treatments-xml/data/03/EF/20/03EF20375870FFBEED37FF1BFDC15FAA.xml

<document id="748D1ED5BBA44EBA83F3508AD1761DF2" ID-CLB-Dataset="8152" ID-DOI="10.2478/aemnp-2018-0035" ID-GBIF-Dataset="92fd172c-9d32-419f-9ea4-924f1cecb1b5" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488739" ID-ZooBank="B1327F4F-90E6-4BE1-83CA-0D5BE068E6C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612217885138" checkinUser="felipe" docAuthor="Huang, Meng-Chi, Li, Li-Zhen &amp; Yin, Zi-Wei" docDate="2018" docId="03EF20375870FFBEED37FF1BFDC15FAA" docLanguage="en" docName="ActaEntMusNatPra.58.2.457-478.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Pselaphodes paraculeus Huang &amp; Li &amp; Yin 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="470" masterDocId="FFD6584F587BFFB3EC22FFA6FF8B5E7F" masterDocTitle="Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species" masterLastPageNumber="478" masterPageNumber="457" pageNumber="468" updateTime="1698926012850" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="A71E56F7FC8AF8A4A35B5DC84CDD8456">Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species</mods:title>
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<mods:namePart id="6B76D17AE72B7F3713E07B8A97825B83">Huang, Meng-Chi</mods:namePart>
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<mods:namePart id="7B3DBEF032F851DA5B72893184CE38AA">Li, Li-Zhen</mods:namePart>
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<mods:namePart id="07037782849672761EFC6252FB3C6DAE">Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier id="9E173387AE8A3BACF25C908C49A5C7BE" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:title id="58A999D60EF71C78D6D2307AFBFF830D">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="EF1D433B7C9E4F139CB709F9D3E4AC67">2018</mods:date>
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<treatment id="03EF20375870FFBEED37FF1BFDC15FAA" ID-DOI="http://doi.org/10.5281/zenodo.4548437" ID-GBIF-Taxon="177423125" ID-Zenodo-Dep="4548437" LSID="urn:lsid:plazi:treatment:03EF20375870FFBEED37FF1BFDC15FAA" httpUri="http://treatment.plazi.org/id/03EF20375870FFBEED37FF1BFDC15FAA" lastPageId="13" lastPageNumber="470" pageId="11" pageNumber="468">
<subSubSection id="C35CC2AA5870FFB8ED37FF1BFDF15EAA" box="[277,634,189,213]" pageId="11" pageNumber="468" type="nomenclature">
<paragraph id="8BF991215870FFB8ED37FF1BFDF15EAA" blockId="11.[277,634,189,213]" box="[277,634,189,213]" pageId="11" pageNumber="468">
<heading id="D0B1264D5870FFB8ED37FF1BFDF15EAA" box="[277,634,189,213]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="468" reason="3">
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<taxonomicName id="4C46EAA25870FFB8ED37FF1BFD965EAA" ID-CoL="4NF87" authority="Huang &amp; Li &amp; Yin, 2018" authorityName="Huang &amp; Li &amp; Yin" authorityYear="2018" box="[277,541,189,213]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="468" phylum="Arthropoda" rank="species" species="paraculeus" status="sp. nov.">
<emphasis id="B9324D335870FFB8ED37FF1BFD965EAA" bold="true" box="[277,541,189,213]" italics="true" pageId="11" pageNumber="468">Pselaphodes paraculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0485870FFB8EE06FF18FDF15EAA" box="[548,634,190,213]" pageId="11" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
</paragraph>
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<paragraph id="8BF991215870FFB8EDABFF7AFD8D5E90" blockId="11.[151,761,220,540]" box="[393,518,220,239]" pageId="11" pageNumber="468">
(
<figureCitation id="137D8DA45870FFB8EDADFF7AFE6B5E90" box="[399,480,220,239]" captionStart="Fig" captionStartId="10.[151,182,2065,2084]" captionTargetBox="[155,1431,818,2037]" captionTargetId="figure-298@10.[151,1436,815,2051]" captionTargetPageId="10" captionText="Fig. 10. Dorsal habitus of Pselaphodes males. A – P. maolanensissp. nov; B – P. paraculeussp. nov. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488779" httpUri="https://zenodo.org/record/4488779/files/figure.png" pageId="11" pageNumber="468">Figs 10B</figureCitation>
, 
<figureCitation id="137D8DA45870FFB8EDCBFF7AFE745E90" box="[489,511,220,239]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="11" pageNumber="468">12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5870FFB8ECB5FF58FDFE5C63" pageId="11" pageNumber="468" type="materials_examined">
<paragraph id="8BF991215870FFB8ECB5FF58FDFE5C63" blockId="11.[151,761,220,540]" pageId="11" pageNumber="468">
<emphasis id="B9324D335870FFB8ECB5FF58FE985F6E" bold="true" box="[151,275,254,273]" pageId="11" pageNumber="468">Type material</emphasis>
(7 exs). 
