245 lines
34 KiB
XML
245 lines
34 KiB
XML
<document id="E6B37AFE84A1E9F7FE89732343F2B95C" ID-CLB-Dataset="48250" ID-DOI="10.5281/zenodo.195063" ID-GBIF-Dataset="249c9409-851e-4342-9401-19b3cf4fdeca" ID-ISSN="1175-5326" ID-Zenodo-Dep="195063" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460222235973" checkinUser="plazi" docAuthor="Shelley, Rowland M., Richart, Casey H. & Bogan, Arthur E." docDate="2010" docId="C93C87E5855D5C679BF7F97DCD21B608" docLanguage="en" docName="zt02446p064.pdf" docOrigin="Zootaxa 2446" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Octoglena claraqua Shelley, Richart & Bogan, 2010, new species" docType="treatment" docVersion="9" lastPageNumber="60" masterDocId="3505FF9D855F5C629B60FF92CE44B55A" masterDocTitle="Octoglena claraqua, n. sp. (Polyzoniida: Hirudisomatidae), a new milliped from Idaho, USA; first record of the order from the western interior of North America" masterLastPageNumber="64" masterPageNumber="55" pageNumber="57" updateTime="1732481108360" updateUser="ExternalLinkService">
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<mods:title id="E3FFB52D72767DC721706458291C3242">Octoglena claraqua, n. sp. (Polyzoniida: Hirudisomatidae), a new milliped from Idaho, USA; first record of the order from the western interior of North America</mods:title>
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<mods:name id="0B03571E2B34147867C0D1292E66E90F" type="personal">
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<mods:roleTerm id="AFB9C5700F21EFC41803AF43A8F15520">Author</mods:roleTerm>
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<mods:namePart id="3CA0217BFD8BF53798419D868CDF7ECC">Shelley, Rowland M.</mods:namePart>
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<mods:name id="969988F58E77CA0145F200DDF1CEF911" type="personal">
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<mods:roleTerm id="381124FF97508A06E69A756E3C980B80">Author</mods:roleTerm>
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<mods:namePart id="0D70793BF974EB1E32695D6E982D4C1D">Richart, Casey H.</mods:namePart>
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<mods:roleTerm id="6EED2E8AAF276C5EC6C8CC10BB2522A1">Author</mods:roleTerm>
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<mods:namePart id="17B9213881C8C3938604B5ED9788A068">Bogan, Arthur E.</mods:namePart>
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<mods:typeOfResource id="366EAD8B7538705B7D47E03CEE9C6485">text</mods:typeOfResource>
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<mods:title id="9D3DB4F1937FC296EAA23C7B3AC9F525">Zootaxa</mods:title>
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<mods:part id="039E47B05F93664EB7ED7A563B6A7FF1">
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<mods:date id="FB3F7C4B682622891AF292E470B116C7">2010</mods:date>
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<mods:detail id="92268DB2A91A971F8543237F31D40DEA" type="volume">
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<mods:number id="B08AE57F32959E0D3CBCADB432E3AEA6">2446</mods:number>
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<mods:classification id="1648158308AC94BD42221DDED8A768BA">journal article</mods:classification>
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<mods:identifier id="E3FE3DF30C0FFEFC32CDF02C5E6BF9B1" type="CLB-Dataset">48250</mods:identifier>
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<mods:identifier id="3D33CA7F1A0B7D8DC489D7AE83B0D58C" type="DOI">10.5281/zenodo.195063</mods:identifier>
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<mods:identifier id="ADE813C46F806E57ABCF5026B4F6499B" type="GBIF-Dataset">249c9409-851e-4342-9401-19b3cf4fdeca</mods:identifier>
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<mods:identifier id="378E320CFCAC8A432F6B36785C81983F" type="ISSN">1175-5326</mods:identifier>
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<treatment id="C93C87E5855D5C679BF7F97DCD21B608" ID-DOI="http://doi.org/10.5281/zenodo.6212186" ID-GBIF-Taxon="119385507" ID-Zenodo-Dep="6212186" LSID="urn:lsid:plazi:treatment:C93C87E5855D5C679BF7F97DCD21B608" httpUri="http://treatment.plazi.org/id/C93C87E5855D5C679BF7F97DCD21B608" lastPageId="5" lastPageNumber="60" pageId="2" pageNumber="57" scope_family="Hirudisomatidae" scope_order="Polyzoniida">
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<subSubSection id="098F6578855D5C609BF7F97DCF40B275" pageId="2" pageNumber="57" type="nomenclature">
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<paragraph id="412A36F3855D5C609BF7F97DCC60B253" blockId="2.[151,548,1775,1839]" box="[151,548,1775,1801]" pageId="2" pageNumber="57">
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<heading id="1A62819F855D5C609BF7F97DCC60B253" bold="true" box="[151,548,1775,1801]" fontSize="11" level="1" pageId="2" pageNumber="57" reason="1">
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<emphasis id="73E1EAE1855D5C609BF7F97DCC60B253" bold="true" box="[151,548,1775,1801]" pageId="2" pageNumber="57">
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<taxonomicName id="86954D70855D5C609BF7F97DCFC1B253" ID-CoL="CG6QB" box="[151,389,1775,1801]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="claraqua" status="sp. nov.">
