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<document id="3F2E4067DBEF6D5DF5BA9D922B0FA560" ID-DOI="10.26028/cybium/2019-423-004" ID-ISSN="2101-0315" ID-Zenodo-Dep="12711126" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720642888589" checkinUser="felipe" docAuthor="Keith, Philippe, Amick, Pita, Toko, Pagi S. &amp; Lord, Clara" docDate="2019" docId="03CE87F38459FFB7FED4FC5208C9A32B" docLanguage="en" docName="Cybium.43.2.163-167.pdf" docOrigin="Cybium 43 (2)" docSource="http://dx.doi.org/10.26028/cybium/2019-423-004" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Sicyopus beremeensis Keith, Amick, Toko &amp; Lord, 2019, n. sp." docType="treatment" docVersion="2" lastPageNumber="166" masterDocId="FFF7FF8B8458FFB4FFFAFFC60C3CA578" masterDocTitle="A new species of Sicyopus (Teleostei: Gobiidae) from New Britain (Papua New Guinea)" masterLastPageNumber="167" masterPageNumber="163" pageNumber="164" updateTime="1720649285720" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="86FD3EE99203757C15A9DA6B6399ECAE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E5E8C187A40F86BC15FC1C37B4C20B59">
<mods:title id="633B0E5FB365D3F4CD753AC0B34EE8D5">A new species of Sicyopus (Teleostei: Gobiidae) from New Britain (Papua New Guinea)</mods:title>
</mods:titleInfo>
<mods:name id="5785695E84F892B24EF12222DA456610" type="personal">
<mods:role id="080B8F9A98C6CC31C4B77E05E55F6031">
<mods:roleTerm id="0B3C4FD0697F80555DCC170FF8D179DD">Author</mods:roleTerm>
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<mods:namePart id="7385FB58C272234C8651C11AB8F95BC4">Keith, Philippe</mods:namePart>
</mods:name>
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<mods:roleTerm id="7615F1BE8A48133ADD8300D2D406917D">Author</mods:roleTerm>
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<mods:namePart id="1A9573A5FB3BDCB8D05D3115B06AE6B1">Amick, Pita</mods:namePart>
</mods:name>
<mods:name id="7655169E1BCDD7F314E4C87A79DB1D45" type="personal">
<mods:role id="6965FA785F609B61943D6AE53414DD70">
<mods:roleTerm id="D62040768CAF78AB5425288CED97C5FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="059E056E1057242AD09187E267E8407C">Toko, Pagi S.</mods:namePart>
</mods:name>
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<mods:namePart id="ABDEF6446277EC9B85D8699D289F56C8">Lord, Clara</mods:namePart>
</mods:name>
<mods:typeOfResource id="1E7CF3EE19653CA3A9750A3B499EE9C4">text</mods:typeOfResource>
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<mods:title id="783555AF148D49ECB24C8B02A36F9BFE">Cybium</mods:title>
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<mods:part id="E87DCB31763425D5922E11825E5D2468">
<mods:date id="0525C3E39200B6B98FECCA18EFDD6DE2">2019</mods:date>
<mods:detail id="E1F7BB1B4B75E762690C3A75BEDA2DBD" type="volume">
<mods:number id="C7BCC0C97CFA5BC8E972DC1F0AB37E50">43</mods:number>
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<mods:location id="121A387B23BE16980277191650F374E3">
<mods:url id="36730F9CDC35472DAF96FC0ED9032A0F">http://dx.doi.org/10.26028/cybium/2019-423-004</mods:url>
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<mods:classification id="EA925E8BFD6C070E354F3D6DC522A709">journal article</mods:classification>
<mods:identifier id="10F40A2F16D58410099C215046BBFE02" type="DOI">10.26028/cybium/2019-423-004</mods:identifier>
<mods:identifier id="1DF52DD8C8C8E4D296C6F212644F7F1A" type="ISSN">2101-0315</mods:identifier>
<mods:identifier id="8A8C4541C4872E0991A6456E4D35D721" type="Zenodo-Dep">12711126</mods:identifier>
</mods:mods>
<treatment id="03CE87F38459FFB7FED4FC5208C9A32B" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B" httpUri="http://treatment.plazi.org/id/03CE87F38459FFB7FED4FC5208C9A32B" lastPageId="3" lastPageNumber="166" pageId="1" pageNumber="164">
<subSubSection id="C37D656E8459FFB5FED4FC520E61A6D5" box="[302,605,916,941]" pageId="1" pageNumber="164" type="nomenclature">
<paragraph id="8BD836E58459FFB5FED4FC520E61A6D5" blockId="1.[302,605,916,941]" box="[302,605,916,941]" pageId="1" pageNumber="164">
<heading id="D09081898459FFB5FED4FC520E61A6D5" bold="true" box="[302,605,916,941]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="164" reason="2">
<emphasis id="B913EAF78459FFB5FED4FC520E61A6D5" bold="true" box="[302,605,916,941]" pageId="1" pageNumber="164">
<taxonomicName id="4C674D668459FFB5FED4FC520E20A6D4" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[302,540,916,940]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="beremeensis" status="sp. nov.">
<emphasis id="B913EAF78459FFB5FED4FC520E20A6D4" bold="true" box="[302,540,916,940]" italics="true" pageId="1" pageNumber="164">Sicyopus beremeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220578C8459FFB5FDD8FC530E61A6D5" box="[546,605,917,941]" pageId="1" pageNumber="164" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37D656E8459FFB5FE83FC700E2EA6B6" box="[377,530,950,974]" pageId="1" pageNumber="164" type="description">
<paragraph id="8BD836E58459FFB5FE83FC700E2EA6B6" blockId="1.[377,530,950,974]" box="[377,530,950,974]" pageId="1" pageNumber="164">
(
<figureCitation id="135C2A608459FFB5FE7BFC700D83A6B6" box="[385,447,950,974]" captionStart="Figure 1" captionStartId="1.[813,879,980,1002]" captionTargetBox="[813,1456,215,964]" captionTargetId="figure-734@1.[813,1456,588,964]" captionTargetPageId="1" captionText="Figure 1. – Sicyopus beremeensis. A: Male (49 mm); B: Female (45 mm) (photograph by P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711128" httpUri="https://zenodo.org/record/12711128/files/figure.png" pageId="1" pageNumber="164">Fig. 1</figureCitation>
,
<tableCitation id="C6E5035E8459FFB5FE36FC700E35A6B5" box="[460,521,950,974]" captionStart="Table I" captionStartId="2.[132,186,217,239]" captionText="Table I. – Counts and morphomeristics of the new species.A, anal fin rays; D, dorsal fins rays; P, pectoral fin rays; C, caudal fin rays; PD, predorsal midline scales; TRB, transverse series backward scales; TRF, transverse series forward scales; ZZ, zigzag series scales; UJ, upper jaw teeth; LJ, lower jaw teeth; SL, standard Length; PDL, predorsal length (% SL); PAL, preanal length (% SL); HL, head length (% SL); JL, jaw length (% SL); CPL, caudal peduncle length (% SL); CPD, caudal peduncle depth (% SL); BDa, body depth at anus (% SL); BDD1, body depth at first dorsal fin in males (% SL); SDFL, second dorsal fin length (% SL); AFL, anal fin length (% SL); CFL, caudal fin length (% SL))." pageId="1" pageNumber="164">Tab. I</tableCitation>
)
</paragraph>
</subSubSection>
<caption id="DF18666D8459FFB5FCD7FC1208BEA17A" ID-DOI="http://doi.org/10.5281/zenodo.12711128" ID-Zenodo-Dep="12711128" httpUri="https://zenodo.org/record/12711128/files/figure.png" pageId="1" pageNumber="164" startId="1.[813,879,980,1002]" targetBox="[813,1456,215,964]" targetPageId="1" targetType="figure">
<paragraph id="8BD836E58459FFB5FCD7FC1208BEA17A" blockId="1.[813,1455,980,1026]" pageId="1" pageNumber="164">
Figure 1. –
<emphasis id="B913EAF78459FFB5FC5BFC1308B8A691" box="[929,1156,980,1002]" italics="true" pageId="1" pageNumber="164">
<taxonomicName id="4C674D668459FFB5FC5BFC130843A691" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[929,1151,980,1002]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="beremeensis">Sicyopus beremeensis</taxonomicName>
.
</emphasis>
<emphasis id="B913EAF78459FFB5FB70FC1208A7A691" bold="true" box="[1162,1179,980,1001]" pageId="1" pageNumber="164">A</emphasis>
: Male (49 mm);
<emphasis id="B913EAF78459FFB5FAB2FC120964A691" bold="true" box="[1352,1368,980,1001]" pageId="1" pageNumber="164">B</emphasis>
: Female (45 mm) (photograph by P. Amick).
</paragraph>
</caption>
<subSubSection id="C37D656E8459FFB5FF7EFC310CD2A20B" pageId="1" pageNumber="164" type="materials_examined">
<paragraph id="8BD836E58459FFB5FF7EFC310DBCA177" blockId="1.[132,775,1015,1106]" box="[132,384,1015,1039]" pageId="1" pageNumber="164">
<emphasis id="B913EAF78459FFB5FF7EFC310DBCA177" bold="true" box="[132,384,1015,1039]" pageId="1" pageNumber="164">Comparative material</emphasis>
</paragraph>
<paragraph id="8BD836E58459FFB5FF50FBDF0E22A181" blockId="1.[132,775,1015,1106]" lastBlockId="1.[132,775,1192,1374]" pageId="1" pageNumber="164">
The new species is compared with
<taxonomicName id="4C674D668459FFB5FDD3FBDF0DB3A12A" authority="de Beaufort, 1912" authorityName="de Beaufort" authorityYear="1912" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="multisqua-uamatus">
<emphasis id="B913EAF78459FFB5FDD3FBDF0CF8A12A" italics="true" pageId="1" pageNumber="164">Sicyopus multisqua-uamatus</emphasis>
de Beaufort, 1912
</taxonomicName>
, the only other species of
<taxonomicName id="4C674D668459FFB5FD5DFBFC0F3BA129" box="[679,775,1082,1105]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="genus">
<emphasis id="B913EAF78459FFB5FD5DFBFC0F3BA129" box="[679,775,1082,1105]" italics="true" pageId="1" pageNumber="164">Sicyopus</emphasis>
</taxonomicName>
having a first dorsal fin with six spines and a second dorsal fin with one spine and ten segmented rays (D VI-I,10), an anal fin with one spine and ten segmented rays (A I,10), and a pectoral fin with generally 17 rays.
<taxonomicName id="4C674D668459FFB5FDF5FB060C9FA181" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="multisquama-uamatus">
<emphasis id="B913EAF78459FFB5FDF5FB060C9FA181" italics="true" pageId="1" pageNumber="164">Sicyopus multisquama-uamatus</emphasis>
</taxonomicName>
was redescribed by
<bibRefCitation id="EFF64B148459FFB5FE80FB240E26A181" author="WATSON R. E." box="[378,538,1249,1273]" pageId="1" pageNumber="164" pagination="267 - 278" refId="ref4231" refString="WATSON R. E., 1995. - A new species of Sicyopus from Papua New Guinea with redescription of S. multisquamatus (Teleostei: Gobioidei: Sicydiinae). Ichthyol. Expl. Freshw., 6 (3): 267 - 278." type="journal article" year="1995">Watson (1995)</bibRefCitation>
.
</paragraph>
<paragraph id="8BD836E58459FFB5FF50FAC50E54A025" blockId="1.[132,775,1192,1374]" pageId="1" pageNumber="164">
<emphasis id="B913EAF78459FFB5FF50FAC50DDAA062" box="[170,486,1283,1307]" italics="true" pageId="1" pageNumber="164">
<taxonomicName id="4C674D668459FFB5FF50FAC50DF2A063" authorityName="de Beaufort" authorityYear="1912" box="[170,462,1283,1307]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="multisquamatus">Sicyopus multisquamatus</taxonomicName>
. –
</emphasis>
<materialsCitation id="3B0F3CB88459FFB5FE15FAC50E58A025" collectingDate="1910-02-19" collectionCode="ZMA" collectorName="de Beaufort" country="Indonesia" location="Honitetu" municipality="Ceram" pageId="1" pageNumber="164" specimenCode="ZMA 110.982" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBC19E9E8459FFB5FE15FAC50EADA063" box="[495,657,1283,1307]" collectionCode="ZMA" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="1" pageNumber="164">ZMA 110.982</specimenCode>
,
<typeStatus id="54DC88478459FFB5FD67FAC50F3FA063" box="[669,771,1283,1307]" pageId="1" pageNumber="164" type="holotype">holotype</typeStatus>
, female,
<quantity id="4C9F9B008459FFB5FF27FAE20D03A044" box="[221,319,1316,1340]" metricMagnitude="-2" metricUnit="m" metricValue="4.62" pageId="1" pageNumber="164" unit="mm" value="46.2">46.2 mm</quantity>
SL; small mountain creek near
<location id="8EB8603E8459FFB5FD65FAE20F3EA044" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B:8EB8603E8459FFB5FD65FAE20F3EA044" box="[671,770,1316,1340]" country="Indonesia" municipality="Ceram" name="Honitetu" pageId="1" pageNumber="164">Honitetu</location>
,
<collectingMunicipality id="6BBCAC9F8459FFB5FF7EFA800CF2A026" box="[132,206,1350,1374]" pageId="1" pageNumber="164">Ceram</collectingMunicipality>
,
<collectingCountry id="F37076758459FFB5FF22FA830D7EA025" box="[216,322,1349,1373]" name="Indonesia" pageId="1" pageNumber="164">Indonesia</collectingCountry>
,
<date id="FFD910258459FFB5FEB6FA800DE6A026" box="[332,474,1349,1374]" pageId="1" pageNumber="164" value="1910-02-19">
<collectingDate id="EF9DE9CD8459FFB5FEB6FA800DE6A026" box="[332,474,1349,1374]" pageId="1" pageNumber="164" value="1910-02-19">19 Feb. 1910</collectingDate>
</date>
,
<collectorName id="269253338459FFB5FE1EFA830E58A025" box="[484,612,1349,1373]" pageId="1" pageNumber="164">de Beaufort</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD836E58459FFB5FF7EFA410D60A0E7" blockId="1.[132,775,1415,1907]" box="[132,348,1415,1439]" pageId="1" pageNumber="164">
<emphasis id="B913EAF78459FFB5FF7EFA410D60A0E7" bold="true" box="[132,348,1415,1439]" pageId="1" pageNumber="164">Material examined</emphasis>
</paragraph>
<paragraph id="8BD836E58459FFB5FF50FA6E0D96A35D" blockId="1.[132,775,1415,1907]" pageId="1" pageNumber="164">
<materialsCitation id="3B0F3CB88459FFB5FF50FA6E0D31A09A" country="Papua New Guinea" location="New Britain" pageId="1" pageNumber="164" specimenCount="11" stateProvince="West New Britain">
<specimenCount id="9D61FD6C8459FFB5FF50FA6E0D51A0B8" box="[170,365,1448,1472]" count="11" pageId="1" pageNumber="164" type="generic">Eleven specimens</specimenCount>
from
<collectingRegion id="49A3F8078459FFB5FE4AFA6F0E09A0B8" box="[432,565,1448,1472]" country="Papua New Guinea" name="West New Britain" pageId="1" pageNumber="164">New Britain</collectingRegion>
(
<collectingCountry id="F37076758459FFB5FDB9FA6F0CA3A09A" name="Papua New Guinea" pageId="1" pageNumber="164">Papua New Guinea</collectingCountry>
), totalling
</materialsCitation>
<specimenCount id="9D61FD6C8459FFB5FEEEFA0C0D5BA09A" box="[276,359,1482,1506]" count="8" pageId="1" pageNumber="164" type="male">8 males</specimenCount>
and
<specimenCount id="9D61FD6C8459FFB5FE61FA0C0E3BA09A" box="[411,519,1482,1506]" count="3" pageId="1" pageNumber="164" type="female">3 females</specimenCount>
; size range
<quantity id="4C9F9B008459FFB5FD7EFA0C0F3BA099" box="[644,775,1482,1506]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" metricValueMax="5.68" metricValueMin="3.9" pageId="1" pageNumber="164" unit="mm" value="47.9" valueMax="56.8" valueMin="39.0">39-56.8 mm</quantity>
SL (
<quantity id="4C9F9B008459FFB5FF4BFA2D0D6CA37B" box="[177,336,1515,1539]" metricMagnitude="-2" metricUnit="m" metricValue="5.865" metricValueMax="6.87" metricValueMin="4.86" pageId="1" pageNumber="164" unit="mm" value="58.65" valueMax="68.7" valueMin="48.6">48.6-68.7 mm</quantity>
, total length), largest male
<quantity id="4C9F9B008459FFB5FD83FA2D0EE6A37B" box="[633,730,1515,1539]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" pageId="1" pageNumber="164" unit="mm" value="56.8">56.8 mm</quantity>
SL, largest female
<quantity id="4C9F9B008459FFB5FEDAF9CA0D41A35C" box="[288,381,1548,1572]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" pageId="1" pageNumber="164" unit="mm" value="50.5">50.5 mm</quantity>
SL.
</paragraph>
<paragraph id="8BD836E58459FFB5FF50F9E80D41A3F1" blockId="1.[132,775,1415,1907]" pageId="1" pageNumber="164">
<materialsCitation id="3B0F3CB88459FFB5FF50F9E80D41A3F1" collectingDate="2015-12-03" collectionCode="MNHN" country="Papua New Guinea" location="Wani Creek" municipality="Amick" pageId="1" pageNumber="164" specimenCode="MNHN 2016-0628" specimenCount="1" stateProvince="West New Britain" typeStatus="holotype">
<emphasis id="B913EAF78459FFB5FF50F9E80D2EA33E" box="[170,274,1582,1606]" italics="true" pageId="1" pageNumber="164">
<typeStatus id="54DC88478459FFB5FF50F9E80D2EA33E" box="[170,274,1582,1606]" pageId="1" pageNumber="164" type="holotype">Holotype</typeStatus>
</emphasis>
. –
<specimenCode id="DBC19E9E8459FFB5FEC2F9E90E2FA33E" box="[312,531,1582,1606]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="164">MNHN 2016-0628</specimenCode>
, male (
<quantity id="4C9F9B008459FFB5FD9DF9E80EF1A33E" box="[615,717,1582,1606]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="1" pageNumber="164" unit="mm" value="55.6">55.6 mm</quantity>
SL),
<location id="8EB8603E8459FFB5FF7EF9890D32A31F" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B:8EB8603E8459FFB5FF7EF9890D32A31F" box="[132,270,1615,1639]" country="Papua New Guinea" municipality="Amick" name="Wani Creek" pageId="1" pageNumber="164" stateProvince="West New Britain">Wani Creek</location>
,
<collectingRegion id="49A3F8078459FFB5FEE1F9960D9BA31F" box="[283,423,1615,1639]" country="Papua New Guinea" name="West New Britain" pageId="1" pageNumber="164">New Britain</collectingRegion>
(
<collectingCountry id="F37076758459FFB5FE4DF9960EA6A31F" box="[439,666,1615,1640]" name="Papua New Guinea" pageId="1" pageNumber="164">Papua New Guinea</collectingCountry>
),
<date id="FFD910258459FFB5FD55F9960C80A3F1" pageId="1" pageNumber="164" value="2015-12-03">
<collectingDate id="EF9DE9CD8459FFB5FD55F9960C80A3F1" pageId="1" pageNumber="164" value="2015-12-03">03 Dec. 2015</collectingDate>
</date>
,
<collectingMunicipality id="6BBCAC9F8459FFB5FF3FF9B70D32A3F1" box="[197,270,1649,1673]" pageId="1" pageNumber="164">Amick</collectingMunicipality>
<emphasis id="B913EAF78459FFB5FEE9F9B40D7EA3F1" box="[275,322,1649,1673]" italics="true" pageId="1" pageNumber="164">et al</emphasis>
. coll.
</materialsCitation>
</paragraph>
<paragraph id="8BD836E58459FFB5FF50F9550CD2A20B" blockId="1.[132,775,1415,1907]" pageId="1" pageNumber="164">
<materialsCitation id="3B0F3CB88459FFB5FF50F9550E1FA3B4" collectingDate="2015-12-03" collectionCode="MNHN" country="Papua New Guinea" location="Same" municipality="Amick" pageId="1" pageNumber="164" specimenCode="MNHN 2016-0629" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="West New Britain" typeStatus="paratype">
<emphasis id="B913EAF78459FFB5FF50F9550D25A3D2" box="[170,281,1683,1706]" italics="true" pageId="1" pageNumber="164">
<typeStatus id="54DC88478459FFB5FF50F9550D25A3D2" box="[170,281,1683,1706]" pageId="1" pageNumber="164" type="paratype">Paratypes</typeStatus>
</emphasis>
. –
<specimenCode id="DBC19E9E8459FFB5FEC6F9550E2CA3D2" box="[316,528,1682,1706]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="164">MNHN 2016-0629</specimenCode>
,
<specimenCount id="9D61FD6C8459FFB5FDE1F9540E54A3D2" box="[539,616,1682,1706]" count="1" pageId="1" pageNumber="164" type="male">1 male</specimenCount>
,
<specimenCount id="9D61FD6C8459FFB5FD89F9540EEEA3D2" box="[627,722,1682,1706]" count="1" pageId="1" pageNumber="164" type="female">1 female</specimenCount>
(
<quantity id="4C9F9B008459FFB5FD1BF9540CDBA3B3" metricMagnitude="-2" metricUnit="m" metricValue="4.68" metricValueMax="4.86" metricValueMin="4.5" pageId="1" pageNumber="164" unit="mm" value="46.8" valueMax="48.6" valueMin="45.0">45- 48.6 mm</quantity>
SL).
<location id="8EB8603E8459FFB5FEDCF9720D5FA3B4" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B:8EB8603E8459FFB5FEDCF9720D5FA3B4" box="[294,355,1716,1740]" name="Same" pageId="1" pageNumber="164">Same</location>
data as holotype
</materialsCitation>
.
<materialsCitation id="3B0F3CB88459FFB5FDD7F9720E61A277" collectingDate="2015-11-12" collectionCode="MNHN" country="Papua New Guinea" location="Elnge Creek" municipality="Amick" pageId="1" pageNumber="164" specimenCode="MNHN 2018-0717" specimenCount="1" specimenCount-male="1" stateProvince="West New Britain" typeStatus="paratype">
<specimenCode id="DBC19E9E8459FFB5FDD7F9720F3FA3B4" box="[557,771,1716,1740]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="164">MNHN 2018-0717</specimenCode>
,
<specimenCount id="9D61FD6C8459FFB5FF7EF9130CF0A395" box="[132,204,1749,1773]" count="1" pageId="1" pageNumber="164" type="male">1 male</specimenCount>
(
<quantity id="4C9F9B008459FFB5FF21F9130D14A395" box="[219,296,1749,1773]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="1" pageNumber="164" unit="mm" value="47.0">47 mm</quantity>
SL),
<location id="8EB8603E8459FFB5FE9EF9130DD1A395" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B:8EB8603E8459FFB5FE9EF9130DD1A395" box="[356,493,1749,1773]" country="Papua New Guinea" municipality="Amick" name="Elnge Creek" pageId="1" pageNumber="164" stateProvince="West New Britain">Elnge Creek</location>
,
<collectingRegion id="49A3F8078459FFB5FE0DF9100E40A395" box="[503,636,1749,1773]" country="Papua New Guinea" name="West New Britain" pageId="1" pageNumber="164">New Britain</collectingRegion>
(
<collectingCountry id="F37076758459FFB5FD70F9130CE7A277" name="Papua New Guinea" pageId="1" pageNumber="164">Papua New Guinea</collectingCountry>
),
<date id="FFD910258459FFB5FF15F9310DB1A276" box="[239,397,1782,1807]" pageId="1" pageNumber="164" value="2015-11-12">
<collectingDate id="EF9DE9CD8459FFB5FF15F9310DB1A276" box="[239,397,1782,1807]" pageId="1" pageNumber="164" value="2015-11-12">12 Nov. 2015</collectingDate>
</date>
,
<collectingMunicipality id="6BBCAC9F8459FFB5FE6DF9310DD8A277" box="[407,484,1783,1807]" pageId="1" pageNumber="164">Amick</collectingMunicipality>
<emphasis id="B913EAF78459FFB5FE16F9310E23A277" box="[492,543,1783,1807]" italics="true" pageId="1" pageNumber="164">et al</emphasis>
. coll
</materialsCitation>
.
<materialsCitation id="3B0F3CB88459FFB5FD93F9310CD6A20B" collectingDate="2015-11-13" collectingDateMax="2015-11-17" collectingDateMin="2015-11-13" collectionCode="MNHN" country="Papua New Guinea" location="Kumkom Creek" municipality="Amick" pageId="1" pageNumber="164" specimenCode="MNHN 2018- 0718" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="West New Britain" typeStatus="paratype">
<specimenCode id="DBC19E9E8459FFB5FD93F9310C81A248" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="164">MNHN 2018- 0718</specimenCode>
,
<specimenCount id="9D61FD6C8459FFB5FF33F8DE0D1CA248" box="[201,288,1816,1840]" count="5" pageId="1" pageNumber="164" type="male">5 males</specimenCount>
,
<specimenCount id="9D61FD6C8459FFB5FED0F8DE0DA8A248" box="[298,404,1816,1840]" count="2" pageId="1" pageNumber="164" type="female">2 females</specimenCount>
(
<quantity id="4C9F9B008459FFB5FE58F8DE0E28A248" box="[418,532,1816,1840]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" metricValueMax="5.7" metricValueMin="3.9" pageId="1" pageNumber="164" unit="mm" value="48.0" valueMax="57.0" valueMin="39.0">39-57 mm</quantity>
SL),
<location id="8EB8603E8459FFB5FDA8F8DE0F3FA248" LSID="urn:lsid:plazi:treatment:03CE87F38459FFB7FED4FC5208C9A32B:8EB8603E8459FFB5FDA8F8DE0F3FA248" box="[594,771,1816,1840]" country="Papua New Guinea" municipality="Amick" name="Kumkom Creek" pageId="1" pageNumber="164" stateProvince="West New Britain">Kumkom Creek</location>
,
<collectingRegion id="49A3F8078459FFB5FF7EF8FC0D34A229" box="[132,264,1849,1873]" country="Papua New Guinea" name="West New Britain" pageId="1" pageNumber="164">New Britain</collectingRegion>
(
<collectingCountry id="F37076758459FFB5FEECF8FC0DD7A229" box="[278,491,1849,1874]" name="Papua New Guinea" pageId="1" pageNumber="164">Papua New Guinea</collectingCountry>
),
<date id="FFD910258459FFB5FE07F8FC0E88A229" box="[509,692,1849,1874]" pageId="1" pageNumber="164" value="2015-11-13" valueMax="2015-11-17" valueMin="2015-11-13">
<collectingDate id="EF9DE9CD8459FFB5FE07F8FC0E88A229" box="[509,692,1849,1874]" pageId="1" pageNumber="164" value="2015-11-13" valueMax="2015-11-17" valueMin="2015-11-13">13-17 Nov. 2015</collectingDate>
</date>
,
<collectingMunicipality id="6BBCAC9F8459FFB5FD44F8FF0F3BA229" box="[702,775,1849,1873]" pageId="1" pageNumber="164">Amick</collectingMunicipality>
<emphasis id="B913EAF78459FFB5FF7EF89A0C8FA20B" box="[132,179,1883,1907]" italics="true" pageId="1" pageNumber="164">et al</emphasis>
. coll
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D656E8459FFB5FCD7FBE2088AA184" pageId="1" pageNumber="164" type="diagnosis">
<paragraph id="8BD836E58459FFB5FCD7FBE20FA7A144" blockId="1.[813,923,1060,1084]" box="[813,923,1060,1084]" pageId="1" pageNumber="164">
<emphasis id="B913EAF78459FFB5FCD7FBE20FA7A144" bold="true" box="[813,923,1060,1084]" pageId="1" pageNumber="164">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD836E58459FFB5FCA8FB88088AA184" blockId="1.[813,1455,1102,1149]" lastBlockId="1.[813,1455,1198,1276]" pageId="1" pageNumber="164">
The new species has a first dorsal fin with six spines in both sexes, a second dorsal fin with one spine and ten segmented rays, an anal fin with one spine and ten segmented rays, 17 pectoral fin rays and preopercular pores M’, N and O’. Scales in lateral series 35-42, in predorsal series 0, and
<quantity id="4C9F9B008459FFB5FCD7FB220FBAA184" box="[813,902,1252,1276]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="1" pageNumber="164" unit="in" value="13.5" valueMax="15.0" valueMin="12.0">12-15 in</quantity>
transverse back scale series.
</paragraph>
</subSubSection>
<subSubSection id="C37D656E8459FFB7FCD7FAE20EA4A735" lastPageId="3" lastPageNumber="166" pageId="1" pageNumber="164" type="description">
<paragraph id="8BD836E58459FFB5FCD7FAE20F8DA044" blockId="1.[813,1455,1316,1404]" box="[813,945,1316,1340]" pageId="1" pageNumber="164">
<emphasis id="B913EAF78459FFB5FCD7FAE20F8DA044" bold="true" box="[813,945,1316,1340]" pageId="1" pageNumber="164">Description</emphasis>
</paragraph>
<paragraph id="8BD836E58459FFB5FCA8FA820910A0C4" blockId="1.[813,1455,1316,1404]" lastBlockId="1.[813,1455,1422,1501]" pageId="1" pageNumber="164">
Scale counts, number of jaw teeth, morphometrics and fin lengths in
<taxonomicName id="4C674D668459FFB5FC31FAA308FEA004" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[971,1218,1380,1404]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="164" phylum="Chordata" rank="species" species="beremeensis" status="sp. nov.">
<emphasis id="B913EAF78459FFB5FC31FAA308FEA004" box="[971,1218,1380,1404]" italics="true" pageId="1" pageNumber="164">Sicyopus beremeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A220578C8459FFB5FB30FAA3091EA004" box="[1226,1314,1381,1404]" pageId="1" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
are given in table I. Below, the
<typeStatus id="54DC88478459FFB5FC0DFA480869A0E5" box="[1015,1109,1422,1437]" pageId="1" pageNumber="164" type="holotype">holotype</typeStatus>
counts are given first, followed, in brackets if different, by the paratypes’ counts.
</paragraph>
<paragraph id="8BD836E58459FFB5FCA8FA08098CA085" blockId="1.[813,1455,1422,1501]" lastBlockId="1.[813,1456,1518,1533]" pageId="1" pageNumber="164">First dorsal fin (D1) with six spines in both sexes, second dorsal fin (D2) with one spine and ten segmented rays</paragraph>
<paragraph id="8BD836E58459FFB5FCD7F9C808DBA245" blockId="1.[813,1455,1550,1565]" lastBlockId="1.[813,1455,1774,1853]" pageId="1" pageNumber="164">(D VI-I,10). Spines 5-6 longest and slightly filamentous in males, less in females. The base of first dorsal fin is not reaching the base of second dorsal fin origin in both sexes; distance between insertion of D1 and origin of D2 is about twice the eye diameter in female, less than the eye diameter in male. The anal fin has one flexible spine and mostly 10 segmented rays (A I,10) and is directly opposite to second dorsal fin. Pectoral fin rays 16(1)-17(10). Caudal fin with 14 branched rays with a posterior margin slightly rounded. Pelvic disc with 1 spine and 5 branched rays.</paragraph>
<caption id="DF18666D845AFFB6FF7EFF1F0EAFA4A7" pageId="2" pageNumber="165" startId="2.[132,186,217,239]" targetType="table">
<paragraph id="8BD836E5845AFFB6FF7EFF1F0EAFA4A7" blockId="2.[132,776,217,479]" pageId="2" pageNumber="165">Table I. – Counts and morphomeristics of the new species. A, anal fin rays; D, dorsal fins rays; P, pectoral fin rays; C, caudal fin rays; PD, predorsal midline scales; TRB, transverse series backward scales; TRF, transverse series forward scales; ZZ, zigzag series scales; UJ, upper jaw teeth; LJ, lower jaw teeth; SL, standard Length; PDL, predorsal length (% SL); PAL, preanal length (% SL); HL, head length (% SL); JL, jaw length (% SL); CPL, caudal peduncle length (% SL); CPD, caudal peduncle depth (% SL); BDa, body depth at anus (% SL); BDD1, body depth at first dorsal fin in males (% SL); SDFL, second dorsal fin length (% SL); AFL, anal fin length (% SL); CFL, caudal fin length (% SL)).</paragraph>
</caption>
<paragraph id="8BD836E5845AFFB6FE63FE280EB7A058" pageId="2" pageNumber="165">
<table id="F967C445845A004BFF10FE280EB7A059" box="[234,651,494,1313]" gridcols="3" gridrows="25" pageId="2" pageNumber="165">
<tr id="355734A7845A004BFF10FE280EB7A77C" box="[234,651,494,516]" gridrow="0" pageId="2" pageNumber="165" rowspan-0="1">
<th id="76865DDB845A004BFE63FE280DCEA77C" box="[409,498,494,516]" gridcol="1" gridrow="0" pageId="2" pageNumber="165">Holotype</th>
<th id="76865DDB845A004BFDD4FE280EB7A77C" box="[558,651,494,516]" gridcol="2" gridrow="0" pageId="2" pageNumber="165">Paratypes</th>
</tr>
<tr id="355734A7845A004BFF10FDC90EB7A75D" box="[234,651,527,549]" gridrow="1" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FDC90D52A75D" box="[234,366,527,549]" gridcol="0" gridrow="1" pageId="2" pageNumber="165">P</th>
<td id="76865DDB845A004BFE63FDC90DCEA75D" box="[409,498,527,549]" gridcol="1" gridrow="1" pageId="2" pageNumber="165">17</td>
<td id="76865DDB845A004BFDD4FDC90EB7A75D" box="[558,651,527,549]" gridcol="2" gridrow="1" pageId="2" pageNumber="165">16-17</td>
</tr>
<tr id="355734A7845A004BFF10FDF70EB7A73F" box="[234,651,561,583]" gridrow="2" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FDF70D52A73F" box="[234,366,561,583]" gridcol="0" gridrow="2" pageId="2" pageNumber="165">D</th>
<td id="76865DDB845A004BFE63FDF70DCEA73F" box="[409,498,561,583]" gridcol="1" gridrow="2" pageId="2" pageNumber="165">VI 10</td>
<td id="76865DDB845A004BFDD4FDF70EB7A73F" box="[558,651,561,583]" gridcol="2" gridrow="2" pageId="2" pageNumber="165">VI 10</td>
</tr>
<tr id="355734A7845A004BFF10FD940EB7A71F" box="[234,651,594,615]" gridrow="3" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FD940D52A71F" box="[234,366,594,615]" gridcol="0" gridrow="3" pageId="2" pageNumber="165">A</th>
<td id="76865DDB845A004BFE63FD940DCEA71F" box="[409,498,594,615]" gridcol="1" gridrow="3" pageId="2" pageNumber="165">I10</td>
<td id="76865DDB845A004BFDD4FD940EB7A71F" box="[558,651,594,615]" gridcol="2" gridrow="3" pageId="2" pageNumber="165">I9-10</td>
</tr>
<tr id="355734A7845A004BFF10FDB50EB7A7F0" box="[234,651,627,648]" gridrow="4" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FDB50D52A7F0" box="[234,366,627,648]" gridcol="0" gridrow="4" pageId="2" pageNumber="165">C</th>
<td id="76865DDB845A004BFE63FDB50DCEA7F0" box="[409,498,627,648]" gridcol="1" gridrow="4" pageId="2" pageNumber="165">14</td>
<td id="76865DDB845A004BFDD4FDB50EB7A7F0" box="[558,651,627,648]" gridcol="2" gridrow="4" pageId="2" pageNumber="165">14</td>
</tr>
<tr id="355734A7845A004BFF10FD520EB7A7D2" box="[234,651,660,682]" gridrow="5" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FD520D52A7D2" box="[234,366,660,682]" gridcol="0" gridrow="5" pageId="2" pageNumber="165">LS</th>
<td id="76865DDB845A004BFE63FD520DCEA7D2" box="[409,498,660,682]" gridcol="1" gridrow="5" pageId="2" pageNumber="165">38</td>
<td id="76865DDB845A004BFDD4FD520EB7A7D2" box="[558,651,660,682]" gridcol="2" gridrow="5" pageId="2" pageNumber="165">35-42</td>
</tr>
<tr id="355734A7845A004BFF10FD730EB7A7B3" box="[234,651,693,715]" gridrow="6" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FD730D52A7B3" box="[234,366,693,715]" gridcol="0" gridrow="6" pageId="2" pageNumber="165">TRB</th>
<td id="76865DDB845A004BFE63FD730DCEA7B3" box="[409,498,693,715]" gridcol="1" gridrow="6" pageId="2" pageNumber="165">13</td>
<td id="76865DDB845A004BFDD4FD730EB7A7B3" box="[558,651,693,715]" gridcol="2" gridrow="6" pageId="2" pageNumber="165">12-15</td>
</tr>
<tr id="355734A7845A004BFF10FD110EB7A794" box="[234,651,727,748]" gridrow="7" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FD110D52A794" box="[234,366,727,748]" gridcol="0" gridrow="7" pageId="2" pageNumber="165">TRF</th>
<td id="76865DDB845A004BFE63FD110DCEA794" box="[409,498,727,748]" gridcol="1" gridrow="7" pageId="2" pageNumber="165">11</td>
<td id="76865DDB845A004BFDD4FD110EB7A794" box="[558,651,727,748]" gridcol="2" gridrow="7" pageId="2" pageNumber="165">11-14</td>
</tr>
<tr id="355734A7845A004BFF10FD3E0EB7A675" box="[234,651,760,781]" gridrow="8" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FD3E0D52A675" box="[234,366,760,781]" gridcol="0" gridrow="8" pageId="2" pageNumber="165">ZZ</th>
<td id="76865DDB845A004BFE63FD3E0DCEA675" box="[409,498,760,781]" gridcol="1" gridrow="8" pageId="2" pageNumber="165">11</td>
<td id="76865DDB845A004BFDD4FD3E0EB7A675" box="[558,651,760,781]" gridcol="2" gridrow="8" pageId="2" pageNumber="165">11-13</td>
</tr>
<tr id="355734A7845A004BFF10FCDF0EB7A657" box="[234,651,793,815]" gridrow="9" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FCDF0D52A657" box="[234,366,793,815]" gridcol="0" gridrow="9" pageId="2" pageNumber="165">UJ</th>
<td id="76865DDB845A004BFE63FCDF0DCEA657" box="[409,498,793,815]" gridcol="1" gridrow="9" pageId="2" pageNumber="165">6</td>
<td id="76865DDB845A004BFDD4FCDF0EB7A657" box="[558,651,793,815]" gridcol="2" gridrow="9" pageId="2" pageNumber="165">7-12</td>
</tr>
<tr id="355734A7845A004BFF10FCFC0EB7A628" box="[234,651,826,848]" gridrow="10" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FCFC0D52A628" box="[234,366,826,848]" gridcol="0" gridrow="10" pageId="2" pageNumber="165">LJ</th>
<td id="76865DDB845A004BFE63FCFC0DCEA628" box="[409,498,826,848]" gridcol="1" gridrow="10" pageId="2" pageNumber="165">6</td>
<td id="76865DDB845A004BFDD4FCFC0EB7A628" box="[558,651,826,848]" gridcol="2" gridrow="10" pageId="2" pageNumber="165">4-6</td>
</tr>
<tr id="355734A7845A004BFF10FC9D0EB7A609" box="[234,651,859,881]" gridrow="11" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FC9D0D52A609" box="[234,366,859,881]" gridcol="0" gridrow="11" pageId="2" pageNumber="165">PDL</th>
<td id="76865DDB845A004BFE63FC9D0DCEA609" box="[409,498,859,881]" gridcol="1" gridrow="11" pageId="2" pageNumber="165">32.4</td>
<td id="76865DDB845A004BFDD4FC9D0EB7A609" box="[558,651,859,881]" gridcol="2" gridrow="11" pageId="2" pageNumber="165">32.7-37.8</td>
</tr>
<tr id="355734A7845A004BFF10FCBA0EB7A6EA" box="[234,651,892,914]" gridrow="12" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FCBA0D52A6EA" box="[234,366,892,914]" gridcol="0" gridrow="12" pageId="2" pageNumber="165">PAL</th>
<td id="76865DDB845A004BFE63FCBA0DCEA6EA" box="[409,498,892,914]" gridcol="1" gridrow="12" pageId="2" pageNumber="165">58.3</td>
<td id="76865DDB845A004BFDD4FCBA0EB7A6EA" box="[558,651,892,914]" gridcol="2" gridrow="12" pageId="2" pageNumber="165">59.7-64.8</td>
</tr>
<tr id="355734A7845A004BFF10FC5B0EB7A6CC" box="[234,651,925,948]" gridrow="13" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FC5B0D52A6CC" box="[234,366,925,948]" gridcol="0" gridrow="13" pageId="2" pageNumber="165">HL</th>
<td id="76865DDB845A004BFE63FC5B0DCEA6CC" box="[409,498,925,948]" gridcol="1" gridrow="13" pageId="2" pageNumber="165">25.5</td>
<td id="76865DDB845A004BFDD4FC5B0EB7A6CC" box="[558,651,925,948]" gridcol="2" gridrow="13" pageId="2" pageNumber="165">23.6-28.7</td>
</tr>
<tr id="355734A7845A004BFF10FC790EB7A6AD" box="[234,651,959,981]" gridrow="14" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FC790D52A6AD" box="[234,366,959,981]" gridcol="0" gridrow="14" pageId="2" pageNumber="165">JL</th>
<td id="76865DDB845A004BFE63FC790DCEA6AD" box="[409,498,959,981]" gridcol="1" gridrow="14" pageId="2" pageNumber="165">13</td>
<td id="76865DDB845A004BFDD4FC790EB7A6AD" box="[558,651,959,981]" gridcol="2" gridrow="14" pageId="2" pageNumber="165">11.4-14.5</td>
</tr>
<tr id="355734A7845A004BFF10FC260EB7A68D" box="[234,651,992,1013]" gridrow="15" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FC260D52A68D" box="[234,366,992,1013]" gridcol="0" gridrow="15" pageId="2" pageNumber="165">CPD</th>
<td id="76865DDB845A004BFE63FC260DCEA68D" box="[409,498,992,1013]" gridcol="1" gridrow="15" pageId="2" pageNumber="165">10.3</td>
<td id="76865DDB845A004BFDD4FC260EB7A68D" box="[558,651,992,1013]" gridcol="2" gridrow="15" pageId="2" pageNumber="165">9-11.4</td>
</tr>
<tr id="355734A7845A004BFF10FBC70EB7A16F" box="[234,651,1025,1047]" gridrow="16" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FBC70D52A16F" box="[234,366,1025,1047]" gridcol="0" gridrow="16" pageId="2" pageNumber="165">CPL</th>
<td id="76865DDB845A004BFE63FBC70DCEA16F" box="[409,498,1025,1047]" gridcol="1" gridrow="16" pageId="2" pageNumber="165">14</td>
<td id="76865DDB845A004BFDD4FBC70EB7A16F" box="[558,651,1025,1047]" gridcol="2" gridrow="16" pageId="2" pageNumber="165">12.8-16.2</td>
</tr>
<tr id="355734A7845A004BFF10FBE40EB7A140" box="[234,651,1058,1080]" gridrow="17" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FBE40D52A140" box="[234,366,1058,1080]" gridcol="0" gridrow="17" pageId="2" pageNumber="165">BDD1</th>
<td id="76865DDB845A004BFE63FBE40DCEA140" box="[409,498,1058,1080]" gridcol="1" gridrow="17" pageId="2" pageNumber="165">12.2</td>
<td id="76865DDB845A004BFDD4FBE40EB7A140" box="[558,651,1058,1080]" gridcol="2" gridrow="17" pageId="2" pageNumber="165">12.2-15</td>
</tr>
<tr id="355734A7845A004BFF10FB820EB7A121" box="[234,651,1092,1113]" gridrow="18" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FB820D52A121" box="[234,366,1092,1113]" gridcol="0" gridrow="18" pageId="2" pageNumber="165">BDa</th>
<td id="76865DDB845A004BFE63FB820DCEA121" box="[409,498,1092,1113]" gridcol="1" gridrow="18" pageId="2" pageNumber="165">10.8</td>
<td id="76865DDB845A004BFDD4FB820EB7A121" box="[558,651,1092,1113]" gridcol="2" gridrow="18" pageId="2" pageNumber="165">10.3-14.9</td>
</tr>
<tr id="355734A7845A004BFF10FBA30EB7A103" box="[234,651,1125,1147]" gridrow="19" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FBA30D52A103" box="[234,366,1125,1147]" gridcol="0" gridrow="19" pageId="2" pageNumber="165">SDFL-male</th>
<td id="76865DDB845A004BFE63FBA30DCEA103" box="[409,498,1125,1147]" gridcol="1" gridrow="19" pageId="2" pageNumber="165">39.6</td>
<td id="76865DDB845A004BFDD4FBA30EB7A103" box="[558,651,1125,1147]" gridcol="2" gridrow="19" pageId="2" pageNumber="165">34.5-41.4</td>
</tr>
<tr id="355734A7845A004BFF10FB400EB7A1E4" box="[234,651,1158,1180]" gridrow="20" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FB400D52A1E4" box="[234,366,1158,1180]" gridcol="0" gridrow="20" pageId="2" pageNumber="165">AFL-male</th>
<td id="76865DDB845A004BFE63FB400DCEA1E4" box="[409,498,1158,1180]" gridcol="1" gridrow="20" pageId="2" pageNumber="165">34.9</td>
<td id="76865DDB845A004BFDD4FB400EB7A1E4" box="[558,651,1158,1180]" gridcol="2" gridrow="20" pageId="2" pageNumber="165">29.9-34.1</td>
</tr>
<tr id="355734A7845A004BFF10FB610EB7A1C5" box="[234,651,1191,1213]" gridrow="21" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FB610D52A1C5" box="[234,366,1191,1213]" gridcol="0" gridrow="21" pageId="2" pageNumber="165">CFL-male</th>
<td id="76865DDB845A004BFE63FB610DCEA1C5" box="[409,498,1191,1213]" gridcol="1" gridrow="21" pageId="2" pageNumber="165">25.9</td>
<td id="76865DDB845A004BFDD4FB610EB7A1C5" box="[558,651,1191,1213]" gridcol="2" gridrow="21" pageId="2" pageNumber="165">19.3-23.2</td>
</tr>
<tr id="355734A7845A004BFF10FB0E0EB7A1A6" box="[234,651,1224,1246]" gridrow="22" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FB0E0D52A1A6" box="[234,366,1224,1246]" gridcol="0" gridrow="22" pageId="2" pageNumber="165">SDFL-female</th>
<td id="76865DDB845A004BFE63FB0E0DCEA1A6" box="[409,498,1224,1246]" gridcol="1" gridrow="22" pageId="2" pageNumber="165">–</td>
<td id="76865DDB845A004BFDD4FB0E0EB7A1A6" box="[558,651,1224,1246]" gridcol="2" gridrow="22" pageId="2" pageNumber="165">26.29.8</td>
</tr>
<tr id="355734A7845A004BFF10FB2C0EB7A078" box="[234,651,1258,1280]" gridrow="23" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FB2C0D52A078" box="[234,366,1258,1280]" gridcol="0" gridrow="23" pageId="2" pageNumber="165">ALF-female</th>
<td id="76865DDB845A004BFE63FB2C0DCEA078" box="[409,498,1258,1280]" gridcol="1" gridrow="23" pageId="2" pageNumber="165">–</td>
<td id="76865DDB845A004BFDD4FB2C0EB7A078" box="[558,651,1258,1280]" gridcol="2" gridrow="23" pageId="2" pageNumber="165">27.6-30.3</td>
</tr>
<tr id="355734A7845A004BFF10FACD0EB7A059" box="[234,651,1291,1313]" gridrow="24" pageId="2" pageNumber="165">
<th id="76865DDB845A004BFF10FACD0D52A059" box="[234,366,1291,1313]" gridcol="0" gridrow="24" pageId="2" pageNumber="165">CLF-female</th>
<td id="76865DDB845A004BFE63FACD0DCEA059" box="[409,498,1291,1313]" gridcol="1" gridrow="24" pageId="2" pageNumber="165">–</td>
<td id="76865DDB845A004BFDD4FACD0EB7A059" box="[558,651,1291,1313]" gridcol="2" gridrow="24" pageId="2" pageNumber="165">22.2-22.7</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD836E5845AFFB6FF50FA8D0F3BA0DD" blockId="2.[170,775,1355,1380]" lastBlockId="2.[132,775,1397,1445]" pageId="2" pageNumber="165">Scales in lateral series 38 (35-42); scales may extend midlaterally slightly anterior to the origin of first dorsal fin in male and female, and posteriorly to the hypural base.</paragraph>
<paragraph id="8BD836E5845AFFB6FF7EFA710F3AA370" blockId="2.[132,775,1463,1478]" lastBlockId="2.[132,775,1496,1544]" pageId="2" pageNumber="165">Scales usually ctenoid from hypural base to origin of the first dorsal fin. A few cycloid scales along dorsum and anal fin base. In males, ctenoid scales on anterior body region ossi-</paragraph>
<paragraph id="8BD836E5845AFFB6FF7EF9DC085FA42A" blockId="2.[132,775,1562,1808]" lastBlockId="2.[813,1456,323,658]" pageId="2" pageNumber="165">fied, each with 3-6 cteni; ctenoid scales on medium part of body with more cteni (6-10). In females, on anterior body region, scales mostly cycloid, as on the inferior part of the flanks. Ctenoid scales each with 3-6 cteni midlaterally. Scales in zigzag series 11 (11-13), transverse back series 13 (12-15), transverse forward series 11 (11-14). Predorsal midline naked. Head, breast, pectoral base and belly naked in most specimens. Lips smooth without cleft. The tongue is free from the floor of the mouth. Upper jaw teeth in one row, mostly conical in females with fewer canines and slightly more teeth (8-12) than in males 6 (6-9), mostly caniniform. Lower jaw teeth conical in females (range 5-6) and males 6 (4-6). Labial teeth (10-14) present along the edge of lower jaw mostly enclosed in a fleshy sheath. Jaw, dorsal, caudal and anal fins longer in males.</paragraph>
<paragraph id="8BD836E5845AFFB6FCA8FE9F0F7BA749" blockId="2.[813,1456,323,658]" pageId="2" pageNumber="165">Cephalic sensory pore system A’, B, C, D, F, H’, K’, L’, M’, N and O’; D single, with all others paired, oculoscapular canal separated into anterior and posterior canals between pores H’ and K’. Cutaneous sensory papillae not well developed and very variable among specimens. Several with papillae between pores M’ and N and between pores F and D.</paragraph>
<paragraph id="8BD836E5845AFFB6FCA8FDFC0F55A7EA" blockId="2.[813,1456,323,658]" pageId="2" pageNumber="165">Urogenital papilla in male long and thin with pointed to rounded tip. Urogenital papilla in female rounded to rectangular.</paragraph>
<paragraph id="8BD836E5845AFFB6FCD7FD7F0816A7A9" blockId="2.[813,1456,697,818]" box="[813,1066,697,721]" pageId="2" pageNumber="165">
<emphasis id="B913EAF7845AFFB6FCD7FD7F0816A7A9" bold="true" box="[813,1066,697,721]" pageId="2" pageNumber="165">Colour in preservative</emphasis>
</paragraph>
<paragraph id="8BD836E5845AFFB6FCA8FD1C0993A6EA" blockId="2.[813,1456,697,818]" lastBlockId="2.[813,1456,835,914]" pageId="2" pageNumber="165">
<emphasis id="B913EAF7845AFFB6FCA8FD1C0FAEA78A" box="[850,914,730,754]" italics="true" pageId="2" pageNumber="165">Male.</emphasis>
– Background cream to tan. Anterior part of head blackish dorsally and ventrally greyish. Body with usually three vertical black bands, including the head; second band below the first dorsal fin (this band is sometimes divided in two black parts slightly paler toward centre) and third band in front of the hypural crease. Lateral midline not well-
</paragraph>
<paragraph id="8BD836E5845AFFB6FCD7FC650998A68A" blockId="2.[813,1456,931,978]" lastBlockId="2.[813,1456,995,1139]" pageId="2" pageNumber="165">marked. Nape brownish. Caudal fin rays greyish. Dorsal and anal fins greyish to blackish, with a lighter basal part. Pelvic disk, pectoral rays and pectoral fin base greyish to blackish.</paragraph>
<paragraph id="8BD836E5845AFFB6FCA8FC3C08C2A1AA" blockId="2.[813,1456,995,1139]" lastBlockId="2.[813,1456,1156,1235]" pageId="2" pageNumber="165">
<emphasis id="B913EAF7845AFFB6FCA8FC3C0F98A16A" box="[850,932,1018,1042]" italics="true" pageId="2" pageNumber="165">Female</emphasis>
. – Mostly yellowish. Head and body greyish to whitish, snout dusky. Lateral midline with a diffuse greyish subcutaneous stripe. Inferior part of body whitish. Above midline body greyish. Dorsal rays and spines greyish. Caudal fin rays greyish as is the membrane and distal margin. Anal fin whitish to greyish. Pelvic disc not pigmented. Pectoral rays and membrane greyish to whitish.
</paragraph>
<paragraph id="8BD836E5845AFFB6FCD7FB3C0825A06A" blockId="2.[813,1455,1274,1555]" box="[813,1049,1274,1298]" pageId="2" pageNumber="165">
<emphasis id="B913EAF7845AFFB6FCD7FB3C0825A06A" bold="true" box="[813,1049,1274,1298]" pageId="2" pageNumber="165">
Colour in life (
<figureCitation id="135C2A60845AFFB6FC2AFB3C082CA06A" box="[976,1040,1274,1298]" captionStart="Figure 1" captionStartId="1.[813,879,980,1002]" captionTargetBox="[813,1456,215,964]" captionTargetId="figure-734@1.[813,1456,588,964]" captionTargetPageId="1" captionText="Figure 1. – Sicyopus beremeensis. A: Male (49 mm); B: Female (45 mm) (photograph by P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711128" httpUri="https://zenodo.org/record/12711128/files/figure.png" pageId="2" pageNumber="165">Fig. 1</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BD836E5845AFFB6FCA8FADC091AA04A" blockId="2.[813,1455,1274,1555]" box="[850,1318,1306,1330]" pageId="2" pageNumber="165">The sexual dichromatism is well developed.</paragraph>
<paragraph id="8BD836E5845AFFB7FCA8FAFC0D74A46C" blockId="2.[813,1455,1274,1555]" lastBlockId="3.[132,775,217,415]" lastPageId="3" lastPageNumber="166" pageId="2" pageNumber="165">
<emphasis id="B913EAF7845AFFB6FCA8FAFC0FB0A02A" box="[850,908,1338,1362]" italics="true" pageId="2" pageNumber="165">Male</emphasis>
(
<figureCitation id="135C2A60845AFFB6FC5AFAFC0FC7A02A" box="[928,1019,1338,1362]" captionStart="Figure 1" captionStartId="1.[813,879,980,1002]" captionTargetBox="[813,1456,215,964]" captionTargetId="figure-734@1.[813,1456,588,964]" captionTargetPageId="1" captionText="Figure 1. – Sicyopus beremeensis. A: Male (49 mm); B: Female (45 mm) (photograph by P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711128" httpUri="https://zenodo.org/record/12711128/files/figure.png" pageId="2" pageNumber="165">Fig. 1A</figureCitation>
).
<emphasis id="B913EAF7845AFFB6FBEEFAFD081EA02A" box="[1044,1058,1339,1362]" italics="true" pageId="2" pageNumber="165">–</emphasis>
Head black from the snout to the nape, both dorsally and ventrally; lips black.
<collectingRegion id="49A3F807845AFFB6FAE5FA9D0954A00B" box="[1311,1384,1371,1395]" country="France" name="Centre" pageId="2" pageNumber="165">Centre</collectingRegion>
of the eye black, while eye circumference is bright orange. Body with orange and black alternating wide bands (3 black bands including the head and three orange bands). Each band visible dorsally, on the flanks and ventrally, forming large alternating rings around the body. First orange band covers the operculum and stretches to the first third of the pectoral fin. Pelvic sucker orange. Then, black band, twice as wide as the first orange band, stretches along entire base of first dorsal fin to anterior base of anal fin. Second orange band along entire length of anal fin. Third black band stretched from posterior base of anal fin to caudal peduncle. Caudal peduncle and first third of the caudal fin orange, forming the last band. Caudal fin black, slightly translucent, with a bright blue border dorsally. Pectoral fins black. Anal fin bright orange with a black border. Posterior border of scales clearly pigmented, dark orange to black on the black bands and bright orange on the orange bands. Male colour can vary in brightness according to courtship ritual.
</paragraph>
<paragraph id="8BD836E5845BFFB7FF50FED90F34A752" blockId="3.[132,775,217,415]" lastBlockId="3.[132,776,539,554]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FF50FED90CC0A44F" box="[170,252,287,311]" italics="true" pageId="3" pageNumber="166">Female</emphasis>
(
<figureCitation id="135C2A60845BFFB7FEF1FED90D61A44F" box="[267,349,287,311]" captionStart="Figure 1" captionStartId="1.[813,879,980,1002]" captionTargetBox="[813,1456,215,964]" captionTargetId="figure-734@1.[813,1456,588,964]" captionTargetPageId="1" captionText="Figure 1. – Sicyopus beremeensis. A: Male (49 mm); B: Female (45 mm) (photograph by P.Amick)." figureDoi="http://doi.org/10.5281/zenodo.12711128" httpUri="https://zenodo.org/record/12711128/files/figure.png" pageId="3" pageNumber="166">Fig. 1B</figureCitation>
).
<emphasis id="B913EAF7845BFFB7FE8FFED90DBFA44E" box="[373,387,287,310]" italics="true" pageId="3" pageNumber="166">–</emphasis>
Female colour less variable than in males. Overall body brownish to greyish. Head dark brown from snout to nape. Lips dark brown.
<collectingRegion id="49A3F807845BFFB7FDDBFEA20E56A404" box="[545,618,356,380]" country="France" name="Centre" pageId="3" pageNumber="166">Centre</collectingRegion>
of eyes black, while circumference is light brown to orange. Body brownish dorsally and rather grey on the flanks. Belly whitish from pelvic sucker to anterior base of anal fin. Black longitudinal stripe stretching from under the eye to the posterior border of the caudal fin. Pectoral fins beige-yellow and translucent.
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFDF80EA4A735" blockId="3.[132,664,574,589]" box="[132,664,574,589]" pageId="3" pageNumber="166">Caudal fin greyish translucent. Anal fin bright red.</paragraph>
</subSubSection>
<subSubSection id="C37D656E845BFFB7FF7EFDB10D56A7AC" pageId="3" pageNumber="166" type="distribution">
<paragraph id="8BD836E5845BFFB7FF7EFDB10D33A7F7" blockId="3.[132,775,631,724]" box="[132,271,631,655]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FF7EFDB10D33A7F7" bold="true" box="[132,271,631,655]" pageId="3" pageNumber="166">Distribution</emphasis>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF50FD5C0D56A7AC" blockId="3.[132,775,631,724]" pageId="3" pageNumber="166">
Currently known from freshwater streams in
<collectingRegion id="49A3F807845BFFB7FD7FFD5C0F3BA7CA" box="[645,775,666,690]" country="Papua New Guinea" name="West New Britain" pageId="3" pageNumber="166">New Britain</collectingRegion>
(
<collectingCountry id="F3707675845BFFB7FF71FD7A0D63A7AC" box="[139,351,700,724]" name="Papua New Guinea" pageId="3" pageNumber="166">Papua New Guinea</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C37D656E845BFFB7FF7EFD390D6CA20A" pageId="3" pageNumber="166" type="biology_ecology">
<paragraph id="8BD836E5845BFFB7FF7EFD390CE2A66F" blockId="3.[132,775,767,965]" box="[132,222,767,791]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FF7EFD390CE2A66F" bold="true" box="[132,222,767,791]" pageId="3" pageNumber="166">Ecology</emphasis>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF50FCE40D27A6BD" blockId="3.[132,775,767,965]" pageId="3" pageNumber="166">
<taxonomicName id="4C674D66845BFFB7FF50FCE40DA5A642" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[170,409,802,826]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="beremeensis">
<emphasis id="B913EAF7845BFFB7FF50FCE40DA5A642" box="[170,409,802,826]" italics="true" pageId="3" pageNumber="166">Sicyopus beremeensis</emphasis>
</taxonomicName>
was collected in small, rapid and boulder-strewn mountain streams with rocky bottoms at altitudes ranging between 300 and
<quantity id="4C9F9B00845BFFB7FE18FCA10E1BA607" box="[482,551,871,895]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="166" unit="m" value="900.0">900 m</quantity>
asl. It is assumed to be amphidromous as the other members of the subfamily (
<bibRefCitation id="EFF64B14845BFFB7FF76FC6A0D33A6BD" author="KEITH P." box="[140,271,940,965]" pageId="3" pageNumber="166" pagination="831 - 847" refId="ref3799" refString="KEITH P., 2003. - Biology and ecology of amphidromous Gobiidae of the Indo-Pacific and the Caribbean regions. J. Fish Biol., 63: 831 - 847." type="journal article" year="2003">Keith, 2003</bibRefCitation>
).
</paragraph>
<caption id="DF18666D845BFFB7FCD7FC660FD8A69E" ID-DOI="http://doi.org/10.5281/zenodo.12711132" ID-Zenodo-Dep="12711132" httpUri="https://zenodo.org/record/12711132/files/figure.png" pageId="3" pageNumber="166" startId="3.[813,877,928,950]" targetBox="[819,1446,226,906]" targetPageId="3" targetType="figure">
<paragraph id="8BD836E5845BFFB7FCD7FC660FD8A69E" blockId="3.[813,1456,928,998]" pageId="3" pageNumber="166">
Figure 2. – Diagrammatic illustration of the head in
<taxonomicName id="4C674D66845BFFB7FAD0FC670FB4A6B6" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="beremeensis">
<emphasis id="B913EAF7845BFFB7FAD0FC670FB4A6B6" italics="true" pageId="3" pageNumber="166">Sicyopus beremeensis</emphasis>
</taxonomicName>
showing head pores.
<emphasis id="B913EAF7845BFFB7FB9EFC7E0849A6B5" bold="true" box="[1124,1141,952,973]" pageId="3" pageNumber="166">A</emphasis>
: Dorsal view;
<emphasis id="B913EAF7845BFFB7FAF7FC7F0921A6B6" bold="true" box="[1293,1309,953,974]" pageId="3" pageNumber="166">B</emphasis>
: Lateral view. Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="8BD836E5845BFFB7FF7EFC290D2DA17F" blockId="3.[132,775,1007,1101]" box="[132,273,1007,1031]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FF7EFC290D2DA17F" bold="true" box="[132,273,1007,1031]" pageId="3" pageNumber="166">Comparison</emphasis>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF50FBD40F3AA135" blockId="3.[132,775,1007,1101]" pageId="3" pageNumber="166">
<taxonomicName id="4C674D66845BFFB7FF50FBD40DA6A152" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[170,410,1042,1066]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="beremeensis">
<emphasis id="B913EAF7845BFFB7FF50FBD40DA6A152" box="[170,410,1042,1066]" italics="true" pageId="3" pageNumber="166">Sicyopus beremeensis</emphasis>
</taxonomicName>
(D VI-I,10 A I,10, and P 17) differs from:
<taxonomicName id="4C674D66845BFFB7FF03FBF30F3AA135" authority="Watson &amp; Kottelat, 1994" authorityName="Watson &amp; Kottelat" authorityYear="1994" box="[249,774,1077,1101]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="auxilimentus">
<emphasis id="B913EAF7845BFFB7FF03FBF30DD1A135" box="[249,493,1077,1101]" italics="true" pageId="3" pageNumber="166">Sicyopus auxilimentus</emphasis>
Watson &amp; Kottelat, 1994
</taxonomicName>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFBA70F3BA1EA" blockId="3.[132,775,1121,1205]" pageId="3" pageNumber="166">
(D VI-I,9 A I,9 and P 14-15) from South
<collectingCountry id="F3707675845BFFB7FDC3FBA70E4AA108" box="[569,630,1121,1136]" name="Japan" pageId="3" pageNumber="166">Japan</collectingCountry>
to
<collectingCountry id="F3707675845BFFB7FD62FBA70F38A108" box="[664,772,1121,1136]" name="Indonesia" pageId="3" pageNumber="166">Indonesia</collectingCountry>
;
<taxonomicName id="4C674D66845BFFB7FF7EFBBC0EBBA1EA" authority="Keith, Marquet &amp; Taillebois, 2011" authorityName="Keith, Marquet &amp; Taillebois" authorityYear="2011" box="[132,647,1146,1170]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="lord">
<emphasis id="B913EAF7845BFFB7FF7EFBBC0D28A1EA" box="[132,276,1146,1170]" italics="true" pageId="3" pageNumber="166">Sicyopus lord</emphasis>
<bibRefCitation id="EFF64B14845BFFB7FEE1FB450EBBA1EA" author="KEITH P. &amp; MARQUET G. &amp; TAILLEBOIS L." box="[283,647,1155,1170]" pageId="3" pageNumber="166" pagination="43 - 44" refId="ref3890" refString="KEITH P., MARQUET G. &amp; TAILLEBOIS L., 2011. - Discovery of the freshwater genus Sicyopus (Teleostei: Gobioidei: Sicydiinae) in Madagascar, with a description of a new species and comments about regional dispersal. J. Nat. Hist., 45: 43 - 44, 2725 - 2746." type="journal article" year="2011">Keith, Marquet &amp; Taillebois, 2011</bibRefCitation>
</taxonomicName>
(D VI-I,9 A
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFB5B0F3BA1CD" blockId="3.[132,775,1121,1205]" box="[132,775,1181,1205]" pageId="3" pageNumber="166">
I,9 and P 15) from
<collectingCountry id="F3707675845BFFB7FEB5FB5B0DE8A1CD" box="[335,468,1181,1205]" name="Madagascar" pageId="3" pageNumber="166">Madagascar</collectingCountry>
;
<taxonomicName id="4C674D66845BFFB7FE25FB5B0F3BA1CD" authority="Keith, Hadiaty, Busson &amp; Hubert, 2014" authorityName="Keith" authorityYear="2014" box="[479,775,1181,1205]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="rubicundus">
<emphasis id="B913EAF7845BFFB7FE25FB5B0E82A1CD" box="[479,702,1181,1205]" italics="true" pageId="3" pageNumber="166">Sicyopus rubicundus</emphasis>
<bibRefCitation id="EFF64B14845BFFB7FD3EFB5B0F3BA1CD" author="KEITH P. &amp; HADIATY R. &amp; BUSSON F. &amp; HUBERT N." box="[708,775,1181,1205]" pageId="3" pageNumber="166" pagination="173 - 178" refId="ref3951" refString="KEITH P., HADIATY R., BUSSON F. &amp; HUBERT N., 2014. - A new species of Sicyopus (Teleostei: Gobioidei) from Indonesia. Cybium, 38 (3): 173 - 178." type="journal article" year="2014">Keith,</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFB0F0F3BA182" blockId="3.[132,775,1225,1274]" pageId="3" pageNumber="166">
Hadiaty, Busson &amp; Hubert, 2014 (D VI-I,9 A I,9 and P 15) from Sunda Islands;
<taxonomicName id="4C674D66845BFFB7FE9AFB240F3BA182" authority="(Bleeker, 1856)" baseAuthorityName="Bleeker" baseAuthorityYear="1856" box="[352,775,1250,1274]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="zosterophorus">
<emphasis id="B913EAF7845BFFB7FE9AFB240E67A182" box="[352,603,1250,1274]" italics="true" pageId="3" pageNumber="166">Sicyopus zosterophorus</emphasis>
(Bleeker, 1856)
</taxonomicName>
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFAC80F3BA038" blockId="3.[132,775,1294,1656]" pageId="3" pageNumber="166">
(D VI-I,9 A I,9-10 and P 15-16) widespread in the Pacific;
<taxonomicName id="4C674D66845BFFB7FF7EFAEE0E1AA038" authority="Watson, 1995" authorityName="Watson" authorityYear="1995" box="[132,550,1319,1344]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="discordipinnis">
<emphasis id="B913EAF7845BFFB7FF7EFAEE0DB5A047" box="[132,393,1319,1343]" italics="true" pageId="3" pageNumber="166">Sicyopus discordipinnis</emphasis>
<bibRefCitation id="EFF64B14845BFFB7FE75FAF70E1AA038" author="WATSON R. E." box="[399,550,1329,1344]" pageId="3" pageNumber="166" pagination="267 - 278" refId="ref4231" refString="WATSON R. E., 1995. - A new species of Sicyopus from Papua New Guinea with redescription of S. multisquamatus (Teleostei: Gobioidei: Sicydiinae). Ichthyol. Expl. Freshw., 6 (3): 267 - 278." type="journal article" year="1995">Watson, 1995</bibRefCitation>
</taxonomicName>
(D V-I,9 A I,9 and P
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EFA8C0ED8A0DF" blockId="3.[132,775,1294,1656]" pageId="3" pageNumber="166">
mostly 15) from
<collectingCountry id="F3707675845BFFB7FEBAFA8C0D90A01A" box="[320,428,1354,1378]" name="Indonesia" pageId="3" pageNumber="166">Moluccas</collectingCountry>
to Solomon; and
<taxonomicName id="4C674D66845BFFB7FD8BFA8D0D82A0FD" authority="Klausewitz &amp; Henrich" authorityName="Klausewitz &amp; Henrich" authorityYear="1986" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="jon-onklaasi">
<emphasis id="B913EAF7845BFFB7FD8BFA8D0CF8A0FD" italics="true" pageId="3" pageNumber="166">Sicyopus jon-onklaasi</emphasis>
Klausewitz &amp; Henrich
</taxonomicName>
,
<superScript id="7C129BAD845BFFB7FE33FAAB0E3BA003" attach="left" box="[457,519,1386,1413]" fontSize="7" pageId="3" pageNumber="166">19861</superScript>
(D V-I,9 A I,9-10 and P mostly 16) from
<collectingCountry id="F3707675845BFFB7FECCFA560D9EA0D0" box="[310,418,1424,1448]" name="Sri Lanka" pageId="3" pageNumber="166">Sri Lanka</collectingCountry>
. See
<bibRefCitation id="EFF64B14845BFFB7FE2DFA560EA8A0D0" author="KEITH P. &amp; HADIATY R. &amp; BUSSON F. &amp; HUBERT N." box="[471,660,1424,1448]" pageId="3" pageNumber="166" pagination="173 - 178" refId="ref3951" refString="KEITH P., HADIATY R., BUSSON F. &amp; HUBERT N., 2014. - A new species of Sicyopus (Teleostei: Gobioidei) from Indonesia. Cybium, 38 (3): 173 - 178." type="journal article" year="2014">
Keith
<emphasis id="B913EAF7845BFFB7FDE2FA560E7BA0D0" box="[536,583,1424,1448]" italics="true" pageId="3" pageNumber="166">et al</emphasis>
. (2014
</bibRefCitation>
,
<bibRefCitation id="EFF64B14845BFFB7FD64FA490EE4A0DF" author="KEITH P. &amp; LORD C. &amp; MAEDA K." box="[670,728,1423,1447]" pageId="3" pageNumber="166" refId="ref3994" refString="KEITH P., LORD C. &amp; MAEDA K., 2015. - Indo-Pacific Sicydiine Gobies: Biodiversity, Life Traits and Conservation. Societe Francaise d'Ichtyologie (SFI), Paris, 256 p." type="book" year="2015">2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BD836E5845BFFB7FF50FA750D38A300" blockId="3.[132,775,1294,1656]" pageId="3" pageNumber="166">
<taxonomicName id="4C674D66845BFFB7FF50FA750D9DA0B2" authorityName="Keith &amp; Amick &amp; Toko &amp; Lord" authorityYear="2019" box="[170,417,1458,1482]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="beremeensis">
<emphasis id="B913EAF7845BFFB7FF50FA750D9DA0B2" box="[170,417,1458,1482]" italics="true" pageId="3" pageNumber="166">Sicyopus beremeensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C674D66845BFFB7FDC2FA750F3BA0B2" authorityName="de Beaufort" authorityYear="1912" box="[568,775,1458,1482]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="multisquamatus">
<emphasis id="B913EAF7845BFFB7FDC2FA750F3BA0B2" box="[568,775,1458,1482]" italics="true" pageId="3" pageNumber="166">S. multisquamatus</emphasis>
</taxonomicName>
in having fewer scales in lateral series (35-42
<emphasis id="B913EAF7845BFFB7FD8EFA100EB6A095" box="[628,650,1494,1517]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 47), transverse back series (12-15
<emphasis id="B913EAF7845BFFB7FE6DFA3E0D92A377" box="[407,430,1528,1551]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 16) and predorsal series (0
<emphasis id="B913EAF7845BFFB7FD13FA3E0F3CA377" box="[745,768,1528,1551]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 4), smaller predorsal length (32-38
<emphasis id="B913EAF7845BFFB7FDFCF9DD0E20A34A" box="[518,540,1563,1586]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 40 %SL) and caudal peduncle length (13-16
<emphasis id="B913EAF7845BFFB7FE7FF9F80DA7A32D" box="[389,411,1598,1621]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 18) and fewer teeth in lower jaw (4-6
<emphasis id="B913EAF7845BFFB7FF4DF9A70CF1A300" box="[183,205,1633,1656]" italics="true" pageId="3" pageNumber="166">vs</emphasis>
. 10).
</paragraph>
<paragraph id="8BD836E5845BFFB7FF7EF9660D6CA20A" blockId="3.[132,775,1696,1907]" pageId="3" pageNumber="166">
<superScript id="7C129BAD845BFFB7FF7EF9660CB0A3D6" attach="left" box="[132,140,1696,1710]" fontSize="6" pageId="3" pageNumber="166">1</superScript>
Note concerning
<taxonomicName id="4C674D66845BFFB7FEC5F9640DCEA3CF" authorityName="Klausewizt &amp; Henrich" authorityYear="1986" box="[319,498,1698,1719]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="jonklaasi">
<emphasis id="B913EAF7845BFFB7FEC5F9640DCEA3CF" box="[319,498,1698,1719]" italics="true" pageId="3" pageNumber="166">Sicyopus jonklaasi</emphasis>
</taxonomicName>
: in the description of
<emphasis id="B913EAF7845BFFB7FD3BF9640CC5A3AA" italics="true" pageId="3" pageNumber="166">
<taxonomicName id="4C674D66845BFFB7FD3BF9640CCAA3AA" authorityName="Axelrod" authorityYear="1972" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="jonklaasi">Gobius jonklaasi</taxonomicName>
,
</emphasis>
the text and pictures of
<bibRefCitation id="EFF64B14845BFFB7FE21F97B0E53A3AA" author="AXELROD H. R." box="[475,623,1725,1747]" pageId="3" pageNumber="166" refId="ref3692" refString="AXELROD H. R. et al. (eds), 1972. - Exotic Tropical Fishes. Looseleaf edition. TFH Publications, Jersey City, NJ. Insert No. 168. Fish F- 300.10 - 13 (4 p.)." type="book" year="1972">Axelrod (1972)</bibRefCitation>
(pages F300.10 to 13) clearly refers to
<taxonomicName id="4C674D66845BFFB7FE87F91E0E46A394" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[381,634,1751,1773]" family="Gobiidae" genus="Sicyopterus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="lagocephalus">
<emphasis id="B913EAF7845BFFB7FE87F91E0E46A394" box="[381,634,1751,1773]" italics="true" pageId="3" pageNumber="166">Sicyopterus lagocephalus</emphasis>
</taxonomicName>
, which would suggest
<taxonomicName id="4C674D66845BFFB7FF16F9340D99A27F" authorityName="Klausewizt &amp; Henrich" authorityYear="1986" box="[236,421,1778,1799]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="jonklaasi">
<emphasis id="B913EAF7845BFFB7FF16F9340D99A27F" box="[236,421,1778,1799]" italics="true" pageId="3" pageNumber="166">Sicyopus jonklaasi</emphasis>
</taxonomicName>
is a different species and should be attributed to Klausewizt and Henrich, 1986 and not to
<bibRefCitation id="EFF64B14845BFFB7FD4CF8CB0CE2A245" author="AXELROD H. R." pageId="3" pageNumber="166" refId="ref3692" refString="AXELROD H. R. et al. (eds), 1972. - Exotic Tropical Fishes. Looseleaf edition. TFH Publications, Jersey City, NJ. Insert No. 168. Fish F- 300.10 - 13 (4 p.)." type="book" year="1972">Axelrod (1972)</bibRefCitation>
. So,
<taxonomicName id="4C674D66845BFFB7FEF7F8EE0EE7A245" authority="Klausewizt &amp; Henrich, 1986" authorityName="Klausewizt &amp; Henrich" authorityYear="1986" box="[269,731,1831,1853]" family="Gobiidae" genus="Sicyopus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="166" phylum="Chordata" rank="species" species="jonklaasi">
<emphasis id="B913EAF7845BFFB7FEF7F8EE0DFDA244" box="[269,449,1831,1853]" italics="true" pageId="3" pageNumber="166">Sicyopus jonklaasi</emphasis>
Klausewizt &amp; Henrich, 1986
</taxonomicName>
was not incorrectly treated as valid by several authors as stated by
<bibRefCitation id="EFF64B14845BFFB7FF61F89B0D77A20A" author="FRICKE R. &amp; ESCHMEYER W. N. &amp; VAN DER LAAN R." box="[155,331,1885,1907]" pageId="3" pageNumber="166" refId="ref3738" refString="FRICKE R., ESCHMEYER W. N. &amp; VAN DER LAAN R. (eds), 2018. - Catalog of Fishes: Genera, Species, References. (http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp). Electronic version accessed 01 Nov. 2018." type="url" year="2018">
Fricke
<emphasis id="B913EAF7845BFFB7FF24F89B0D3BA20A" box="[222,263,1885,1906]" italics="true" pageId="3" pageNumber="166">et al</emphasis>
., 2018
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37D656E845BFFB7FCD7FB9A08C9A32B" pageId="3" pageNumber="166" type="etymology">
<paragraph id="8BD836E5845BFFB7FCD7FB9A0F94A10C" blockId="3.[813,1455,1116,1236]" box="[813,936,1116,1140]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FCD7FB9A0F94A10C" bold="true" box="[813,936,1116,1140]" pageId="3" pageNumber="166">Etymology</emphasis>
</paragraph>
<paragraph id="8BD836E5845BFFB7FCA8FBBA0F49A1AC" blockId="3.[813,1455,1116,1236]" pageId="3" pageNumber="166">The new species is named after Bereme village, to thank the local community for its warm welcome during our field- works.</paragraph>
<paragraph id="8BD836E5845BFFB7FCD7FAC308C9A32B" blockId="3.[813,1456,1285,1619]" pageId="3" pageNumber="166">
<emphasis id="B913EAF7845BFFB7FCD7FAC30FC4A063" bold="true" box="[813,1016,1285,1307]" pageId="3" pageNumber="166">Acknowledgements</emphasis>
. – We wish to thank Joe Penny and Brendan Kilme of Ainbul Village in the Whiteman Range. The New
<collectingCountry id="F3707675845BFFB7FA8CFAE00F7FA033" name="Guinea-Bissau" pageId="3" pageNumber="166">Guinea</collectingCountry>
Binatang Research
<collectingRegion id="49A3F807845BFFB7FBF4FAF0086DA033" box="[1038,1105,1334,1355]" country="France" name="Centre" pageId="3" pageNumber="166">Centre</collectingRegion>
(BRC) provided logistical support throughout the 2015 survey, and fieldwork was financed by UNDP GEF-SGP. The present study and the 2018 trip were funded by the MNHN (UMR 7208 BOREA), the French Ichthyological Society (SFI) and the Fondation de
<collectingCountry id="F3707675845BFFB7FBC9FA50084BA0D3" box="[1075,1143,1430,1451]" name="France" pageId="3" pageNumber="166">France</collectingCountry>
. Thanks to G. Kaipu (
<collectingCountry id="F3707675845BFFB7FABDFA500945A0D3" box="[1351,1401,1430,1451]" name="Papua New Guinea" pageId="3" pageNumber="166">PNG</collectingCountry>
NRI) for the research permit, N. Gowep (CEPA) for the export permit and our friends B. Ruli (Live &amp; Learn), J. Anamiato (National Museum and Art gallery of
<collectingCountry id="F3707675845BFFB7FBC5FA20084EA08C" box="[1087,1138,1510,1524]" name="Papua New Guinea" pageId="3" pageNumber="166">PNG</collectingCountry>
) and D. Vaghelo (WNB Provin- cial Govt, Environment section) for their help during the 2018 field support. Finally, we thank for the loan of specimens: G. Moore, M. Allen, S. Morrison (WAM); F. Krupp, S. Dorow (SMF); R. de Ruiter (RMNH); R. Causse, Z. Gabsi (MNHN).
</paragraph>
</subSubSection>
</treatment>
</document>
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