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:♁, 
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<collectingCountry id="F351D1B15870FFB8EDE9FF58FD9F5F6E" box="[459,532,254,273]" name="China" pageId="11" pageNumber="468">CHINA</collectingCountry>
:
</emphasis>
‘ 
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: 
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, Wuming County (
<emphasis id="B9324D335870FFB8ECC7FEB2FEAE5F56" bold="true" box="[229,293,276,297]" pageId="11" pageNumber="468">ĒḆữ</emphasis>
), Daming Shan (
<emphasis id="B9324D335870FFB8ED9CFEB1FE755F54" bold="true" box="[446,510,279,299]" pageId="11" pageNumber="468">†fiƜ</emphasis>
), 
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, 
<geoCoordinate id="EE72F7E65870FFB8EEA6FEBFFD7F5F53" box="[644,756,281,300]" degrees="108" direction="east" minutes="26" orientation="longitude" pageId="11" pageNumber="468" precision="15" seconds="06" value="108.435">108°26′06″E</geoCoordinate>
, mixed leaf litter, sifted, 
<quantity id="4CBE3CC45870FFB8ED46FE95FE2C5F39" box="[356,423,307,326]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="468" unit="m" value="1500.0">1500 m</quantity>
, 
<date id="FFF8B7E15870FFB8ED92FE95FD985F39" box="[432,531,307,326]" pageId="11" pageNumber="468" value="2012-07-29">29.vii.2012</date>
, Jia-Yao Hu leg.’ (
<collectionCode id="ED5709E45870FFB8EE93FE95FD645F39" box="[689,751,307,326]" pageId="11" pageNumber="468">SNUC</collectionCode>
). 
<typeStatus id="54FD2F835870FFB8ECB5FEE8FF645F20" box="[151,239,334,353]" pageId="11" pageNumber="468" type="paratype">PARATYPES</typeStatus>
: 
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<collectingCountry id="F351D1B15870FFB8ECDEFEE8FECD5F1E" box="[252,326,334,353]" name="China" pageId="11" pageNumber="468">CHINA</collectingCountry>
:
</emphasis>
2 ♁♁, same label data as the 
<typeStatus id="54FD2F835870FFB8EE7BFEE8FD225F1E" box="[601,681,334,353]" pageId="11" pageNumber="468" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED5709E45870FFB8EE93FEE8FD645F1E" box="[689,751,334,353]" pageId="11" pageNumber="468">SNUC</collectionCode>
); 2 ♁♁, also from Daming Shan, except ‘ 
<geoCoordinate id="EE72F7E65870FFB8EE24FECFFDFE5F03" box="[518,629,361,380]" degrees="23" direction="north" minutes="30" orientation="latitude" pageId="11" pageNumber="468" precision="15" seconds="57" value="23.515833">23°30′57″N</geoCoordinate>
, 
<geoCoordinate id="EE72F7E65870FFB8EEA3FECFFD7F5F03" box="[641,756,361,380]" degrees="108" direction="east" minutes="26" orientation="longitude" pageId="11" pageNumber="468" precision="15" seconds="13" value="108.43694">108°26′13″E</geoCoordinate>
, 
<quantity id="4CBE3CC45870FFB8ECB5FE25FE9A5FE9" box="[151,273,387,406]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="11" pageNumber="468" unit="m" value="1200.0" valueMax="1250.0" valueMin="1150.0">1150–1250 m</quantity>