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<emphasis id="73E1EAE1855D5C609BF7F97DCFC1B253" bold="true" box="[151,389,1775,1801]" italics="true" pageId="2" pageNumber="57">Octoglena claraqua</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="68D2579A855D5C609AF4F97DCC60B253" box="[404,548,1775,1801]" pageId="2" pageNumber="57" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="412A36F3855D5C609BF7F887CF40B275" blockId="2.[151,548,1775,1839]" box="[151,260,1813,1839]" pageId="2" pageNumber="57">
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<figureCitation id="D9AE2A76855D5C609BF7F887CF40B275" box="[151,260,1813,1839]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 8" captionStartId-0="3.[151,270,1731,1755]" captionStartId-1="4.[151,269,993,1017]" captionTargetBox-0="[306,1281,197,1707]" captionTargetBox-1="[605,971,306,965]" captionTargetId-0="figure@3.[306,1281,197,1707]" captionTargetId-1="figure@4.[604,983,306,972]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." captionText-1="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri-0="https://zenodo.org/record/195064/files/figure.png" httpUri-1="https://zenodo.org/record/195065/files/figure.png" pageId="2" pageNumber="57">Figs. 1–8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="098F6578855D5C609BF7F8CDCDC6B2B0" pageId="2" pageNumber="57" type="materials_examined">
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<paragraph id="412A36F3855D5C609BF7F8CDCDC6B2B0" blockId="2.[151,1436,1887,2026]" pageId="2" pageNumber="57">
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<emphasis id="73E1EAE1855D5C609BF7F8CDCF12B223" bold="true" box="[151,342,1887,1913]" pageId="2" pageNumber="57">
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<typeStatus id="9E2E8851855D5C609BF7F8CDCE90B223" box="[151,212,1887,1913]" pageId="2" pageNumber="57">Type</typeStatus>
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specimen.
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</emphasis>
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3
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<typeStatus id="9E2E8851855D5C609A1AF8F2CFA4B220" box="[378,480,1888,1914]" pageId="2" pageNumber="57" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="2784AE36855D5C609A8FF8F2CC01B220" box="[495,581,1888,1914]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="57">NCSM</collectionCode>
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) collected by C.H. Richart,
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<date id="352B1033855D5C6098ECF8F2CA62B220" box="[908,1062,1888,1914]" pageId="2" pageNumber="57" value="2008-07-20">20 July 2008</date>
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,
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<quantity id="866D9B16855D5C609F51F8F2CA38B220" box="[1073,1148,1888,1914]" metricMagnitude="2" metricUnit="m" metricValue="6.437376" pageId="2" pageNumber="57" unit="mi" value="0.4">0.4 mi</quantity>
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(
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<quantity id="866D9B16855D5C609FEBF8F2CA9BB220" box="[1163,1247,1888,1914]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="57" unit="km" value="0.6">0.6 km</quantity>
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) NW
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<collectingCountry id="39827663855D5C609E46F8F2CB0FB220" box="[1318,1355,1888,1914]" name="United States of America" pageId="2" pageNumber="57">US</collectingCountry>
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Forest Service Rd. 720 and
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<quantity id="866D9B16855D5C609AC0F817CFB5B2C5" box="[416,497,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.1265408" pageId="2" pageNumber="57" unit="mi" value="0.7">0.7 mi</quantity>
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(
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<quantity id="866D9B16855D5C609963F817CC1AB2C5" box="[515,606,1925,1951]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="57" unit="km" value="1.1">1.1 km</quantity>
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) WNW
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<collectionCode id="2784AE36855D5C6099A6F817CD54B2C5" box="[710,784,1925,1951]" pageId="2" pageNumber="57">USFS</collectionCode>
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Rd. 250 & Cedars Recreation site, beside unnamed tributary of North Fork Clearwater River, Clearwater National Forest (
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<geoCoordinate id="24A15034855D5C6098B8F838CAE7B29E" box="[984,1187,1962,1988]" direction="north" orientation="latitude" pageId="2" pageNumber="57" precision="1" value="46.877594">N 46° 52' 39.34"</geoCoordinate>
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,
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<geoCoordinate id="24A15034855D5C609FD0F838CBCBB29E" box="[1200,1423,1962,1988]" direction="west" orientation="longitude" pageId="2" pageNumber="57" precision="1" value="-115.088905">W 115° 05' 20.05"</geoCoordinate>
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), ca.
