treatments-xml/data/03/CE/87/03CE87C3FFD2E169FF41FB126CB2EDAF.xml

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<mods:title id="C5BE70AC9D9F52F553A3E3E9FCF81D78">Begonia benitotanii (section Petermannia, Begoniaceae) a new species endemic to the Philippine island of Bucas Grande</mods:title>
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<mods:namePart id="60A8FB637D1994482667399D4EAA102E">Rubite, Rosario Rivera</mods:namePart>
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<mods:affiliation id="200C0D68B2807EE1329D39A39C4C8095">Department of Biological Sciences, College of Science; Research Center for the Natural and Applied Sciences, Thomas Aquinas Research Center, University of Santo Tomas, España Boulevard, Sampaloc, Manila 1015, Philippines &amp; cibanag @ ust. edu. phmailto; https: // orcid. org / 0000 - 0003 - 1490 - 2702</mods:affiliation>
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<emphasis id="B913EAC7FFD2E16EFF41FB126C64E851" bold="true" box="[136,364,1556,1583]" italics="true" pageId="2" pageNumber="259">Begonia benitotanii</emphasis>
Rubite, Tandang &amp; C.W. Lin
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<emphasis id="B913EAC7FFD2E16EFD13FB126E23E850" bold="true" box="[730,811,1556,1582]" italics="true" pageId="2" pageNumber="259">sp. nov</emphasis>
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(
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&amp; 
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elevation 
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1003
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Monoecious, perennial, glabrous herb. 
<emphasis id="B913EAC7FFD2E16EFDFEFBCE6F7CE89C" bold="true" box="[567,628,1736,1762]" pageId="2" pageNumber="259">Stem</emphasis>
erect, 
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tall, 
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in diameter, dark green some becoming brown, internodes 
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long, nodes slightly swollen. 
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deciduous, green, oblong, 25–30 × 
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, slightly keeled, glabrous, margin entire, apex mucronate. 
<emphasis id="B913EAC7FFD2E16EFD63FA166E0EE954" bold="true" box="[682,774,1808,1834]" pageId="2" pageNumber="259">Petioles</emphasis>
green, terete, 
<quantity id="4C9F9B30FFD2E16EFC6BFA166EFCE955" box="[930,1012,1808,1835]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="2" pageNumber="259" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 cm</quantity>
long, 
<emphasis id="B913EAC7FFD2E16EFBF0FA176952E954" box="[1081,1114,1809,1834]" italics="true" pageId="2" pageNumber="259">ca.</emphasis>
<quantity id="4C9F9B30FFD2E16EFBA9FA1669BDE955" box="[1120,1205,1808,1835]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="259" unit="mm" value="2.5">2.5 mm</quantity>
in diameter, glabrous. 
<emphasis id="B913EAC7FFD2E16EFF41FA326DD2E930" bold="true" box="[136,218,1844,1870]" pageId="2" pageNumber="259">Leaves</emphasis>
glabrous, obovately oblong, asymmetric, base obliquely cordate, apex acuminate, margin irregularly serrate and undulate, 8–21 × 
<quantity id="4C9F9B30FFD2E16EFEB3FA5E6CD7E90D" box="[378,479,1880,1907]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="2" pageNumber="259" unit="cm" value="9.0" valueMax="12.0" valueMin="6.0">6–12 cm</quantity>
; adaxially uniformly emerald green, sometimes with sparse, large, white spots between veins, veins sulcate; abaxially pale green to pinkish, veins emergent; venation palmate-pinnate, primary veins 7 or 8. 
<emphasis id="B913EAC7FFD2E16EFF41FAA66C2EE9C4" bold="true" box="[136,294,1952,1978]" pageId="2" pageNumber="259">Inflorescence</emphasis>
terminal opposite a leaf, separate inflorescence between male and female; female inflorescence with short peduncle (
<quantity id="4C9F9B30FFD2E16EFEF6FAC26CA5E9A1" box="[319,429,1988,2015]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="2" pageNumber="259" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
) flowers basal, borne in pairs; male inflorescence with longer peduncle (
<quantity id="4C9F9B30FFD2E16EFB26FAC2684FE9A1" box="[1263,1351,1988,2015]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="259" unit="cm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 cm</quantity>
) flowers apical on short cymes branching 3–4 times; protogynous. 
<emphasis id="B913EAC7FFD2E16EFCD5FAEE6E62E67C" bold="true" box="[796,874,2024,2050]" pageId="2" pageNumber="259">Bracts</emphasis>
deciduous, light green, basal pair lanceolate, 9–11 × 
<quantity id="4C9F9B30FFD2E16EFF69F50A6C08E659" box="[160,256,2060,2087]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="2" pageNumber="259" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, margin entire. 
<emphasis id="B913EAC7FFD2E16EFE78F50A6F76E658" bold="true" box="[433,638,2060,2086]" pageId="2" pageNumber="259">Staminate flower</emphasis>
: pedicel 
<quantity id="4C9F9B30FFD2E16EFD2EF50A6E4DE659" box="[743,837,2060,2087]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="259" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
long; tepals 2, pink, orbicular, 4–5 × 
<quantity id="4C9F9B30FFD2E16EFB39F50A6858E659" box="[1264,1360,2060,2087]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="259" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
, margin entire, apex rounded; androecium actinomorphic; stamens 15–20, golden yellow; filaments 
<quantity id="4C9F9B30FFD3E16FFBB7FD9869F9EEC7" box="[1150,1265,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="3" pageNumber="260" unit="mm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 mm</quantity>
, slightly fused at base; anthers very widely obovate, apex truncate to round, 
<emphasis id="B913EAC7FFD3E16FFCD6FDC56E48EEA2" box="[799,832,195,220]" italics="true" pageId="3" pageNumber="260">ca.</emphasis>
<quantity id="4C9F9B30FFD3E16FFC8EFDC46EA9EEA3" box="[839,929,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="260" unit="mm" value="1.5">1.5 mm</quantity>
. 
<emphasis id="B913EAC7FFD3E16FFC65FDC4696FEEA2" bold="true" box="[940,1127,194,220]" pageId="3" pageNumber="260">Pistillate flower</emphasis>
: pedicel 
<quantity id="4C9F9B30FFD3E16FFB07FDC4682AEEA3" box="[1230,1314,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="3" pageNumber="260" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
long; tepals 5, pink, obovate to elliptic, 11–15 × 
<quantity id="4C9F9B30FFD3E16FFDE4FDE06F93EF7F" box="[557,667,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="3" pageNumber="260" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
, margin entire, apex acute; ovary light green to pink, elliptic, 10–14 × 
<quantity id="4C9F9B30FFD3E16FFF68FC0C6C05EF5B" box="[161,269,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="3" pageNumber="260" unit="mm" value="10.0" valueMax="11.0" valueMin="9.0">9–11 mm</quantity>
(wings excluded), 3-locular, placentation axile; 3-winged surrounding and extending beyond the ovary, wings pink, equal; styles 3, golden yellow, 
<emphasis id="B913EAC7FFD3E16FFDAAFC296F8CEF36" box="[611,644,303,328]" italics="true" pageId="3" pageNumber="260">ca.</emphasis>
<quantity id="4C9F9B30FFD3E16FFD43FC286FC1EF37" box="[650,713,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="260" unit="mm" value="3.0">3 mm</quantity>
long, apically 2-cleft; stigmas in a spiral band. 
<emphasis id="B913EAC7FFD3E16FFB1BFC28683AEF36" bold="true" box="[1234,1330,302,328]" pageId="3" pageNumber="260">Capsule</emphasis>
trigonouselliptic, 30–38 × 
<quantity id="4C9F9B30FFD3E16FFE8CFC546CBCEF13" box="[325,436,338,365]" metricMagnitude="-1" metricUnit="m" metricValue="2.8" metricValueMax="3.6" metricValueMin="2.0" pageId="3" pageNumber="260" unit="cm" value="28.0" valueMax="36.0" valueMin="20.0">20–36 cm</quantity>
including wings; pedicel 
<quantity id="4C9F9B30FFD3E16FFD1BFC546E2DEF13" box="[722,805,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="260" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 cm</quantity>
long; wings 3, equal, truncate to rounded distally, cuneate proximally, 
<quantity id="4C9F9B30FFD3E16FFEC6FC706C80EFEF" box="[271,392,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="3" pageNumber="260" unit="mm" value="34.0" valueMax="38.0" valueMin="30.0">30–38 mm</quantity>
long, 
<quantity id="4C9F9B30FFD3E16FFE07FC706F4FEFEF" box="[462,583,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.8" metricValueMin="1.0" pageId="3" pageNumber="260" unit="mm" value="14.0" valueMax="18.0" valueMin="10.0">10–18 mm</quantity>
wide.
</paragraph>
</subSubSection>
<caption id="DF18665DFFD3E16FFF41FAB96EF1E663" ID-DOI="http://doi.org/10.5281/zenodo.5312594" ID-Zenodo-Dep="5312594" httpUri="https://zenodo.org/record/5312594/files/figure.png" pageId="3" pageNumber="260" startId="3.[136,229,1983,2005]" targetBox="[277,1314,439,1944]" targetPageId="3">
<paragraph id="8BD836D5FFD3E16FFF41FAB96EF1E663" blockId="3.[136,1452,1983,2077]" pageId="3" pageNumber="260">
<emphasis id="B913EAC7FFD3E16FFF41FAB96DFFE9AA" bold="true" box="[136,247,1983,2005]" pageId="3" pageNumber="260">FIGURE 2</emphasis>
. 
<taxonomicName id="4C674D56FFD3E16FFECBFAB96E41E9AB" authority="Rubite, Tandang &amp; C. W. Lin. A. Habit" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[258,841,1983,2005]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="260" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD3E16FFECBFAB96FE2E9AB" bold="true" box="[258,746,1983,2005]" pageId="3" pageNumber="260">
<emphasis id="B913EAC7FFD3E16FFECBFAB96CCDE9AB" bold="true" box="[258,453,1983,2005]" italics="true" pageId="3" pageNumber="260">Begonia benitotanii</emphasis>
Rubite, Tandang &amp; C.W.Lin
</emphasis>
. A. Habit
</taxonomicName>
; B. Stipule; C-C''. Bracts; D, D'. Staminate flower, face and side views; E, E'. Stamens, dorsal and ventral views; F, F'. Pistillate flower, face and side views; G, G'. Style and stigmatic band, ventral and dorsal views; H. Capsule; I. Cross section of an immature capsule. All from 
<emphasis id="B913EAC7FFD3E16FFCADF50E6EFBE662" box="[868,1011,2055,2077]" italics="true" pageId="3" pageNumber="260">R. Rubite 1003</emphasis>
.
</paragraph>
</caption>
<caption id="DF18665DFFD4E168FF41FA896DB6E993" ID-DOI="http://doi.org/10.5281/zenodo.5312598" ID-Zenodo-Dep="5312598" httpUri="https://zenodo.org/record/5312598/files/figure.png" pageId="4" pageNumber="261" startId="4.[136,229,1935,1957]" targetBox="[151,1436,190,1911]" targetPageId="4">
<paragraph id="8BD836D5FFD4E168FF41FA896DB6E993" blockId="4.[136,1452,1935,2029]" pageId="4" pageNumber="261">
<emphasis id="B913EAC7FFD4E168FF41FA896C08E9DA" bold="true" box="[136,256,1935,1957]" pageId="4" pageNumber="261">FIGURE 3.</emphasis>
<taxonomicName id="4C674D56FFD4E168FEC1FA896E77E9DB" authority="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[264,895,1935,1957]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="261" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD4E168FEC1FA896FF7E9DB" bold="true" box="[264,767,1935,1957]" pageId="4" pageNumber="261">
<emphasis id="B913EAC7FFD4E168FEC1FA896CC6E9DB" bold="true" box="[264,462,1935,1957]" italics="true" pageId="4" pageNumber="261">Begonia benitotanii</emphasis>
Rubite, Tandang &amp; C.W.Lin
</emphasis>
. A, B. Habit
</taxonomicName>
and habitats; C. Stipule; D. Inflorescence; E. Staminate flower; F. Pistillate flower, face and side views; G. Immature capsules; H. Cross section of an immature capsule. All from 
<emphasis id="B913EAC7FFD4E168FA99FAB26DB0E993" italics="true" pageId="4" pageNumber="261">R. Rubite 1003</emphasis>
.
</paragraph>
</caption>
<subSubSection id="C37D655EFFD5E169FF74FD986EFEEEA2" pageId="5" pageNumber="262" type="etymology">
<paragraph id="8BD836D5FFD5E169FF74FD986EFEEEA2" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="262">
<emphasis id="B913EAC7FFD5E169FF74FD986C6CEEC6" bold="true" box="[189,356,158,184]" pageId="5" pageNumber="262">Etymology:—</emphasis>
This species is named in honor of Professor Benito C. Tan, a distinguished Filipino muscologist, who inspired a lot of students from different parts of the world to study plants.
</paragraph>
</subSubSection>
<subSubSection id="C37D655EFFD5E169FF74FDE06CB2EDAF" pageId="5" pageNumber="262" type="distribution">
<paragraph id="8BD836D5FFD5E169FF74FDE06C21EF36" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="262">
<emphasis id="B913EAC7FFD5E169FF74FDE06F07EF7E" bold="true" box="[189,527,230,256]" pageId="5" pageNumber="262">Distribution and habitat:—</emphasis>
<taxonomicName id="4C674D56FFD5E169FDC6FDE16FF9EF7E" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[527,753,230,256]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD5E169FDC6FDE16FF9EF7E" box="[527,753,230,256]" italics="true" pageId="5" pageNumber="262">Begonia benitotanii</emphasis>
</taxonomicName>
is endemic to 
<collectingRegion id="49A3F837FFD5E169FC6EFDE069E0EF7E" box="[935,1256,230,257]" country="Philippines" name="Surigao del Norte" pageId="5" pageNumber="262">Surigao del Norte Province</collectingRegion>
, Caraga Region, Mindanao, where it is currently known only from its 
<typeStatus id="54DC8877FFD5E169FD21FC0D6E10EF5B" box="[744,792,267,293]" pageId="5" pageNumber="262">type</typeStatus>
locality. The plants grow on soil slopes with coconut and banana plants.
</paragraph>
<paragraph id="8BD836D5FFD5E169FF74FC546E8BEC17" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="262">
<emphasis id="B913EAC7FFD5E169FF74FC546C2EEF12" bold="true" box="[189,294,338,364]" pageId="5" pageNumber="262">Notes:—</emphasis>
<taxonomicName id="4C674D56FFD5E169FEEFFC556CB6EF12" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[294,446,338,364]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD5E169FEEFFC556CB6EF12" box="[294,446,338,364]" italics="true" pageId="5" pageNumber="262">B. benitotanii</emphasis>
</taxonomicName>
is similar to 
<taxonomicName id="4C674D56FFD5E169FD98FC556E00EF12" box="[593,776,338,364]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="balangcodiae">
<emphasis id="B913EAC7FFD5E169FD98FC556E00EF12" box="[593,776,338,364]" italics="true" pageId="5" pageNumber="262">B. balangcodiae</emphasis>
</taxonomicName>
in having upright stems, adaxial leaf green, sometimes with sparse, large, white spots, two-tepaled staminate flowers and five-tepaled pistillate flowers, and capsule with 3 equal wings. However, the new species differs in having longer stems, petioles, pistillate flower pedicels, larger lamina, stipules, ovaries and capsules; stipules oblong (vs. ovate-triangular); leaf shape obovately oblong (vs. lanceolate) apex acuminate (vs. acute), adaxial leaf green along the veins, veins sulcate (vs. reddish along veins, veins flat), abaxial leaf pale green to pinkish, emergent veins (vs. crimson, flat veins); pink, orbicular staminate tepals, 4–5 × 
<quantity id="4C9F9B30FFD5E169FA84FF0068A4EC5F" box="[1357,1452,518,545]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="5" pageNumber="262" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
(vs. white, sometimes greenish at apex, ovate, 12–18 × 
<quantity id="4C9F9B30FFD5E169FD26FF2C6E63EC3B" box="[751,875,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="262" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
); pink wings surrounding and extending beyond the ovary (vs. reddish, does not surround and extend beyond the ovary).