, 
<date id="FFF8B7E15870FFB8ED39FE25FEF55FE9" box="[283,382,387,406]" pageId="11" pageNumber="468" value="2012-07-31">31.vii.2012</date>
, Hu &amp; Song leg.’ (
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); 1 ♁, also from 
<collectingRegion id="49825FC35870FFB8ECB5FE38FF685FCE" box="[151,227,414,433]" country="China" name="Guangxi" pageId="11" pageNumber="468">Guangxi</collectingRegion>
, except ‘Guilin City (
<emphasis id="B9324D335870FFB8EDBBFE3CFE525FD0" bold="true" box="[409,473,410,431]" pageId="11" pageNumber="468">ẂŴffl</emphasis>
), Xingan County (
<emphasis id="B9324D335870FFB8EE5AFE3CFD335FD0" bold="true" box="[632,696,410,431]" pageId="11" pageNumber="468">
<collectionCode id="ED5709E45870FFB8EE5AFE3CFD335FD0" box="[632,696,410,431]" pageId="11" pageNumber="468">KÊR</collectionCode>
</emphasis>
), Maoer Shan (
<emphasis id="B9324D335870FFB8ECC4FE12FEAD5FB6" bold="true" box="[230,294,436,457]" pageId="11" pageNumber="468">
<collectionCode id="ED5709E45870FFB8ECC4FE12FEAD5FB6" box="[230,294,436,457]" country="Isle of Man" lsid="urn:lsid:biocol.org:col:13908" name="Ramsey Public Library" pageId="11" pageNumber="468" type="Herbarium">ŘĀƜ</collectionCode>
</emphasis>
), 
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, 
<geoCoordinate id="EE72F7E65870FFB8ED88FE1FFD925FB3" box="[426,537,441,460]" degrees="110" direction="east" minutes="29" orientation="longitude" pageId="11" pageNumber="468" precision="15" seconds="04" value="110.48444">110°29′04″E</geoCoordinate>
, 
<quantity id="4CBE3CC45870FFB8EE00FE1FFD0D5FB3" box="[546,646,441,460]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="11" pageNumber="468" unit="m" value="550.0" valueMax="650.0" valueMin="450.0">450–650 m</quantity>
, mixed litter, sifted, 
<date id="FFF8B7E15870FFB8ECEDFE75FEBA5F99" box="[207,305,467,486]" pageId="11" pageNumber="468" value="2012-07-25">25.vii.2012</date>
, Hu &amp; Song leg.’ (
<collectionCode id="ED5709E45870FFB8EDEBFE75FD8D5F99" box="[457,518,467,486]" pageId="11" pageNumber="468">SNUC</collectionCode>
); 1♁, ‘ 
<collectingCountry id="F351D1B15870FFB8EE1CFE75FDFC5F99" box="[574,631,467,486]" name="China" pageId="11" pageNumber="468">China</collectingCountry>
: 
<collectingRegion id="49825FC35870FFB8EE5CFE75FD425F99" box="[638,713,467,486]" country="China" name="Guizhou" pageId="11" pageNumber="468">Guizhou</collectingRegion>
, Libo Hsien, Maolan 
<taxonomicNameLabel id="A201F0485870FFB8ED3BFE48FECF5C7E" box="[281,324,494,513]" pageId="11" pageNumber="468" rank="subSpecies">
N. 
<collectionCode id="ED5709E45870FFB8ED0DFE48FECB5C7E" box="[303,320,494,513]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="468">R</collectionCode>
.