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<quantity id="866D9B16855D5C609BDFF842CF47B2B0" box="[191,259,2000,2026]" metricMagnitude="5" metricUnit="m" metricValue="1.07826048" pageId="2" pageNumber="57" unit="mi" value="67.0">67 mi</quantity>
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(
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<quantity id="866D9B16855D5C609A73F842CF2AB2B0" box="[275,366,2000,2026]" metricMagnitude="5" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="57" unit="km" value="107.0">107 km</quantity>
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||
) NE Pierce, Clearwater County, Idaho,
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<collectingCountry id="39827663855D5C609820F842CD3BB2B0" box="[832,895,2000,2026]" name="United States of America" pageId="2" pageNumber="57">USA</collectingCountry>
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.
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</paragraph>
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||
</subSubSection>
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||
<caption id="15EA667B855C5C619BF7F951CD23B240" httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="3" pageNumber="58" targetBox="[306,1281,197,1707]" targetPageId="3">
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<paragraph id="412A36F3855C5C619BF7F951CD23B240" blockId="3.[151,1436,1731,1818]" pageId="3" pageNumber="58">
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<emphasis id="73E1EAE1855C5C619BF7F951CF07B381" bold="true" box="[151,323,1731,1755]" pageId="3" pageNumber="58">FIGURES 1–4.</emphasis>
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<taxonomicName id="86954D70855C5C619A2BF951CC65B380" box="[331,545,1731,1754]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="claraqua">
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<emphasis id="73E1EAE1855C5C619A2BF951CC65B380" box="[331,545,1731,1754]" italics="true" pageId="3" pageNumber="58">Octoglena claraqua</emphasis>
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</taxonomicName>
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holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end;
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<emphasis id="73E1EAE1855C5C619F1EF971CADDB3A1" bold="true" box="[1150,1177,1763,1787]" pageId="3" pageNumber="58">ag</emphasis>
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, anterior gonopods,
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<emphasis id="73E1EAE1855C5C619E19F971CBD1B3A1" bold="true" box="[1401,1429,1763,1787]" pageId="3" pageNumber="58">pg</emphasis>
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, posterior gonopods. 4, anterior view of head, antennae, and collum.
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</paragraph>
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</caption>
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<subSubSection id="098F6578855C5C669BA6F8D1CCFEB5A6" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="diagnosis">
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<paragraph id="412A36F3855C5C619BA6F8D1CDD1B2F2" blockId="3.[151,1436,1859,2035]" pageId="3" pageNumber="58">
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<emphasis id="73E1EAE1855C5C619BA6F8D1CF02B207" bold="true" box="[198,326,1859,1885]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
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Prozona transversely banded. Gonopods
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<emphasis id="73E1EAE1855C5C61983CF8D6CDEBB207" box="[860,943,1860,1885]" italics="true" pageId="3" pageNumber="58">in situ</emphasis>
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recumbent, directed anteriad. Anterior gonopod with coxal lobe directed laterad; 5th podomere with sinuate lobe arising mediad from caudal surface and directed distad; ultimate podomere in form of terminal calyx.
