treatments-xml/data/A7/CE/A5/A7CEA5BD61E2E9280A98DDDE2C01FAB9.xml

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<mods:title id="641835667E74C6B6536B6F4E77E0E446">A review of the Cercyon Leach (Coleoptera, Hydrophilidae, Sphaeridiinae) of the Greater Antilles</mods:title>
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<subSubSection id="227CF6BAD3F92DDC283F5F8E9E209F9D" pageId="18" pageNumber="57" type="nomenclature">
<paragraph id="9EE198332C21CF19A6E9DE620F08590A" pageId="18" pageNumber="57">
<taxonomicName id="AD3A1F10239A8EC89781790051D1C30D" ID-CoL="SVY3" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insularis">Cercyon insularis Chevrolat, 1863</taxonomicName>
Figures 4 
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, 12 
<normalizedToken id="C4C6E1207B6D5C4ECC3BDEDEE4CDB8D4" originalValue="a–i">a-i</normalizedToken>
, 16c
</paragraph>
</subSubSection>
<subSubSection id="5A34D2B7A81573EC3A2350FFBFD54E66" pageId="18" pageNumber="57" type="reference_group">
<paragraph id="105016DCCBB16C8200938FB037BE5345" pageId="18" pageNumber="57">
<taxonomicName id="86C56AC765BFC058DE147B4A075BA486" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insulare" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insulare">Cercyon insulare</taxonomicName>
Chevrolat, 1863: 208.
</paragraph>
<paragraph id="D77C8BA1A2C5A0E842AB30CB0521D048" pageId="18" pageNumber="57">
<taxonomicName id="99CFCDDDCFEE18EE52BDEDAEDBE0E2E8" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insulare" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insulare">Cercyon insulare</taxonomicName>
<bibRefCitation id="06ECBE6A9F93CC6E98A451288CD369A4" author="Gundlach, J" journalOrPublisher="Alvarez Comp., Habana" pageId="31" pageNumber="70" title="Constribucion a la entomologia Cubana. Tomo III. A." year="1891">Gundlach (1891</bibRefCitation>
: 50, redescription).
</paragraph>
</subSubSection>
<subSubSection id="D2D7CE00642135864DCB8370C9ACCDD2" pageId="18" pageNumber="57" type="dna barcodes">
<paragraph id="73FD8ED12DD66CDF90465DBFB53A5965" pageId="18" pageNumber="57">DNA barcodes.</paragraph>
<paragraph id="04EB021B86793E6B6AF1ABE191657FB7" pageId="18" pageNumber="57">GANTC001-16, GANTC011-17, GANTC012-17</paragraph>
</subSubSection>
<subSubSection id="2367103011351746B6704149C1DDC8AB" pageId="18" pageNumber="57" type="bin id">
<paragraph id="36487F75FEAF7F5FBE94D6F1C6E57D58" pageId="18" pageNumber="57">BIN ID.</paragraph>
<paragraph id="12AAF370F2BE976F4FA86DC7BA3D1D7E" pageId="18" pageNumber="57">BOLD:ADC9388.</paragraph>
</subSubSection>
<subSubSection id="A18DFCCE0CCA3428ECA05E981F3356BB" pageId="18" pageNumber="57" type="figures in flickr">
<paragraph id="66D9837703D16FAE2F2A808516D77688" pageId="18" pageNumber="57">Figures in Flickr.</paragraph>
<paragraph id="0ABF139E8707CA5892A80EF849538FF4" pageId="18" pageNumber="57">www.flickr.com/photos/142655814@N07/albums/72157669492393134</paragraph>
</subSubSection>
<subSubSection id="4683D6DA542633AB2B0A38181566CE28" pageId="18" pageNumber="57" type="type locality">
<paragraph id="D5AC4243B45994A2088B7CD0D708ACAF" pageId="18" pageNumber="57">Type locality.</paragraph>
<paragraph id="4C8FC29050CEB43849584B4756270E7F" pageId="18" pageNumber="57">Cuba: Havana.</paragraph>
</subSubSection>
<subSubSection id="F54CEF388FED8F2D1A127CEDAE986CC9" pageId="18" pageNumber="57" type="type material">
<paragraph id="14200DA0EC337C22263EA100421A9689" pageId="18" pageNumber="57">Type material.</paragraph>
<paragraph id="3426ED40F5ADB4E239933C9C858DB9E0" pageId="18" pageNumber="57">
Holotype (unsexed specimen): &quot;Havana, D. Poey // 
<taxonomicName id="5D7765D7EF1C1E031329694CBA42B02A" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insulare" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insulare">Cercyon insulare</taxonomicName>
, Chev Cuba, [illegible] // TYPE [red label]&quot; (MNHN).
</paragraph>
<caption id="90AFE5439D035E2E2B70641633ADDAA6" pageId="18" pageNumber="57">
<paragraph id="3C6379F68371EB02B6B73BE945EBB1BE" pageId="18" pageNumber="57">
Figure 4. 
<taxonomicName id="E29E560448A7BF4D88EF8AFF51105A75" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="insularis">Cercyon insularis</taxonomicName>
Chevrolat and 
<taxonomicName id="F2464623308748360E470C455C739245" lsidName="C. variegatus" pageId="18" pageNumber="57" rank="species" species="variegatus">C. variegatus</taxonomicName>
Sharp. 
<normalizedToken id="8BEFDAD4923C2B87FF438953AA101611" originalValue="a–i">a-i</normalizedToken>
<taxonomicName id="8053DF8BB2C113C64E538808511709FB" lsidName="C. insularis" pageId="18" pageNumber="57" rank="species" species="insularis">C. insularis</taxonomicName>
: 
<normalizedToken id="876FB844B31AB0CEF0265DD8004D6B42" originalValue="a–c">a-c</normalizedToken>
dorsal, ventral and lateral habitus of the non-type specimen from Cuba d habitus of the holotype e labels of the holotype 
<normalizedToken id="4ADA5CA5C340A7B8D79CEBD84E4A24F9" originalValue="f–i">f-i</normalizedToken>
male genitalia of non-type specimen from Dominican Republic. 
<normalizedToken id="69C4B984D996ABFB004906BCDF264A4A" originalValue="j–o">j-o</normalizedToken>
lectotype of 
<taxonomicName id="E4B0C9B33CA1AFFA9DA145A367D632FF" lsidName="C. variegatus" pageId="18" pageNumber="57" rank="species" species="variegatus">C. variegatus</taxonomicName>
: j dorsal habitus k labels 
<normalizedToken id="A3EC8172E017EA48FF8E6C50AB9A6A92" originalValue="l–o">l-o</normalizedToken>
male genitalia. Genital parts illustrated: f, l tegmen of aedeagus g, m median lobe of aedeagus h, n detail of apex of median lobe; i, o 9th sternite.
</paragraph>
</caption>
<caption id="A122AD878EBB4E36F76CF990F742A99E" pageId="18" pageNumber="57">
<paragraph id="2031B167D47E17EE4CAF289E9A452775" pageId="18" pageNumber="57">
Figure 5. 
<taxonomicName id="841AB951AB3EBA27818ED73AA4EEC38F" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="genus">Cercyon</taxonomicName>
spp. n. genitalia 
<normalizedToken id="915CB8E90E6A6E19C5B4151127C4FEC8" originalValue="a–d">a-d</normalizedToken>
<taxonomicName id="2AF0A906FF76926E25E79037642539FB" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon gimmeli" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="gimmeli">Cercyon gimmeli</taxonomicName>
sp. n. 
