bolivar" pageId="155" pageNumber="156" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E030C152FFEDAAF89C7667066A" box="[189,363,1851,1877]" name="Venezuela" pageId="155" pageNumber="156">VENEZUELA</collectingCountry>
<emphasis id="A78D1B6230C152FFEC94F89E76D3066B" bold="true" box="[387,479,1849,1876]" pageId="155" pageNumber="156">
<collectingRegion id="573D099230C152FFEC94F89E76D3066B" box="[387,479,1849,1876]" country="Venezuela" name="Bolivar" pageId="155" pageNumber="156">Bolívar</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C152FFECE3F89C754A066A" box="[500,582,1851,1877]" count="1" pageId="155" pageNumber="156" type="generic">1 spec.</specimenCount>
;
<collectorName id="380CA2A630C152FFEF47F89D75D5066A" box="[592,729,1850,1877]" pageId="155" pageNumber="156">El Manteco</collectorName>
;
<date id="E147E1B030C152FFEFF4F89C747E066B" box="[739,882,1850,1877]" pageId="155" pageNumber="156" value="2006-08-02">
<collectingDate id="F103185830C152FFEFF4F89C747E066B" box="[739,882,1850,1877]" pageId="155" pageNumber="156" value="2006-08-02">2Aug. 2006</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C152FFEE6BF89C74C3066A" box="[892,975,1851,1877]" pageId="155" pageNumber="156">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30C152FFEEFBF89E749D0647" ID-GBIF-Occurrence="4979307371" collectingDate="1995-04-13" collectionCode="CEMT" collectorName="La Muchuy &amp; Parque Nacional Sierra Nevada" country="Venezuela" elevation="2400" location="La Muchuy" pageId="155" pageNumber="156" specimenCount="1" specimenCount-female="1" stateProvince="Merida">
<emphasis id="A78D1B6230C152FFEEFBF89E7345066B" bold="true" box="[1004,1097,1849,1876]" pageId="155" pageNumber="156">
<collectingRegion id="573D099230C152FFEEFBF89E7345066B" box="[1004,1097,1849,1876]" country="Venezuela" name="Merida" pageId="155" pageNumber="156">Mérida</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C152FFE94AF89C7384066A" box="[1117,1160,1851,1877]" count="1" pageId="155" pageNumber="156" type="female">1 ♀</specimenCount>
;
<location id="902691AB30C152FFE982F89C7210066B" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C152FFE982F89C7210066B" box="[1173,1308,1850,1877]" country="Venezuela" name="La Muchuy" pageId="155" pageNumber="156" stateProvince="Merida">La Muchuy</location>
,
<location id="902691AB30C152FFE832F89C76D00647" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C152FFE832F89C76D00647" country="Venezuela" name="Parque Nacional Sierra Nevada" pageId="155" pageNumber="156" stateProvince="Merida">Parque Nacional Sierra Nevada</location>
;
<elevation id="1ED4204330C152FFECFEF8F975890647" box="[489,645,1886,1912]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="155" pageNumber="156" unit="m" value="2400.0">
<quantity id="52016A9530C152FFECFEF8F975480647" box="[489,580,1886,1912]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="155" pageNumber="156" unit="m" value="2400.0">2400 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030C152FFEF83F8F9743F0647" box="[660,819,1885,1912]" pageId="155" pageNumber="156" value="1995-04-13">
<collectingDate id="F103185830C152FFEF83F8F9743F0647" box="[660,819,1885,1912]" pageId="155" pageNumber="156" value="1995-04-13">13 Apr. 1995</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C152FFEE57F8F9749D0647" box="[832,913,1886,1912]" pageId="155" pageNumber="156">CEMT</collectionCode>
</materialsCitation>
<materialsCitation id="2591CD2D30C152FFEEBAF8F9742606A4" ID-GBIF-Occurrence="4979307449" collectingDate="2009-04-08" collectionCode="CEMT" elevation="2235" latitude="8.660847" location="Jaji" longLatPrecision="1" longitude="-71.39884" municipality="Sucre" pageId="155" pageNumber="156" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="83FF0CF930C152FFEEBAF8F974D10647" box="[941,989,1886,1912]" count="1" pageId="155" pageNumber="156" type="male">1 ♂</specimenCount>
,
<specimenCount id="83FF0CF930C152FFEEFCF8F973150647" box="[1003,1049,1886,1912]" count="1" pageId="155" pageNumber="156" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="75225D0A30C152FFE93EF8F973630647" box="[1065,1135,1886,1912]" pageId="155" pageNumber="156">Sucre</collectingMunicipality>
,
<location id="902691AB30C152FFE96BF8F973A50647" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C152FFE96BF8F973A50647" box="[1148,1193,1886,1912]" latitude="8.660847" longLatPrecision="1" longitude="-71.39884" municipality="Sucre" name="Jaji" pageId="155" pageNumber="156">Jají</location>