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</paragraph>
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<paragraph id="412A36F3855C5C669BA6F821CCFEB5A6" blockId="3.[151,1436,1859,2035]" lastBlockId="4.[151,1437,152,252]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
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<emphasis id="73E1EAE1855C5C619BA6F821CF9CB297" bold="true" box="[198,472,1971,1997]" pageId="3" pageNumber="58">Preserved coloration.</emphasis>
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Head cream-colored; ocelli black, situated in angular gray to blackish band extending just below basalmost ocelli (
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<figureCitation id="D9AE2A76855C5C61991FF84BCC8AB2A9" box="[639,718,2009,2035]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="3" pageNumber="58">Fig. 4</figureCitation>
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). Prozona of segments 2-26 with distinct, pinkish-brown, speckled bands (
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<figureCitation id="D9AE2A76855B5C669A03FF0ACFEFB5E8" box="[355,427,152,178]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
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), collum and metaterga 2-5 with additional, scattered, light speckling (
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<figureCitation id="D9AE2A76855B5C669E7BFF0ACBC8B5E8" box="[1307,1420,152,178]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Figs. 1-4</figureCitation>
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), remaining metaterga light cream-colored; telson colorless and transparent, revealing underlying paraprocts (
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<figureCitation id="D9AE2A76855B5C669BFFFF70CEA2B5A6" box="[159,230,226,252]" captionStart="FIGURES 5 – 8" captionStartId="4.[151,269,993,1017]" captionTargetBox="[605,971,306,965]" captionTargetId="figure@4.[604,983,306,972]" captionTargetPageId="4" captionText="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59">Fig. 5</figureCitation>
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); venter uniformly light cream-colored.
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</paragraph>
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</subSubSection>
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<caption id="15EA667B855B5C669BF7FC73CC47B102" httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59" targetBox="[605,971,306,965]" targetPageId="4">
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||
<paragraph id="412A36F3855B5C669BF7FC73CC47B102" blockId="4.[151,1436,993,1112]" pageId="4" pageNumber="59">
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<emphasis id="73E1EAE1855B5C669BF7FC73CF06B6A3" bold="true" box="[151,322,993,1017]" pageId="4" pageNumber="59">FIGURES 5–8.</emphasis>
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<taxonomicName id="86954D70855B5C669A28FC70CC59B6A3" box="[328,541,994,1017]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="claraqua">
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<emphasis id="73E1EAE1855B5C669A28FC70CC59B6A3" box="[328,541,994,1017]" italics="true" pageId="4" pageNumber="59">Octoglena claraqua</emphasis>
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</taxonomicName>
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, somatic and gonopodal features. 5, telson and segments 25–27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25–28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6–8 = 0.05 mm.
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</paragraph>
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</caption>
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<subSubSection id="098F6578855B5C679BA6FB13CF35B58D" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59" type="materials_examined">
|
||
<paragraph id="412A36F3855B5C669BA6FB13CDBEB301" blockId="4.[151,1437,1153,2038]" pageId="4" pageNumber="59">
|
||
<emphasis id="73E1EAE1855B5C669BA6FB13CF04B1C1" bold="true" box="[198,320,1153,1179]" pageId="4" pageNumber="59">
|
||
<typeStatus id="9E2E8851855B5C669BA6FB13CF7FB1C1" box="[198,315,1153,1179]" pageId="4" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Length ca. 4.0 mm, maximum width ca.
|
||
<quantity id="866D9B16855B5C66987EFB13CD3EB1C1" box="[798,890,1153,1179]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="59" unit="mm" value="1.2">1.2 mm</quantity>
|
||
, W/L ratio 30%; 28 segments including telson, last two legless. Antennae directed sublaterad (
|
||
<figureCitation id="D9AE2A76855B5C66999DFB35CD0AB19B" box="[765,846,1191,1217]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Fig. 4</figureCitation>
|
||
), reaching back only to caudal margin of 2nd metatergite, relative lengths of antennomeres 6>3>5>2>4>1>7. Caudal metatergal margins strongly upturned and transparent (
|
||
<figureCitation id="D9AE2A76855B5C669A3AFB63CF8FB051" box="[346,459,1265,1291]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Figs. 1–2</figureCitation>
|
||
), revealing pigmented band on underlying prozona; collum semilunar, overhanging epicranium and uppermost ocelli, anterior margin gently convex and flared (
|
||
<figureCitation id="D9AE2A76855B5C669F49FA85CADAB06B" box="[1065,1182,1303,1329]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Figs. 3–4</figureCitation>
|
||
); anteriolateral tergal corners blunt, caudolateral corners rounded, produced very slightly caudad on segments 13–23, more strongly prolonged on segments 26–27 (
|
||
<figureCitation id="D9AE2A76855B5C669977FAF3CC26B021" box="[535,610,1377,1403]" captionStart="FIGURES 5 – 8" captionStartId="4.[151,269,993,1017]" captionTargetBox="[605,971,306,965]" captionTargetId="figure@4.[604,983,306,972]" captionTargetPageId="4" captionText="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59">Fig. 5</figureCitation>
|
||
); ozopores opening laterad but positioned submarginally on minute elevations visible only in profile, arising on segment 5 and extending to around segment 25. Telson transparent and broadly semilunar, extending caudad well beyond margins of paraprocts, over twice as broad as long but overhanging and extending well beyond margins of paraprocts, expanding laterad for entire breadth of body (
|
||
<figureCitation id="D9AE2A76855B5C669A0BFA65CFA4B34B" box="[363,480,1527,1553]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="D9AE2A76855B5C669A8FFA65CC44B34B" box="[495,512,1527,1553]" captionStart="FIGURES 5 – 8" captionStartId="4.[151,269,993,1017]" captionTargetBox="[605,971,306,965]" captionTargetId="figure@4.[604,983,306,972]" captionTargetPageId="4" captionText="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59">5</figureCitation>
|
||
); paraprocts small and indistinct, visible through telson; hypoproct minute. Sterna concave. Legs extending through segment 26, pairs 1–4 becoming progressively longer and less crassate, reaching maximum length and consistent thickness on leg pair 5.
|
||
</paragraph>
|
||
<paragraph id="412A36F3855B5C679BA4F9F5CF35B58D" blockId="4.[151,1437,1153,2038]" lastBlockId="5.[151,1437,152,850]" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
|
||
Gonopods
|
||
<emphasis id="73E1EAE1855B5C669A29F9F5CFD2B3DA" box="[329,406,1639,1664]" italics="true" pageId="4" pageNumber="59">in situ</emphasis>
|
||
recumbent (
|
||
<figureCitation id="D9AE2A76855B5C669950F9F5CCE3B3DB" box="[560,679,1639,1665]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="4" pageNumber="59">Figs. 2–3</figureCitation>
|
||
), apices directed anteriad, anterior surface facing dorsad, and caudal surface facing ventrad; appearing disproportionately large in relation to overall body-size and rhomboid, completely filling sternal region of body between medial pleural margins, expanding body longitudinally around segments 7–8. Anterior gonopods (
|
||
<figureCitation id="D9AE2A76855B5C669838F945CD8AB3AB" box="[856,974,1751,1777]" captionStart="FIGURES 5 – 8" captionStartId="4.[151,269,993,1017]" captionTargetBox="[605,971,306,965]" captionTargetId="figure@4.[604,983,306,972]" captionTargetPageId="4" captionText="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59">Figs. 6–7</figureCitation>
|
||
) upright in relation to sternum, distal podomeres positioned parallel to sternal lobes rather than overhanging them; sternal lobes moderately long, relatively widely separated; coxal lobe apically rounded and expanded, directed laterad, arising behind (caudal to) sternum, origin obscured in anterior view; 5th podomere with long, broad, lightly sinuate process arising around midlength on medial side of caudal surface, directed distomediad and lying subparallel to ultimate podomere; latter in form of calyx with open side facing anteriad, also with hirsute, basal lobe anteriolaterad. 3rd podomere of posterior gonopod with broad, rounded, sublaminate lobe mediad (
|
||
<figureCitation id="D9AE2A76855B5C669E22F825CBC8B28B" box="[1346,1420,1975,2001]" captionStart="FIGURES 5 – 8" captionStartId="4.[151,269,993,1017]" captionTargetBox="[605,971,306,965]" captionTargetId="figure@4.[604,983,306,972]" captionTargetPageId="4" captionText="FIGURES 5 – 8. Octoglena claraqua, somatic and gonopodal features. 5, telson and segments 25 – 27, dashed line denotes the area of the paraprocts and anus; scale line = 0.5 mm. 6, left anterior gonopod and sternum, anterior view. 7, right gonopods, caudal view. 8, right posterior gonopod, sublateral view. 25 – 28, segment numbers. Scale line for fig. 5 = 0.5 mm; line for figs. 6 – 8 = 0.05 mm." httpUri="https://zenodo.org/record/195065/files/figure.png" pageId="4" pageNumber="59">Fig. 8</figureCitation>
|
||
); ultimate podomere extending distad on anterior side of projection on 5th anterior gonopod podomere, terminating beside ultimate podomere of anterior gonopod, tapering smoothly and subcontinuously to blunt, angled, barbed tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="098F6578855A5C679BA4FF70CA53B436" pageId="5" pageNumber="60" type="etymology">
|
||
<paragraph id="412A36F3855A5C679BA4FF70CC1DB478" blockId="5.[151,1437,152,850]" pageId="5" pageNumber="60">
|
||
<emphasis id="73E1EAE1855A5C679BA4FF70CF15B5A6" bold="true" box="[196,337,226,252]" pageId="5" pageNumber="60">Etymology.</emphasis>
|
||
The species name is a latinization of "clearwater," the name of the Idaho county and drainage in which the
|
||
<typeStatus id="9E2E8851855A5C679A4FFE9ACFD1B478" box="[303,405,264,290]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
was discovered.
|
||
</paragraph>
|
||
<paragraph id="412A36F3855A5C679BA6FEBFCA53B436" blockId="5.[151,1437,152,850]" pageId="5" pageNumber="60">
|
||
<emphasis id="73E1EAE1855A5C679BA6FEBFCF6BB41D" bold="true" box="[198,303,301,327]" pageId="5" pageNumber="60">Ecology.</emphasis>
|
||
The
|
||
<typeStatus id="9E2E8851855A5C679A0BFEBFCF95B41D" box="[363,465,301,327]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
was discovered in streamside woody debris in a predominantly Englemann Spruce (
|
||
<taxonomicName id="86954D70855A5C679BC1FEC0CC15B436" authority="Parry ex Engelm." authorityName="Parry ex Engelm." box="[161,593,338,364]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="60" phylum="Tracheophyta" rank="species" species="englemannii">
|
||
<emphasis id="73E1EAE1855A5C679BC1FEC0CF38B431" box="[161,380,338,363]" italics="true" pageId="5" pageNumber="60">Picea englemannii</emphasis>
|
||
Parry ex Engelm.
|
||
</taxonomicName>
|
||
) forest at
|
||
<quantity id="866D9B16855A5C6799A7FEC0CD65B436" box="[711,801,338,364]" metricMagnitude="3" metricUnit="m" metricValue="1.1399519999999999" pageId="5" pageNumber="60" unit="ft" value="3740.0">3,740 ft</quantity>
|
||
(
|
||
<quantity id="866D9B16855A5C679851FEC0CDD0B436" box="[817,916,338,364]" metricMagnitude="3" metricUnit="m" metricValue="1.14" pageId="5" pageNumber="60" unit="m" value="1140.0">1,140 m</quantity>
|
||
) elevation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="098F6578855A5C679BA6FEE5CA25B716" pageId="5" pageNumber="60" type="distribution">
|
||
<paragraph id="412A36F3855A5C679BA6FEE5CA25B716" blockId="5.[151,1437,152,850]" pageId="5" pageNumber="60">
|
||
<emphasis id="73E1EAE1855A5C679BA6FEE5CF23B4CB" bold="true" box="[198,359,375,401]" pageId="5" pageNumber="60">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="9E2E8851855A5C67991CFEEACCEAB4C8" box="[636,686,376,402]" pageId="5" pageNumber="60">type</typeStatus>
|
||
locality, which is in the northeastern corner of Clearwater Co. and uncommonly remote. It is actually closer to (ca.