</taxonomicNameLabel>
, 
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, 
<geoCoordinate id="EE72F7E65870FFB8ED99FE48FDA05C7E" box="[443,555,494,513]" degrees="107" direction="east" minutes="54" orientation="longitude" pageId="11" pageNumber="468" precision="15" seconds="26" value="107.90723">107°54′26″E</geoCoordinate>
, mixed leaf litter, sifted, 
<quantity id="4CBE3CC45870FFB8ECB5FDAFFF715C63" box="[151,250,521,540]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="11" pageNumber="468" unit="m" value="800.0" valueMax="850.0" valueMin="750.0">750–850 m</quantity>
, 
<date id="FFF8B7E15870FFB8ED20FDAFFEEE5C63" box="[258,357,521,540]" pageId="11" pageNumber="468" value="2015-07-20">20.vii.2015</date>
, He, Hu &amp; Wang leg.’ (
<collectionCode id="ED5709E45870FFB8EE0CFDAFFDE75C63" box="[558,620,521,540]" pageId="11" pageNumber="468">SNUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5870FFB8EF1EFF18FAE05F2A" pageId="11" pageNumber="468" type="diagnosis">
<paragraph id="8BF991215870FFB8EF1EFF18FAE05F2A" blockId="11.[828,1443,190,533]" pageId="11" pageNumber="468">
<emphasis id="B9324D335870FFB8EF1EFF18FB895EAA" bold="true" box="[828,1026,190,213]" pageId="11" pageNumber="468">Diagnosis of male.</emphasis>
Length 
<quantity id="4CBE3CC45870FFB8E876FF18FB665EAA" box="[1108,1261,190,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.82" metricValueMin="2.52" pageId="11" pageNumber="468" unit="mm" value="2.67" valueMax="2.82" valueMin="2.52">2.52–2.82 mm</quantity>
; antennomere IX with disc-shaped process near apex; metaventral processes long, with blunt spine at apical third; protibia with distinct, blunt projection near middle; mesotibia with small apical spine; metatibia with preapical setose tuft and spine.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5870FFBEEF1EFEF8FD725F0A" lastPageId="13" lastPageNumber="470" pageId="11" pageNumber="468" type="description">
<paragraph id="8BF991215870FFBFEF1EFEF8FC705FEA" blockId="11.[828,1443,190,533]" lastBlockId="12.[828,1443,190,533]" lastPageId="12" lastPageNumber="469" pageId="11" pageNumber="468">
<emphasis id="B9324D335870FFB8EF1EFEF8FC435F0A" bold="true" box="[828,968,350,373]" pageId="11" pageNumber="468">Description.</emphasis>
<emphasis id="B9324D335870FFB8EFF0FEF8FB825F0A" box="[978,1033,350,373]" italics="true" pageId="11" pageNumber="468">Male</emphasis>
(
<figureCitation id="137D8DA45870FFB8E83EFEF8FB095F0A" box="[1052,1154,350,373]" captionStart="Fig" captionStartId="10.[151,182,2065,2084]" captionTargetBox="[155,1431,818,2037]" captionTargetId="figure-298@10.[151,1436,815,2051]" captionTargetPageId="10" captionText="Fig. 10. Dorsal habitus of Pselaphodes males. A – P. maolanensissp. nov; B – P. paraculeussp. nov. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488779" httpUri="https://zenodo.org/record/4488779/files/figure.png" pageId="11" pageNumber="468">Fig. 10B</figureCitation>
). Body yellowish-brown, BL 
<quantity id="4CBE3CC45870FFB8EF47FED8FB8F5FEA" box="[869,1028,382,405]" metricMagnitude="-3" metricUnit="m" metricValue="2.67" metricValueMax="2.82" metricValueMin="2.52" pageId="11" pageNumber="468" unit="mm" value="2.67" valueMax="2.82" valueMin="2.52">2.52–2.82 mm</quantity>
. Head wider than long, HL 
<quantity id="4CBE3CC45870FFB8E910FED8FCE35FCA" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="11" pageNumber="468" unit="mm" value="0.485" valueMax="0.52" valueMin="0.45">0.45–0.52 mm</quantity>
, HW 
<quantity id="4CBE3CC45870FFB8EF87FE38FBC85FCA" box="[933,1091,414,437]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" metricValueMax="5.5" metricValueMin="4.9" pageId="11" pageNumber="468" unit="mm" value="0.52" valueMax="0.55" valueMin="0.49">0.49–0.55 mm</quantity>
; each eye composed of about 25 facets; with well-developed ocular canthus.Antennomeres IX–XI (
<figureCitation id="137D8DA45870FFB8EFB2FE78FC795F8A" box="[912,1010,478,501]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="11" pageNumber="468">Fig. 12A</figureCitation>
) forming distinct club, antennomere IX with disc-shaped process at apex. Pronotum (