</paragraph>
<paragraph id="8BD836D5FFD5E169FF74FF746CB2EDAF" blockId="5.[136,1452,158,977]" pageId="5" pageNumber="262">
<emphasis id="B913EAC7FFD5E169FF74FF746CBEECF2" bold="true" box="[189,438,626,652]" pageId="5" pageNumber="262">Conservation status:</emphasis>
—In 2011, there were two populations observed, each of about 300 individuals in a 
<quantity id="4C9F9B30FFD5E169FA48FF746D96ECCF" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="262" unit="m" value="100.0">100 m</quantity>
area. During our recent fieldwork, one population was found totally decimated while the other was reduced to just about 100 individuals. The site is being converted into coconut, banana and rice plantations. The area is not currently protected under the country’s National Integrated Protected Areas System by the Department of Environment and Natural Resources. These observations of 
<taxonomicName id="4C674D56FFD5E169FD93FE056FF9ED62" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[602,753,770,796]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD5E169FD93FE056FF9ED62" box="[602,753,770,796]" italics="true" pageId="5" pageNumber="262">B. benitotanii</emphasis>
</taxonomicName>
satisfy the IUCN Red List CR (Critically Endangered) Criteria B1ab(ii,iii,v)+2ab(i,ii,iii,v) (
<bibRefCitation id="EFF64B24FFD5E169FE0FFE206F51ED3E" author="IUCN Standards &amp; Petitions Committee" box="[454,601,806,832]" pageId="5" pageNumber="262" refId="ref2834" refString="IUCN Standards and Petitions Committee (2019) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: ttp: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed 11 May 2021)" type="book" year="2019">IUCN, 2019</bibRefCitation>
), i.e. the species has an EOO &lt;
<quantity id="4C9F9B30FFD5E169FC0CFE206916ED3E" box="[965,1054,806,832]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="262" unit="km" value="100.0">100 km</quantity>
2 and is known from only a single location with inferred and observed declines in area of occupancy, quality of habitat and number of mature individuals, and an AOO &lt;
<quantity id="4C9F9B30FFD5E169FEE7FE686C7FEDF6" box="[302,375,878,904]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="262" unit="km" value="10.0">10 km</quantity>
2 at the single location with a reduction in area of occurrence, occupancy, extent and quality of habitat, and number of mature individuals. Therefore, 
<taxonomicName id="4C674D56FFD5E169FD25FE956E8DEDD2" authorityName="Rubite, Tandang &amp; C. W. Lin. A, B. Habit" authorityYear="2021" box="[748,901,914,940]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="262" phylum="Tracheophyta" rank="species" species="benitotanii">
<emphasis id="B913EAC7FFD5E169FD25FE956E8DEDD2" box="[748,901,914,940]" italics="true" pageId="5" pageNumber="262">B. benitotanii</emphasis>
</taxonomicName>
is hereby proposed to be placed under Critically Endangered (CR) category.
</paragraph>
</subSubSection>
</treatment>
</document>