|
||
<quantity id="866D9B16855A5C67987AFE0FCD24B4ED" box="[794,864,413,439]" metricMagnitude="4" metricUnit="m" metricValue="5.7936384" pageId="5" pageNumber="60" unit="mi" value="36.0">36 mi</quantity>
|
||
[
|
||
<quantity id="866D9B16855A5C679810FE0FCD86B4ED" box="[880,962,413,439]" metricMagnitude="4" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="60" unit="km" value="58.0">58 km</quantity>
|
||
] SW) Superior, Mineral Co., Montana, from which it is reachable via Diamond and Trout Creek rds., and then USFS 250 after crossing into Idaho over Hoodoo Pass. Cedars USFS Campsite and Recreation Area is at the juncture of USFS Rds. 250 & 720. While it may be restricted to the western part of the Bitterroot Mountains,
|
||
<taxonomicName id="86954D70855A5C679F7CFD9FCAE8B77C" box="[1052,1196,525,550]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="claraqua">
|
||
<emphasis id="73E1EAE1855A5C679F7CFD9FCAE8B77C" box="[1052,1196,525,550]" italics="true" pageId="5" pageNumber="60">
|
||
<collectingCountry id="39827663855A5C679F7CFD9FCA71B77C" box="[1052,1077,525,550]" name="Iceland" pageId="5" pageNumber="60">O</collectingCountry>
|
||
. claraqua
|
||
</emphasis>
|
||
</taxonomicName>
|
||
conceivably occurs east of the crest of the Range in Mineral Co., Montana, ~
|
||
<quantity id="866D9B16855A5C679853FDA0CD33B716" box="[819,887,562,588]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="5" pageNumber="60" unit="mi" value="14.0">14 mi</quantity>
|
||
(
|
||
<quantity id="866D9B16855A5C6798E6FDA0CD97B716" box="[902,979,562,588]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="60" unit="km" value="22.0">22 km</quantity>
|
||
) to the east.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="098F6578855A5C679BA6FDC5CD21B608" pageId="5" pageNumber="60" type="discussion">
|
||
<paragraph id="412A36F3855A5C679BA6FDC5CB12B7CD" blockId="5.[151,1437,152,850]" pageId="5" pageNumber="60">
|
||
<emphasis id="73E1EAE1855A5C679BA6FDC5CF04B72B" bold="true" box="[198,320,599,625]" pageId="5" pageNumber="60">Remarks.</emphasis>
|
||
The smallest representative of the genus,
|
||
<taxonomicName id="86954D70855A5C67985EFDCACD8BB72B" box="[830,975,600,625]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="claraqua">
|
||
<emphasis id="73E1EAE1855A5C67985EFDCACD8BB72B" box="[830,975,600,625]" italics="true" pageId="5" pageNumber="60">
|
||
<collectingCountry id="39827663855A5C67985EFDCACD12B72B" box="[830,854,600,625]" name="Iceland" pageId="5" pageNumber="60">O</collectingCountry>
|
||
. claraqua
|
||
</emphasis>
|
||
</taxonomicName>
|
||
constitutes the first record of order & family from Idaho, the western interior of North
|
||
<collectingCountry id="39827663855A5C6799B3FDEFCD7FB7CD" box="[723,827,637,663]" name="United States of America" pageId="5" pageNumber="60">America</collectingCountry>
|
||
, and the Rocky Mountains in the broad sense.
|
||
</paragraph>
|
||
<paragraph id="412A36F3855A5C679BA6FD30CD21B608" blockId="5.[151,1437,152,850]" pageId="5" pageNumber="60">
|
||
<emphasis id="73E1EAE1855A5C679BA6FD30CF56B7E1" box="[198,274,674,699]" italics="true" pageId="5" pageNumber="60">In situ</emphasis>
|
||
, the gonopods appear disproportionately large in relation to overall body size (
|
||
<figureCitation id="D9AE2A76855A5C679FDDFD30CB75B7E6" box="[1213,1329,674,700]" captionStart="FIGURES 1 – 4" captionStartId="3.[151,270,1731,1755]" captionTargetBox="[306,1281,197,1707]" captionTargetId="figure@3.[306,1281,197,1707]" captionTargetPageId="3" captionText="FIGURES 1 – 4. Octoglena claraqua holotype. 1, lateral view of entire specimen showing color pattern. 2, ventrolateral view of entire specimen showing recumbent gonopod complex. 3, the same of anterior end; ag, anterior gonopods, pg, posterior gonopods. 4, anterior view of head, antennae, and collum." httpUri="https://zenodo.org/record/195064/files/figure.png" pageId="5" pageNumber="60">Figs. 2–3</figureCitation>
|
||
); viewed from a directly ventral perspective, they appear rhomboid and distort the body slightly in this region. However, this dramatically large size is primarily an illusion and a function of their recumbent, rather than upright, orientation. It results from the lengthwise orientation of their long, rather than narrow, dimension, the latter occurring in other Western Hemisphere hirudisomatids.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |