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<mods:title id="2442F1F8839976C384499E40C7BBDE4F">Diversity and taxonomy of the intertidal patellogastropod limpets of the mainland Indian coastline</mods:title>
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<mods:namePart id="56A260A15B3FA34B5A9C656FF86E6CF4">Vakani, Bhavik</mods:namePart>
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<mods:namePart id="97D108F4D398FEFE93DD9FDBF8D71FFB">Nakano, Tomoyuki</mods:namePart>
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<paragraph id="0800133B5D43A20D80AB81B7FE6CFA6C" blockId="4.[151,484,1461,1524]" box="[151,484,1461,1488]" pageId="4" pageNumber="215">
<heading id="5348A4575D43A20D80AB81B7FE6CFA6C" bold="true" box="[151,484,1461,1488]" fontSize="11" level="1" pageId="4" pageNumber="215" reason="1">
<taxonomicName id="CFBF68B85D43A20D80AB81B7FE6CFA6C" ID-CoL="S3PF" authority="Born, 1778" authorityName="Born" authorityYear="1778" box="[151,484,1461,1488]" class="Gastropoda" family="Nacellidae" genus="Cellana" kingdom="Animalia" pageId="4" pageNumber="215" phylum="Mollusca" rank="species" species="radiata">
<emphasis id="3ACBCF295D43A20D80AB81B7FE6CFA6C" bold="true" box="[151,484,1461,1488]" pageId="4" pageNumber="215">
<emphasis id="3ACBCF295D43A20D80AB81B7FEDBFA73" bold="true" box="[151,339,1461,1488]" italics="true" pageId="4" pageNumber="215">Cellana radiata</emphasis>
<bibRefCitation id="6C2E6ECA5D43A20D816781B4FE6CFA6C" author="Born, I." box="[347,484,1461,1488]" pageId="4" pageNumber="215" pagination="1 - 458" refId="ref6926" refString="Born, I. (1778) Vol. Index rerum naturalium Musei Caesarei Vindobonensis. Pars I. ma. Testacea. Verzeichniss der naturlichen Seltenheiten des k. k. Naturalien Cabinets zu Wien. Erster Theil. Schalthiere, 1 - 40, 1 - 458. [1 - 82] https: // doi. org / 10.5962 / bhl. title. 11581" type="journal article" year="1778">Born, 1778</bibRefCitation>
</emphasis>
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<subSubSection id="40A540B05D43A20D80AB81DBFE98F800" pageId="4" pageNumber="215" type="description">
<paragraph id="0800133B5D43A20D80AB81DBFEC0FA57" blockId="4.[151,484,1461,1524]" box="[151,328,1497,1524]" pageId="4" pageNumber="215">
(
<figureCitation id="90840FBE5D43A20D809C81DBFF6BFA50" box="[160,227,1497,1523]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1209]" captionTargetBox="[151,1428,373,1164]" captionTargetId="figure@5.[151,1428,373,1164]" captionTargetPageId="5" captionText="FIGURE 3. Cellana radiata. a (dorsal), b (ventral), c (ventral empty shell), d (lateral). ad (ZMGDNC51.3a), ej variants of Cellana radiata—e (ZMGDNC51.3e), f (ZMGDNC51.18), g (ZMGDNC41.3g), h (ZMGDNC41.3h), i (ZMGDNC34.3i), j (ZMGDNC34.3j)." figureDoi="http://doi.org/10.5281/zenodo.3621139" httpUri="https://zenodo.org/record/3621139/files/figure.png" pageId="4" pageNumber="215">Fig. 3</figureCitation>
,
<figureCitation id="90840FBE5D43A20D80CC81DBFEC8FA57" box="[240,320,1497,1524]" captionStart="FIGURE 9" captionStartId="11.[151,250,1747,1772]" captionTargetBox="[152,1435,625,1728]" captionTargetId="figure@11.[151,1435,625,1728]" captionTargetPageId="11" captionText="FIGURE 9. Radular teeth of patellogastropods (x 50). a: Cellana karachiensis, b: Cellana radiata, c: Cellana rota, d: Cellana conciliata, e: Cellana cylindrica, f: Patelloida lentiginosa, g: Lottia sp. (0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3621151" httpUri="https://zenodo.org/record/3621151/files/figure.png" pageId="4" pageNumber="215">Fig. 9b</figureCitation>
)
</paragraph>
<paragraph id="0800133B5D43A20D80AB8222FAF1F8B0" blockId="4.[151,1437,1568,2028]" pageId="4" pageNumber="215">
<emphasis id="3ACBCF295D43A20D80AB8222FEA1F998" bold="true" box="[151,297,1568,1595]" pageId="4" pageNumber="215">Description.</emphasis>
Shell up to
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in length, ovate, and broader to anterior to narrow posterior and moderately elevated. Apex is straight, pointed and sub-central. Sculpture consisting of numerous, narrow, radial riblets around 62 to 72, in some large specimens, up to 100. Coloration varies from greyish to black, with 4 to 5 bifid white lighter radial stripes observed on brown background, inner surface of shell has pearl white coloration. Foot is creamy-white in colour. Coloration difference also observed in foot, which is creamy white to white in colour. Same coloration observed at tip of cephalic tentacles, which covers one third of their length. Cephalic tentacles with radial maculations, bifurcated towards the margin, some specimens have radial maculations that join up to form a few broad bands.
