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<document id="60CCF121F82B559C01B8AAD530D4102C" ID-DOI="10.11646/zootaxa.5353.4.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="8430765" ID-ZooBank="DBCD0660-BFF4-4507-8BC7-F6339450B011" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697020339292" checkinUser="plazi" docAuthor="Pati, Sameer Kumar, Thackeray, Tejas &amp; Pawar, Swapnil" docDate="2023" docId="03CEB61969762B2EFF1981F2FE4B8BC5" docLanguage="en" docName="zootaxa.5353.4.4.pdf" docOrigin="Zootaxa 5353 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5353.4.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.14:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="14" docTitle="Ghatiana sanguinolenta Pati &amp; Thackeray &amp; Pawar 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="377" masterDocId="FFF7CE6169742B2BFF8E8045FFD28C30" masterDocTitle="Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India" masterLastPageNumber="378" masterPageNumber="372" pageNumber="374" updateTime="1697031112029" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="25427FB271CB264C923325F2F58FE6B7">Ghatiana sanguinolenta, a new species of freshwater crab (Brachyura: Gecarcinucidae) from the Central Western Ghats of India</mods:title>
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<mods:namePart id="5C6A5A862697B4BCD054111E5CF6626B">Pati, Sameer Kumar</mods:namePart>
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<mods:namePart id="90A349C76FED59DF629026ADE1246012">Thackeray, Tejas</mods:namePart>
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<mods:nameIdentifier id="3577C96E8BE4AC88822E2695C8891CB9" type="email">tejasthackeray@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FDD56757B41D66C76C305AB02ACF70A0">Pawar, Swapnil</mods:namePart>
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<mods:nameIdentifier id="77F3206175C166C5447DE1B10CD0D6B8" type="email">tejasthackeray@gmail.com</mods:nameIdentifier>
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<treatment id="03CEB61969762B2EFF1981F2FE4B8BC5" LSID="urn:lsid:plazi:treatment:03CEB61969762B2EFF1981F2FE4B8BC5" httpUri="http://treatment.plazi.org/id/03CEB61969762B2EFF1981F2FE4B8BC5" lastPageId="5" lastPageNumber="377" pageId="2" pageNumber="374">
<subSubSection id="C37D548469762B29FF1981F2FDF08DE1" box="[151,546,439,466]" pageId="2" pageNumber="374" type="nomenclature">
<paragraph id="8BD8070F69762B29FF1981F2FDF08DE1" blockId="2.[151,546,439,537]" box="[151,546,439,466]" pageId="2" pageNumber="374">
<heading id="D090B06369762B29FF1981F2FDF08DE1" bold="true" box="[151,546,439,466]" fontSize="11" level="1" pageId="2" pageNumber="374" reason="1">
<emphasis id="B913DB1D69762B29FF1981F2FDF08DE1" bold="true" box="[151,546,439,466]" pageId="2" pageNumber="374">
<taxonomicName id="4C677C8C69762B29FF1981F2FE6C8DE2" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[151,446,439,466]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69762B29FF1981F2FE6C8DE2" bold="true" box="[151,446,439,466]" italics="true" pageId="2" pageNumber="374">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220666669762B29FE4B81F2FDF08DE1" box="[453,546,439,465]" pageId="2" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C37D548469762B29FF19819EFE448DC5" box="[151,406,475,501]" pageId="2" pageNumber="374" type="vernacular_names">
<paragraph id="8BD8070F69762B29FF19819EFE448DC5" blockId="2.[151,546,439,537]" box="[151,406,475,501]" pageId="2" pageNumber="374">[Blood-red Ghat Crab]</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B29FF1981BAFEFC8E29" box="[151,302,511,537]" pageId="2" pageNumber="374" type="description">
<paragraph id="8BD8070F69762B29FF1981BAFEFC8E29" blockId="2.[151,546,439,537]" box="[151,302,511,537]" pageId="2" pageNumber="374">
(
<figureCitation id="135C1B8A69762B29FF1181BAFF3D8E29" box="[159,239,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2</figureCitation>
,
<figureCitation id="135C1B8A69762B29FF7581BAFED88E29" box="[251,266,511,537]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3</figureCitation>
,
<figureCitation id="135C1B8A69762B29FE9981BAFEF48E29" box="[279,294,511,537]" captionStart="FIGURE 4" captionStartId="5.[151,250,1060,1084]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure-30@5.[151,1436,181,1037]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana sanguinolenta sp. nov., colour in life, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354." figureDoi="http://doi.org/10.5281/zenodo.8430773" httpUri="https://zenodo.org/record/8430773/files/figure.png" pageId="2" pageNumber="374">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B29FF198202FE208F25" pageId="2" pageNumber="374" type="materials_examined">
<paragraph id="8BD8070F69762B29FF198202FC688E99" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF198202FE918E51" bold="true" box="[151,323,583,609]" pageId="2" pageNumber="374">Type material.</emphasis>
<materialsCitation id="3B0F0D5269762B29FEC68202FC688E99" collectingDate="2021-09-01" collectionCode="CW" collectorName="Swapnil Pawar" country="India" county="Uttara Kannada district" elevation="413" latitude="14.527" location="Balekoppa" longLatPrecision="76" longitude="74.596" municipality="Devimane on Sirsi-Kumta" pageId="2" pageNumber="374" specimenCode="ZSI-WRC C.2353" specimenCount="1" stateProvince="Karnataka state" typeStatus="holotype">
<typeStatus id="54DCB9AD69762B29FEC68202FE328E51" box="[328,480,583,609]" pageId="2" pageNumber="374">HOLOTYPE</typeStatus>
:
<specimenCode id="DBC1AF7469762B29FE678202FD648E52" box="[489,694,583,610]" pageId="2" pageNumber="374">ZSI-WRC C.2353</specimenCode>
, adult
<specimenCount id="9D61CC8669762B29FD738202FCE18E51" box="[765,819,583,609]" pageId="2" pageNumber="374" type="male">male</specimenCount>
(CW
<quantity id="4C9FAAEA69762B29FCFA8202FC348E52" box="[884,998,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="2.448" pageId="2" pageNumber="374" unit="mm" value="24.48">24.48 mm</quantity>
, CL
<quantity id="4C9FAAEA69762B29FB968202FB588E52" box="[1048,1162,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.538" pageId="2" pageNumber="374" unit="mm" value="15.38">15.38 mm</quantity>
, CH
<quantity id="4C9FAAEA69762B29FB4E8202FAE18E52" box="[1216,1331,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.028" pageId="2" pageNumber="374" unit="mm" value="10.28">10.28 mm</quantity>
, FW
<quantity id="4C9FAAEA69762B29FAE58202FF158EB5" metricMagnitude="-3" metricUnit="m" metricValue="9.04" pageId="2" pageNumber="374" unit="mm" value="9.04">9.04 mm</quantity>
),
<location id="8EB851D469762B29FF55822EFE848EB5" LSID="urn:lsid:plazi:treatment:03CEB61969762B2EFF1981F2FE4B8BC5:8EB851D469762B29FF55822EFE848EB5" box="[219,342,619,645]" country="India" county="Uttara Kannada district" latitude="14.527" longLatPrecision="76" longitude="74.596" municipality="Devimane on Sirsi-Kumta" name="Balekoppa" pageId="2" pageNumber="374" stateProvince="Karnataka state">Balekoppa</location>
, near
<collectingMunicipality id="6BBC9D7569762B29FE14822EFD118EB5" box="[410,707,619,646]" pageId="2" pageNumber="374">Devimane on Sirsi-Kumta</collectingMunicipality>
road,
<collectingCounty id="62B97F8369762B29FC828229FBC58EB5" box="[780,1047,619,646]" pageId="2" pageNumber="374">Uttara Kannada district</collectingCounty>
,
<collectingRegion id="49A3C9ED69762B29FBAC822EFB068EB6" box="[1058,1236,619,646]" country="India" name="Karnataka" pageId="2" pageNumber="374">Karnataka state</collectingRegion>
,
<collectingCountry id="F370479F69762B29FB6E822EFACE8EB5" box="[1248,1308,619,645]" name="India" pageId="2" pageNumber="374">India</collectingCountry>
,
<geoCoordinate id="EE5361C869762B29FAA7822EFA4A8EB6" box="[1321,1432,619,646]" degrees="14.527" direction="north" orientation="latitude" pageId="2" pageNumber="374" precision="55" value="14.527">14.527°N</geoCoordinate>
,
<geoCoordinate id="EE5361C869762B29FF1982CAFED68E9A" box="[151,260,655,682]" degrees="74.596" direction="east" orientation="longitude" pageId="2" pageNumber="374" precision="55" value="74.596">74.596°E</geoCoordinate>
, altitude
<elevation id="004AE03C69762B29FEE782CAFE3F8E99" box="[361,493,655,682]" metricMagnitude="2" metricUnit="m" metricValue="4.13" pageId="2" pageNumber="374" unit="m" value="413.0">
<quantity id="4C9FAAEA69762B29FEE782CAFE628E9A" box="[361,432,655,682]" metricMagnitude="2" metricUnit="m" metricValue="4.13" pageId="2" pageNumber="374" unit="m" value="413.0">413 m</quantity>
a.s.l.
