207 lines
22 KiB
XML
207 lines
22 KiB
XML
<document id="67CA7C0F498E2E66BA023BA3C274506B" ENCODING="UTF-8" ID-GBIF-Dataset="d42acefb-3c32-4b96-8400-e1575aa17bc6" ModsDocID="z01578p041" checkinTime="1247322647604" checkinUser="thomas" docAuthor="Paul H. Skelton" docDate="2007" docId="BF305CA5C27B8E567919934D2C9F3AAE" docLanguage="en" docName="2007_Skelton_gg1.xml" docOrigin="Zootaxa 1578" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:91D85038-3B5D-4493-AE8C-6D2ECA205CAA" docTitle="Doumea sanaga Skelton, 2007, sp. nov." docType="treatment" docVersion="11" lastPageNumber="65" masterDocId="69594900A1F947C1E217DCD0031C8796" masterDocTitle="New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae)." masterLastPageNumber="68" masterPageNumber="41" pageNumber="63" updateTime="1701314704579" updateUser="plazi">
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<mods:mods id="5B55318FDC185B832F4F4B5805D0E5CC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0C71E2EA2C2FF77C6174CE79B4808C14">
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<mods:title id="FFD647FBACC141E5D625F1359616A968">New species of the amphiliid catfish genera Amphilius, Doumea and Phractura and the taxonomy of Paramphilius from West Central Africa (Siluriformes, Amphiliidae).</mods:title>
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</mods:titleInfo>
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<mods:name id="A89A351333A25FBBC1D1F76A7438FF5D" type="personal">
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<mods:role id="13EE00C648D80142C47B27608E874344">
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<mods:roleTerm id="D710A0810DFB01FCD49C9951895C6F68">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="63349ABC55B1879DC4E7ED24AD6F85A0">Paul H. Skelton</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E74B3CC8C17B2619DA02EB0730D0510A">text</mods:typeOfResource>
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<mods:title id="16250D18B9BD42D53A73BDB2681D395D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8B6C8C87C8D3D1A39D91860BCA834A11">
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<mods:date id="A13A71FEE7473B3231FCF100B1FDCF19">2007</mods:date>
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<mods:detail id="4FEB451790CDA6385B2EF40AD24B5919" type="volume">
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<mods:number id="86B480546084E94FE39B239E58F09445">1578</mods:number>
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<mods:extent id="E883136300D8594F36876899CC80F70D" unit="page">
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<mods:start id="A5853D92434513F022B624C65DF4EFB0">41</mods:start>
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<mods:end id="3BBF5F0FDCCF3138E58183AAF2E4CAE9">68</mods:end>
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<mods:location id="0573E6B75A243FADDE226FF469C0DB3B">
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<mods:url id="E03EB1787CB15042B14E894CDB7E1D8B">http://www.zoobank.org/urn:lsid:zoobank.org:pub:91D85038-3B5D-4493-AE8C-6D2ECA205CAA</mods:url>
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</mods:location>
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<mods:classification id="E41B079D5826A07CD5698157E84DB504">journal article</mods:classification>
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<mods:identifier id="15D2E3853634A24C77EE7A306F361D87" type="Plazi-Custom">z01578p041</mods:identifier>
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<mods:identifier id="3C6954887C7DBA8A6CD25BAC6AD0E118" type="ZooBank">91D85038-3B5D-4493-AE8C-6D2ECA205CAA</mods:identifier>
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</mods:mods>
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<treatment id="BF305CA5C27B8E567919934D2C9F3AAE" ID-DOI="http://doi.org/10.5281/zenodo.6236504" ID-GBIF-Taxon="100121965" ID-Zenodo-Dep="6236504" LSID="urn:lsid:plazi:treatment:BF305CA5C27B8E567919934D2C9F3AAE" httpUri="http://treatment.plazi.org/id/BF305CA5C27B8E567919934D2C9F3AAE" lastPageNumber="65" pageNumber="63">
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<subSubSection id="D1350FEE5851D9D8F1D9139D9B7C6E80" type="nomenclature">
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<paragraph id="925E982FF80A94880F9B72DA66E55BDA" pageNumber="63">
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<taxonomicName id="804B4FC7CFB628B037CDE50B7E7F07A2" ID-CoL="37HL2" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="sanaga">Doumea sanaga</taxonomicName>
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<taxonomicNameLabel id="311935052386B0EF8AE5191DCD37ABB2" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9558974FD6DD9F03F0C4B6187A6FE449" type="description">
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<paragraph id="E4369B9D8ECADB1C61E7C624FF0E548F" pageNumber="63">(Fig. 11)</paragraph>
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</subSubSection>
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<subSubSection id="26DC1546B2FDA8C802362C5FF393AF27" type="nomenclature">
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<paragraph id="98027027D231DE77E294828BBC164FDC" pageNumber="63">
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<materialsCitation id="CE3A1528C1AD8E1A0CC94F976AC6C512" ID-GBIF-Occurrence="923906126" country="Cameroon">
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Type material.
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<typeStatus id="3F440FC82AA59815C9021C23DD9A2A73">Holotype</typeStatus>
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,
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<collectionCode id="45D91D822D90FEC3C0E803E4534146B6" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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||
95-054-P-130, SL 79.6 mm, male, Mbal river, at Mbal, Sanaga River system,
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<geoCoordinate id="EBB59F97DBDD393B629E2BE706BDA3B9" direction="north" orientation="latitude" value="5.616667">05°37'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="E2F051BC5A1E0854D62FD9BB41002076" direction="east" orientation="longitude" value="14.05">14°03'E</geoCoordinate>
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,
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<collectingCountry id="BD48DEB5DFA0881ED5D5D29DCF26ABA2">Cameroon</collectingCountry>
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||
, col. E. van den Bergh and Forbin,
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<date id="CA9E11401C84CEEC5FF40D6EB9A22A62" day="04" month="03" value="1994-03-04" year="1994">4 March 1994</date>
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||
</materialsCitation>
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||
;
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<materialsCitation id="D3AB5E5F8B45A6F617BF147F6160B818" ID-GBIF-Occurrence="923906118" country="Cameroon">
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31
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<typeStatus id="25E4C5E772B12F8A699FB43267D9AC3E">paratypes</typeStatus>
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, 1,
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<collectionCode id="A59746719D5A72D4C845D371071E1357" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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95-054-P-131, SL 50 mm, same data as holotype
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</materialsCitation>
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||
;
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<materialsCitation id="30D96D7D244BC35F9971105C7B2598E4" ID-GBIF-Occurrence="923906116" country="Cameroon">
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2,
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<collectionCode id="D4FAB0D02BCEE374F44F6A2F9AA8292E" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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95-054-P-132-133, SL 36-39 mm, Kinkina River, tributary of the Lom River, below the Garoua Boulai bridge, Sanaga River system,
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<geoCoordinate id="F75B85E6020F0F8923B5E18C7837778A" direction="north" orientation="latitude" value="6.0666666">06°04'N</geoCoordinate>
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,
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<geoCoordinate id="5521025E70D62D186C8936D15DBDB997" direction="east" orientation="longitude" value="14.366667">14°22'E</geoCoordinate>
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,
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<collectingCountry id="492FDFFB8CCEF74FA7359E0104F97851">Cameroon</collectingCountry>
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, col. E. van den Bergh and Forbin,
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<date id="D5A5A3DDEF23B0E175A9E80A0B321A8A" day="06" month="03" value="1994-03-06" year="1994">6 March 1994</date>
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||
</materialsCitation>
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;
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<materialsCitation id="5B9C410ECC8F32703A11BDD5FCE14A7D" ID-GBIF-Occurrence="923906120" country="Cameroon">
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3,
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<collectionCode id="F616B35C2EDCA28563390A2EDE4E2374" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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95-054-P-134-136, SL 31-35 mm,
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<normalizedToken id="4CB3A9152F033C897E20FB90773BD35C" originalValue="Sessé">Sesse</normalizedToken>
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River, at the entrance to Massa and Manboya, Sanaga River system,
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<geoCoordinate id="AD204F92D5DE6426E3B0EF8956205483" direction="north" orientation="latitude" value="5.0666666">05°04'N</geoCoordinate>
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,
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<geoCoordinate id="F626C5F884E008F3E890E689DEE4A552" direction="east" orientation="longitude" value="13.5">13°30'E</geoCoordinate>
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, col. E. van den Bergh and Forbin,
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<date id="36821433C17C53E3250AE26BA4888A03" day="03" month="03" value="1994-03-03" year="1994">3 March 1994</date>
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</materialsCitation>
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;
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<materialsCitation id="3D34E37248F6998D4799983DF2C3B328" ID-GBIF-Occurrence="923906110" country="Cameroon">
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||
1,
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<collectionCode id="E8D90FEFC1000B086DF145E9597BC6C8" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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95-054-P-137, SL 42 mm, Lom river, at Wakaso, Sanaga River system,
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<geoCoordinate id="6ED056CB68F6B862AC9F9619EC19AF81" direction="north" orientation="latitude" value="6.3166666">06°19'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="F8956733D08B57A2FF5B7DEA40D22EBA" direction="east" orientation="longitude" value="14.5">14°30'E</geoCoordinate>
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||
,
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||
<collectingCountry id="8C4CD3E1D8F11D693715317514E49EE6">Cameroon</collectingCountry>
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||
, col. E. van den Bergh and Forbin,
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||
<date id="758AA1F0856B3420BAAD8C2DBAC812DE" day="08" month="03" value="1994-03-08" year="1994">8 March 1994</date>
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="A3437203605D9EBB2F658A0EB7563F54" ID-GBIF-Occurrence="923906119" country="Cameroon">
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||
8,
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||
<collectionCode id="AFFA80BB79C794125E05AEA232E2DB4E" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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95-054-P-138-145, SL 36-44 mm, Lom river, at bridge entrance to Betare Oya and Garoua Boulai, Sanaga River system,
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<geoCoordinate id="8920FB8E443D9FCBE9B376DB234464BE" direction="north" orientation="latitude" value="6.0833335">06°05'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EAA166E3D54A15DDAD8132BF53DD7630" direction="east" orientation="longitude" value="14.383333">14°23'E</geoCoordinate>
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||
,
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||
<collectingCountry id="23AE128B4E4EA5A1D5ACE3E9B08C1A81">Cameroon</collectingCountry>
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||
, col. E. van den Bergh and Forbin
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||
</materialsCitation>
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||
;
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||
<materialsCitation id="D9B9D98B72CB197C098FFDDF7895CBB7" ID-GBIF-Occurrence="923906129" country="Cameroon">
|
||
16,
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||
<collectionCode id="BE9EAA3FB12D90173B4E25393E8109E0" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale">MRAC</collectionCode>
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||
95-054-P-146-151, SL 35-53.6 mm, Mi river, at Tigoum, Sanaga River system,
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||
<geoCoordinate id="6F0B93C722FABC33F2314A9DD9EEB715" direction="north" orientation="latitude" value="6.233333">06°14'N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE1255D14985F80C24B3BE93B055A57E" direction="east" orientation="longitude" value="14.383333">14°23'E</geoCoordinate>
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||
,
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||
<collectingCountry id="72F5AFB38C07394FF7491B28F9C2047B">Cameroon</collectingCountry>
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||
, col. E.van den Bergh and Forbin,
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||
<date id="7E26EBDABC3732A7F4DA2065E68399D7" day="06" month="03" value="1994-03-06" year="1994">6 March 1994</date>
|
||
</materialsCitation>
|
||
;
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||
<materialsCitation id="6A76648E264E2FDC95E6E0A2CAC8F411" ID-GBIF-Occurrence="923906132" country="Cameroon">
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||
5,
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<collectionCode id="EF7CFF06F18379882B71D99DB53902BC" collectionName="USA, New York, New York, American Museum of Natural History">AMNH</collectionCode>
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240491 (ex MRAC 95-054-P-158-162), 38.7-47.1 mm SL, same data as MRAC 95-054-P-146-151
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</materialsCitation>
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||
;
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<materialsCitation id="C54B3A5903CB7D792BD38483E0EA71EC" ID-GBIF-Occurrence="923906115" country="Cameroon">
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||
6,
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||
<collectionCode id="9098DA620685BF28E5F0608A6F592773" collectionName="SAIAB">SAIAB</collectionCode>
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79511 (ex MRAC 95-054-P-152-157), 39-46.1 mm SL, same data as MRAC 95-054-P-146-151
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="DB4991AB7A45F0FB482927CED51E5A0C" type="diagnosis">
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<paragraph id="F59EDC66FFBF94A0DFD3B84BBB2D62ED" pageNumber="63">
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Diagnosis.
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<taxonomicName id="2B68AE1CCA682EF0BE7A9DA9D9FD6B95" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="sanaga">D. sanaga</taxonomicName>
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is a relatively small species of
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<taxonomicName id="49D3B1BF850E0D7075BB1FFC9A0CC23F" LSID-ZBK="urn:lsid:zoobank.org:act:1148424C-8833-4034-9EDB-303E964EE1DB" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea Sauvage 1879:96 [7]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="genus">Doumea</taxonomicName>
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(maximum size recorded is SL 79.6 mm) with a relatively long, (3.4-3.6 times in SL), slender (depth into length 9-11.5 times) caudal peduncle. The species is most similar to
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<taxonomicName id="0D4DEB791C7B7BA5D23ECC6D047E36E9" LSID-ZBK="urn:lsid:zoobank.org:act:DD318F95-2F88-40F6-950E-7AC858150E2D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea typica Sauvage 1879:97 [8]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="typica">D. typica</taxonomicName>
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and
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<taxonomicName id="1A80C81D6803A8A2782955B48CC69890" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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but differs from both these species in several features including: the shape of the head, which has a broad round snout and anterior profile vs. a triangular and pointed profile in
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<taxonomicName id="738825E45D8D4727CE53C460E04A2B72" LSID-ZBK="urn:lsid:zoobank.org:act:DD318F95-2F88-40F6-950E-7AC858150E2D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea typica Sauvage 1879:97 [8]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="typica">D. typica</taxonomicName>
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and
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<taxonomicName id="DE58538B9610B5574D1E940020F98573" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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; the edge of the branchiostegal membrane forms a distinct shallow ‘V across the ventral surface vs.a straight or nearly straight edge in
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<taxonomicName id="F15442BA277FB8FE860E9A102FD27D89" LSID-ZBK="urn:lsid:zoobank.org:act:DD318F95-2F88-40F6-950E-7AC858150E2D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea typica Sauvage 1879:97 [8]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="typica">D. typica</taxonomicName>
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and
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<taxonomicName id="BB1F6AC219311D6D7913D3C8EC631B47" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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; the leading dorsal (nuchal) pterygiophore intercepting at least the second or third post Weberian vertebra vs.in
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<taxonomicName id="3A9E18B1F7B4BFE90C8177B38E38B374" LSID-ZBK="urn:lsid:zoobank.org:act:DD318F95-2F88-40F6-950E-7AC858150E2D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea typica Sauvage 1879:97 [8]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="typica">D. typica</taxonomicName>
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and
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<taxonomicName id="1C2F29BB4D1AD7AFBCC5E8257D449EB7" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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the dorsal leading pterygiophore intercepts the first post-Weberian vertebra; in the relative position of the pelvic fins, the origin of the pelvic fins is beneath the base of the dorsal fin vs.behind the base of the dorsal in
|
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<taxonomicName id="71136CF002B78691D8C7637E4E39375C" LSID-ZBK="urn:lsid:zoobank.org:act:DD318F95-2F88-40F6-950E-7AC858150E2D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea typica Sauvage 1879:97 [8]" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="typica">D. typica</taxonomicName>
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and
|
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<taxonomicName id="5609294FF158FF17E0018F3FC7D30B2B" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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. It differs from
|
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<taxonomicName id="BE25852724E6CD4AD74D3AFE161A6E4B" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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<taxonomicNameLabel id="D6D6CBCF98830F01A967F822909BDC49" rank="species">sp. nov.</taxonomicNameLabel>
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in the length of the caudal peduncle (
|
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<taxonomicName id="BA1CCEAF701CA8CAE81D6377B88DD421" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="gracila">D. gracila</taxonomicName>
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caudal peduncle in SL 2.9-3.2 vs. 3.4-3.6
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<taxonomicName id="63FADE9EF466C469833C3D9B53F11C6C" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="sanaga">D. sanaga</taxonomicName>
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). It differs from
|
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<taxonomicName id="20181A9BD65C25E44A28AF37CAA09C94" LSID-ZBK="urn:lsid:zoobank.org:act:E1C58DFC-DB30-419E-A65F-DDA497253198" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea thysi Skelton 1989:6" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="thysi">D. thysi</taxonomicName>
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in the length and depth of the caudal peduncle (28-30% SL vs.in
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<taxonomicName id="E49810CC7F2624284D2F56132A563821" LSID-ZBK="urn:lsid:zoobank.org:act:E1C58DFC-DB30-419E-A65F-DDA497253198" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea thysi Skelton 1989:6" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="thysi">D. thysi</taxonomicName>
|
||
the caudal peduncle is short 20% SL, and stout, 9-11 times long as deep vs.2.5-4.0 times long as deep in
|
||
<taxonomicName id="4F21EEDC717B5240738EF7F676A45923" LSID-ZBK="urn:lsid:zoobank.org:act:E1C58DFC-DB30-419E-A65F-DDA497253198" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea thysi Skelton 1989:6" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="thysi">D. thysi</taxonomicName>
|
||
),as well as in color pattern (
|
||
<taxonomicName id="391EBEAC01702624DB26F86629E4169D" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="sanaga">D. sanaga</taxonomicName>
|
||
has a series of light saddles that are not characteristic of
|
||
<taxonomicName id="182F6C53E7C62A16D6EAEE2D0C172417" LSID-ZBK="urn:lsid:zoobank.org:act:E1C58DFC-DB30-419E-A65F-DDA497253198" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" lsidName-ZBK="Doumea thysi Skelton 1989:6" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="thysi">D. thysi</taxonomicName>
|
||
, and the latter has a distinctly demarcated white band along the lateral line that is not characteristic of
|
||
<taxonomicName id="9FD09274759F8596A103F5BFC4B9F739" class="Actinopterygii" family="Amphiliidae" genus="Doumea" kingdom="Animalia" order="Siluriformes" pageNumber="63" phylum="Chordata" rank="species" species="sanaga">D. sanaga</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="593449EE6DB3EAEA159D9D6ECC8ED07A" type="description">
|
||
<paragraph id="265179D4C7EC363C0B748873A45D93EB" pageNumber="64">Description. Measurements and fin ray counts given in Table 8. Body firm, slender, moderately depressed, strongly tapered to caudal base, with a mid dorsal ridge from dorsal to adipose, and thin bi-lateral ridges along caudal peduncle. Skin is smooth or in larger individuals the head and upper body is covered with short vermiculate white unculi. Head short 4.8-5.5 times in SL, obtusely pointed, flat below, rounded above; snout longer than postorbit, 1.5-1.9 times in head length; nares separated, on anterior half of snout; mouth ovoid, ventral, upper lips fleshy and papillose, lower lip divided by medial triangular pad; barbels short, tapered, papillose, maxillaries longest less than 2/3 head length, extend from lateral edge of upper lip not reaching edge of branchiostegal membrane; outer mandibular barbels from lateral corner of mouth to about 2/ 3 distance to edge of branchiostegal membrane; inner mandibular barbels from medial position on ventral side of mandible separated by medial mandibular pad, reaching less than halfway to edge of branchiostegal membrane; eyes small, orbits bony, widely separated by distance 1.9-2.4 times orbit diameter; supraoccipital process slender, half length of snout, separated from nuchal shield by gap; obscure short humeral process above gill openings; branchiostegal membrane united ventrally, margin with shallow medial notch. Caudal peduncle long, 3.4-3.6 times in SL, slender, length 8.7-11.1 times depth, and bony. Fins large and pointed; origin of dorsal less than one head length behind head, dorsal entirely in advance of pelvic fins, dorsal fin pointed with concave hind margin; pectoral fins falcate, not reaching bases of pelvic fins, leading ray curved, pectinate and padded, outer rays lie horizontal, inner rays vertical against body; origin of pelvic fins beneath posterior edge of dorsal base, pelvic fins falcate, reaching base of anal fin, leading ray curved, pectinate and padded; anal fin large, trailing edge straight; caudal fin with compact base, forked with ventral lobe slightly longer than dorsal; adipose fin triangulate, positioned over posterior base of anal. Ano-genital pores placed midway between bases of pelvic fins and origin of anal fin, males with conical genital papillae. Skin smooth or rugose with small white unculi on predorsal and dorsal surface of head in larger specimens. Lateral line complete, with short dorsal and ventral branches to pores along its length, visible as a thin white line in preserved specimens. Reaches 80 mm SL.</paragraph>
|
||
<paragraph id="E52464C5E4844B441F1325C675A29F0E" pageNumber="65">Coloration. Dark brown above, light cream below, flanks light brown, lateral line highlighted in cream, head with dark pre-orbital stripes and light patches above, a series of 5-6 lighter saddles along back, dark brown patches above pelvic fin bases, fins with brown rays and dark mid-distal band, outer rays of caudal brown. Lateral line visible as thin white line along mid-flanks.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A82A708FF38FFE9405BC17D7823308C3" type="distribution">
|
||
<paragraph id="3905652E95F71769BD142A0C07D7F84C" pageNumber="65">Distribution. Upper reaches of the Sanaga River, in Cameroon (Fig. 10).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DA5CA016FCF3B7192C77F11F7CDA0061" type="etymology">
|
||
<paragraph id="7DF0F8DA0CECB3160D0873454A86AB5B" pageNumber="65">Etymology. From ‘Sanaga’ the name of the river system where the species occurs. Used as a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |