treatments-xml/data/03/EF/BB/03EFBB36FFA78059CFC8FED54256F987.xml

<document id="FF1FD4C04E7FB8EAB327CF3AF6D51F1C" ID-DOI="10.5281/zenodo.5341574" ID-ISSN="2345-7600" ID-Zenodo-Dep="5341574" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190967238" checkinUser="diego" docAuthor="Cai, Yixiong, Choy, Satish &amp; Ng, Peter K. L." docDate="2009" docId="03EFBB36FFA78059CFC8FED54256F987" docLanguage="en" docName="57rbz065-089.pdf" docOrigin="Raffles Bulletin of Zoology 57 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Caridina liaoi Cai &amp; Choy &amp; Ng 2009, new species" docType="treatment" docVersion="4" lastPageNumber="75" masterDocId="FFD6C34EFFA08053CC18FFEE406CFFD5" masterDocTitle="Epigean And Hypogean Freshwater Shrimps Of Bohol Island, Central Philippines (Crustacea: Decapoda: Caridea)" masterLastPageNumber="89" masterPageNumber="65" pageNumber="72" updateTime="1699181529684" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A672049FAF6C240DFF64C5ED6CAF8E9B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EAF2AF8FD77904C0E6E7ACC698F2B2C2">
<mods:title id="C95ED2A018039A2D97510A9767A2961F">Epigean And Hypogean Freshwater Shrimps Of Bohol Island, Central Philippines (Crustacea: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="6DC3532949EA66F9586F6319C21B7DAF" type="personal">
<mods:role id="32D2B66E9EE8BCE1F6A2FB88C9D01638">
<mods:roleTerm id="D248BE4A8F364AD84BFB584BC0774927">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A7FB2310BCACAE19EB6FBCFE81B0FF6D">Cai, Yixiong</mods:namePart>
</mods:name>
<mods:name id="A6AEB706C403D4FB42AA107E452C8B5F" type="personal">
<mods:role id="C2C89C4BBBEF587EC06E66B05BDDECAC">
<mods:roleTerm id="2476D9F39123CFDFAE8C1A2FBDC3A3C6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="67F389AE858787A777CFD021894F546F">Choy, Satish</mods:namePart>
</mods:name>
<mods:name id="8FE80B57D832F4030384F8CD087A093D" type="personal">
<mods:role id="1323282102DBAC65A9635FC16B098440">
<mods:roleTerm id="9C1ED35E08EFBCA901FF269B2892AB2A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A61642FD0B709E44811C15C8E842C79E">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="209D6E50B4433A44587686CADF850679">text</mods:typeOfResource>
<mods:relatedItem id="A7F19DA512ACEB1F9AE14BD7D3F55B50" type="host">
<mods:titleInfo id="AD8799CD31FC9387F8D3E0E15040AF72">
<mods:title id="218F36A20ACB85E5B8B4E26126F10E12">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="CB5D680F6E3B2613BE9A3E6711415186">
<mods:date id="83F9CD48320DE57D902CD4D61359AFAC">2009</mods:date>
<mods:detail id="F356A228339F77CF42F343CC52B765EE" type="pubDate">
<mods:number id="45DE52D9BF0FE4CA2C456D903694AF6A">2009-02-28</mods:number>
</mods:detail>
<mods:detail id="8D50824E871869E4FB84DD9BFD86223C" type="volume">
<mods:number id="2532302DA163F4DCA08192866EB58AF9">57</mods:number>
</mods:detail>
<mods:detail id="8049CF26EA745E580117DAF1F0545FC5" type="issue">
<mods:number id="39D188E5CB52723C0A62E3C21858EEA8">1</mods:number>
</mods:detail>
<mods:extent id="2276A1139AA141521B224E32BF757CFA" unit="page">
<mods:start id="4640E517B9D62E651E32A49DB9F0A811">65</mods:start>
<mods:end id="95EE5A881EA1A6FFED18CD079CD77997">89</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F37FEA852FCD7E56BC22A8BE618DCC1A">journal article</mods:classification>
<mods:identifier id="DEDF9C62BC306F1D6292E89F4CE2CEC6" type="DOI">10.5281/zenodo.5341574</mods:identifier>
<mods:identifier id="E4ED0E5A59585F514AC0BDD83A0EB169" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="40633F43A7239F29B74E77324D47C41A" type="Zenodo-Dep">5341574</mods:identifier>
</mods:mods>
<treatment id="03EFBB36FFA78059CFC8FED54256F987" ID-DOI="http://doi.org/10.5281/zenodo.5444222" ID-Zenodo-Dep="5444222" LSID="urn:lsid:plazi:treatment:03EFBB36FFA78059CFC8FED54256F987" httpUri="http://treatment.plazi.org/id/03EFBB36FFA78059CFC8FED54256F987" lastPageId="10" lastPageNumber="75" pageId="7" pageNumber="72">
<subSubSection id="C35C59ABFFA78054CFC8FED5456CFE86" box="[976,1280,315,339]" pageId="7" pageNumber="72" type="nomenclature">
<paragraph id="8BF90A20FFA78054CFC8FED5456CFE86" blockId="7.[976,1280,315,370]" box="[976,1280,315,339]" pageId="7" pageNumber="72">
<heading id="D0B1BD4CFFA78054CFC8FED5456CFE86" bold="true" box="[976,1280,315,339]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="72" reason="2">
<emphasis id="B932D632FFA78054CFC8FED5456CFE86" bold="true" box="[976,1280,315,339]" pageId="7" pageNumber="72">
<taxonomicName id="4C4671A3FFA78054CFC8FED54407FE86" ID-CoL="8VK9F" authority="Cai &amp; Choy &amp; Ng, 2009" authorityName="Cai &amp; Choy &amp; Ng" authorityYear="2009" box="[976,1131,315,339]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="72" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="B932D632FFA78054CFC8FED54407FE86" bold="true" box="[976,1131,315,339]" italics="true" pageId="7" pageNumber="72">Caridina liaoi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2016B49FFA78054C863FED5456CFE86" box="[1147,1280,315,339]" pageId="7" pageNumber="72" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35C59ABFFA78054C833FEB544C8FEA7" box="[1067,1188,347,370]" pageId="7" pageNumber="72" type="description">
<paragraph id="8BF90A20FFA78054C833FEB544C8FEA7" blockId="7.[976,1280,315,370]" box="[1067,1188,347,370]" pageId="7" pageNumber="72">
(
<figureCitation id="137D16A5FFA78054C82BFEB544ECFEA7" box="[1075,1152,347,370]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="7" pageNumber="72">Figs. 5</figureCitation>
, 
<figureCitation id="137D16A5FFA78054C896FEB544F0FEA7" box="[1166,1180,347,370]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C59ABFFA78054CF30FE7443F3FDEA" pageId="7" pageNumber="72" type="materials_examined">
<paragraph id="8BF90A20FFA78054CF30FE744535FE19" blockId="7.[808,1448,410,460]" pageId="7" pageNumber="72">
<emphasis id="B932D632FFA78054CF30FE744469FE7A" bold="true" box="[808,1029,410,431]" italics="true" pageId="7" pageNumber="72">Material examined. –</emphasis>
<typeStatus id="54FDB482FFA78054C816FE744401FE7A" box="[1038,1133,410,431]" pageId="7" pageNumber="72" type="holotype">Holotype</typeStatus>
: Ovigerous female, cl 
<quantity id="4CBEA7C5FFA78054C94CFE7445C8FE7A" box="[1364,1444,410,431]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="7" pageNumber="72" unit="mm" value="4.1">4.1 mm</quantity>
, USC, Bilar River, Bilar, coll. Y. Cai et al., 
<date id="FFF82CE0FFA78054C8CDFE594539FE19" box="[1237,1365,439,460]" pageId="7" pageNumber="72" value="2000-12-19">19 Dec.2000</date>
.
</paragraph>
<paragraph id="8BF90A20FFA78054CF30FE1E43F3FDEA" blockId="7.[808,1448,496,575]" pageId="7" pageNumber="72">
<typeStatus id="54FDB482FFA78054CF30FE1E43E5FDD0" box="[808,905,496,517]" pageId="7" pageNumber="72" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D40C1A9FFA78054CF8BFE1E4380FDD0" box="[915,1004,496,517]" pageId="7" pageNumber="72" type="male">82 males</specimenCount>
, cl 
<quantity id="4CBEA7C5FFA78054C808FE1E44EBFDD0" box="[1040,1159,496,517]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="4.3" metricValueMin="2.8" pageId="7" pageNumber="72" unit="mm" value="3.55" valueMax="4.3" valueMin="2.8">2.8–4.3 mm</quantity>
, 
<materialsCitation id="3B2E007DFFA78054C88AFE1E43F7FDEA" collectionCode="ZRC" pageId="7" pageNumber="72" specimenCode="ZRC 2007.0284" specimenCount="112" specimenCount-female="112" typeStatus="paratype">
<specimenCount id="9D40C1A9FFA78054C88AFE1E4566FDD0" box="[1170,1290,496,517]" pageId="7" pageNumber="72" type="female">112 females</specimenCount>
, cl 3.7–5.0 mm, 26 ovigerous females, cl 4.2–5.0 mm, 
<specimenCode id="DBE0A25BFFA78054C886FDE34551FDF7" box="[1182,1341,525,546]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="7" pageNumber="72">ZRC 2007.0284</specimenCode>
, data same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C59ABFFA78059CF30FD86411EFC67" lastPageId="10" lastPageNumber="75" pageId="7" pageNumber="72" type="description">
<paragraph id="8BF90A20FFA78054CF30FD864542FD0A" blockId="7.[808,1449,616,735]" pageId="7" pageNumber="72">
<emphasis id="B932D632FFA78054CF30FD8643AFFD55" bold="true" box="[808,963,616,640]" italics="true" pageId="7" pageNumber="72">Description. –</emphasis>
Rostrum (
<figureCitation id="137D16A5FFA78054C820FD8644F5FDAA" box="[1080,1177,616,639]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="7" pageNumber="72">Figs. 5A</figureCitation>
, 
<figureCitation id="137D16A5FFA78054C8BDFD8644A4FDAA" box="[1189,1224,616,639]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">6A</figureCitation>
) long, reaching near to or slightly beyond end of scaphocerite; rostral formula: 3–4+13–14/13–15. Antennal spine fused with inferior orbital angle. Pterygostomian margin broadly rounded.
</paragraph>
<paragraph id="8BF90A20FFA78054CF30FCE6448EFC0A" blockId="7.[808,1448,776,991]" pageId="7" pageNumber="72">
Sixth abdominal somite 0.51 times of carapace, 1.7 times as long as fifth somite, distinctly shorter than telson. Telson (
<figureCitation id="137D16A5FFA78054CF28FCA643C0FC8A" box="[816,940,840,863]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">Figs. 6B, C</figureCitation>
) 3.5 times as long as wide, not terminating in a projection, with 3 or 4 pairs of dorsal spinules and 1 pair of dorsolateral spinules; lateral pair of spines distinctly longer than intermediate pairs of spiniform setae. Preanal carina (
<figureCitation id="137D16A5FFA78054CF28FC2643EAFC0A" box="[816,902,968,991]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">Fig. 6K</figureCitation>
) high, triangular, lacking spine.
</paragraph>
<paragraph id="8BF90A20FFA78054CF30FBE6441BFACA" blockId="7.[808,1448,1032,1311]" pageId="7" pageNumber="72">
Eyes well developed, anterior end reaching to 0.8 times length of basal segment of antennular peduncle. Antennular peduncle 0.74 times as long as carapace; basal segment of antennular peduncle longer than both second and third segment lengths, anterolateral angle reaching 0.20 times length of second segment, second segment distinctly longer than third segment. Stylocerite reaching to 0.8 times length of basal segment of antennular peduncle. Scaphocerite (
<figureCitation id="137D16A5FFA78054C965FB064325FACA" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">Fig. 6D</figureCitation>
) 3.4 times as long as wide.
</paragraph>
<paragraph id="8BF90A20FFA78054CF30FAA643E4F9AA" blockId="7.[808,1448,1352,1663]" pageId="7" pageNumber="72">Incisor process of mandible ending in irregular teeth, molar process truncated. Lower lacinia of maxillula broadly rounded, upper lacinia elongated, with a number of distinct teeth on inner margin, palp slender. Upper endites of maxilla subdivided, palp short, scaphognathite tapering posteriorly with some long, curved setae at posterior end. Palp of first maxilliped broadly triangular. Second maxilliped typical of genus. Third maxilliped reaching to end of antennular peduncle, with ultimate segment as long as penultimate segment.</paragraph>
<paragraph id="8BF90A20FFA78059CF30F94642C9FE67" blockId="7.[808,1449,1704,2079]" lastBlockId="10.[132,771,187,434]" lastPageId="10" lastPageNumber="75" pageId="7" pageNumber="72">
Epipods on first 4 pereiopods. First pereiopod (
<figureCitation id="137D16A5FFA78054C902F946451AF96A" box="[1306,1398,1704,1727]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="7" pageNumber="72">Figs. 5B</figureCitation>
, 
<figureCitation id="137D16A5FFA78054C999F94645CCF96A" box="[1409,1440,1704,1727]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">6E</figureCitation>
) reaching to end of basal segment of antennular peduncle; merus 1.9–2.2 times as long as broad, shorter than carpus; carpus excavated anteriorly, shorter than chela, 1.3–1.7 times as long as high; chela 2.3–3.5 time as long as broad; fingers slightly shorter, as long as or distinctly longer than palm. Second pereiopod (
<figureCitation id="137D16A5FFA78054CFE1F886443BF8AA" box="[1017,1111,1896,1919]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="7" pageNumber="72">Figs. 5C</figureCitation>
, 
<figureCitation id="137D16A5FFA78054C87CF88644EDF8AA" box="[1124,1153,1896,1919]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">6F</figureCitation>
) reaching to end of second segment of antennular peduncle; merus as long as carpus, 4.2–5.0 times as long as broad; carpus 1.1 times as long as chela, 2.3–2.4 times as long as high; chela 2.3–2.4 times as long as broad; fingers 1.6 times as long as palm. Third pereiopod (
<figureCitation id="137D16A5FFA78054CFB8F7E64477F7CA" box="[928,1051,2056,2079]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="7" pageNumber="72">Figs. 5D, E</figureCitation>
, 
<figureCitation id="137D16A5FFA78054C83DF7E6440AF7CA" box="[1061,1126,2056,2079]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="7" pageNumber="72">6G, H</figureCitation>
) reaching to end of antennular peduncle, propodus 11–12 times as long as broad, 4.5–4.7 times as long as dactylus; dactylus 3.1–3.3 times as long as wide (spines included), with 4–6 accessory spines on flexor margin. Fifth pereiopod reaching to end of second segment of antennular peduncle, propodus 14–18 times as long as broad, 3.6–3.8 times as long as dactylus, dactylus 4.0–4.4 times as long as wide (spinules included), terminating in 1 large claw, with 60–65 spinules on flexor margin.
</paragraph>
<caption id="DF395AA8FFA8805BCC9CF83C4264F7E8" ID-DOI="http://doi.org/10.5281/zenodo.5341584" ID-Zenodo-Dep="5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="8" pageNumber="73" startId="8.[132,167,2002,2023]" targetBox="[180,1408,205,1957]" targetPageId="8">
<paragraph id="8BF90A20FFA8805BCC9CF83C4264F7E8" blockId="8.[132,1469,2002,2109]" pageId="8" pageNumber="73">
Fig. 5. 
<taxonomicName id="4C4671A3FFA8805BCCD4F83C413AF832" authority="Cai &amp; Choy &amp; Ng, 2009" authorityName="Cai &amp; Choy &amp; Ng" authorityYear="2009" box="[204,342,2002,2023]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="73" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="B932D632FFA8805BCCD4F83C413AF832" box="[204,342,2002,2023]" italics="true" pageId="8" pageNumber="73">Caridina liaoi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2016B49FFA8805BCD7CF83C41B0F832" box="[356,476,2002,2023]" pageId="8" pageNumber="73" rank="species">new species</taxonomicNameLabel>
. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river).
</paragraph>
</caption>
<caption id="DF395AA8FFA9805ACC77F83F423EF7E8" ID-DOI="http://doi.org/10.5281/zenodo.5341586" ID-Zenodo-Dep="5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="9" pageNumber="74" startId="9.[111,146,2001,2022]" targetBox="[143,1381,178,1973]" targetPageId="9">
<paragraph id="8BF90A20FFA9805ACC77F83F423EF7E8" blockId="9.[111,1449,2001,2109]" pageId="9" pageNumber="74">
Fig. 6. 
<taxonomicName id="4C4671A3FFA9805ACCA1F83C4129F832" authority="Cai &amp; Choy &amp; Ng, 2009" authorityName="Cai &amp; Choy &amp; Ng" authorityYear="2009" box="[185,325,2002,2023]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="74" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">
<emphasis id="B932D632FFA9805ACCA1F83C4129F832" box="[185,325,2002,2023]" italics="true" pageId="9" pageNumber="74">Caridina liaoi</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A2016B49FFA9805ACD4DF83F41A2F833" box="[341,462,2001,2022]" pageId="9" pageNumber="74" rank="species">new species</taxonomicNameLabel>
, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River).
</paragraph>
</caption>
<paragraph id="8BF90A20FFAA8059CC9CFE35410EFDA7" blockId="10.[132,771,475,626]" pageId="10" pageNumber="75">
Endopod of male first pleopod (
<figureCitation id="137D16A5FFAA8059CDF3FE35422EFE27" box="[491,578,475,498]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="10" pageNumber="75">Fig. 5H</figureCitation>
) sub-rectangular, two-fifth length of exopod, no appendix interna. Appendix masculina of male second pleopod (
<figureCitation id="137D16A5FFAA8059CE3DFDF54218FDE7" box="[549,628,539,562]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="10" pageNumber="75">Fig. 5I</figureCitation>
) half length of endopod, with appendix interna reaching near end of appendix masculina.
</paragraph>
<paragraph id="8BF90A20FFAA8059CC9CFD75408FFD07" blockId="10.[132,769,667,722]" pageId="10" pageNumber="75">
Uropodal diaeresis (
<figureCitation id="137D16A5FFAA8059CD90FD754185FD67" box="[392,489,667,690]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="10" pageNumber="75">Fig. 6L</figureCitation>
) with 14–16 movable spinules.
</paragraph>
<paragraph id="8BF90A20FFAA8059CC9CFD1540D9FCE7" blockId="10.[132,771,763,818]" pageId="10" pageNumber="75">
Ovigerous females with eggs sized 0.92–0.95 × 
<quantity id="4CBEA7C5FFAA8059CE80FD1540DDFCE7" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="10" pageNumber="75" unit="mm" value="0.575" valueMax="0.6" valueMin="0.55">0.55–0.60 mm</quantity>
.
</paragraph>
<paragraph id="8BF90A20FFAA8059CC9CFCB5411EFC67" blockId="10.[132,771,859,946]" pageId="10" pageNumber="75">
<emphasis id="B932D632FFAA8059CC9CFCB541F9FCA7" box="[132,405,859,883]" italics="true" pageId="10" pageNumber="75">
<emphasis id="B932D632FFAA8059CC9CFCB5409FFCA6" bold="true" box="[132,243,859,883]" italics="true" pageId="10" pageNumber="75">Habitat. –</emphasis>
<taxonomicName id="4C4671A3FFAA8059CCE4FCB541F9FCA7" authority="Cai &amp; Choy &amp; Ng, 2009" authorityName="Cai &amp; Choy &amp; Ng" authorityYear="2009" box="[252,405,859,882]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">Caridina liaoi</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="A2016B49FFAA8059CDBCFCB54245FCA7" box="[420,553,859,882]" pageId="10" pageNumber="75" rank="species">new species</taxonomicNameLabel>
, was collected from a tributary of the Bilar River near Bilar town, 
<collectingRegion id="4982C4C2FFAA8059CE61FC9542D6FC47" box="[633,698,891,914]" country="Philippines" name="Bohol" pageId="10" pageNumber="75">Bohol</collectingRegion>
Island in central 
<collectingCountry id="F3514AB0FFAA8059CCEBFC754103FC67" box="[243,367,923,946]" name="Philippines" pageId="10" pageNumber="75">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35C59ABFFAA8059CC9CFC3540A9FB87" pageId="10" pageNumber="75" type="etymology">
<paragraph id="8BF90A20FFAA8059CC9CFC3540A9FB87" blockId="10.[132,771,987,1106]" pageId="10" pageNumber="75">
<emphasis id="B932D632FFAA8059CC9CFC35417FFC26" bold="true" box="[132,275,987,1011]" italics="true" pageId="10" pageNumber="75">Etymology. –</emphasis>
The new species is named after Dr. Lawrence Liao, who has been instrumental in helping the first author making collections in the 
<collectingCountry id="F3514AB0FFAA8059CDDBFBF5422AFBE7" box="[451,582,1051,1074]" name="Philippines" pageId="10" pageNumber="75">Philippines</collectingCountry>
for the present study.
</paragraph>
</subSubSection>
<subSubSection id="C35C59ABFFAA8059CC9CFB954256F987" pageId="10" pageNumber="75" type="discussion">
<paragraph id="8BF90A20FFAA8059CC9CFB954256F987" blockId="10.[132,771,1147,1618]" pageId="10" pageNumber="75">
<emphasis id="B932D632FFAA8059CC9CFB9541CBFB47" box="[132,423,1147,1171]" italics="true" pageId="10" pageNumber="75">
<emphasis id="B932D632FFAA8059CC9CFB954096FB46" bold="true" box="[132,250,1147,1171]" italics="true" pageId="10" pageNumber="75">Remarks.-</emphasis>
<taxonomicName id="4C4671A3FFAA8059CD1DFB9541CBFB47" authority="Cai &amp; Choy &amp; Ng, 2009" authorityName="Cai &amp; Choy &amp; Ng" authorityYear="2009" box="[261,423,1147,1170]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="liaoi" status="sp. nov.">Caridina liaoi</taxonomicName>
</emphasis>
, 
<taxonomicNameLabel id="A2016B49FFAA8059CDA1FB954229FB47" box="[441,581,1147,1170]" pageId="10" pageNumber="75" rank="species">new species</taxonomicNameLabel>
, is similar to 
<taxonomicName id="4C4671A3FFAA8059CEF2FB9541D4FB67" authority="Cai &amp; Shokita, 2006 a" authorityName="Cai &amp; Shokita" authorityYear="2006" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="buhi">
<emphasis id="B932D632FFAA8059CEF2FB9540D8FB67" italics="true" pageId="10" pageNumber="75">C. buhi</emphasis>
Cai &amp; Shokita, 2006a
</taxonomicName>
, from Luzon, by the form of pereiopods, but it differs from 
<taxonomicName id="4C4671A3FFAA8059CDD5FB554270FB07" authorityName="Cai &amp; Shokita" authorityYear="2006" box="[461,540,1211,1234]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="buhi">
<emphasis id="B932D632FFAA8059CDD5FB554270FB07" box="[461,540,1211,1234]" italics="true" pageId="10" pageNumber="75">C. buhi</emphasis>
</taxonomicName>
by its longer rostrum (reaches to or slightly beyond the anterior margin of the scaphocerite vs. reaches to the end of the second segment of the antennular peduncle or to the end of the antennular peduncle); the more ventral rostral teeth (13–15 vs. 3–7); the telson does not terminate in a projection vs. terminate in a projection; the bigger appendix interna on the male second pleopod (reaches near the end of the appendix masculina vs. reaches the base of the distal two-third length of the appendix masculina) and the distal spines on the telson (lateral spines much longer than the intermediate pairs of spiniform setae vs. lateral spines subequal to intermediate pairs) (
<figureCitation id="137D16A5FFAA8059CEB7F9F54292F9E7" box="[687,766,1563,1586]" captionStart="Fig" captionStartId="8.[132,167,2002,2023]" captionTargetBox="[180,1408,205,1957]" captionTargetId="figure-8@8.[162,1438,146,1985]" captionTargetPageId="8" captionText="Fig. 5. Caridina liaoi, new species. A: cephalothorax and cephalic appendages, lateral view; B, first pereiopod; C, second pereiopod; D, third pereiopod; E, dactylus of third pereiopod; F, fifth pereiopod; G, dactylus of fifth pereiopod; H, endopod of male first pleopod; I, appendix masculina and appendix interna of male second pleopod. Scale bars: A–D, F = 1 mm; E, G = 0.1 mm; H, I = 0.2 mm (male, cl 4.0 mm, paratype, ZRC, Bilar river)." figureDoi="http://doi.org/10.5281/zenodo.5341584" httpUri="https://zenodo.org/record/5341584/files/figure.png" pageId="10" pageNumber="75">Figs. 5</figureCitation>
, 
<figureCitation id="137D16A5FFAA8059CC9CF9D540F8F987" box="[132,148,1595,1618]" captionStart="Fig" captionStartId="9.[111,146,2001,2022]" captionTargetBox="[143,1381,178,1973]" captionTargetId="figure-92@9.[124,1432,167,1976]" captionTargetPageId="9" captionText="Fig. 6. Caridina liaoi, new species, A: cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scales: A, C–G, I = 1 mm; B, L = 0.2 mm; H, J = 0.1 mm; K = 0.5 mm. (ovig. females, cl 4.6 mm, paratype, ZRC, Bilar River)." figureDoi="http://doi.org/10.5281/zenodo.5341586" httpUri="https://zenodo.org/record/5341586/files/figure.png" pageId="10" pageNumber="75">6</figureCitation>
; cf. Cai &amp; Shokita, 2006a: 
<figureCitation id="137D16A5FFAA8059CDDBF9D54263F987" box="[451,527,1595,1618]" captionStart="Fig" captionStartId="5.[109,144,2000,2021]" captionTargetBox="[149,1407,199,1959]" captionTargetId="figure-9@5.[134,1433,177,1981]" captionTargetPageId="5" captionText="Fig. 3. Caridina lobocensis, new species: A, cephalothorax and cephalic appendages, lateral view; B, distal portion of telson; C, first pereiopod; D, second pereiopod; E, third pereiopod; F, dactylus of third pereiopod; G, fifth pereiopod; H, dactylus of fifth pereiopod; I, preanal carina; J, uropodal diaeresis. Scale bars: A = 2 mm, B, F, H, J = 0.2 mm; C–E, G = 1 mm; I = 0.5 mm (male, cl 5.7 mm, paratype, ZRC, Bohol Island, Philippines)." figureDoi="http://doi.org/10.5281/zenodo.5341580" httpUri="https://zenodo.org/record/5341580/files/figure.png" pageId="10" pageNumber="75">Figs. 3</figureCitation>
, 
<figureCitation id="137D16A5FFAA8059CE05F9D54241F987" box="[541,557,1595,1618]" captionStart="Fig" captionStartId="6.[132,167,2002,2023]" captionTargetBox="[208,1420,196,1964]" captionTargetId="figure-91@6.[182,1430,176,1978]" captionTargetPageId="6" captionText="Fig. 4. Caridina lobocensis, new species: A, telson; B, endopod of male first pleopod; C, appendix masculina and appendix interna of male second pleopod; D, first pereiopod; E, second pereiopod; F, third pereiopod; G, dactylus of third pereiopod; H, fifth pereiopod; I, dactylus of fifth pereiopod. Scale bars: A, D–F, H = 1 mm; B, C, G, I = 0.2 mm (A–C, male, cl 5.7 mm, paratype, ZRC, Bohol Island, Philippines; D–I, female, cl 6.9 mm, paratype, ZRC, Bohol Island, Philippines)." figureDoi="http://doi.org/10.5281/zenodo.5341582" httpUri="https://zenodo.org/record/5341582/files/figure.png" pageId="10" pageNumber="75">4</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>