</elevation>
, coll.
<collectorName id="269262D969762B29FDBF82CAFD0E8E9A" box="[561,732,655,682]" pageId="2" pageNumber="374">Swapnil Pawar</collectorName>
,
<date id="FFD921CF69762B29FD6982CAFC648E99" box="[743,950,655,681]" pageId="2" pageNumber="374" value="2021-09-01">
<collectingDate id="EF9DD82769762B29FD6982CAFC648E99" box="[743,950,655,681]" pageId="2" pageNumber="374" value="2021-09-01">1 September 2021</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD8070F69762B29FF4982F6FE208F25" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<materialsCitation id="3B0F0D5269762B29FF4982F6FE3C8F25" collectingDate="2021-09-01" collectionCode="CW" collectorName="Swapnil Pawar" country="Switzerland" county="Uttara Kannada district" elevation="413" latitude="14.527" location="Balekoppa" longLatPrecision="76" longitude="74.596" municipality="Devimane on Sirsi-Kumta" pageId="2" pageNumber="374" specimenCode="ZSI-WRC C.2354" specimenCount="3" specimenCount-adult="2" stateProvince="Karnataka" typeStatus="paratype">
<typeStatus id="54DCB9AD69762B29FF4982F6FEB48EFD" box="[199,358,691,717]" pageId="2" pageNumber="374">PARATYPES</typeStatus>
:
<specimenCode id="DBC1AF7469762B29FEFC82F6FD938EFE" box="[370,577,691,718]" pageId="2" pageNumber="374">ZSI-WRC C.2354</specimenCode>
,
<specimenCount id="9D61CC8669762B29FDC382F6FD118EFD" box="[589,707,691,717]" pageId="2" pageNumber="374" type="adult">adult male</specimenCount>
(CW
<quantity id="4C9FAAEA69762B29FC8482F6FCAD8EFE" box="[778,895,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="2.853" pageId="2" pageNumber="374" unit="mm" value="28.53">28.53 mm</quantity>
, CL
<quantity id="4C9FAAEA69762B29FC3882F6FBF98EFE" box="[950,1067,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" pageId="2" pageNumber="374" unit="mm" value="17.85">17.85 mm</quantity>
, CH
<quantity id="4C9FAAEA69762B29FBE882F6FB098EFE" box="[1126,1243,691,718]" metricMagnitude="-2" metricUnit="m" metricValue="1.282" pageId="2" pageNumber="374" unit="mm" value="12.82">12.82 mm</quantity>
, FW
<quantity id="4C9FAAEA69762B29FA9782F6FA428EFD" box="[1305,1424,691,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="374" unit="mm" value="10.4">10.40 mm</quantity>
),
<specimenCount id="9D61CC8669762B29FF198292FE958EC1" box="[151,327,727,753]" pageId="2" pageNumber="374">2 adult females</specimenCount>
(CW
<quantity id="4C9FAAEA69762B29FE1E8292FD868EC2" box="[400,596,727,754]" metricMagnitude="-2" metricUnit="m" metricValue="2.742" metricValueMax="2.8449999999999998" metricValueMin="2.639" pageId="2" pageNumber="374" unit="mm" value="27.42" valueMax="28.45" valueMin="26.39">26.3928.45 mm</quantity>
, CL
<quantity id="4C9FAAEA69762B29FD018292FC808EC2" box="[655,850,727,754]" metricMagnitude="-2" metricUnit="m" metricValue="1.755" metricValueMax="1.826" metricValueMin="1.684" pageId="2" pageNumber="374" unit="mm" value="17.55" valueMax="18.26" valueMin="16.84">16.8418.26 mm</quantity>
, CH
<quantity id="4C9FAAEA69762B29FC1F8292FB868EC2" box="[913,1108,727,754]" metricMagnitude="-2" metricUnit="m" metricValue="1.2545" metricValueMax="1.271" metricValueMin="1.238" pageId="2" pageNumber="374" unit="mm" value="12.545" valueMax="12.71" valueMin="12.38">12.3812.71 mm</quantity>
, FW
<quantity id="4C9FAAEA69762B29FB188292FA9F8EC1" box="[1174,1357,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="1.009" metricValueMax="1.039" metricValueMin="0.9789999999999999" pageId="2" pageNumber="374" unit="mm" value="10.09" valueMax="10.39" valueMin="9.79">9.7910.39 mm</quantity>
), same collection data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B29FF49835AFA518965" pageId="2" pageNumber="374" type="diagnosis">
<paragraph id="8BD8070F69762B29FF49835AFEB38901" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF49835AFE908F09" bold="true" box="[199,322,799,825]" pageId="2" pageNumber="374">Diagnosis.</emphasis>
Carapace in adult proportionately broad (CW/CL = 1.6) (
<figureCitation id="135C1B8A69762B29FC54835AFB928F09" box="[986,1088,799,825]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A</figureCitation>
,
<figureCitation id="135C1B8A69762B29FBC0835AFB478F09" box="[1102,1173,799,825]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
), strongly arched (
<collectingCountry id="F370479F69762B29FAE2835AFA468F09" box="[1388,1428,799,825]" name="Switzerland" pageId="2" pageNumber="374">CH</collectingCountry>
/ CL = 0.7) (
<figureCitation id="135C1B8A69762B29FE918306FEA98F6D" box="[287,379,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2B</figureCitation>
); lateral margins strongly convex (
<figureCitation id="135C1B8A69762B29FC9A8306FCA98F6D" box="[788,891,835,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A</figureCitation>
,
<figureCitation id="135C1B8A69762B29FC058306FC078F6D" box="[907,981,835,861]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
); anterolateral margins relatively short, subcristate (
<figureCitation id="135C1B8A69762B29FE908322FE508FB1" box="[286,386,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A</figureCitation>
,
<figureCitation id="135C1B8A69762B29FE008322FE078FB1" box="[398,469,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
); epibranchial tooth visible as weak notch (
<figureCitation id="135C1B8A69762B29FC338322FBF38FB1" box="[957,1057,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A</figureCitation>
,
<figureCitation id="135C1B8A69762B29FBA38322FBA18FB1" box="[1069,1139,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
); branchial regions rugose (
<figureCitation id="135C1B8A69762B29FF1183CEFED68F95" box="[159,260,907,933]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A</figureCitation>
,
<figureCitation id="135C1B8A69762B29FE9F83CEFE8B8F95" box="[273,345,907,933]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
); frontal margin relatively close to anterior margin of epistome, almost hiding antennular fossae (
<figureCitation id="135C1B8A69762B29FF1183EAFF2A8FF9" box="[159,248,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2B</figureCitation>
). Eyes relatively small as compared to orbital space; each eye with relatively slenderer eyestalk (
<figureCitation id="135C1B8A69762B29FAB983EAFA5D8FF9" box="[1335,1423,943,969]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2B</figureCitation>
). Maxilliped 1, 2 each with short flagellum on exopod (
<figureCitation id="135C1B8A69762B29FD738396FC8A8FDD" box="[765,856,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2D</figureCitation>
); maxilliped 3 exopod without flagellum (
<figureCitation id="135C1B8A69762B29FAB98396FA428FDD" box="[1335,1424,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2E</figureCitation>
). Major chela in adult male with pointed fingertips; palm in adult male relatively stout; ventral margin of fixed finger and distal half of palm in adult male strongly concave (
<figureCitation id="135C1B8A69762B29FC85845EFCB08805" box="[779,866,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2F</figureCitation>
). Ambulatory legs relatively long (P3 length/CL = ca. 2.5) (
<figureCitation id="135C1B8A69762B29FF76847AFEA88869" box="[248,378,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2A, C</figureCitation>
,
<figureCitation id="135C1B8A69762B29FE0B847AFE188869" box="[389,458,1087,1113]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3A, C</figureCitation>
). Male pleonal somite 6 subquadrate (
<figureCitation id="135C1B8A69762B29FCE1847AFC208869" box="[879,1010,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2C, G</figureCitation>
,
<figureCitation id="135C1B8A69762B29FC72847AFBCD8869" box="[1020,1055,1087,1113]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3B</figureCitation>
). Male telson elongated (
<figureCitation id="135C1B8A69762B29FAB8847AFF7C884D" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Figs. 2C, G</figureCitation>
,
<figureCitation id="135C1B8A69762B29FF358426FF0C884D" box="[187,222,1123,1149]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">3B</figureCitation>
). G1 relatively slender, medially gently curved outwards; ultimate article relatively slender, distally gently curved outwards, relatively long, ca. 0.4 times length of penultimate article; penultimate article relatively slender (
<figureCitation id="135C1B8A69762B29FF1184EEFEC388F5" box="[159,273,1195,1221]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2H, I</figureCitation>
). G2 very short, with very short penultimate article (
<figureCitation id="135C1B8A69762B29FCE984EEFC6B88F5" box="[871,953,1195,1221]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2J</figureCitation>
). Female pleon and telson in adult broadly subtriangular, with lateral margins of telson strongly concave (
<figureCitation id="135C1B8A69762B29FCC4848AFC7188D9" box="[842,931,1231,1257]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">Fig. 3D</figureCitation>
). Vulvae in adult relatively closely positioned (VD/SW = ca. 0.25), each subovate in shape, relatively large, occupying ca. 0.5 times length of s6, positioned close to s5/s6 (
<figureCitation id="135C1B8A69762B29FF738552FE868901" box="[253,340,1303,1329]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">Fig. 3E</figureCitation>
).
</paragraph>
<paragraph id="8BD8070F69762B29FF49857EFA518965" blockId="2.[151,1437,583,1978]" box="[199,1411,1339,1366]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF49857EFEBF8965" bold="true" box="[199,365,1339,1365]" pageId="2" pageNumber="374">Colour in life.</emphasis>
Crabs have dark blood-red coloured carapace and pereiopods, with blanched fingers (
<figureCitation id="135C1B8A69762B29FABE857EFAA48965" box="[1328,1398,1339,1365]" captionStart="FIGURE 4" captionStartId="5.[151,250,1060,1084]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure-30@5.[151,1436,181,1037]" captionTargetPageId="5" captionText="FIGURE 4. Ghatiana sanguinolenta sp. nov., colour in life, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354." figureDoi="http://doi.org/10.5281/zenodo.8430773" httpUri="https://zenodo.org/record/8430773/files/figure.png" pageId="2" pageNumber="374">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B29FF49851AFBC289F1" pageId="2" pageNumber="374" type="etymology">
<paragraph id="8BD8070F69762B29FF49851AFD1789AD" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF49851AFE9C8949" bold="true" box="[199,334,1375,1401]" pageId="2" pageNumber="374">Etymology.</emphasis>
The specific epithet, sanguinolenta meaning blood-red, is a Latin adjective in the nominative singular, which alludes to the crab's colour in life.
</paragraph>
<paragraph id="8BD8070F69762B29FF4985E2FBC289F1" blockId="2.[151,1437,583,1978]" box="[199,1040,1447,1474]" pageId="2" pageNumber="374">The proposed common name of the new species is “Blood-red Ghat Crab”.</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B29FF49858EFC6D8A1E" pageId="2" pageNumber="374" type="biology_ecology">
<paragraph id="8BD8070F69762B29FF49858EFC6D8A1E" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF49858EFE4289D5" bold="true" box="[199,400,1483,1509]" pageId="2" pageNumber="374">Ecological notes.</emphasis>
<taxonomicName id="4C677C8C69762B29FE16858EFD7B89D5" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[408,681,1483,1509]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69762B29FE16858EFD7B89D5" box="[408,681,1483,1509]" italics="true" pageId="2" pageNumber="374">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69762B29FD3F858EFCDE89D5" bold="true" box="[689,780,1483,1509]" pageId="2" pageNumber="374">
<taxonomicNameLabel id="A220666669762B29FD3F858EFCDE89D5" box="[689,780,1483,1509]" pageId="2" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dwells in the forests of the Central Western Ghats. These crabs live in tree holes during the evening and night-time. Juveniles were found living with adults in some tree holes. Individuals can also forage on open ground in non-forested areas.