<figureCitation id="137D8DA45870FFB8E910FE58FA125C6A" box="[1330,1433,510,533]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="11" pageNumber="468">Fig. 12B</figureCitation>
) as long as wide, PL 
<quantity id="4CBE3CC45877FFBFED55FF18FD9C5EAA" box="[375,535,190,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.6" metricValueMin="4.699999999999999" pageId="12" pageNumber="469" unit="mm" value="0.515" valueMax="0.56" valueMin="0.47">0.47–0.56 mm</quantity>
, PW 
<quantity id="4CBE3CC45877FFBFEE76FF18FD785EAA" box="[596,755,190,213]" metricMagnitude="-4" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="12" pageNumber="469" unit="mm" value="0.505" valueMax="0.53" valueMin="0.48">0.48–0.53 mm</quantity>
, lateral margins narrowed anteriorly at apical third. Elytra wider than long, EL 
<quantity id="4CBE3CC45877FFBFED5AFF58FD9C5F6A" box="[376,535,254,277]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="8.1" metricValueMin="7.7" pageId="12" pageNumber="469" unit="mm" value="0.79" valueMax="0.81" valueMin="0.77">0.77–0.81 mm</quantity>
, EW 
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. Metaventral processes (
<figureCitation id="137D8DA45877FFBFEDB5FEB8FE725F4A" box="[407,505,286,309]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12C</figureCitation>
) long, with blunt spine near apical third, and two spines at apex. Protrochanter (
<figureCitation id="137D8DA45877FFBFECBDFEF8FE8B5F0A" box="[159,256,350,373]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12D</figureCitation>
) with acute ventral spine, profemur with large blunt ventral spine; protibia(
<figureCitation id="137D8DA45877FFBFEDE8FED8FDAE5FEA" box="[458,549,382,405]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12E</figureCitation>
) with distinct, blunt projection near middle; mesotrochanter (
<figureCitation id="137D8DA45877FFBFEE1AFE38FD1B5FCA" box="[568,656,414,437]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12F</figureCitation>
) with thin but distinct ventral spine, mesofemur broadly thickened ventrally; mesotibia (
<figureCitation id="137D8DA45877FFBFED53FE78FE465F8A" box="[369,461,478,501]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12G</figureCitation>
) with small apical spine; metatrochanter and metafemur (
<figureCitation id="137D8DA45877FFBFEDECFE58FDBB5C6A" box="[462,560,510,533]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12H</figureCitation>
) simple; metatibia (
<figureCitation id="137D8DA45877FFBFEF66FF18FC115EAA" box="[836,922,190,213]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12I</figureCitation>
) with preapical setose tuft and spine. Abdomen broad at base and narrowed apically, AL 
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, AW 
<quantity id="4CBE3CC45877FFBFEF4EFF58FB805F6A" box="[876,1035,254,277]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="9.8" metricValueMin="9.0" pageId="12" pageNumber="469" unit="mm" value="0.94" valueMax="0.98" valueMin="0.9">0.90–0.98 mm</quantity>
. Sternite IX (
<figureCitation id="137D8DA45877FFBFE8B8FF58FB795F6A" box="[1178,1266,254,277]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Fig. 12J</figureCitation>
) semi-membranous. Length of aedeagus 
<quantity id="4CBE3CC45877FFBFE87CFEB8FB485F4A" box="[1118,1219,286,309]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="469" unit="mm" value="0.5">0.50 mm</quantity>
; median lobe broad and nearly symmetric, parameres elongate, symmetric, endophallus (
<figureCitation id="137D8DA45877FFBFEFE8FEF8FB925F0A" box="[970,1049,350,373]" captionStart="Fig" captionStartId="12.[150,181,1981,2000]" captionTargetBox="[191,1402,564,1966]" captionTargetId="figure-188@12.[184,1404,559,1969]" captionTargetPageId="12" captionText="Fig. 12. Male diagnostic features of Pselaphodes paraculeus sp. nov. A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G." figureDoi="http://doi.org/10.5281/zenodo.4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469">Figs 12</figureCitation>
K–M) composed of several elongate or broad sclerites.
</paragraph>
<caption id="DF39C1A95870FFB8ECB5F87EFBF7565F" ID-DOI="http://doi.org/10.5281/zenodo.4488783" ID-Zenodo-Dep="4488783" httpUri="https://zenodo.org/record/4488783/files/figure.png" pageId="11" pageNumber="468" startId="11.[151,182,2008,2027]" targetBox="[176,1416,593,1983]" targetPageId="11">
<paragraph id="8BF991215870FFB8ECB5F87EFBF7565F" blockId="11.[151,1437,2008,2080]" pageId="11" pageNumber="468">
Fig. 11. Male diagnostic features of 
<taxonomicName id="4C46EAA25870FFB8EDE6F87EFD145994" authority="Huang &amp; Li &amp; Yin, 2018" authorityName="Huang &amp; Li &amp; Yin" authorityYear="2018" box="[452,671,2008,2027]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="468" phylum="Arthropoda" rank="species" species="maolanensis" status="sp. nov.">
<emphasis id="B9324D335870FFB8EDE6F87EFD145994" box="[452,671,2008,2027]" italics="true" pageId="11" pageNumber="468">Pselaphodes maolanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0485870FFB8EE81F87EFD6E5994" box="[675,741,2008,2027]" pageId="11" pageNumber="468" rank="species">sp. nov.</taxonomicNameLabel>
A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – apex of protibia; F – mesotrochanter and mesofemur; G – metatrochanter and metafemur; H – sternite IX; I–K – aedeagus, in dorsal (I), lateral (J), and ventral (K) view. Scale bar: 0.3 mm in A–B, D, F–G; 0.2 mm in C, I–K; 0.1 mm in E, H.
</paragraph>
</caption>
<paragraph id="8BF991215877FFBFEF78FE38FBA35FCA" blockId="12.[828,1443,190,533]" box="[858,1064,414,437]" pageId="12" pageNumber="469">
<emphasis id="B9324D335877FFBFEF78FE38FC215FCA" box="[858,938,414,437]" italics="true" pageId="12" pageNumber="469">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8BF991215877FFBEEF1EFE18FD725F0A" blockId="12.[828,1443,190,533]" lastBlockId="13.[151,761,190,469]" lastPageId="13" lastPageNumber="470" pageId="12" pageNumber="469">
<emphasis id="B9324D335877FFBFEF1EFE18FB975FAA" bold="true" box="[828,1052,446,469]" pageId="12" pageNumber="469">Comparative notes.</emphasis>
The new species is most similar to 
<taxonomicName id="4C46EAA25877FFBFEF1EFE78FC2D5F8A" authorityName="Yin, Li &amp; Zhao" authorityYear="2010" box="[828,934,478,501]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="469" phylum="Arthropoda" rank="species" species="aculeus">
<emphasis id="B9324D335877FFBFEF1EFE78FC2D5F8A" box="[828,934,478,501]" italics="true" pageId="12" pageNumber="469">P. aculeus</emphasis>
</taxonomicName>
in sharingthe presence of a disc-like process of antennomere IX, and a preapical spine of the metatibia. The latter species is widely distributed from eastern to southern 
<collectingCountry id="F351D1B15876FFBEECB5FF78FF525E8A" box="[151,217,222,245]" name="China" pageId="13" pageNumber="470">China</collectingCountry>
, with considerable variations of male external features (although the combination is always stable). However, the aedeagal median lobe of all populations of 
<taxonomicName id="4C46EAA25876FFBEEEAEFEB8FD7C5F4A" authorityName="Yin, Li &amp; Zhao" authorityYear="2010" box="[652,759,286,309]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="470" phylum="Arthropoda" rank="species" species="aculeus">
<emphasis id="B9324D335876FFBEEEAEFEB8FD7C5F4A" box="[652,759,286,309]" italics="true" pageId="13" pageNumber="470">P. aculeus</emphasis>
</taxonomicName>
is deeply and broadly concave in the middle of the apex, while that of the new species is simply rounded at the apex.
</paragraph>
</subSubSection>
<caption id="DF39C1A95877FFBFECB4F81BFEFD565F" ID-DOI="http://doi.org/10.5281/zenodo.4488787" ID-Zenodo-Dep="4488787" httpUri="https://zenodo.org/record/4488787/files/figure.png" pageId="12" pageNumber="469" startId="12.[150,181,1981,2000]" targetBox="[191,1402,564,1966]" targetPageId="12">
<paragraph id="8BF991215877FFBFECB4F81BFEFD565F" blockId="12.[150,1437,1981,2080]" pageId="12" pageNumber="469">
Fig. 12. Male diagnostic features of 
<taxonomicName id="4C46EAA25877FFBFEDF3F81BFD2959AF" authority="Huang &amp; Li &amp; Yin, 2018" authorityName="Huang &amp; Li &amp; Yin" authorityYear="2018" box="[465,674,1981,2000]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="469" phylum="Arthropoda" rank="species" species="paraculeus" status="sp. nov.">
<emphasis id="B9324D335877FFBFEDF3F81BFD2959AF" box="[465,674,1981,2000]" italics="true" pageId="12" pageNumber="469">Pselaphodes paraculeus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201F0485877FFBFEE8AF81BFD6759AF" box="[680,748,1981,2000]" pageId="12" pageNumber="469" rank="species">sp. nov.</taxonomicNameLabel>
A – antennal club; B – pronotum; C – metaventral process, in lateral view; D – protrochanter and profemur; E – protibia; F – mesotrochanter and mesofemur; G – apex of mesotibia; H – metatrochanter and metafemur; I – apex of metatibia; J – sternite IX; K–M – aedeagus, in dorsal (K), lateral (L), and ventral (M) view. Scale bar: 0.3 mm in A–B, D–F, H; 0.2 mm in C, K–M; 0.1 mm in I, J; 0.05 mm in G.
</paragraph>
</caption>
<subSubSection id="C35CC2AA5876FFBEECB5FED8FDD45FCA" pageId="13" pageNumber="470" type="etymology">
<paragraph id="8BF991215876FFBEECB5FED8FDD45FCA" blockId="13.[151,761,190,469]" pageId="13" pageNumber="470">
<emphasis id="B9324D335876FFBEECB5FED8FE9F5FEA" bold="true" box="[151,276,382,405]" pageId="13" pageNumber="470">Etymology.</emphasis>
The species name indicates a close relationship of the new species to 
<taxonomicName id="4C46EAA25876FFBEED5FFE38FE625FCA" authorityName="Yin, Li &amp; Zhao" authorityYear="2010" box="[381,489,414,437]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="470" phylum="Arthropoda" rank="species" species="aculeus">
<emphasis id="B9324D335876FFBEED5FFE38FE625FCA" box="[381,489,414,437]" italics="true" pageId="13" pageNumber="470">P. aculeus</emphasis>
</taxonomicName>
; adjective.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5876FFBEECB5FE18FDC15FAA" box="[151,586,446,469]" pageId="13" pageNumber="470" type="distribution">
<paragraph id="8BF991215876FFBEECB5FE18FDC15FAA" blockId="13.[151,761,190,469]" box="[151,586,446,469]" pageId="13" pageNumber="470">
<emphasis id="B9324D335876FFBEECB5FE18FEA15FAA" bold="true" box="[151,298,446,469]" pageId="13" pageNumber="470">Distribution.</emphasis>
<collectingCountry id="F351D1B15876FFBEED12FE18FEFF5FAA" box="[304,372,446,469]" name="China" pageId="13" pageNumber="470">China</collectingCountry>
: 
<collectingRegion id="49825FC35876FFBEED5CFE18FE565FAA" box="[382,477,446,469]" country="China" name="Guangxi" pageId="13" pageNumber="470">Guangxi</collectingRegion>
, 
<collectingRegion id="49825FC35876FFBEEDC5FE18FDCD5FAA" box="[487,582,446,469]" country="China" name="Guizhou" pageId="13" pageNumber="470">Guizhou</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>