<normalizedToken id="C6DD90367F1BC4C54EFD69F75C160498" originalValue="e–h">e-h</normalizedToken>
<taxonomicName id="872F96AF5C00D06305B6BE4267104A68" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon taino" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="taino">Cercyon taino</taxonomicName>
sp. n. 
<normalizedToken id="53E2CF6CD085CEDBF696C8C95A6ECC37" originalValue="i–l">i-l</normalizedToken>
<taxonomicName id="98AC5690878D2A13F31FD08678BC60C5" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon armatipenis" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="armatipenis">Cercyon armatipenis</taxonomicName>
sp. n. 
<normalizedToken id="BAC55D560251D8BFAC6EFFEF60493209" originalValue="m–p">m-p</normalizedToken>
<taxonomicName id="4A8CB9FE687E14C36E67AABAFDED8654" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon sklodowskae" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="sklodowskae">Cercyon sklodowskae</taxonomicName>
sp. n. a, e, i, m tegmen of aedeagus b, f, j, n median lobe of aedeagus c, g, k, o 9th sternite.
</paragraph>
</caption>
<caption id="DD8F0FF1BF4B1ED7216215FBA9100313" pageId="18" pageNumber="57">
<paragraph id="2D0C9410535681DA7DE4A3D77DA8D711" pageId="18" pageNumber="57">
Figure 6. 
<taxonomicName id="5FD3BF039C21584E76A64B0BF1F15388" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="genus">Cercyon</taxonomicName>
spp. n. genitalia 
<normalizedToken id="C473AA6F00A553878EF1AAD03723BCFD" originalValue="a–d">a-d</normalizedToken>
<taxonomicName id="AC2C22C0A6DD78BEFC1A3EE1F6B7E809" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon praetextatus" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="praetextatus">Cercyon praetextatus</taxonomicName>
Say 
<normalizedToken id="D07B08DB8DEFC722A9A797ACA3F65AB9" originalValue="e–h">e-h</normalizedToken>
<taxonomicName id="9C70B92E16A672BF2784760FA36A9B1C" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon spiniventris" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="spiniventris">Cercyon spiniventris</taxonomicName>
sp. n. 
<normalizedToken id="827C3A6C7C57D6C0D9B8A000F6DDB27F" originalValue="i–k">i-k</normalizedToken>
<taxonomicName id="A87DE62E98D49DA00D683E2919E06E43" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon nigriceps" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="nigriceps">Cercyon nigriceps</taxonomicName>
Marsham 
<normalizedToken id="EDFCBD9E67DA708119529C6A0BDB47E1" originalValue="l–n">l-n</normalizedToken>
<taxonomicName id="DB653929348DFD1ECC750170D8755FCA" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon quisquilius" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="quisquilius">Cercyon quisquilius</taxonomicName>
Linnaeus a, e, i, l tegmen of aedeagus b, f, j, m median lobe of aedeagus c, g 9th sternite.
</paragraph>
</caption>
<caption id="65927F5035DBA7D153C74BD73F20C492" pageId="18" pageNumber="57">
<paragraph id="F0AC9B2E158C878BA0F28F2391BBEB6A" pageId="18" pageNumber="57">
Figure 7. 
<taxonomicName id="F6BE735690E3FED76E4E8987E5BF2BE9" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon floridanus" order="Coleoptera" pageId="18" pageNumber="57" phylum="Arthropoda" rank="species" species="floridanus">Cercyon floridanus</taxonomicName>
Horn a tegmen of aedeagus b median lobe of aedeagus c 9th sternite d ventral view of pterothorax.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="58568F42C619F128CC7BF50EDCD068F9" lastPageId="21" lastPageNumber="60" pageId="18" pageNumber="57" type="materials_examined">
<paragraph id="073D30CE9055672840DBE5ED7343A80B" pageId="18" pageNumber="57">Additional material examined.</paragraph>
<paragraph id="4B22FDEF77232A132FFC677E8C7ED8CC" lastPageId="21" lastPageNumber="60" pageId="18" pageNumber="57">
CUBA: 
<normalizedToken id="A1A0A2A661FC8160B2F45D124DCDE62F" originalValue="Camagüey">Camagueey</normalizedToken>
: Sierra de Cubitas municipality, Limones-Tuabaquey, 
<geoCoordinate id="E3285691C3F6D143C16D8F9F88C8EECE" direction="north" orientation="latitude" precision="1" value="21.597807">21°35'52.10&quot;N</geoCoordinate>
, 
<geoCoordinate id="85F6D5149E8905E8B61692083CAFA408" direction="west" orientation="longitude" precision="1" value="-77.78823">77°47'17.62&quot;W</geoCoordinate>
, 16.v.2013, leg. R. Anderson (1 spec.: NMPC). 
<normalizedToken id="32C8F17A402CC66D1D9ECF9B8C720DB3" originalValue="Guantánamo">Guantanamo</normalizedToken>
: El Yunque, 3.2 km SW of campismo popular, at right tributary of Duabe river, secondary evergreen forest, cow excrement, 
<geoCoordinate id="97ACD6A0FBFDCEBCE1E17C6A0A602B44" direction="north" orientation="latitude" precision="925" value="20.316668">20°19'N</geoCoordinate>
, 
<geoCoordinate id="B99631594BE65ADD49E435981D52A918" direction="west" orientation="longitude" precision="925" value="-74.566666">74°34'W</geoCoordinate>
, 150 m a.s.l., 13.vi.2012, leg. 
<normalizedToken id="2A95E9DF7DC7944382E3BB90B6A45BA4" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="7547A682E2B6ADC42FE1BE5EC9B025EC" originalValue="Fikáček">Fikacek</normalizedToken>
(MF09) (7 spec.: NMPC); El Yunque, ca. 1.4 km W of campismo popular, cocoa plantations shaded by palms, cow excrement, 
<geoCoordinate id="33532CA4BB09C1C2FE9F3E5B255D8438" direction="north" orientation="latitude" precision="92" value="20.336666">20°20.2'N</geoCoordinate>
, 
<geoCoordinate id="BA3AE8F8B97C3235EBE751279C82286E" direction="west" orientation="longitude" precision="92" value="-74.56167">74°33.7'W</geoCoordinate>
, 60-150 m a.s.l., 11.vi.2012, 
<normalizedToken id="B50CEC085FC3831D721C6FC886D0C3A6" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="19292D30D17E7205A13DD7ECBFCD3B66" originalValue="Fikáček">Fikacek</normalizedToken>
(MF03) (16 spec.: NMPC). Santiago de Cuba: El Vivero, 1.6 km E of Dos Caminos, cow excrements on pasture, 
<geoCoordinate id="0A1191AC8468078CC2BE884760E5B457" direction="north" orientation="latitude" precision="92" value="20.18">20°10.8'N</geoCoordinate>
, 
<geoCoordinate id="703DE1CAE46DDE233E05B396F3E3CADA" direction="west" orientation="longitude" precision="92" value="-75.77333">75°46.4'W</geoCoordinate>
, 150 m, 20-21.vi.2012, leg. 
<normalizedToken id="9A8F0CF09AD4E04B56B39FBC4BC15043" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="BECCE798484449A0143EA50DF50226A7" originalValue="Fikáček">Fikacek</normalizedToken>
(MF18) (1 spec.: NMPC); San Luis Municipality, Dos Caminos, 
<geoCoordinate id="AA43C4E891AF001E431DC6B6EF4D0173" direction="north" orientation="latitude" precision="1" value="20.182728">20°10'57.82&quot;N</geoCoordinate>
, 
<geoCoordinate id="B8B61D01EE82C36B9F5F88D181CAA434" direction="west" orientation="longitude" precision="1" value="-75.778015">75°46'40.84&quot;W</geoCoordinate>
, 3.x.2012, leg. 
<normalizedToken id="42662BFCACB899A6DA55DB16F12AF9E0" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
(12 spec.: NMPC). Granma: PN Turquino, around La Platica, 
<geoCoordinate id="878DF65709BBB2106386EED7035AC893" direction="north" orientation="latitude" precision="92" value="20.011667">20°0.7'N</geoCoordinate>
, 
<geoCoordinate id="8CD4B25619AB95B11DF1C944F2FEE7F5" direction="west" orientation="longitude" precision="92" value="-76.89">76°53.4'W</geoCoordinate>
, 880 m, 25-26.vi.2012, leg. 
<normalizedToken id="067458D20F0806535971F41A3041B54D" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="CB6672135176344BC944309CD12B0774" originalValue="Fikáček">Fikacek</normalizedToken>
(MF24) (12 spec.: NMPC); PN Turquino, on the trail up to 0.5 km S of La Platica, 
<geoCoordinate id="BEC3F27CE0B4BF6774D0AE6DEE847B05" direction="north" orientation="latitude" precision="92" value="20.008333">20°0.5'N</geoCoordinate>
, 
<geoCoordinate id="82434C42AC8724CFE1344A4975794EC6" direction="west" orientation="longitude" precision="92" value="-76.888336">76°53.3'W</geoCoordinate>
, 920 m, 23-27.vi.2012, leg. 
<normalizedToken id="AADF4910D994DFD7CC2E4164746D2199" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="942C8F081BAF6EA2834F91D6FE7FFE50" originalValue="Fikáček">Fikacek</normalizedToken>
(MF20) (2 spec.: NMPC); PN Turquino, La Siguapa, ca. 1.5 km SE of La Platica, sifting leaf litter in evergreen forest, 
<geoCoordinate id="72C60F0EE10C2B668C007CC7883A8C72" direction="north" orientation="latitude" precision="92" value="20.003334">20°0.2'N</geoCoordinate>
, 
<geoCoordinate id="2491A5263FE13B248DC3EF9F742CAD14" direction="west" orientation="longitude" precision="92" value="-76.88">76°52.8'W</geoCoordinate>
, 1290 m, 25.vi.2012, leg. F. Cala-Riquelme (MF25) (1 spec.: NMPC) [DNA extract at NMPC] (1: NMPC). Artemisa: 
<normalizedToken id="CAE0EF00BB164F5B14CB005671E5AADE" originalValue="Cañon">Canon</normalizedToken>
de Santa Cruz, 
<normalizedToken id="38A791507B9A11725AD7F695EAFF7835" originalValue="Río">Rio</normalizedToken>
de Santa Cruz, 
<geoCoordinate id="426558AA5D87715E126D0CAF5E1ED1ED" direction="north" orientation="latitude" precision="1" value="22.750359">22°45'1.29&quot;N</geoCoordinate>
<geoCoordinate id="A78A8589F5320CE821F77DD75BC03A9D" direction="west" orientation="longitude" precision="1" value="-83.14899">83°08'56.36&quot;W</geoCoordinate>
, 199 m a.s.l., 16.vii.2016, leg. A. 
<normalizedToken id="ECDBB0E761F963CF083D38B1A733DFC2" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
(1 spec.: NMPC) [DNA extraction: MF1749]. DOMINICAN REPUBLIC: La Vega: 7.0 km W of Manabao, side of a stony stream in a valley with scattered houses and plantations surrounded by montane forest, in cow excrements, 
<geoCoordinate id="574381880C386C800EDF9F5B2D4ED208" direction="north" orientation="latitude" precision="9" value="19.076">19°4.56'N</geoCoordinate>
, 
<geoCoordinate id="729FC33F1627B87BCB908DC8E8560B09" direction="west" orientation="longitude" precision="9" value="-70.857666">70°51.46'W</geoCoordinate>
, 1185 m a.s.l., 23.viii.2014, leg. 
<normalizedToken id="A7C2439DD81D889254A9D1EE0963DFAE" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
, 
<normalizedToken id="F838958BA601680AA3E3D0EAE3851ADB" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Gimmel (DR18) (3 spec.: NMPC); at S margin of Manabao, 
<geoCoordinate id="8DB711288A8290B2A7B2B86124A285E2" direction="north" orientation="latitude" precision="9" value="19.064167">19°3.85'N</geoCoordinate>
, 
<geoCoordinate id="0547C1A39D1E57BB365321047C4D3776" direction="west" orientation="longitude" precision="9" value="-70.7935">70°47.61'W</geoCoordinate>
, 912 m a.s.l., 27.viii.2014, leg. 
<normalizedToken id="4CB5A05C303E9BF7A8E9BC53965C35A7" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
&amp; 
<normalizedToken id="98BB27AEAD380C11C5F0E928D85A2871" originalValue="Fikáček">Fikacek</normalizedToken>
(DR23) (3 spec.: NMPC). 
<normalizedToken id="6FA5E36C41ECF4CDAA461B7834A680E3" originalValue="Samaná">Samana</normalizedToken>
: MN Salto El 
<normalizedToken id="B5DF4139376F716D6F171AAA9341CB06" originalValue="Limón">Limon</normalizedToken>
2.8 km SSW of El 
<normalizedToken id="497F85CFBA00E24DB41C22485B549B0C" originalValue="Limón">Limon</normalizedToken>
, secondary vegetation and tiny remnants of forests among coffee plantations and pastures, cow excrements, 
<geoCoordinate id="35AF55090A20F45E5DFCB2A86FEF0B3F" direction="north" orientation="latitude" precision="9" value="19.276">19°16.56'N</geoCoordinate>
, 
<geoCoordinate id="B45DF2C226AB2253D3AA57460000390F" direction="west" orientation="longitude" precision="9" value="-69.44117">69°26.47'W</geoCoordinate>
, 2.ix.2014, leg. 
<normalizedToken id="43405772D2C7C87B7D275846009EF0EF" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
, 
<normalizedToken id="B0E3FA5B8D3DEF8CD196B1D90D0CEEC5" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Gimmel (DR29a) (16 spec.: NMPC). 
<normalizedToken id="E9F7C82D792BFEBD5EADEFCEF76C1AE3" originalValue="Monseñor">Monsenor</normalizedToken>
Nouel: PN La Humeadora; 11.6 km SSW, of Piedra Blanca, in horse excrement in moist broad-leaf forest in a valley of a small stony stream, 
<geoCoordinate id="EC909259A722AB4581A7B65B8C49C793" direction="north" orientation="latitude" precision="9" value="18.748667">18°44.92'N</geoCoordinate>
, 
<geoCoordinate id="C088D996CC1B210F9B4BC4E2CA8CF499" direction="west" orientation="longitude" precision="9" value="-70.3605">70°21.63'W</geoCoordinate>
, 636 m a.s.l., 8.ix.2014, leg. Deler, 
<normalizedToken id="DFFFAB6EFA59CA4F9DCF9B5D584CA6BC" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Gimmel (DR41) (4 spec.: NMPC) [DNA 
<pageBreakToken id="AD45A186010CCD5B9C2C52AFE2D4C94D" pageId="19" pageNumber="58" start="start">extracts</pageBreakToken>
: MF1214.1, MF1214.2]. PUERTO RICO: Naguabo: El Yunque National Forest (southern part), 3.45 km N of 
<normalizedToken id="0AFFA7FE16D7FE2E3F9B2C8C4DFB738D" originalValue="Río">Rio</normalizedToken>
Blanco at road PR191, in horse excrements on exposed small pasture on the slope of El Yunque massive, 
<geoCoordinate id="1018F8A4E6EFF2505E9D8FD9E5447C19" direction="north" orientation="latitude" precision="92" value="18.246666">18°14.8'N</geoCoordinate>
, 
<geoCoordinate id="45990D2E66F3F22CC4D9EA40138EF709" direction="west" orientation="longitude" precision="92" value="-65.79833">65°47.9'W</geoCoordinate>
, 
<pageBreakToken id="4E9FC1831A64816985AA1AA2AB89B024" pageId="20" pageNumber="59" start="start">170</pageBreakToken>
m a.s.l., 24.vi.2016, leg. 
<normalizedToken id="D371B99CB4F6FADADCF5C31020AACDBF" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
, 
<normalizedToken id="9A96916879D7FD922EE51AAF77B7181E" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Seidel (PR2a) (19 spec.: NMPC) [DNA extraction: MF1731]; El Yunque National Forest (southern part), 4.9 km N iof 
<normalizedToken id="C22FFB68B9ABECC64E6AA430B117DFC8" originalValue="Río">Rio</normalizedToken>
Blanco, margin of the rainforest in an area with many flowering 
<taxonomicName id="3E35CA8071E895AE0014B21D4425597F" class="Liliopsida" family="Zingiberaceae" genus="Etlingera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Etlingera elatior" order="Zingiberales" pageId="20" pageNumber="59" phylum="Tracheophyta" rank="species" species="elatior">Etlingera elatior</taxonomicName>
plants, FIT, 
<geoCoordinate id="4A44D58BE04BAC65A2DFE7AA3B37CAE6" direction="north" orientation="latitude" precision="92" value="18.263334">18°15.8'N</geoCoordinate>
, 
<geoCoordinate id="82EEBCE7BD721EABFA814C446DE96314" direction="west" orientation="longitude" precision="92" value="-65.78833">65°47.3'W</geoCoordinate>
, 495 m a.s.l., 24.vi.-2.vii.2016, leg. 
<normalizedToken id="2E62BDED4B03C8D037E7B5EB97214C16" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Seidel (PR11) (3 spec.: NMPC). Arecibo: small settlement in Bosque Estatal 
<normalizedToken id="CA016B04AD2CB02494387C0F957180CE" originalValue="Río">Rio</normalizedToken>
Abajo, in the middle of the lowland forest, horse excrement, 
<geoCoordinate id="B62C89738C54B0377DB25B6D5280ACF4" direction="north" orientation="latitude" precision="92" value="18.328333">18°19.7'N</geoCoordinate>
, 
<geoCoordinate id="FCD5359B181880A484A2ADC6CB830FC2" direction="west" orientation="longitude" precision="92" value="-66.70167">66°42.1'W</geoCoordinate>
, 340 m a.s.l., 27.vi.2016, leg. 
<normalizedToken id="0F4DC69A977F9FAD299F315447356839" originalValue="Deler-Hernández">Deler-Hernandez</normalizedToken>
, 
<normalizedToken id="C71384BAAC4311B087ED792B8B9A7712" originalValue="Fikáček">Fikacek</normalizedToken>
&amp; Seidel (PR15) (1 spec.: NMPC). Lesser Antilles: DOMINICA: Springfield Estate, mature secondary forest, FIT, 
<geoCoordinate id="1DEBBD39C0886715F7B8F7CF6BC1593B" direction="north" orientation="latitude" precision="1" value="15.3466">15°20.796'N</geoCoordinate>
, 
<geoCoordinate id="F6EBE7D5C0213A96EE6DEA028A0D57FF" direction="west" orientation="longitude" precision="1" value="-61.369034">61°22.142'W</geoCoordinate>
, 30.v.-16.vi.2004, S. &amp; J. Peck (04-86) (3 spec.: SBP). GRENADA: Grand Etang Forest Reserve, FIT in rainforest, 
<geoCoordinate id="D17C58E151C43CC30416937705271CC1" direction="north" orientation="latitude" precision="1" value="12.080767">12°04.846'N</geoCoordinate>
<geoCoordinate id="C7846F00E70DBFBAA385F00A08BF1B2F" direction="west" orientation="longitude" precision="1" value="-61.70555">61°42.333'W</geoCoordinate>
, 360 m a.s.l., leg. S. Peck (10-61) (2 spec.: SBP); Grand Etang Forest Reserve, FIT in rainforest, 
<geoCoordinate id="96A0BB4BE0496046CA978AFF64A8A3AA" direction="north" orientation="latitude" precision="1" value="12.069366">12°04.162'N</geoCoordinate>
<geoCoordinate id="8405199D771C6D5706A35F3847206EBB" direction="west" orientation="longitude" precision="1" value="-61.7027">61°42.162'W</geoCoordinate>
, 400 m a.s.l., 
<pageBreakToken id="BE16700115F844C39E5AE6FA444C522C" pageId="21" pageNumber="60" start="start">leg</pageBreakToken>
. S. Peck (10-63) (2 spec.: SBP); St. Andrew, Mirabeu Agriculture Lab, light trap, 19.iv.1990, leg. J. Telesford (2 spec.: SBP). SAINT LUCIA: Mon Repos, 6.5 km W Fox Grove Inn, submontane forest, carrion traps, 
<geoCoordinate id="31BDB2E3CCBB55B8AACC5DDF6B49334D" direction="north" orientation="latitude" precision="92" value="13.875">13°52.5'N</geoCoordinate>
, 
<geoCoordinate id="864D72714CD8350E813844A10CC180AA" direction="west" orientation="longitude" precision="92" value="-60.94">60°56.4'W</geoCoordinate>
, 300 m a.s.l., leg. S. &amp; J. Peck (0759) (2 spec.: SBP).
</paragraph>
</subSubSection>
<subSubSection id="3D08090E1A1E3F26B6F223BDA523E482" pageId="21" pageNumber="60" type="published records">
<paragraph id="4BC0324B95E93E3115B585A30A1C8A2A" pageId="21" pageNumber="60">Published records</paragraph>
<paragraph id="586E86AB73D3C278021F560952BD9E5E" pageId="21" pageNumber="60">
(as 
<taxonomicName id="1D478EE88771E6CDD7B6A17211D6EC03" lsidName="C. variegatus" pageId="21" pageNumber="60" rank="species" species="variegatus">C. variegatus</taxonomicName>
). JAMAICA: without precise locality (
<bibRefCitation id="FF69B84679E32BEFA4EF8B9CC85C0556" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="32" pageNumber="71" pagination="1 - 292" title="Revision of the subfamily Sphaeridiinae of America north of Mexico (Coleoptera: Hydrophilidae)." url="https://doi.org/10.4039/entm110105fv" volume="105" year="1978">Smetana 1978</bibRefCitation>
). PUERTO RICO: without precise locality (
<bibRefCitation id="64B5ECCA536CDE9525893F94F56D0F08" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="32" pageNumber="71" pagination="1 - 292" title="Revision of the subfamily Sphaeridiinae of America north of Mexico (Coleoptera: Hydrophilidae)." url="https://doi.org/10.4039/entm110105fv" volume="105" year="1978">Smetana 1978</bibRefCitation>
). DOMINICA: without precise locality (
<bibRefCitation id="5840A504E234A85DC7A0B45D731FBEE3" author="Leng, CW" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="31" pageNumber="70" pagination="191 - 220" title="Supplement to preliminary list of the Coleoptera of the West Indies." volume="37" year="1917">Leng and Mutchler 1917</bibRefCitation>
; 
<bibRefCitation id="77DE07E485CCEB9D057E5539B99748C6" author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="31" pageNumber="70" pagination="165 - 209" title="The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): diversity and distribution." volume="20" year="2006">Peck 2006</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4DCE4D369603B1ED8D494B82EAA360FE" pageId="21" pageNumber="60" type="diagnosis">
<paragraph id="8A96F245874219B3D6F32AB9C144F55C" pageId="21" pageNumber="60">Diagnosis.</paragraph>
<paragraph id="A740A786198B88FB4B00F213BC5F900C" pageId="21" pageNumber="60">
Body size 2.4-3.4 mm; dorsal surface of head (Fig. 4a) black with a pair of small pale spots on vertex (sometimes fused in one central spot), pronotum yellowish to dark reddish-brown with a large blackish central spot and two small round blackish spots at sides (sometimes obscured); elytra yellowish with black humeral spot, in dark specimens whole elytral base and suture darkened; medial ridge of prosternum not projected ventrally (Fig. 12c); mesoventral plate narrow, 5.8 
<normalizedToken id="92F3A0788C37E78C9F9AA95053C905AF" originalValue="×">x</normalizedToken>
as long as wide (Fig. 12f); metaventrite (Fig. 12g) without femoral lines, with raised pentagonal area as long as wide; first abdominal ventrite without spine-like process in both sexes (Fig. 12h); apex of fifth abdominal ventrite (Fig. 12i) without apical projection in both sexes; aedeagus narrow, parameres 0.7 
<normalizedToken id="001AEACC3AC3406E02AB5D8FB2174F50" originalValue="×">x</normalizedToken>
as long as phallobase (Fig. 4f), rounded at apex; median lobe (Fig. 4g) subparallel throughout except for acuminate apex, without subapical spines.
</paragraph>
</subSubSection>
<subSubSection id="D0F7404D83187E90FB42CC8995915D2A" lastPageId="22" lastPageNumber="61" pageId="21" pageNumber="60" type="redescription">
<paragraph id="3F122D7BC7A41A572E6A386F552B0C26" pageId="21" pageNumber="60">Redescription.</paragraph>
<paragraph id="54794C38AD55B5EF3F857F8A944605E5" pageId="21" pageNumber="60">
Body. (Fig. 4 
<normalizedToken id="87EB3D6AE7CCF03F98C3D3DB6C4AFA04" originalValue="a–d">a-d</normalizedToken>
) 2.4-3.4 mm long (length of holotype: 2.8 mm); moderately elongate oval, 1.7 
<normalizedToken id="84D750DA759F7E0362BACABBE00BCC65" originalValue="–1.8×">-1.8x</normalizedToken>
as long as wide, widest at basal fifth of elytra; moderately convex, 2.6 
<normalizedToken id="D31738F5C20BBE265096581E775A23C6" originalValue="–2.8×">-2.8x</normalizedToken>
as long as high (height of holotype: 1.0 mm). Coloration. Dorsal surface of head black with a pair of small rufotestaceous spots on vertex. Antennal scape and flagellum and ventral surface of head including mouthparts light-brown, antennal club and mentum dark brown. Pronotum yellowish to dark reddish-brown, with a large blackish central spot and two small round blackish spots at its sides, sometimes connected with central spot. Prosternum yellowish to light brown, hypomeron slightly darkened. Elytra with elongate blackish spot posterior to humeri, elytral base and suture darkened, elytral epipleura uniformly pale. Ventral surface of mesothorax blackish. Metepisternum brown. Metaventrite blackish, darker at medial elevation. Abdomen yellowish to reddish-brown. Legs yellowish to light brown.
</paragraph>
<paragraph id="2D7311AC00472698925DA8B4BE0AB0A9" lastPageId="22" lastPageNumber="61" pageId="21" pageNumber="60">
Head. Clypeus with moderately dense and shallow punctation consisting of small transverse punctures; interstices without microsculpture. Anterior margin of clypeus with narrow bead. Frontoclypeal suture conspicuous as a zone without punctuation, vanished mesally. Frons with punctation similar to that on clypeus, punctures sparser on sides; interstices without microsculpture. Eyes rather small, interocular distance about 6 
<normalizedToken id="B7821C60D6B03E1B134714E2A3683FAA" originalValue="×">x</normalizedToken>
the width of one eye in dorsal view. Labrum membranous, nearly completely concealed under clypeus, only with narrowly exposed sinuate anterior margin. Mentum (Fig. 12a) subtrapezoid, widest at posterior fourth, about 2 
<normalizedToken id="3C97192A4506BFE2BC259C30548A44E8" originalValue="×">x</normalizedToken>
wider than long, 1.5 
<normalizedToken id="400B0519CDDE1122D441B9BB79E231C0" originalValue="×">x</normalizedToken>
wider at widest part than at anterior margin, strongly concave in anterior half, anterior margin not emarginate; surface almost glabrous, punctures small, shallow and 
<pageBreakToken id="7AD65CC515C2815A9DB84BF86110BAB6" pageId="22" pageNumber="61" start="start">sparse</pageBreakToken>
, almost vanishing anteromesally, interstices without microsculpture. Antenna with 9 antennomeres, scapus ca. 1.8 
<normalizedToken id="F8C81B7573D263DDA26F92EE830AC60E" originalValue="×">x</normalizedToken>
as long antennomeres 2-6 combined; antennal club moderately elongate, about twice as long as wide, as long as scapus; antennomere 9 acuminate at apex.
</paragraph>
<paragraph id="CBA176F2C00D7A714C84F23BE1E9A4C8" pageId="22" pageNumber="61">
Prothorax. Pronotum transverse, widest at base 2.1 
<normalizedToken id="FB4FD2E18CE45586C4A93D19B299F309" originalValue="–2.2×">-2.2x</normalizedToken>
wider than long; 1.7 
<normalizedToken id="521F86415F85FC22F4115F229B95CBF4" originalValue="–1.8×">-1.8x</normalizedToken>
wider at base than between front angles, 1.8 
<normalizedToken id="7FF67C59AEB9E2D1539330E4DFE27A90" originalValue="×">x</normalizedToken>
wider than head including eyes, as convex as pronotum in lateral view. Punctation rather dense and moderately deep, consisting of crescent-shaped punctures intermixed with denser, slightly smaller and rather transverse punctures; punctures slightly feebler on sides. Prosternum (Fig. 12b) strongly tectiform medially, median ridge (Fig. 12c) with the same width throughout, anterior apex not projecting ventrally. Antennal grooves distinct, with lateral margin curved.
</paragraph>
<paragraph id="A5EE0CF123EAB8FF36DDE9611FCA0541" pageId="22" pageNumber="61">
Pterothorax. Scutellar shield 1.2 
<normalizedToken id="73FA3932DAB75EF803ADE5D40B77BA81" originalValue="×">x</normalizedToken>
as long as wide, sparsely punctured. Elytra widest at anterior fifth, 1.0 
<normalizedToken id="9592C958EED4800B349648E989C2DA4C" originalValue="–1.1×">-1.1x</normalizedToken>
longer than wide, 2.6 
<normalizedToken id="5216F3BBFB7C921C2F875164624CACE1" originalValue="–2.8×">-2.8x</normalizedToken>
as long as pronotum, 1.2 
<normalizedToken id="C1E7EB9B19C69CAC7FE7B418AF893BEF" originalValue="–1.3×">-1.3x</normalizedToken>
as wide as pronotum, surface glabrous, with 10 series of punctures; series 6, 8 and 9 not reaching anterior margin, surface glabrous (Fig. 12d), serial punctures getting slightly smaller lateraly; intervals moderately convex; punctation on interval 1 and odd intervals composed of crescent-shaped setiferous punctures, close to striae denser and intermixed with smaller, transverse non-setiferous punctures; even intervals with non-setiferous punctures only; all interstices without microsculpture. Humeral bulge indistinct. Mesoventral plate (Fig. 12f) narrowly elongate, ca. 5.8 
<normalizedToken id="05F08B6FC9B976BDFD599A38A0C5F3FF" originalValue="×">x</normalizedToken>
as long as wide, widest at midlength, symmetrically narrowing to both apices, anterior apex pointed, posterior apex rounded, posterior tip slightly overlapping over anterior portion of metaventrite; surface with few sparse punctures. Metaventrite (Fig. 12g) with raised pentagonal area ca. as wide as long, weakly, sparsely, uniformly punctated, without visible setae, bare part not reaching anterior margin of metaventrite; femoral lines absent; lateral parts of metaventrite densely covered by short pubescence.
</paragraph>
<paragraph id="1053289820B3B8007400B602DC5F3D20" pageId="22" pageNumber="61">
Legs. Femora with sparse shallow punctures ventrally, interstices with weak microsculpture consisting of longitudinal lines; tibial grooves distinct. Tibiae with rather small lateral spines. Metatibiae moderately broad and long, straight, 0.33 
<normalizedToken id="3E8461D69C29492BB7ADE6AA183C4346" originalValue="×">x</normalizedToken>
as long as elytra, 5 
<normalizedToken id="3CA4198D5941984C05BBB2DA364BE44B" originalValue="×">x</normalizedToken>
as long as wide. Metatarsus long, 0.86 
<normalizedToken id="4EA8056214A82DCB381CEC1462D64D47" originalValue="–0.89×">-0.89x</normalizedToken>
as long as metatibia, with just a few short rather stout setae ventrally.
</paragraph>
<paragraph id="6CA574A53C1C0D81061B8B93F7BB97BD" pageId="22" pageNumber="61">Abdomen with five ventrites, first abdominal ventrite (Fig. 12h) about as long as the second and third ventrites together, with distinct median longitudinal carina narrowing posteriad, not projecting posteriorly in both sexes; fifth ventrite with acuminate apex and weakly bulged in both sexes (Fig. 12i).</paragraph>
<paragraph id="C5165391429B312121077423D1720E46" pageId="22" pageNumber="61">
Male genitalia. Median projection of sternite 9 (Fig. 4i) rounded apically, with a pair of subapical setae, base symmetrical. Phallobase (Fig. 4f) almost 1.4 
<normalizedToken id="C94ED935E0557BD3B0C83A66A5BEFD49" originalValue="×">x</normalizedToken>
longer than parameres, narrow, parallel sided, base widely rounded, manubrium indistinct. Parameres nearly of the same width in basal 3/4, divergent near apex, rounded and weakly narrowing apically. Median lobe (Fig. 4g) narrow and subparallel throughout, pointed at apex (Fig. 4h), gonopore moderately large, basal portion of median lobe with dorsal plate narrow and simply bifid basally.
</paragraph>
</subSubSection>
<subSubSection id="ECC792690D6F1ACBCCE2108F5FD72804" pageId="23" pageNumber="62" type="variability">
<paragraph id="14AFC0A81F4F52C0A58D1A43CE58B05F" pageId="23" pageNumber="62">
<pageBreakToken id="B5ED761708BF4BDEB80E0F7BB4063BA8" pageId="23" pageNumber="62" start="start">Variability</pageBreakToken>
.
</paragraph>
<paragraph id="F2800FB0C503212CBEDDB307333AB3AA" pageId="23" pageNumber="62">The general dorsal coloration of the pronotum and elytra varies from yellow to dark reddish-brown. In dark specimens, lateral pronotal spots join the large central spot, and the whole anterior part of elytra and the elytral suture are distinctly darkened, with pale areas maintained in humeral area and at sides of scutellar shield. In pale specimens, the lateral pronotal spots are rather small and sometimes very vague and indistinct, and the elytra are completely yellow except base, sutural interval and the posthumeral dark spots.</paragraph>
</subSubSection>
<subSubSection id="CD5AAC8880B70FAA3E074896936164FE" pageId="23" pageNumber="62" type="distribution">
<paragraph id="DAE673B87B1345674042B68325F1461C" pageId="23" pageNumber="62">Distribution.</paragraph>
<paragraph id="4ADF6C4DF2C9DED9B5E2F62787767139" pageId="23" pageNumber="62">
<taxonomicName id="6C9E703D4F8D2B79B6A87BAE8DD009E7" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="23" pageNumber="62" phylum="Arthropoda" rank="species" species="insularis">Cercyon insularis</taxonomicName>
seems to be widely distributed across Greater and Lesser Antilles, here we are recording it from Cuba, Dominican Republic, Puerto Rico, Grenada, Saint Lucia and Dominica. It seems that all records of 
<taxonomicName id="108A05C8661E8CA9D4CD06A052DD38CC" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
from the Caribbean (Jamaica, Puerto Rico: 
<bibRefCitation id="A1BB732DCBB96720E1A3B47F865E9954" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="32" pageNumber="71" pagination="1 - 292" title="Revision of the subfamily Sphaeridiinae of America north of Mexico (Coleoptera: Hydrophilidae)." url="https://doi.org/10.4039/entm110105fv" volume="105" year="1978">Smetana 1978</bibRefCitation>
; Dominica: 
<bibRefCitation id="29205A9593AA29965135FEC57E9BBA93" author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="31" pageNumber="70" pagination="165 - 209" title="The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): diversity and distribution." volume="20" year="2006">Peck 2006</bibRefCitation>
) actually concern 
<taxonomicName id="2BF952F77CA0A68F04CF0A484CC89BBE" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
, as we failed to find the true 
<taxonomicName id="8037F28C819CC23AA2E72189D2DD78D9" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
in the material examined. For that reason we consider 
<taxonomicName id="40BD310365192B1E4BA24522A2D4799E" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
to occur in Jamaica, although we did not examine any specimens from Jamaica ourselves.
</paragraph>
</subSubSection>
<subSubSection id="D8D1CB5205DC30587365DC4E717AF424" pageId="23" pageNumber="62" type="bionomics">
<paragraph id="3450C45DF2C0681BB65E69523670F461" pageId="23" pageNumber="62">Bionomics.</paragraph>
<paragraph id="90A51FD7035AF01887F0A19DC6316408" pageId="23" pageNumber="62">Most of the specimens were collected in cow and horse dung on pastures, in coffee plantations and in tropical forests; few were collected using flight intercept traps.</paragraph>
</subSubSection>
<subSubSection id="EE68D0AF94F8D08B51C4641B2E8563EC" lastPageId="24" lastPageNumber="63" pageId="23" pageNumber="62" type="discussion">
<paragraph id="82B22E57C8E6A6EAF2558A2F579E051D" pageId="23" pageNumber="62">Discussion.</paragraph>
<paragraph id="5CF99F09510ABF56BB6099DBE063C656" pageId="23" pageNumber="62">
<bibRefCitation id="7E50AC1AE026B4542A369E77EEE69C40" author="Chevrolat, A" journalOrPublisher="Annales de la Societe entomologique de France" pageId="30" pageNumber="69" pagination="183 - 210" title="Coleopteres de I'lle de Cuba. Notes, synonym ieset descriptionsd'especesnouve ll es. Families des Cicindeietes, Carabiques, Dytiscides, Gyrinides et Palpicornes" volume="4" year="1863">Chevrolat (1863)</bibRefCitation>
described 
<taxonomicName id="8310094A6EC0ADC99AF5D0E145FEBD6C" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
based on a single specimen from Cuba collected by D. F. 
<normalizedToken id="2E27D8F10EFC5273609D99FA5F8DA026" originalValue="Poëy">Poey</normalizedToken>
and deposited in Chevrolat's collection. On our request to loan this specimen, we received a single specimen standing under the name 
<taxonomicName id="EBA535E4EFDDE6CA2823E185C3E84C16" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
in the Chevrolat collection, corresponding well with the original description and marked as a type. In contrast to the data mentioned by 
<bibRefCitation id="EC9E5ED458C5B9C53FA6485E6C00435A" author="Chevrolat, A" journalOrPublisher="Annales de la Societe entomologique de France" pageId="30" pageNumber="69" pagination="183 - 210" title="Coleopteres de I'lle de Cuba. Notes, synonym ieset descriptionsd'especesnouve ll es. Families des Cicindeietes, Carabiques, Dytiscides, Gyrinides et Palpicornes" volume="4" year="1863">Chevrolat (1863)</bibRefCitation>
, the specimen also bears a label indicating Habana as the place of its origin. Since there is no reason to doubt the type identity of this specimen, we correct the type locality of 
<taxonomicName id="AAE591CB16CDE76652575CDE582D6A0E" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
to Habana, in agreement with the label data of the holotype.
</paragraph>
<paragraph id="794FFF29963F9157A6E5E7B758C18962" pageId="23" pageNumber="62">
<taxonomicName id="E13169B1B00F8BE0EE5D1D306FF8B445" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="23" pageNumber="62" phylum="Arthropoda" rank="species" species="insularis">Cercyon insularis</taxonomicName>
was only briefly mentioned once by 
<bibRefCitation id="BF0189BB0CEB606840DE12A44285E10D" author="Gundlach, J" journalOrPublisher="Alvarez Comp., Habana" pageId="31" pageNumber="70" title="Constribucion a la entomologia Cubana. Tomo III. A." year="1891">Gundlach (1891)</bibRefCitation>
and its type was not reexamined, therefore its identity remained unclear. Our inspection of the type revealed it corresponds by coloration with what was recorded from the Caribbean as 
<taxonomicName id="52CD03045DBD0A034AEA97FA191AA2AF" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon variegatus" order="Coleoptera" pageId="23" pageNumber="62" phylum="Arthropoda" rank="species" species="variegatus">Cercyon variegatus</taxonomicName>
Sharp, 1882, by 
<bibRefCitation id="D5DD913CAF55AD0506F1DF1FC4B3A32C" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="32" pageNumber="71" pagination="1 - 292" title="Revision of the subfamily Sphaeridiinae of America north of Mexico (Coleoptera: Hydrophilidae)." url="https://doi.org/10.4039/entm110105fv" volume="105" year="1978">Smetana (1978)</bibRefCitation>
, 
<bibRefCitation id="EF624500193D21E9941F39D18336995A" author="Hansen, M" journalOrPublisher="Apollo Books, Stenstrup" pageId="31" pageNumber="70" title="World Catalogue of Insects 2: Hydrophiloidea (Coleoptera)." year="1999">Hansen (1999)</bibRefCitation>
and 
<bibRefCitation id="AE8D5BE45B395B3F7C33A70459EB9F01" author="Peck, SB" journalOrPublisher="Insecta Mundi" pageId="31" pageNumber="70" pagination="165 - 209" title="The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): diversity and distribution." volume="20" year="2006">Peck (2006)</bibRefCitation>
. In order to determine the identity of the species present in the Caribbean we studied the lectotype of 
<taxonomicName id="7BFEDFB7BEE7ED3541F8CC5155E92468" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
(Fig. 4 
<normalizedToken id="106F069737280049EBBF5001B3374223" originalValue="j–o">j-o</normalizedToken>
, deposited in BMNH) and compared it to the type of 
<taxonomicName id="568BD47DF0C953BE4B594C8CDAB576B8" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
and additional recently collected material from Cuba. Based on this comparison, it became clear that Cuban specimens are not conspecific with 
<taxonomicName id="4A786E577519907BA98F8DDDD35FB99F" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
, but belong to a different species indistinguishable from it by external morphology: all dissected Cuban specimens were conspecific, differed from 
<taxonomicName id="D70B8D0ED1E5EA5A4C058FA7031FCEA2" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
by genital morphology, and no other species of the same external coloration was found. We hence consider the Cuban specimens conspecific with the type specimen of 
<taxonomicName id="FE5B3B0B6761636DD32E7CC8F7475038" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
, even though it cannot be dissected because of its poor condition. Both median lobe (Fig. 4g) as well as phallobase and parameres (Fig. 4f) are narrower in 
<taxonomicName id="FA929B5C491BE39AF9D952A7FFEF3FE1" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
than in 
<taxonomicName id="AAF6DF210732FCED6ACF982295205E63" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
. The apices of parameres are rounded in 
<taxonomicName id="E3ACE55642951C928DB52763638D6123" lsidName="C. insularis" pageId="23" pageNumber="62" rank="species" species="insularis">C. insularis</taxonomicName>
, while they are more acuminate in 
<taxonomicName id="0B50680827463B75162D4D967BC477AF" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
(Fig. 4l). Moreover, the apex of the median lobe of 
<taxonomicName id="B13DD9E8279A617E603FC3895EEDD460" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
has a small flank on each side (Fig. 4n) (Full set of pictures of the lectotype of 
<taxonomicName id="07AD574693878E76E5064224E40B2B25" lsidName="C. variegatus" pageId="23" pageNumber="62" rank="species" species="variegatus">C. variegatus</taxonomicName>
in www.flickr.com/photos/142655814@N07/albums/72157676248390724).
</paragraph>
<paragraph id="6C8B4D6EE6F1DB9452A2B5D530B85973" pageId="24" pageNumber="63">
<taxonomicName id="2D5B4ADE0DCC6A5E1EE3675F23C117C2" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="insularis">
<pageBreakToken id="6C16BB6BE9F7E8A6126853D24075AA57" pageId="24" pageNumber="63" start="start">Cercyon</pageBreakToken>
insularis
</taxonomicName>
and 
<taxonomicName id="73A0D14ACAD72750B77505CFDB039D57" lsidName="C. variegatus" pageId="24" pageNumber="63" rank="species" species="variegatus">C. variegatus</taxonomicName>
belong to a species complex corresponding to the 
<taxonomicName id="413AB20A107C3A375E2F2EDBF05BA040" lsidName="C. variegatus" pageId="24" pageNumber="63" rank="species" species="variegatus">C. variegatus</taxonomicName>
group of 
<bibRefCitation id="AA4E38E9B53541E76331E00DFFDC5206" author="Smetana, A" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="32" pageNumber="71" pagination="1 - 292" title="Revision of the subfamily Sphaeridiinae of America north of Mexico (Coleoptera: Hydrophilidae)." url="https://doi.org/10.4039/entm110105fv" volume="105" year="1978">Smetana (1978)</bibRefCitation>
, distributed from the southern USA to Argentina; the species within this complex can be only distinguished by the morphology of the male genitalia (Arriaga-Varela and 
<normalizedToken id="1946184067F2377AF5EB241B320F0C68" originalValue="Fikáček">Fikacek</normalizedToken>
, pers. observation). Only two species of this species complex have been formally described (
<taxonomicName id="107997C204ADD71831FDA35B988FDC71" lsidName="C. variegatus" pageId="24" pageNumber="63" rank="species" species="variegatus">C. variegatus</taxonomicName>
and 
<taxonomicName id="AE1F22B2C4335ADA0C11FD1D44D36884" lsidName="C. insularis" pageId="24" pageNumber="63" rank="species" species="insularis">C. insularis</taxonomicName>
) and the group requires a detailed revision. The species recorded from Suriname as &quot; 
<taxonomicName id="2BBBB0A335C6166F28F0F636A6BF14C0" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon rishwani" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="rishwani">Cercyon rishwani</taxonomicName>
&quot; by 
<bibRefCitation id="AF5642D8CC82ABBD60629B588CDA152C" author="Makhan, D" journalOrPublisher="Calodema" pageId="31" pageNumber="70" pagination="11 - 26" title="Hydrochidae of the World, Dryopidae and Hydrophilidae (Coleoptera)." volume="2" year="2004">Makhan (2004)</bibRefCitation>
also belongs to this species complex based on color pattern of the pronotum and the general shape of the aedeagus, but a more detailed comparison with 
<taxonomicName id="F47247F94EF428229CEA4C839F398F38" lsidName="C. insularis" pageId="24" pageNumber="63" rank="species" species="insularis">C. insularis</taxonomicName>
and 
<taxonomicName id="D09545B34D413CAF65670F6B41883328" lsidName="C. variegatus" pageId="24" pageNumber="63" rank="species" species="variegatus">C. variegatus</taxonomicName>
is impossible based on the description and illustrations provided. &quot; 
<taxonomicName id="2B06BA5C9CB32571F716306F8FBCC383" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon rishwani" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="rishwani">Cercyon rishwani</taxonomicName>
Makhan, 2004&quot; is moreover considered a nomen nudum (see Short and Hebauer, 2006 for details).
</paragraph>
<caption id="33451478682F9FAD507CC224E9204C5B" pageId="24" pageNumber="63">
<paragraph id="163D4126A9ABEE419B83246221B9262F" pageId="24" pageNumber="63">
Figure 8. 
<taxonomicName id="AF43E3F07207146BFB15EE0B8D246EE2" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon gimmeli" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="gimmeli">Cercyon gimmeli</taxonomicName>
sp. n. a mentum b prosternum c lateral view of median ridge of prosternum d detail of elytral surface f mesoventral plate g metaventrite h median ridge of first abdominal ventrite i fifth abdominal ventrite.
</paragraph>
</caption>
<caption id="EF61709A506C7A33E40041C81C6E2503" pageId="24" pageNumber="63">
<paragraph id="FB6E95B4C86E7D630A8442851386750C" pageId="24" pageNumber="63">
Figure 9. 
<taxonomicName id="64369E3404A8CEF539B55CFE4ECC5844" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="genus">Cercyon</taxonomicName>
spp. n. 
<normalizedToken id="4174F632FAEA103F2FE79142C8198E36" originalValue="a–d">a-d</normalizedToken>
<taxonomicName id="82379163DE08B41ACE1F52EA37E0096D" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon armatipenis" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="armatipenis">Cercyon armatipenis</taxonomicName>
sp. n. 
<normalizedToken id="E37F6F48BADA603A2ACB3424C0D2BBCB" originalValue="f–i">f-i</normalizedToken>
<taxonomicName id="E9C71382D5BCFD384F48A7DD7BE66F1D" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon taino" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="taino">Cercyon taino</taxonomicName>
sp. n. a, f mentum b detail of pronotal surface c, h ventral view of pterothorax d, i detail of elytral surface g lateral view of median ridge of prosternum.
</paragraph>
</caption>
<caption id="CF266FBAA89764175821C7C3530483D7" pageId="24" pageNumber="63">
<paragraph id="9742D0850D027B25B05FB24A5A5B90FE" pageId="24" pageNumber="63">
Figure 10. 
<taxonomicName id="04DDB4074D6CF920068FCC61FF0C4BA3" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon sklodowskae" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="sklodowskae">Cercyon sklodowskae</taxonomicName>
sp. n. a mentum b detail of pronotal surface c lateral view of median ridge of prosternum d detail of elytral surface f mesoventral plate g metaventrite h median ridge of first abdominal ventrite i female fifth abdominal ventrite.
</paragraph>
</caption>
<caption id="9116BFC427954BAE2A73F14F122B512E" pageId="24" pageNumber="63">
<paragraph id="81CE1464B56644ABA5C696E0418745F4" pageId="24" pageNumber="63">
Figure 11. 
<taxonomicName id="8B75522026B65DFC6A997519F974C653" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon spiniventris" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="spiniventris">Cercyon spiniventris</taxonomicName>
sp. n. a mentum b prosternum c lateral view of median ridge of prosternum d detail of elytral surface f mesoventral plate g metaventrite h median ridge of female first abdominal ventrite i fifth abdominal ventrite.
</paragraph>
</caption>
<caption id="0B324F995124338350EE55F0A2C5E364" pageId="24" pageNumber="63">
<paragraph id="0C512AFB9362E18860EF30D1F546883C" pageId="24" pageNumber="63">
Figure 12. 
<taxonomicName id="4E4C61E2AD0C497E829CB6B57C60895C" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon insularis" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="insularis">Cercyon insularis</taxonomicName>
Chevrolat a mentum b prosternum c lateral view of median ridge of prosternum d detail of elytral surface f mesoventral plate g metaventrite h median ridge of female first abdominal ventrite i fifth abdominal ventrite.
</paragraph>
</caption>
<caption id="F0EAEF593F7AB15F0966074090CB3A0B" pageId="24" pageNumber="63">
<paragraph id="65112CC7542F4AEED1C0AF0FDE60108E" pageId="24" pageNumber="63">
Figure 13. 
<taxonomicName id="8D0730C228AE7324E4DF25F805D860F2" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="genus">Cercyon</taxonomicName>
spp. 
<normalizedToken id="E19605A7105BB77AA2DE378C48B9BF1C" originalValue="a–b">a-b</normalizedToken>
<taxonomicName id="B2BF82A58B7FF2B1E4BC5D3612085969" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon praetextatus" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="praetextatus">Cercyon praetextatus</taxonomicName>
Say 
<normalizedToken id="ECDA8525280BCFE8EFF5014B7BFD3E6E" originalValue="c–d">c-d</normalizedToken>
<taxonomicName id="C1DC2A6116CC7EFE8D97FB52FDA24F80" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon nigriceps" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="nigriceps">Cercyon nigriceps</taxonomicName>
Marsham 
<normalizedToken id="BBE489ADD07670580032768BC5CA6467" originalValue="e–f">e-f</normalizedToken>
<taxonomicName id="FA1EE524B7EC465EA282190D5CC0E1D5" class="Insecta" family="Hydrophilidae" genus="Cercyon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cercyon quisquilius" order="Coleoptera" pageId="24" pageNumber="63" phylum="Arthropoda" rank="species" species="quisquilius">Cercyon quisquilius</taxonomicName>
Linnaeus a, c, e mesoventral plate b, d, f metaventrite.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>