</paragraph>
<paragraph id="0800133B5D43A20D80FB831FFE98F800" blockId="4.[151,1437,1568,2028]" pageId="4" pageNumber="215">
<emphasis id="3ACBCF295D43A20D80FB831FFE41F894" bold="true" box="[199,457,1821,1847]" pageId="4" pageNumber="215">Radular morphology.</emphasis>
(
<figureCitation id="90840FBE5D43A20D81E5831FFDA4F89B" box="[473,556,1821,1848]" captionStart="FIGURE 9" captionStartId="11.[151,250,1747,1772]" captionTargetBox="[152,1435,625,1728]" captionTargetId="figure@11.[151,1435,625,1728]" captionTargetPageId="11" captionText="FIGURE 9. Radular teeth of patellogastropods (x 50). a: Cellana karachiensis, b: Cellana radiata, c: Cellana rota, d: Cellana conciliata, e: Cellana cylindrica, f: Patelloida lentiginosa, g: Lottia sp. (0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.3621151" httpUri="https://zenodo.org/record/3621151/files/figure.png" pageId="4" pageNumber="215">Fig. 9b</figureCitation>
). Radula
<typeStatus id="D704AD995D43A20D82A6831CFD5EF89B" box="[666,726,1822,1848]" pageId="4" pageNumber="215">Type</typeStatus>
: Docoglossan, Formula: 1 + D1 + 1 + D1 +1. The radular ribbon of
<taxonomicName id="CFBF68B85D43A20D80DB8340FED1F8F8" authorityName="Born" authorityYear="1778" box="[231,345,1857,1884]" class="Gastropoda" family="Nacellidae" genus="Cellana" kingdom="Animalia" pageId="4" pageNumber="215" phylum="Mollusca" rank="species" species="radiata">
<emphasis id="3ACBCF295D43A20D80DB8340FED1F8F8" box="[231,345,1857,1884]" italics="true" pageId="4" pageNumber="215">C. radiata</emphasis>
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is similar to that of
<emphasis id="3ACBCF295D43A20D82008340FD7AF8F8" box="[572,754,1857,1884]" italics="true" pageId="4" pageNumber="215">
<taxonomicName id="CFBF68B85D43A20D82008340FD66F8F8" authorityName="Winckworth" authorityYear="1930" box="[572,750,1857,1884]" class="Gastropoda" family="Nacellidae" genus="Cellana" kingdom="Animalia" pageId="4" pageNumber="215" phylum="Mollusca" rank="species" species="karachiensis">C. karachiensis</taxonomicName>
.
</emphasis>
In every row, there is a spear-shaped very small rachidian or middle tooth and one bifurcated lateral tooth on each side. One dominant marginal tooth on each side that is also bifurcated.
</paragraph>
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<subSubSection id="40A540B05D43A20C80FB83AEFE0DFF7B" lastPageId="5" lastPageNumber="216" pageId="4" pageNumber="215" type="materials_examined">
<paragraph id="0800133B5D43A20C80FB83AEFE0DFF7B" blockId="4.[151,1437,1568,2028]" lastBlockId="5.[151,1436,152,359]" lastPageId="5" lastPageNumber="216" pageId="4" pageNumber="215">
<emphasis id="3ACBCF295D43A20D80FB83AEFE42F864" bold="true" box="[199,458,1964,1991]" pageId="4" pageNumber="215">Materials examined.</emphasis>
Specimens were collected from Uttambar coast,
<collectingRegion id="CA7BDDD95D43A20D846883AFFB68F864" box="[1108,1248,1965,1991]" country="India" name="Maharashtra" pageId="4" pageNumber="215">Maharashtra</collectingRegion>
(
<geoCoordinate id="6D8B75FC5D43A20D84C183AEFA14F864" box="[1277,1436,1964,1991]" degrees="17" direction="north" minutes="53" orientation="latitude" pageId="4" pageNumber="215" precision="1" seconds="33.1" value="17.892529">17°5333.1”N</geoCoordinate>
<geoCoordinate id="6D8B75FC5D43A20D80AB83D3FEBBF848" box="[151,307,2001,2027]" degrees="73" direction="east" minutes="03" orientation="longitude" pageId="4" pageNumber="215" precision="1" seconds="40.8" value="73.06133">73°0340.8”E</geoCoordinate>
) and Konkan coastline,
<collectingRegion id="CA7BDDD95D43A20D827F83D3FD32F848" box="[579,698,2001,2027]" country="India" name="Karnataka" pageId="4" pageNumber="215">Karnataka</collectingRegion>
. Four specimens (length 14.230.3 ±
<quantity id="CF47BEDE5D43A20D846283D2FB4CF84F" box="[1118,1220,2000,2028]" metricMagnitude="-3" metricUnit="m" metricValue="5.77" pageId="4" pageNumber="215" unit="mm" value="5.77">5.77 mm</quantity>
, width 12.125.8 ±
<quantity id="CF47BEDE5D42A20C80AB849AFF74FF10" box="[151,252,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="5" pageNumber="216" unit="mm" value="4.95">4.95 mm</quantity>
and Height 10.14 ± 2.46 were examined. One specimen is deposited in the Museum (ZMGDNC51.18, coll. Bhavik Vakani).
</paragraph>
</subSubSection>
<subSubSection id="40A540B05D42A20C80FB84E3FBF9FEE0" pageId="5" pageNumber="216" type="discussion">
<paragraph id="0800133B5D42A20C80FB84E3FB84FEBC" blockId="5.[151,1436,152,359]" pageId="5" pageNumber="216">
<emphasis id="3ACBCF295D42A20C80FB84E3FEB3FF58" bold="true" box="[199,315,225,251]" pageId="5" pageNumber="216">Remarks.</emphasis>
<bibRefCitation id="6C2E6ECA5D42A20C817884E3FE6CFF58" author="Powell, A. W. B." box="[324,484,225,251]" pageId="5" pageNumber="216" pagination="75 - 206" refId="ref9107" refString="Powell, A. W. B. (1973) The Patellid limpets of the world (Patellidae). Indo-Pacific Mollusca, 3 (15), 75 - 206." type="journal article" year="1973">Powell (1973)</bibRefCitation>
indicated that this species is common in the Indo-Pacific. In our study, this species was found from all over the west coast of
<collectingCountry id="70A853AB5D42A20C82528507FD20FEBC" box="[622,680,261,287]" name="India" pageId="5" pageNumber="216">India</collectingCountry>
except of the
<collectingRegion id="CA7BDDD95D42A20C837A8507FC12FEBC" box="[838,922,261,287]" country="India" name="Gujarat" pageId="5" pageNumber="216">Gujarat</collectingRegion>
coastline.
</paragraph>
<paragraph id="0800133B5D42A20C80FB852AFBF9FEE0" blockId="5.[151,1436,152,359]" box="[199,1137,296,323]" pageId="5" pageNumber="216">
<emphasis id="3ACBCF295D42A20C80FB852AFEA2FEE0" bold="true" box="[199,298,296,323]" pageId="5" pageNumber="216">Habitat.</emphasis>
Puddles and crevices of middle to upper littoral area of the intertidal zone.
</paragraph>
</subSubSection>
<subSubSection id="40A540B05D42A20C80FB854EFB34FEC4" box="[199,1212,332,359]" pageId="5" pageNumber="216" type="distribution">
<paragraph id="0800133B5D42A20C80FB854EFB34FEC4" blockId="5.[151,1436,152,359]" box="[199,1212,332,359]" pageId="5" pageNumber="216">
<emphasis id="3ACBCF295D42A20C80FB854EFEE9FEC4" bold="true" box="[199,353,332,359]" pageId="5" pageNumber="216">Distribution.</emphasis>
Konkan to Malabar coastline of the west coast of mainland Indian coastline.
</paragraph>
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