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<mods:title id="3B41599D84C351F8642AEEB0EC90D691">The reclassification of Sophiodela and other tiger beetles (Coleoptera, Cicindelidae) based on the structure of the everted internal sac of the male genitalia</mods:title>
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<treatment id="1D1C8783DF718E0D698637C689E9F9FD" ID-DOI="http://doi.org/10.5281/zenodo.4337364" ID-GBIF-Taxon="167293602" ID-Zenodo-Dep="4337364" LSID="urn:lsid:plazi:treatment:1D1C8783DF718E0D698637C689E9F9FD" httpUri="http://treatment.plazi.org/id/1D1C8783DF718E0D698637C689E9F9FD" lastPageNumber="292" pageId="21" pageNumber="292">
<subSubSection id="DDAF651EDF718E0D698637C68B06FC6F" box="[151,606,900,927]" pageId="21" pageNumber="292" type="nomenclature">
<paragraph id="950A3695DF718E0D698637C68B06FC6F" blockId="21.[151,606,900,963]" box="[151,606,900,927]" pageId="21" pageNumber="292">
<heading id="CE4281F9DF718E0D698637C68B06FC6F" box="[151,606,900,927]" level="2" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D698637C68B06FC6F" ID-CoL="PXDY" authority="(Schaum, 1863)" baseAuthorityName="Schaum" baseAuthorityYear="1863" box="[151,606,900,927]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="species" species="lacrymans">
<emphasis id="A7C1EA87DF718E0D698637C688CDFC6E" box="[151,405,901,927]" italics="true" pageId="21" pageNumber="292">Calochroa lacrymans</emphasis>
<emphasis id="A7C1EA87DF718E0D688D37C78B06FC6F" bold="true" box="[412,606,900,926]" pageId="21" pageNumber="292">
(
<bibRefCitation id="F1244B64DF718E0D68B537C78B0EFC6F" author="Schaum, H. R." box="[420,598,900,926]" pageId="21" pageNumber="292" pagination="57 - 73" refId="ref20340" refString="Schaum, H. R., (1863) Contributions to the knowledge of the Cicindelidae of tropical Asia, containing descriptions of new species, a list of those hitherto described, and synonymical notes. The Journal of Entomology. Descriptive and Geographical, 2, 57 - 73." type="journal article" year="1863">Schaum, 1863</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DDAF651EDF718E0D698637EB8819FC32" box="[151,321,936,963]" pageId="21" pageNumber="292" type="description">
<paragraph id="950A3695DF718E0D698637EB8819FC32" blockId="21.[151,606,900,963]" box="[151,321,936,963]" pageId="21" pageNumber="292">
(
<figureCitation id="0D8E2A10DF718E0D698E37EB89A4FC33" box="[159,252,936,962]" captionStart="FIGURE 6" captionStartId="10.[151,250,1872,1897]" captionTargetBox="[152,1418,183,1827]" captionTargetId="figure@10.[151,1435,175,1854]" captionTargetPageId="10" captionText="FIGURE 6. Habitus of Calochroa. A, Calochroa bicolor bicolor (Fabricius); B, Calochroa flavomaculata flavomaculata (Hope); C, Calochroa harmandi (Fleutiaux); D, Calochroa whithillii Hope; EF, Calochroa pseudosiamensis (Horn); G, Calochroa assamensis (Parry); H, Calochroa lacrymans (Schaum)." figureDoi="http://doi.org/10.5281/zenodo.3998540" httpUri="https://zenodo.org/record/3998540/files/figure.png" pageId="21" pageNumber="292">Figs 6H</figureCitation>
,
<figureCitation id="0D8E2A10DF718E0D681937EA8862FC32" box="[264,314,937,963]" captionStart="FIGURE 10" captionStartId="24.[151,250,1945,1970]" captionTargetBox="[151,1435,181,1919]" captionTargetId="figure@24.[151,1435,181,1919]" captionTargetPageId="24" captionText="FIGURE 10. Everted internal sacs of Calochroa and Cylindera. A, Calochroa harmandi (Fleutiaux); B, C. whithillii Hope; C, C. schillhammeri Wiesner; D, C. lacrymans (Schaum); E, Cylindera (Eugrapha) elisae elisae (Motschulsky). Scales: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3998548" httpUri="https://zenodo.org/record/3998548/files/figure.png" pageId="21" pageNumber="292">10D</figureCitation>
)
</paragraph>
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<subSubSection id="DDAF651EDF718E0D698637AE8D27FB39" pageId="21" pageNumber="292" type="reference_group">
<paragraph id="950A3695DF718E0D698637AE8B70FBF4" blockId="21.[151,830,1004,1152]" box="[151,552,1004,1029]" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D698637AE8B7DFBF4" ID-CoL="V86T" authority="Schaum, 1863: 57" authorityName="Schaum" authorityPageNumber="57" authorityYear="1863" box="[151,549,1004,1029]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="species" species="lacrymans">
<emphasis id="A7C1EA87DF718E0D698637AE8832FBF4" box="[151,362,1005,1029]" italics="true" pageId="21" pageNumber="292">Cicindela lacrymans</emphasis>
<bibRefCitation id="F1244B64DF718E0D686137AF8B7DFBF4" author="Schaum, H. R." box="[368,549,1004,1029]" pageId="21" pageNumber="292" pagination="57 - 73" refId="ref20340" refString="Schaum, H. R., (1863) Contributions to the knowledge of the Cicindelidae of tropical Asia, containing descriptions of new species, a list of those hitherto described, and synonymical notes. The Journal of Entomology. Descriptive and Geographical, 2, 57 - 73." type="journal article" year="1863">Schaum, 1863:57</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="950A3695DF718E0D698630488BF0FBD5" blockId="21.[151,830,1004,1152]" box="[151,680,1035,1060]" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D698630488BFCFBD5" ID-CoL="PXDY" authority=": Fowler 1912: 390" authorityName="Fowler" authorityPageNumber="390" authorityYear="1912" box="[151,676,1035,1060]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="subSpecies" species="discrepans" subSpecies="lacrymans">
<emphasis id="A7C1EA87DF718E0D6986304888BEFBD2" box="[151,486,1035,1059]" italics="true" pageId="21" pageNumber="292">Calochroa discrepans lacrymans</emphasis>
:
<bibRefCitation id="F1244B64DF718E0D68E230488BFCFBD5" author="Fowler, W. W." box="[499,676,1035,1060]" pageId="21" pageNumber="292" refId="ref18243" refString="Fowler, W. W. (1912) The fauna of British India including Ceylon and Burma. Coleoptera. General introduction and Cicindelidae and Paussidae, Taylor and Francis, London, 529 pp. https: // doi. org / 10.5962 / bhl. title. 83566" type="book" year="1912">Fowler 1912:390</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="950A3695DF718E0D698630698B6EFBB3" blockId="21.[151,830,1004,1152]" box="[151,566,1065,1090]" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D698630698B6AFBB3" ID-CoL="PXDY" authority=": Naviaux 1986: 61" authorityName="Naviaux" authorityPageNumber="61" authorityYear="1986" box="[151,562,1065,1090]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="species" species="lacrymans">
<emphasis id="A7C1EA87DF718E0D69863069882BFBB3" box="[151,371,1066,1090]" italics="true" pageId="21" pageNumber="292">Calochroa lacrymans</emphasis>
:
<bibRefCitation id="F1244B64DF718E0D6891306A8B6AFBB3" author="Naviaux, R." box="[384,562,1065,1090]" pageId="21" pageNumber="292" pagination="58 - 69" refId="ref19780" refString="Naviaux, R. (1986) Coleoptera, Cicindelidae. Premier complement aux Cicindeles du Sri Lanka. Revue Scientifique Du Bourbonnais, 58 - 69." type="book chapter" year="1986">Naviaux 1986:61</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="950A3695DF718E0D6986300A8A66FB90" blockId="21.[151,830,1004,1152]" box="[151,830,1096,1121]" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D6986300A8A62FB90" ID-CoL="V86T" authority=": Acciavatti &amp; Pearson 1989: 129" authorityName="Acciavatti &amp; Pearson" authorityPageNumber="129" authorityYear="1989" box="[151,826,1096,1121]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="species" species="lacrymans" subGenus="Calochroa">
<emphasis id="A7C1EA87DF718E0D6986300A89A2FB90" box="[151,250,1097,1121]" italics="true" pageId="21" pageNumber="292">Cicindela</emphasis>
(
<emphasis id="A7C1EA87DF718E0D6819300A882BFB90" box="[264,371,1097,1121]" italics="true" pageId="21" pageNumber="292">Calochroa</emphasis>
)
<emphasis id="A7C1EA87DF718E0D6892300A88B5FB90" box="[387,493,1097,1121]" italics="true" pageId="21" pageNumber="292">lacrymans</emphasis>
:
<bibRefCitation id="F1244B64DF718E0D68E9300B8A62FB90" author="Acciavatti, R. E. &amp; Pearson, D. L." box="[504,826,1096,1121]" pageId="21" pageNumber="292" pagination="77 - 353" refId="ref17273" refString="Acciavatti, R. E. &amp; Pearson, D. L. (1989) The tiger beetle Genus Cicindela (Coleoptera, Insecta) from the indian subcontinent. Annals of Carnegie Museum, 58, 77 - 353." type="journal article" year="1989">Acciavatti &amp; Pearson 1989:129</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="950A3695DF718E0D698630248B18FB71" blockId="21.[151,830,1004,1152]" box="[151,576,1127,1152]" pageId="21" pageNumber="292">
<taxonomicName id="52B54D16DF718E0D698630248B64FB71" ID-CoL="PXDY" authority=": Wiesner 1992: 108" authorityName="Wiesner" authorityPageNumber="108" authorityYear="1992" box="[151,572,1127,1152]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="species" species="lacrymans">
<emphasis id="A7C1EA87DF718E0D69863024882BFB8E" box="[151,371,1127,1151]" italics="true" pageId="21" pageNumber="292">Calochroa lacrymans</emphasis>
:
<bibRefCitation id="F1244B64DF718E0D686E30248B64FB71" author="Wiesner, J." box="[383,572,1127,1152]" pageId="21" pageNumber="292" refId="ref20965" refString="Wiesner, J. (1992) Checklist of the Tiger Beetles of the World. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">Wiesner 1992:108</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="950A3695DF718E0D698630EE8D27FB39" blockId="21.[151,1436,1197,1548]" box="[151,1151,1197,1224]" pageId="21" pageNumber="292">
<emphasis id="A7C1EA87DF718E0D698630EE88CEFB39" bold="true" box="[151,406,1197,1224]" pageId="21" pageNumber="292">Specimens examined.</emphasis>
(N = 1)
<materialsCitation id="25DD3CC8DF718E0D68E630EE8D24FB39" ID-GBIF-Occurrence="2848419316" box="[503,1148,1197,1224]" collectingDate="1990-08" country="Sri Lanka" location="Sri Lanka" pageId="21" pageNumber="292" specimenCount="1" specimenCount-male="1">
<emphasis id="A7C1EA87DF718E0D68E630EE8BDAFB39" bold="true" box="[503,642,1197,1224]" pageId="21" pageNumber="292">
[
<collectingCountry id="EDA27605DF718E0D68EF30EE8B23FB39" box="[510,635,1197,1224]" name="Sri Lanka" pageId="21" pageNumber="292">Sri Lanka</collectingCountry>
]
</emphasis>
<specimenCount id="83B3FD1CDF718E0D6B9830ED8BF4FB39" box="[649,684,1198,1224]" pageId="21" pageNumber="292" type="male">1♂</specimenCount>
(
<collectionCode id="F3A4AE50DF718E0D6BAA30ED8BA0FB39" box="[699,760,1198,1224]" pageId="21" pageNumber="292">MHJ</collectionCode>
), Kandy,
<date id="E10B1055DF718E0D6A7330ED8A83FB39" box="[866,987,1198,1224]" pageId="21" pageNumber="292" value="1990-08">
<collectingDate id="F14FE9BDDF718E0D6A7330ED8A83FB39" box="[866,987,1198,1224]" pageId="21" pageNumber="292" value="1990-08">VIII. 1990</collectingDate>
</date>
, unknown leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDAF651EDF718E0D69D6309089E9F9FD" pageId="21" pageNumber="292" type="description">
<paragraph id="950A3695DF718E0D69D630908A47FAE1" blockId="21.[151,1436,1197,1548]" pageId="21" pageNumber="292">
<emphasis id="A7C1EA87DF718E0D69D630908813FB1D" box="[199,331,1235,1260]" italics="true" pageId="21" pageNumber="292">Description</emphasis>
. Internal sac everted dorso-apically in lateral view. DA largest in bladders; VA the second largest; VLR, VB, DLL slightly swollen; BLR and B not swollen.
</paragraph>
<paragraph id="950A3695DF718E0D69A23159888AFA35" blockId="21.[151,1436,1197,1548]" pageId="21" pageNumber="292">VA shortly and roundly everted right dorso-apically, vested with short and weakly chitinous spines. VLR roundly everted dorsally. DA semicircular swollen right latero-basally. DLL roundly swollen, vested numerously with spines, which are weakly chitinous. VB slightly and roundly swollen left latero-ventrally, vested numerously with long spines, which are forming a spot. Flagellum very long and thin, S-shaped. Medial tooth extending apically, weakly curved at apical 1/3.</paragraph>
<paragraph id="950A3695DF718E0D69D6318D89E9F9FD" blockId="21.[151,1436,1197,1548]" pageId="21" pageNumber="292">
<emphasis id="A7C1EA87DF718E0D69D6318D8869FA19" box="[199,305,1486,1512]" italics="true" pageId="21" pageNumber="292">Remarks.</emphasis>
This species is clearly distinguished from other members of
<taxonomicName id="52B54D16DF718E0D6AC8318D8D08FA19" authorityName=", Hope" authorityYear="1838" box="[985,1104,1486,1512]" class="Insecta" family="Carabidae" genus="Calochroa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="292" phylum="Arthropoda" rank="genus">
<emphasis id="A7C1EA87DF718E0D6AC8318D8D08FA19" box="[985,1104,1486,1512]" italics="true" pageId="21" pageNumber="292">Calochroa</emphasis>
</taxonomicName>
by the characteristics lacking B.
</paragraph>
</subSubSection>
</treatment>
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