</paragraph>
</subSubSection>
<subSubSection id="C37D548469762B2EFF498672FBD78BBA" lastPageId="5" lastPageNumber="377" pageId="2" pageNumber="374" type="discussion">
<paragraph id="8BD8070F69762B29FF498672FAED8A8D" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
<emphasis id="B913DB1D69762B29FF498672FEE98A61" bold="true" box="[199,315,1591,1617]" pageId="2" pageNumber="374">Remarks.</emphasis>
In diagnostic features, the
<typeStatus id="54DCB9AD69762B29FD08867DFD348A62" box="[646,742,1592,1618]" pageId="2" pageNumber="374">paratype</typeStatus>
male (ZSI-WRC C.2354) of
<taxonomicName id="4C677C8C69762B29FBC5867DFAC08A61" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[1099,1298,1591,1618]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69762B29FBC5867DFAC08A61" box="[1099,1298,1591,1618]" italics="true" pageId="2" pageNumber="374">G. sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69762B29FA908672FAAF8A61" bold="true" box="[1310,1405,1591,1617]" pageId="2" pageNumber="374">
<taxonomicNameLabel id="A220666669762B29FA908672FAAF8A61" box="[1310,1405,1591,1617]" pageId="2" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is indistinguishable from the
<typeStatus id="54DCB9AD69762B29FE48861EFDFE8A45" box="[454,556,1627,1653]" pageId="2" pageNumber="374">holotype</typeStatus>
. The male
<typeStatus id="54DCB9AD69762B29FD268619FCDA8A46" box="[680,776,1628,1654]" pageId="2" pageNumber="374">paratype</typeStatus>
differs from the
<typeStatus id="54DCB9AD69762B29FC46861EFBF88A45" box="[968,1066,1627,1653]" pageId="2" pageNumber="374">holotype</typeStatus>
only by the shape of the pleonal somite 6, which is slightly longer than broad (
<figureCitation id="135C1B8A69762B29FD18863AFD398AA9" box="[662,747,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="2" pageNumber="374">Fig. 3B</figureCitation>
) (vs. slightly broader than long in the
<typeStatus id="54DCB9AD69762B29FB1B863AFB298AA9" box="[1173,1275,1663,1689]" pageId="2" pageNumber="374">holotype</typeStatus>
;
<figureCitation id="135C1B8A69762B29FA8B863AFA8C8AA9" box="[1285,1374,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2G</figureCitation>
). The female
<typeStatus id="54DCB9AD69762B29FF6486E1FE878A8E" box="[234,341,1700,1726]" pageId="2" pageNumber="374">paratypes</typeStatus>
(ZSI-WRC C.2354) are consistent with the
<typeStatus id="54DCB9AD69762B29FCCA86E6FC748A8D" box="[836,934,1699,1725]" pageId="2" pageNumber="374">holotype</typeStatus>
in non-sexual diagnostic characters.
</paragraph>
<paragraph id="8BD8070F69762B29FF498682FC408B89" blockId="2.[151,1437,583,1978]" pageId="2" pageNumber="374">
The dark blood-red colour-in-life of
<taxonomicName id="4C677C8C69762B29FD0C8682FC4A8AD1" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[642,920,1735,1761]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69762B29FD0C8682FC4A8AD1" box="[642,920,1735,1761]" italics="true" pageId="2" pageNumber="374">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69762B29FC288682FBD48AD1" bold="true" box="[934,1030,1735,1761]" pageId="2" pageNumber="374">
<taxonomicNameLabel id="A220666669762B29FC288682FBD48AD1" box="[934,1030,1735,1761]" pageId="2" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unique in the genus, which is convenient for identifying the species in field. The outwardly curved ultimate article of the G1 is characteristic of
<taxonomicName id="4C677C8C69762B29FA0F86A9FEE48B19" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69762B29FA0F86A9FEE48B19" italics="true" pageId="2" pageNumber="374">G. sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69762B29FEB3874AFE448B19" bold="true" box="[317,406,1807,1833]" pageId="2" pageNumber="374">
<taxonomicNameLabel id="A220666669762B29FEB3874AFE448B19" box="[317,406,1807,1833]" pageId="2" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="135C1B8A69762B29FE2B874AFE2D8B19" box="[421,511,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2H</figureCitation>
), whereas the remaining species of
<taxonomicName id="4C677C8C69762B29FC03874AFC258B19" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[909,1015,1807,1833]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="genus">
<emphasis id="B913DB1D69762B29FC03874AFC258B19" box="[909,1015,1807,1833]" italics="true" pageId="2" pageNumber="374">Ghatiana</emphasis>
</taxonomicName>
possess a straight or inwardly curved G1 ultimate article (see
<bibRefCitation id="EFF67AFE69762B29FE2D8776FD788B7D" author="Pati, S. K. &amp; Thackeray, T." box="[419,682,1843,1869]" pageId="2" pageNumber="374" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: figs. 3D, 4D, 5D, 6I, 7D, 8I, 9I, 10D;
<bibRefCitation id="EFF67AFE69762B29FBD48776FAB38B7D" author="Pati, S. K. &amp; Thackeray, T." box="[1114,1377,1843,1869]" pageId="2" pageNumber="374" pagination="627 - 647" refId="ref3583" refString="Pati, S. K. &amp; Thackeray, T. (2021) Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014 and Sahyadriana Pati &amp; Thackeray, 2018 from India (Decapoda: Brachyura: Gecarcinucidae). Zoosystema, 43, 627 - 647. https: // doi. org / 10.5252 / zoosystema 2021 v 43 a 26" type="journal article" year="2021">Pati &amp; Thackeray 2021</bibRefCitation>
: figs. 2D, 5D;
<bibRefCitation id="EFF67AFE69762B29FF778712FE618B41" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[249,435,1879,1906]" pageId="2" pageNumber="374" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69762B29FEA5871DFE8E8B41" box="[299,348,1879,1905]" italics="true" pageId="2" pageNumber="374">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 4D). The new species otherwise most closely resembles
<taxonomicName id="4C677C8C69762B29FBED871DFAD38B41" baseAuthorityName="Klaus, Fernandez &amp; Yeo" baseAuthorityYear="2014" box="[1123,1281,1879,1906]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B913DB1D69762B29FBED871DFAD38B41" box="[1123,1281,1879,1906]" italics="true" pageId="2" pageNumber="374">G. basalticola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C677C8C69762B29FA81871DFA448B41" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[1295,1430,1879,1906]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69762B29FA81871DFA448B41" box="[1295,1430,1879,1906]" italics="true" pageId="2" pageNumber="374">G. dvivarna</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C677C8C69762B29FF488739FD828BA5" authority="Pati &amp; Thackeray, 2018" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[198,592,1915,1942]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="374" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69762B29FF488739FEED8BA5" box="[198,319,1915,1942]" italics="true" pageId="2" pageNumber="374">G. pulchra</emphasis>
<bibRefCitation id="EFF67AFE69762B29FEC8873EFD828BA5" author="Pati, S. K. &amp; Thackeray, T." box="[326,592,1915,1941]" pageId="2" pageNumber="374" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray, 2018</bibRefCitation>
</taxonomicName>
, in having a proportionately broader carapace (CW/CL = 1.61.9) (
<figureCitation id="135C1B8A69762B29FAB0873EFA4A8BA5" box="[1342,1432,1915,1941]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="2" pageNumber="374">Fig. 2A</figureCitation>
; see
<bibRefCitation id="EFF67AFE69762B29FF4C87DAFE188B89" author="Pati, S. K. &amp; Thackeray, T." box="[194,458,1951,1977]" pageId="2" pageNumber="374" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: figs. 5A, 8A;
<bibRefCitation id="EFF67AFE69762B29FDE087DAFCFB8B89" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[622,809,1951,1978]" pageId="2" pageNumber="374" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69762B29FD2E87E5FD008B89" box="[672,722,1951,1977]" italics="true" pageId="2" pageNumber="374">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 3A).
</paragraph>
<caption id="DF18578769772B28FF19872DFE778BDC" ID-DOI="http://doi.org/10.5281/zenodo.8430769" ID-Zenodo-Dep="8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="3" pageNumber="375" startId="3.[151,250,1896,1920]" targetBox="[151,1436,181,1872]" targetPageId="3" targetType="figure">
<paragraph id="8BD8070F69772B28FF19872DFE778BDC" blockId="3.[151,1436,1896,2028]" pageId="3" pageNumber="375">
<emphasis id="B913DB1D69772B28FF19872DFEC68BB0" bold="true" box="[151,276,1896,1920]" pageId="3" pageNumber="375">FIGURE 2.</emphasis>
<taxonomicName id="4C677C8C69772B28FE95872DFDC08BB0" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[283,530,1896,1920]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="375" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69772B28FE95872DFDC08BB0" box="[283,530,1896,1920]" italics="true" pageId="3" pageNumber="375">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69772B28FD97872DFDB88BB0" bold="true" box="[537,618,1896,1920]" pageId="3" pageNumber="375">
<taxonomicNameLabel id="A220666669772B28FD97872DFDB88BB0" box="[537,618,1896,1920]" pageId="3" pageNumber="375" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ).
</paragraph>
</caption>
<caption id="DF18578769702B2FFF198564FBA789B1" ID-DOI="http://doi.org/10.5281/zenodo.8430771" ID-Zenodo-Dep="8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="4" pageNumber="376" startId="4.[151,250,1313,1337]" targetBox="[151,1436,181,1289]" targetPageId="4" targetType="figure">
<paragraph id="8BD8070F69702B2FFF198564FBA789B1" blockId="4.[151,1436,1313,1409]" pageId="4" pageNumber="376">
<emphasis id="B913DB1D69702B2FFF198564FEC78909" bold="true" box="[151,277,1313,1337]" pageId="4" pageNumber="376">FIGURE 3.</emphasis>
<taxonomicName id="4C677C8C69702B2FFE908564FDC58909" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[286,535,1313,1337]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69702B2FFE908564FDC58909" box="[286,535,1313,1337]" italics="true" pageId="4" pageNumber="376">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69702B2FFD918564FDA08909" bold="true" box="[543,626,1313,1337]" pageId="4" pageNumber="376">
<taxonomicNameLabel id="A220666669702B2FFD918564FDA08909" box="[543,626,1313,1337]" pageId="4" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354. A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E).
</paragraph>
</caption>
<paragraph id="8BD8070F69702B2FFF498596FB478A4E" blockId="4.[151,1437,1491,1949]" pageId="4" pageNumber="376">
<taxonomicName id="4C677C8C69702B2FFF498596FE0589DD" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[199,471,1491,1517]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69702B2FFF498596FE0589DD" box="[199,471,1491,1517]" italics="true" pageId="4" pageNumber="376">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69702B2FFE508596FDEA89DD" bold="true" box="[478,568,1491,1517]" pageId="4" pageNumber="376">
<taxonomicNameLabel id="A220666669702B2FFE508596FDEA89DD" box="[478,568,1491,1517]" pageId="4" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C677C8C69702B2FFDC98596FD1F89DD" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[583,717,1491,1517]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69702B2FFDC98596FD1F89DD" box="[583,717,1491,1517]" italics="true" pageId="4" pageNumber="376">G. dvivarna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C677C8C69702B2FFC8A8596FCAC89DD" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[772,894,1491,1517]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69702B2FFC8A8596FCAC89DD" box="[772,894,1491,1517]" italics="true" pageId="4" pageNumber="376">G. pulchra</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C677C8C69702B2FFB3E8596FA9D89DD" baseAuthorityName="Klaus, Fernandez &amp; Yeo" baseAuthorityYear="2014" box="[1200,1359,1491,1517]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B913DB1D69702B2FFB3E8596FA9D89DD" box="[1200,1359,1491,1517]" italics="true" pageId="4" pageNumber="376">G. basalticola</emphasis>
</taxonomicName>
by the G1, which is almost straight or medially gently curved outwards, with the ultimate article being relatively longer, ca. 0.40.5 times the length of the penultimate article (
<figureCitation id="135C1B8A69702B2FFD7D865EFC9E8A05" box="[755,844,1563,1589]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="4" pageNumber="376">Fig. 2H</figureCitation>
; see
<bibRefCitation id="EFF67AFE69702B2FFC0E865EFB548A05" author="Pati, S. K. &amp; Thackeray, T." box="[896,1158,1563,1589]" pageId="4" pageNumber="376" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 8I;
<bibRefCitation id="EFF67AFE69702B2FFB6F865EFA4B8A05" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[1249,1433,1563,1590]" pageId="4" pageNumber="376" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69702B2FFA9D8659FA918A05" box="[1299,1347,1563,1589]" italics="true" pageId="4" pageNumber="376">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 4D) (vs. G1 medially distinctly curved outwards, with the ultimate article being relatively shorter, ca. 0.3 times the length of the penultimate article in
<taxonomicName id="4C677C8C69702B2FFDC48626FD3A8A4D" baseAuthorityName="Klaus, Fernandez &amp; Yeo" baseAuthorityYear="2014" box="[586,744,1635,1661]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B913DB1D69702B2FFDC48626FD3A8A4D" box="[586,744,1635,1661]" italics="true" pageId="4" pageNumber="376">G. basalticola</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69702B2FFCAC8626FBF88A4D" author="Pati, S. K. &amp; Thackeray, T." box="[802,1066,1635,1661]" pageId="4" pageNumber="376" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 5D).
</paragraph>
<paragraph id="8BD8070F69702B2FFF4986C2FC688BAD" blockId="4.[151,1437,1491,1949]" pageId="4" pageNumber="376">
<taxonomicName id="4C677C8C69702B2FFF4986C2FE058A91" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[199,471,1671,1697]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69702B2FFF4986C2FE058A91" box="[199,471,1671,1697]" italics="true" pageId="4" pageNumber="376">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69702B2FFE5186C2FDEB8A91" bold="true" box="[479,569,1671,1697]" pageId="4" pageNumber="376">
<taxonomicNameLabel id="A220666669702B2FFE5186C2FDEB8A91" box="[479,569,1671,1697]" pageId="4" pageNumber="376" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C677C8C69702B2FFDFE86C2FD258A91" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[624,759,1671,1697]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69702B2FFDFE86C2FD258A91" box="[624,759,1671,1697]" italics="true" pageId="4" pageNumber="376">G. dvivarna</emphasis>
</taxonomicName>
are immediately differentiated from
<taxonomicName id="4C677C8C69702B2FFB1986C2FAC38A91" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[1175,1297,1671,1697]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69702B2FFB1986C2FAC38A91" box="[1175,1297,1671,1697]" italics="true" pageId="4" pageNumber="376">G. pulchra</emphasis>
</taxonomicName>
by the form of the frontal margin, which is relatively close to the anterior margin of the epistome and conceals the antennular fossae (
<figureCitation id="135C1B8A69702B2FFF60868AFE9A8AD9" box="[238,328,1743,1769]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="4" pageNumber="376">Fig. 2B</figureCitation>
; see
<bibRefCitation id="EFF67AFE69702B2FFE0D868AFD918AD9" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[387,579,1743,1770]" pageId="4" pageNumber="376" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69702B2FFE398695FE388AD9" box="[439,490,1743,1769]" italics="true" pageId="4" pageNumber="376">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 3B) (vs. frontal margin some distance from the anterior margin of the epistome exposing the antennular fossae in
<taxonomicName id="4C677C8C69702B2FFDF786B6FD238B3D" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[633,753,1779,1805]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69702B2FFDF786B6FD238B3D" box="[633,753,1779,1805]" italics="true" pageId="4" pageNumber="376">G. pulchra</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69702B2FFCA686B6FBFF8B3D" author="Pati, S. K. &amp; Thackeray, T." box="[808,1069,1779,1805]" pageId="4" pageNumber="376" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 8B); the eye being relatively small as compared to the orbital space, with the relatively slender eyestalk (
<figureCitation id="135C1B8A69702B2FFC638752FB948B01" box="[1005,1094,1815,1841]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="4" pageNumber="376">Fig. 2B</figureCitation>
; see
<bibRefCitation id="EFF67AFE69702B2FFBF08752FAE88B01" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[1150,1338,1815,1842]" pageId="4" pageNumber="376" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69702B2FFB3F875DFB308B01" box="[1201,1250,1815,1841]" italics="true" pageId="4" pageNumber="376">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 3B) (vs. eye relatively large as compared to the orbital space, with the relatively stout eyestalk in
<taxonomicName id="4C677C8C69702B2FFB35877EFAE78B65" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[1211,1333,1851,1877]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69702B2FFB35877EFAE78B65" box="[1211,1333,1851,1877]" italics="true" pageId="4" pageNumber="376">G. pulchra</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69702B2FFAFF877EFEBF8B49" author="Pati, S. K. &amp; Thackeray, T." pageId="4" pageNumber="376" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 8B); and the G1 ultimate article being comparatively more slender (
<figureCitation id="135C1B8A69702B2FFB2E871AFB288B49" box="[1184,1274,1887,1913]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="4" pageNumber="376">Fig. 2H</figureCitation>
; see
<bibRefCitation id="EFF67AFE69702B2FFABF871AFF328BAD" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." pageId="4" pageNumber="376" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69702B2FFAEA8725FA478B49" box="[1380,1429,1887,1913]" italics="true" pageId="4" pageNumber="376">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 4D) than in
<taxonomicName id="4C677C8C69702B2FFE1087C6FDC58BAD" authorityName="Pati &amp; Thackeray" authorityYear="2018" box="[414,535,1923,1949]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="376" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B913DB1D69702B2FFE1087C6FDC58BAD" box="[414,535,1923,1949]" italics="true" pageId="4" pageNumber="376">G. pulchra</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF67AFE69702B2FFDDC87C6FC888BAD" author="Pati, S. K. &amp; Thackeray, T." box="[594,858,1923,1949]" pageId="4" pageNumber="376" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 8I).
</paragraph>
<caption id="DF18578769712B2EFF198461FA91880C" ID-DOI="http://doi.org/10.5281/zenodo.8430773" ID-Zenodo-Dep="8430773" box="[151,1347,1060,1085]" httpUri="https://zenodo.org/record/8430773/files/figure.png" pageId="5" pageNumber="377" startId="5.[151,250,1060,1084]" targetBox="[151,1436,181,1037]" targetPageId="5" targetType="figure">
<paragraph id="8BD8070F69712B2EFF198461FA91880C" blockId="5.[151,1347,1060,1085]" box="[151,1347,1060,1085]" pageId="5" pageNumber="377">
<emphasis id="B913DB1D69712B2EFF198461FEC1880C" bold="true" box="[151,275,1060,1084]" pageId="5" pageNumber="377">FIGURE 4.</emphasis>
<taxonomicName id="4C677C8C69712B2EFE978460FDDD880D" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[281,527,1061,1085]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFE978460FDDD880D" box="[281,527,1061,1085]" italics="true" pageId="5" pageNumber="377">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFD9B8460FDB4880D" bold="true" box="[533,614,1061,1085]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFD9B8460FDB4880D" box="[533,614,1061,1085]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, colour in life, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.
</paragraph>
</caption>
<paragraph id="8BD8070F69712B2EFF4984C4FDFD8A78" blockId="5.[151,1437,1153,2037]" pageId="5" pageNumber="377">
The morphological resemblance of the new species to
<taxonomicName id="4C677C8C69712B2EFCA484C7FC6388AB" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[810,945,1153,1180]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69712B2EFCA484C7FC6388AB" box="[810,945,1153,1180]" italics="true" pageId="5" pageNumber="377">G. dvivarna</emphasis>
</taxonomicName>
notwithstanding,
<taxonomicName id="4C677C8C69712B2EFBF484C7FAEE88AB" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[1146,1340,1153,1180]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFBF484C7FAEE88AB" box="[1146,1340,1153,1180]" italics="true" pageId="5" pageNumber="377">G. sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFACD84C4FA4E88AB" bold="true" box="[1347,1436,1153,1179]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFACD84C4FA4E88AB" box="[1347,1436,1153,1179]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is separated from
<taxonomicName id="4C677C8C69712B2EFEE684E0FE23888F" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[360,497,1189,1215]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69712B2EFEE684E0FE23888F" box="[360,497,1189,1215]" italics="true" pageId="5" pageNumber="377">G. dvivarna</emphasis>
</taxonomicName>
in having a relatively stout palm of the major chela of the adult males, with the strongly concave ventral margin of the fixed finger and the distal half of the palm (
<figureCitation id="135C1B8A69712B2EFBC6848CFB4E88D3" box="[1096,1180,1225,1251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="5" pageNumber="377">Fig. 2F</figureCitation>
) (vs. major chela with relatively slender palm in adult males, with gently concave ventral margin of the fixed finger and the distal half of the palm; see
<bibRefCitation id="EFF67AFE69712B2EFEBA8555FE23891A" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[308,497,1296,1323]" pageId="5" pageNumber="377" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69712B2EFEE98554FE4B891A" box="[359,409,1296,1322]" italics="true" pageId="5" pageNumber="377">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 4B); the outwardly curved and relatively shorter ultimate article of the G1, ca. 0.4 times the length of the penultimate article (
<figureCitation id="135C1B8A69712B2EFD128571FD21897E" box="[668,755,1332,1358]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="5" pageNumber="377">Fig. 2H</figureCitation>
) (vs. G1 ultimate article straight and relatively longer, ca. 0.5 times the length of the penultimate article; see
<bibRefCitation id="EFF67AFE69712B2EFD29851DFCB18942" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[679,867,1368,1394]" pageId="5" pageNumber="377" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69712B2EFD54851DFCD98942" box="[730,779,1368,1394]" italics="true" pageId="5" pageNumber="377">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 4D); and the strongly concave lateral margins of the female telson (
<figureCitation id="135C1B8A69712B2EFE0D853EFE0889A6" box="[387,474,1403,1430]" captionStart="FIGURE 3" captionStartId="4.[151,250,1313,1337]" captionTargetBox="[151,1436,181,1289]" captionTargetId="figure-20@4.[151,1436,181,1289]" captionTargetPageId="4" captionText="FIGURE 3. Ghatiana sanguinolenta sp. nov.: A, B, paratype male (28.53 × 17.85 mm), ZSI-WRC C.2354; CE, paratype female (28.45 × 18.26 mm), ZSI-WRC C.2354.A, C, overall dorsal view; B, pleonal somites 36 and telson; D, pleonal somites 46 and telson; E, thoracic sternites showing vulvae. Scale bars = 10 mm (A, C), 5 mm (B, D, E)." figureDoi="http://doi.org/10.5281/zenodo.8430771" httpUri="https://zenodo.org/record/8430771/files/figure.png" pageId="5" pageNumber="377">Fig. 3D</figureCitation>
) (vs. female telson with almost straight lateral margins; see
<bibRefCitation id="EFF67AFE69712B2EFBF4853EFAE789A6" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[1146,1333,1403,1430]" pageId="5" pageNumber="377" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69712B2EFB238539FB0C89A5" box="[1197,1246,1403,1429]" italics="true" pageId="5" pageNumber="377">et al</emphasis>
. 2022a
</bibRefCitation>
: fig. 5C). Both
<taxonomicName id="4C677C8C69712B2EFF5A85E5FE478989" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[212,405,1439,1466]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFF5A85E5FE478989" box="[212,405,1439,1466]" italics="true" pageId="5" pageNumber="377">G. sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFE1585DAFE278989" bold="true" box="[411,501,1439,1465]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFE1585DAFE278989" box="[411,501,1439,1465]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C677C8C69712B2EFDA485E5FD628989" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[554,688,1439,1466]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69712B2EFDA485E5FD628989" box="[554,688,1439,1466]" italics="true" pageId="5" pageNumber="377">G. dvivarna</emphasis>
</taxonomicName>
are known only from the Western Ghats of Uttara Kannada district in
<collectingRegion id="49A3C9ED69712B2EFF3B8586FEFA89ED" box="[181,296,1475,1501]" country="India" name="Karnataka" pageId="5" pageNumber="377">Karnataka</collectingRegion>
(
<bibRefCitation id="EFF67AFE69712B2EFEB98586FE2189ED" author="Pati, S. K. &amp; Thackeray, T. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[311,499,1475,1501]" pageId="5" pageNumber="377" pagination="2022019" refId="ref3872" refString="Pati, S. K., Thackeray, T., Bajantri, P. P. &amp; Hegde, G. D. (2022 a) A new species of the freshwater crab genus Ghatiana Pati and Sharma, 2014 (Brachyura: Gecarcinucidae) from the Central Western Ghats, India. Nauplius, 30, e 2022019. https: // doi. org / 10.1590 / 2358 - 2936 e 2022019" type="journal article" year="2022">
Pati
<emphasis id="B913DB1D69712B2EFEE28581FE4F89ED" box="[364,413,1475,1501]" italics="true" pageId="5" pageNumber="377">et al</emphasis>
. 2022a
</bibRefCitation>
; present study). Although the
<typeStatus id="54DCB9AD69712B2EFCC88586FCA489ED" box="[838,886,1475,1501]" pageId="5" pageNumber="377">type</typeStatus>
locality of
<taxonomicName id="4C677C8C69712B2EFC778586FB6889ED" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[1017,1210,1475,1501]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFC778586FB6889ED" box="[1017,1210,1475,1501]" italics="true" pageId="5" pageNumber="377">G. sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFB4C8586FACE89ED" bold="true" box="[1218,1308,1475,1501]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFB4C8586FACE89ED" box="[1218,1308,1475,1501]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is some
<quantity id="4C9FAAEA69712B2EFA0E8586FF698A30" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="377" unit="km" value="30.0">30 km</quantity>
away from the
<typeStatus id="54DCB9AD69712B2EFEE785A2FE4B8A31" box="[361,409,1511,1537]" pageId="5" pageNumber="377">type</typeStatus>
locality of
<taxonomicName id="4C677C8C69712B2EFD9685A2FD4F8A30" authorityName="Pati, Thackeray, Bajantri &amp; Hegde" authorityYear="2022" box="[536,669,1510,1537]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="dvivarna">
<emphasis id="B913DB1D69712B2EFD9685A2FD4F8A30" box="[536,669,1510,1537]" italics="true" pageId="5" pageNumber="377">G. dvivarna</emphasis>
</taxonomicName>
, their respective
<typeStatus id="54DCB9AD69712B2EFCD285A2FC5E8A31" box="[860,908,1511,1537]" pageId="5" pageNumber="377">type</typeStatus>
localities are situated in the isolated mountains separated by the Gangavali River and the deep valley, which could represent the geographic barriers in the form of “sky islands” (see
<bibRefCitation id="EFF67AFE69712B2EFEE9866BFDF18A78" author="Pati, S. K. &amp; Bajantri, P. P. &amp; Hegde, G. D." box="[359,547,1582,1609]" pageId="5" pageNumber="377" pagination="161 - 175" refId="ref3729" refString="Pati, S. K., Bajantri, P. P. &amp; Hegde, G. D. (2023 b) A new genus and new species of freshwater crab (Decapoda: Brachyura: Gecarcinucidae) from the Central Western Ghats of India. Zootaxa, 5285 (1), 161 - 175. https: // doi. org / 10.11646 / zootaxa. 5285.1.7" type="journal article" year="2023">
Pati
<emphasis id="B913DB1D69712B2EFE17866AFE198A78" box="[409,459,1582,1608]" italics="true" pageId="5" pageNumber="377">et al</emphasis>
. 2023b
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD8070F69712B2EFF498617FBD78BBA" blockId="5.[151,1437,1153,2037]" pageId="5" pageNumber="377">
<taxonomicName id="4C677C8C69712B2EFF498617FE008A5C" authorityName="Pati, Thackeray &amp; Khaire" authorityYear="2016" box="[199,466,1618,1644]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="atropurpurea">
<emphasis id="B913DB1D69712B2EFF498617FE008A5C" box="[199,466,1618,1644]" italics="true" pageId="5" pageNumber="377">Ghatiana atropurpurea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C677C8C69712B2EFD9A8617FDA78A5C" authorityName="Pati &amp; Thackeray" authorityYear="2021" box="[532,629,1618,1644]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="rouxi">
<emphasis id="B913DB1D69712B2EFD9A8617FDA78A5C" box="[532,629,1618,1644]" italics="true" pageId="5" pageNumber="377">G. rouxi</emphasis>
</taxonomicName>
are likely to be encountered along with the new species.
<taxonomicName id="4C677C8C69712B2EFABC8617FEE48ABF" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFABC8617FEE48ABF" italics="true" pageId="5" pageNumber="377">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFEB08633FE4A8AA0" bold="true" box="[318,408,1654,1680]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFEB08633FE4A8AA0" box="[318,408,1654,1680]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is easily differentiated from them mainly by the proportionately broader carapace, CW/CL = 1.6 (
<figureCitation id="135C1B8A69712B2EFF6E86DCFEEA8A83" box="[224,312,1689,1715]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="5" pageNumber="377">Fig. 2A</figureCitation>
) (vs. carapace proportionately narrower, CW/CL =
<quantity id="4C9FAAEA69712B2EFCF686DCFC3B8A83" box="[888,1001,1689,1716]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="5" pageNumber="377" unit="in" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 in</quantity>
<taxonomicName id="4C677C8C69712B2EFC6186DFFB748A83" authorityName="Pati, Thackeray &amp; Khaire" authorityYear="2016" box="[1007,1190,1690,1716]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="atropurpurea">
<emphasis id="B913DB1D69712B2EFC6186DFFB748A83" box="[1007,1190,1690,1716]" italics="true" pageId="5" pageNumber="377">G. atropurpurea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C677C8C69712B2EFB5286DFFAEA8A83" authorityName="Pati &amp; Thackeray" authorityYear="2021" box="[1244,1336,1690,1716]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="rouxi">
<emphasis id="B913DB1D69712B2EFB5286DFFAEA8A83" box="[1244,1336,1690,1716]" italics="true" pageId="5" pageNumber="377">G. rouxi</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69712B2EFAFF86DCFEBF8AE7" author="Pati, S. K. &amp; Thackeray, T." pageId="5" pageNumber="377" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 3A; 2021: 4A); the relatively small eye as compared to the orbital space, with the eyestalk relatively slender (
<figureCitation id="135C1B8A69712B2EFEE286A4FE118ACB" box="[364,451,1761,1787]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="5" pageNumber="377">Fig. 2B</figureCitation>
) (vs. eye relatively large as compared to the orbital space, with the eyestalk relatively stout in
<taxonomicName id="4C677C8C69712B2EFF618740FE748B2E" authorityName="Pati, Thackeray &amp; Khaire" authorityYear="2016" box="[239,422,1797,1823]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="atropurpurea">
<emphasis id="B913DB1D69712B2EFF618740FE748B2E" box="[239,422,1797,1823]" italics="true" pageId="5" pageNumber="377">G. atropurpurea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C677C8C69712B2EFE548740FDE78B2E" authorityName="Pati &amp; Thackeray" authorityYear="2021" box="[474,565,1797,1823]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="rouxi">
<emphasis id="B913DB1D69712B2EFE548740FDE78B2E" box="[474,565,1797,1823]" italics="true" pageId="5" pageNumber="377">G. rouxi</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69712B2EFDE38741FCA08B2F" author="Pati, S. K. &amp; Thackeray, T." box="[621,882,1796,1823]" pageId="5" pageNumber="377" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 3B; 2021: 4C); and the outwardly curved and relatively shorter ultimate article of the G1, which is about 0.4 times the length of the penultimate article (
<figureCitation id="135C1B8A69712B2EFAB2876DFA468B72" box="[1340,1428,1832,1858]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1436,181,1872]" captionTargetId="figure-30@3.[151,1436,181,1872]" captionTargetPageId="3" captionText="FIGURE 2. Ghatiana sanguinolenta sp. nov., holotype male (24.48 × 15.38 mm), ZSI-WRC C.2353. A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view; D, mouth parts exposed; E, left maxilliped 3; F, outer view of major or left chela; G, pleon; H, dorsal view of left G1; I, ventral view of left G1; J, left G2. Scale bars = 10 mm (AC, F), 5 mm (G), 2 mm (D, E), 1 mm (HJ)." figureDoi="http://doi.org/10.5281/zenodo.8430769" httpUri="https://zenodo.org/record/8430769/files/figure.png" pageId="5" pageNumber="377">Fig. 2H</figureCitation>
) (vs. G1 ultimate article inwardly curved and relatively longer, ca. 0.5 times the length of the penultimate article in
<taxonomicName id="4C677C8C69712B2EFF198735FE9C8BB9" authorityName="Pati, Thackeray &amp; Khaire" authorityYear="2016" box="[151,334,1904,1930]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="atropurpurea">
<emphasis id="B913DB1D69712B2EFF198735FE9C8BB9" box="[151,334,1904,1930]" italics="true" pageId="5" pageNumber="377">G. atropurpurea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C677C8C69712B2EFE0B8735FE338BB9" authorityName="Pati &amp; Thackeray" authorityYear="2021" box="[389,481,1904,1930]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="rouxi">
<emphasis id="B913DB1D69712B2EFE0B8735FE338BB9" box="[389,481,1904,1930]" italics="true" pageId="5" pageNumber="377">G. rouxi</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFF67AFE69712B2EFD948735FCF08BBA" author="Pati, S. K. &amp; Thackeray, T." box="[538,802,1904,1930]" pageId="5" pageNumber="377" pagination="1 - 73" refId="ref3512" refString="Pati, S. K. &amp; Thackeray, T. (2018) The freshwater crab genera Ghatiana Pati &amp; Sharma, Gubernatoriana Bott, and Inglethelphusa Bott (Crustacea: Decapoda: Brachyura: Gecarcinucidae) revisited, with descriptions of a new genus and eleven new species. Zootaxa, 4440, 1 - 73. https: // doi. org / 10.11646 / zootaxa. 4440.1.1" type="journal article" year="2018">Pati &amp; Thackeray 2018</bibRefCitation>
: fig. 3D; 2021: 5D).
</paragraph>
</subSubSection>
<subSubSection id="C37D548469712B2EFF4987D6FE4B8BC5" pageId="5" pageNumber="377" type="distribution">
<paragraph id="8BD8070F69712B2EFF4987D6FE4B8BC5" blockId="5.[151,1437,1153,2037]" pageId="5" pageNumber="377">
<emphasis id="B913DB1D69712B2EFF4987D6FDD48B9D" bold="true" box="[199,518,1939,1965]" pageId="5" pageNumber="377">Geographical distribution.</emphasis>
<taxonomicName id="4C677C8C69712B2EFD8087D6FCCD8B9D" authority="Pati &amp; Thackeray &amp; Pawar, 2023" authorityName="Pati &amp; Thackeray &amp; Pawar" authorityYear="2023" box="[526,799,1939,1965]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="377" phylum="Arthropoda" rank="species" species="sanguinolenta" status="sp. nov.">
<emphasis id="B913DB1D69712B2EFD8087D6FCCD8B9D" box="[526,799,1939,1965]" italics="true" pageId="5" pageNumber="377">Ghatiana sanguinolenta</emphasis>
</taxonomicName>
<emphasis id="B913DB1D69712B2EFCA987D1FC508B9E" bold="true" box="[807,898,1940,1966]" pageId="5" pageNumber="377">
<taxonomicNameLabel id="A220666669712B2EFCA987D1FC508B9E" box="[807,898,1940,1966]" pageId="5" pageNumber="377" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently known only from the
<typeStatus id="54DCB9AD69712B2EFA8587D1FAE98B9E" box="[1291,1339,1940,1966]" pageId="5" pageNumber="377">type</typeStatus>
locality, i.e.,
<location id="8EB851D469712B2EFF4787F2FDC98BE1" LSID="urn:lsid:plazi:treatment:03CEB61969762B2EFF1981F2FE4B8BC5:8EB851D469712B2EFF4787F2FDC98BE1" box="[201,539,1975,2002]" country="India" county="Central Western Ghats" municipality="Uttara Kannada district" name="Balekoppa in Uttara Kannada" pageId="5" pageNumber="377" stateProvince="Karnataka">
Balekoppa in
<collectingMunicipality id="6BBC9D7569712B2EFEE687FDFDC98BE1" box="[360,539,1975,2002]" pageId="5" pageNumber="377">Uttara Kannada</collectingMunicipality>
</location>
district of
<collectingRegion id="49A3C9ED69712B2EFD1487F2FC9E8BE2" box="[666,844,1975,2002]" country="India" name="Karnataka" pageId="5" pageNumber="377">Karnataka state</collectingRegion>
. The
<typeStatus id="54DCB9AD69712B2EFC0287FDFC6E8BE2" box="[908,956,1976,2002]" pageId="5" pageNumber="377">type</typeStatus>
locality is situated in the
<collectingCounty id="62B97F8369712B2EFB6987F2FF0A8BC5" pageId="5" pageNumber="377">Central Western Ghats</collectingCounty>
of
<collectingCountry id="F370479F69712B2EFF70879EFEEA8BC5" box="[254,312,2011,2037]" name="India" pageId="5" pageNumber="377">India</collectingCountry>
(
<figureCitation id="135C1B8A69712B2EFEC9879EFE5A8BC5" box="[327,392,2011,2037]" captionStart="FIGURE 1" captionStartId="1.[151,250,1800,1824]" captionTargetBox="[253,1334,182,1777]" captionTargetId="figure-29@1.[253,1334,181,1777]" captionTargetPageId="1" captionText="FIGURE 1. Map of India and Western Ghats showing the type locality of Ghatiana sanguinolenta sp. nov." figureDoi="http://doi.org/10.5281/zenodo.8430767" httpUri="https://zenodo.org/record/8430767/files/figure.png" pageId="5" pageNumber="377">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>