;
<geoCoordinate id="F0CDA1B730C152FFE9A1F8FB727E0647" box="[1206,1394,1884,1912]" degrees="08" direction="north" minutes="39" orientation="latitude" pageId="155" pageNumber="156" precision="1" seconds="39.05" value="8.660847">08°3939.05″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730C152FFEDAAF8D8767306A4" box="[189,383,1919,1947]" degrees="71" direction="west" minutes="23" orientation="longitude" pageId="155" pageNumber="156" precision="1" seconds="55.85" value="-71.39884">71°2355.85″ W</geoCoordinate>
;
<quantity id="52016A9530C152FFEC9BF82676EB06A4" box="[396,487,1921,1948]" metricMagnitude="3" metricUnit="m" metricValue="2.235" pageId="155" pageNumber="156" unit="m" value="2235.0">
<elevation id="1ED4204330C152FFEC9BF82676EB06A4" box="[396,487,1921,1948]" metricMagnitude="3" metricUnit="m" metricValue="2.235" pageId="155" pageNumber="156" unit="m" value="2235.0">2235 m</elevation>
</quantity>
a.s.l.;
<date id="E147E1B030C152FFEF2FF82675C506A4" box="[568,713,1921,1947]" pageId="155" pageNumber="156" value="2009-04-08">
<collectingDate id="F103185830C152FFEF2FF82675C506A4" box="[568,713,1921,1947]" pageId="155" pageNumber="156" value="2009-04-08">8 Apr. 2009</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C152FFEFC0F826742606A4" box="[727,810,1921,1947]" pageId="155" pageNumber="156">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30C152FFEE5AF82776F00681" ID-GBIF-Occurrence="4979307393" collectingDate="1989-02-13" collectionCode="CEMT" collectorName="Pregonero" country="Venezuela" elevation="1100" location="La Honda" pageId="155" pageNumber="156" specimenCount="1" specimenCount-female="1" stateProvince="Tachira">
<emphasis id="A78D1B6230C152FFEE5AF82774BC06A4" bold="true" box="[845,944,1920,1947]" pageId="155" pageNumber="156">
<collectingRegion id="573D099230C152FFEE5AF82774BC06A4" box="[845,944,1920,1947]" country="Venezuela" name="Tachira" pageId="155" pageNumber="156">Táchira</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C152FFEEDAF82674F006A4" box="[973,1020,1921,1947]" count="1" pageId="155" pageNumber="156" type="female">1 ♀</specimenCount>
;
<collectorName id="380CA2A630C152FFE91BF826738406A4" box="[1036,1160,1921,1947]" pageId="155" pageNumber="156">Pregonero</collectorName>
,
<location id="902691AB30C152FFE982F826720306A4" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C152FFE982F826720306A4" box="[1173,1295,1921,1947]" country="Venezuela" name="La Honda" pageId="155" pageNumber="156" stateProvince="Tachira">La Honda</location>
;
<elevation id="1ED4204330C152FFE80AF82677FA0681" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="155" pageNumber="156" unit="m" value="1100.0">
<quantity id="52016A9530C152FFE80AF826727B06A4" box="[1309,1399,1921,1947]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="155" pageNumber="156" unit="m" value="1100.0">1100 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030C152FFEC15F80376920681" box="[258,414,1956,1982]" pageId="155" pageNumber="156" value="1989-02-13">
<collectingDate id="F103185830C152FFEC15F80376920681" box="[258,414,1956,1982]" pageId="155" pageNumber="156" value="1989-02-13">13 Feb. 1989</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C152FFECBEF80376F00681" box="[425,508,1956,1982]" pageId="155" pageNumber="156">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30C152FFEF0CF803734C0681" ID-GBIF-Occurrence="4979307524" box="[539,1088,1956,1983]" collectionCode="CEMT" collectorName="Trujillo" country="Venezuela" location="Trujillo" pageId="155" pageNumber="156" specimenCount="1" specimenCount-female="1" stateProvince="Trujillo">
<emphasis id="A78D1B6230C152FFEF0CF80375770681" bold="true" box="[539,635,1956,1982]" pageId="155" pageNumber="156">
<collectingRegion id="573D099230C152FFEF0CF80375770681" box="[539,635,1956,1982]" country="Venezuela" name="Trujillo" pageId="155" pageNumber="156">Trujillo</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C152FFEF85F80375B30680" box="[658,703,1956,1983]" count="1" pageId="155" pageNumber="156" type="female">1 ♀</specimenCount>
;
<collectorName id="380CA2A630C152FFEFDBF80374250681" box="[716,809,1956,1982]" pageId="155" pageNumber="156">Trujillo</collectorName>
; no further data;
<collectionCode id="F3E85FB530C152FFEEFAF803734C0681" box="[1005,1088,1956,1982]" pageId="155" pageNumber="156">CEMT</collectionCode>
</materialsCitation>
.
<materialsCitation id="2591CD2D30C152F8E948F803734A001C" ID-GBIF-Occurrence="4979307451" collectingDate="2009-07-19" collectionCode="CEMT" country="Venezuela" elevation="1415" lastPageId="156" lastPageNumber="157" latitude="10.385028" location="Bolivar" longLatPrecision="1" longitude="-68.979294" pageId="155" pageNumber="156" specimenCount="1" specimenCount-male="1" stateProvince="Yaracuy">
<emphasis id="A78D1B6230C152FFE948F80373CB0681" bold="true" box="[1119,1223,1956,1982]" pageId="155" pageNumber="156">
<collectingRegion id="573D099230C152FFE948F80373CB0681" box="[1119,1223,1956,1982]" country="Venezuela" name="Yaracuy" pageId="155" pageNumber="156">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C152FFE9C9F80372010680" box="[1246,1293,1956,1983]" count="1" pageId="155" pageNumber="156" type="male">1 ♂</specimenCount>
;
<location id="902691AB30C152FFE80FF803727E0681" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C152FFE80FF803727E0681" box="[1304,1394,1956,1982]" country="Venezuela" latitude="10.385028" longLatPrecision="1" longitude="-68.979294" name="Bolivar" pageId="155" pageNumber="156" stateProvince="Yaracuy">Bolívar</location>
,
<location id="902691AB30C652F8EDAAFEAE77F7001C" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C652F8EDAAFEAE77F7001C" box="[189,251,265,291]" country="Venezuela" latitude="10.385028" longLatPrecision="1" longitude="-68.979294" name="Aroa" pageId="156" pageNumber="157" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B730C652F8EC1DFEA076B4001C" box="[266,440,263,291]" degrees="10" direction="north" minutes="23" orientation="latitude" pageId="156" pageNumber="157" precision="1" seconds="06.1" value="10.385028">10°2306.1″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B730C652F8ECD0FEA07586001C" box="[455,650,263,291]" degrees="68" direction="west" minutes="58" orientation="longitude" pageId="156" pageNumber="157" precision="1" seconds="45.45" value="-68.979294">68°5845.45″ W</geoCoordinate>
;
<elevation id="1ED4204330C652F8EF8FFEAE743B001C" box="[664,823,265,292]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="156" pageNumber="157" unit="m" value="1415.0">
<quantity id="52016A9530C652F8EF8FFEAE75F8001C" box="[664,756,265,292]" metricMagnitude="3" metricUnit="m" metricValue="1.415" pageId="156" pageNumber="157" unit="m" value="1415.0">1415 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B030C652F8EE50FEAE74EC001C" box="[839,992,265,291]" pageId="156" pageNumber="157" value="2009-07-19">
<collectingDate id="F103185830C652F8EE50FEAE74EC001C" box="[839,992,265,291]" pageId="156" pageNumber="157" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C652F8EEF8FEAE734E001C" box="[1007,1090,265,291]" pageId="156" pageNumber="157">CEMT</collectionCode>
.
</materialsCitation>
<materialsCitation id="2591CD2D30C652F8E947FEAE76E60079" ID-GBIF-Occurrence="4979307327" collectingDate="2005-11-28" collectingDateMax="2005-11-30" collectingDateMin="2005-11-28" collectionCode="CEMT" country="Venezuela" elevation="1415" latitude="10.385028" location="Miranda" longLatPrecision="1" longitude="-68.979294" pageId="156" pageNumber="157" specimenCount="1" specimenCount-male="1" stateProvince="Zulia">
<emphasis id="A78D1B6230C652F8E97EFEAE73A0001C" bold="true" box="[1129,1196,265,291]" pageId="156" pageNumber="157">
<collectingRegion id="573D099230C652F8E97EFEAE73A0001C" box="[1129,1196,265,291]" country="Venezuela" name="Zulia" pageId="156" pageNumber="157">Zulia</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930C652F8E9DDFEAE73F0001C" box="[1226,1276,265,291]" count="1" pageId="156" pageNumber="157" type="male">1 ♂</specimenCount>
;
<location id="902691AB30C652F8E81CFEAE727E001C" LSID="urn:lsid:plazi:treatment:1D50766630C352F8EF32F94472410256:902691AB30C652F8E81CFEAE727E001C" box="[1291,1394,265,291]" country="Venezuela" latitude="10.385028" longLatPrecision="1" longitude="-68.979294" name="Miranda" pageId="156" pageNumber="157" stateProvince="Zulia">Miranda</location>
;
<date id="E147E1B030C652F8EDAAFE8B76870078" box="[189,395,300,327]" pageId="156" pageNumber="157" value="2005-11-28" valueMax="2005-11-30" valueMin="2005-11-28">
<collectingDate id="F103185830C652F8EDAAFE8B76870078" box="[189,395,300,327]" pageId="156" pageNumber="157" value="2005-11-28" valueMax="2005-11-30" valueMin="2005-11-28">2830 Nov. 2005</collectingDate>
</date>
;
<collectionCode id="F3E85FB530C652F8EC80FE8B76E60079" box="[407,490,300,326]" pageId="156" pageNumber="157">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30C652F8EDAAFECA74610375" pageId="156" pageNumber="157" type="distribution">
<paragraph id="9546C77030C652F8EDAAFECA765A00B8" blockId="156.[189,342,365,391]" box="[189,342,365,391]" pageId="156" pageNumber="157">
<heading id="CE0E701C30C652F8EDAAFECA765A00B8" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="156" pageNumber="157" reason="3">
<emphasis id="A78D1B6230C652F8EDAAFECA765A00B8" bold="true" box="[189,342,365,391]" pageId="156" pageNumber="157">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030C652F8EDAAFE3B73570089" blockId="156.[189,1115,412,439]" box="[189,1115,412,439]" pageId="156" pageNumber="157">
<collectingCountry id="EDEE87E030C652F8EDAAFE3B76380089" box="[189,308,412,438]" name="Colombia" pageId="156" pageNumber="157">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030C652F8EC28FE3B76B70089" box="[319,443,412,438]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E030C652F8ECD2FE3B75B10089" box="[453,701,412,438]" name="Trinidad and Tobago" pageId="156" pageNumber="157">Trinidad and Tobago</collectingCountry>
,
<collectingCountry id="EDEE87E030C652F8EFDFFE3B74200089" box="[712,812,412,438]" name="Ecuador" pageId="156" pageNumber="157">Ecuador</collectingCountry>
, and
<collectingCountry id="EDEE87E030C652F8EE7EFE3A74930088" box="[873,927,413,439]" name="Peru" pageId="156" pageNumber="157">Peru</collectingCountry>
(
<bibRefCitation id="F168BA8130C652F8EEB9FE3B73420089" author="Rossini M." box="[942,1102,412,438]" pageId="156" pageNumber="157" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini 2021</bibRefCitation>
).
</paragraph>
<paragraph id="9546C77030C652F8EDAAFE7A76E100C7" blockId="156.[189,493,477,504]" box="[189,493,477,504]" pageId="156" pageNumber="157">
<heading id="CE0E701C30C652F8EDAAFE7A76E100C7" bold="true" box="[189,493,477,504]" fontSize="11" level="3" pageId="156" pageNumber="157" reason="3">
<emphasis id="A78D1B6230C652F8EDAAFE7A76E100C7" bold="true" box="[189,493,477,504]" pageId="156" pageNumber="157">
Subregions of
<collectingCountry id="EDEE87E030C652F8EC78FE7976E100C7" box="[367,493,478,504]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030C652F8EDAAFDAB74610375" blockId="156.[188,1398,524,586]" pageId="156" pageNumber="157">
System of hills and low sierras Lara-Falcón, Andes mountains, Sierra de
<collectingRegion id="573D099230C652F8E918FDAB737A0319" box="[1039,1142,524,550]" country="Argentina" name="San Luis" pageId="156" pageNumber="157">San Luis</collectingRegion>
and Cerro
<collectingRegion id="573D099230C652F8E9EFFDAB727E0319" box="[1272,1394,524,550]" country="El Salvador" name="Santa Ana" pageId="156" pageNumber="157">Santa Ana</collectingRegion>
, Central
<collectingRegion id="573D099230C652F8EC0DFD8876510376" box="[282,349,559,585]" country="New Zealand" name="West Coast" pageId="156" pageNumber="157">Coast</collectingRegion>
Mountain Range, and
<collectingRegion id="573D099230C652F8EF7EFD8875C50376" box="[617,713,559,585]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="156" pageNumber="157">Oriental</collectingRegion>
<collectingRegion id="573D099230C652F8EFC7FD88741F0376" box="[720,787,559,585]" country="New Zealand" name="West Coast" pageId="156" pageNumber="157">Coast</collectingRegion>
Range.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30C652F8EDAAFDD772410256" pageId="156" pageNumber="157" type="reference_group">
<paragraph id="9546C77030C652F8EDAAFDD776A803B4" blockId="156.[189,420,624,651]" box="[189,420,624,651]" pageId="156" pageNumber="157">
<heading id="CE0E701C30C652F8EDAAFDD776A803B4" bold="true" box="[189,420,624,651]" fontSize="11" level="3" pageId="156" pageNumber="157" reason="3">
<emphasis id="A78D1B6230C652F8EDAAFDD776A803B4" bold="true" box="[189,420,624,651]" pageId="156" pageNumber="157">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030C652F8EDAAFD3872410256" blockId="156.[187,1399,671,874]" pageId="156" pageNumber="157">
<bibRefCitation id="F168BA8130C652F8EDAAFD3876890386" author="Harold E." box="[189,389,671,698]" pageId="156" pageNumber="157" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold 1880: 29</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8EC8FFD3875140386" box="[408,536,671,697]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). — Bates 18861890: 66 (
<collectingCountry id="EDEE87E030C652F8EE7CFD3874E70386" box="[875,1003,671,697]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130C652F8E93DFD0772370386" author="Boucomont A." box="[1066,1339,671,698]" pageId="156" pageNumber="157" pagination="293 - 332" refId="ref112419" refString="Boucomont A. 1932. Synopsis des Onthophagus d'Amerique du Sud (Col. Scarab.). Annales de la Societe entomologique de France 101: 293 - 332. https: // doi. org / 10.1080 / 21686351.1932.12280217" type="journal article" year="1932">Boucomont 1932: 308</bibRefCitation>
, 323 (
<collectingCountry id="EDEE87E030C652F8EDD2FD65764903E3" box="[197,325,706,732]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130C652F8EC6CFD65758603E2" author="Blackwelder R. E." box="[379,650,706,733]" pageId="156" pageNumber="157" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 211</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8EF8EFD65741503E3" box="[665,793,706,732]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130C652F8EE58FD6474F403E2" author="Roze J. A." box="[847,1016,706,733]" pageId="156" pageNumber="157" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 45</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8E910FD65738A03E3" box="[1031,1158,706,732]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230C652F8E987FD65724A03E3" box="[1168,1350,706,732]" country="Venezuela" name="Distrito Capital" pageId="156" pageNumber="157">Distrito Capital</collectingRegion>
and
<collectingRegion id="573D099230C652F8EDABFD41761A023F" box="[188,278,742,768]" country="Venezuela" name="Merida" pageId="156" pageNumber="157">Mérida</collectingRegion>
). —
<bibRefCitation id="F168BA8130C652F8EC58FD417532023F" author="Blanco J." box="[335,574,742,768]" pageId="156" pageNumber="157" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 4647</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8EF5AFD4175C0023F" box="[589,716,742,768]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230C652F8EFC1FD417439023F" box="[726,821,742,768]" country="Venezuela" name="Tachira" pageId="156" pageNumber="157">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA8130C652F8EE7AFD417342023F" author="Havranek D." box="[877,1102,742,768]" pageId="156" pageNumber="157" pagination="55 - 64" refId="ref123886" refString="Havranek D. 1989. Coleoptera coprofagos y necrofagos (Scarabaeidae, Silphidae) atraidos a cebos en bosques venezolanos. UNET 3: 55 - 64." type="journal article" year="1989">Havranek 1989: 61</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8E949FD4173D1023F" box="[1118,1245,742,768]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230C652F8E9F0FD41724A023F" box="[1255,1350,742,768]" country="Venezuela" name="Tachira" pageId="156" pageNumber="157">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA8130C652F8EDAAFCAE7564021C" author="Pulido-Herrera L. A. &amp; Zunino M." box="[189,616,777,803]" pageId="156" pageNumber="157" refId="ref136342" refString="Pulido-Herrera L. A. &amp; Zunino M. 2007. Catalogo preliminar de los Onthophagini de America (Coleoptera: Scarabaeine [sic]). In: Zunino M. &amp; Melic A. (eds) Escarabajos, Diversidad y Conservacion Biologica. Ensayos en Homenaje a Gonzalo Halffter: 93 - 129. Monografias 3 ercer Milenio m 3 m Vol. 7, Sociedad Entomologica Aragonesa, Zaragoza." type="book" year="2007">Pulido-Herrera &amp; Zunino 2007: 101</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8EF6FFCAE75F4021C" box="[632,760,777,803]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130C652F8EE26FCAE7373021C" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[817,1151,777,803]" pageId="156" pageNumber="157" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B6230C652F8EED2FCAD730C021C" box="[965,1024,777,803]" italics="true" pageId="156" pageNumber="157">et al.</emphasis>
2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8E998FCAE7202021C" box="[1167,1294,777,803]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230C652F8E80DFCAE727F021C" box="[1306,1395,777,803]" country="Venezuela" name="Merida" pageId="156" pageNumber="157">Mérida</collectingRegion>
: La Azulita-Jají and
<collectingRegion id="573D099230C652F8ECB7FC8B75040279" box="[416,520,812,838]" country="Venezuela" name="Miranda" pageId="156" pageNumber="157">Miranda</collectingRegion>
:Altos de Pipe). —
<bibRefCitation id="F168BA8130C652F8EFF3FC8B73180279" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." box="[740,1044,812,838]" pageId="156" pageNumber="157" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">
Chamorro
<emphasis id="A78D1B6230C652F8EE48FC8A74940279" box="[863,920,812,838]" italics="true" pageId="156" pageNumber="157">et al.</emphasis>
2019: 186
</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8E936FC8B73AD0279" box="[1057,1185,812,838]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA8130C652F8E9C1FC8B77EB0256" author="Rossini M." pageId="156" pageNumber="157" pagination="262 - 274" refId="ref137534" refString="Rossini M. 2021. Additional mislabeling in African Onthophagus Latreille, 1802 (Coleoptera: Scarabaeidae: Scarabaeinae): the case of Onthophagus viviensis and Onthophagus laevatus. Zootaxa 5032 (2): 262 - 274. https: // doi. org / 10.11646 / zootaxa. 5032.2.7" type="journal article" year="2021">Rossini 2021: 266</bibRefCitation>
(
<collectingCountry id="EDEE87E030C652F8EDE0FCE8767A0256" box="[247,374,847,873]" name="Venezuela" pageId="156" pageNumber="157">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230C652F8EC68FCE876D40256" box="[383,472,847,873]" country="Venezuela" name="Aragua" pageId="156" pageNumber="157">Aragua</collectingRegion>
,
<collectingRegion id="573D099230C652F8ECF3FCE8753B0256" box="[484,567,847,873]" country="Venezuela" name="Falcon" pageId="156" pageNumber="157">Falcón</collectingRegion>
,, La Guaira,
<collectingRegion id="573D099230C652F8EFC1FCE874220256" box="[726,814,847,873]" country="Venezuela" name="Merida" pageId="156" pageNumber="157">Mérida</collectingRegion>
,
<collectingRegion id="573D099230C652F8EE2EFCF774AB0255" box="[825,935,848,874]" country="Venezuela" name="Monagas" pageId="156" pageNumber="157">Monagas</collectingRegion>
,
<collectingRegion id="573D099230C652F8EEA4FCE874F50256" box="[947,1017,847,873]" country="Venezuela" name="Sucre" pageId="156" pageNumber="157">Sucre</collectingRegion>
,
<collectingRegion id="573D099230C652F8E913FCE8736D0256" box="[1028,1121,847,873]" country="Venezuela" name="Tachira" pageId="156" pageNumber="157">Táchira</collectingRegion>
,
<collectingRegion id="573D099230C652F8E97BFCE873C90256" box="[1132,1221,847,873]" country="Venezuela" name="Trujillo" pageId="156" pageNumber="157">Trujillo</collectingRegion>
and
<collectingRegion id="573D099230C652F8E9E9FCE8724D0256" box="[1278,1345,847,873]" country="Venezuela" name="Zulia" pageId="156" pageNumber="157">Zulia</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>