305 lines
58 KiB
XML
305 lines
58 KiB
XML
<document id="AF3AC06322963BCD99C3942CE0DD323C" ID-DOI="10.11646/zootaxa.3664.2.1" ID-GBIF-Dataset="a50b71f5-48a6-464f-aea1-f29bbc8ee322" ID-ISSN="1175-5326" ID-Zenodo-Dep="220503" ID-ZooBank="105D178E-6DBF-401D-85BB-CDB041B67E91" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460615203542" checkinUser="plazi" docAuthor="Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin & Zhou, Hongzhang" docDate="2013" docId="03AA1008564AFFFBC1D8FDD2FDC1FB91" docLanguage="en" docName="zt03664p135.pdf" docOrigin="Zootaxa 3664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) stictopleurus subsp. cangshanensis Shi & Sciaky, new subspecies" docType="treatment" docVersion="9" lastPageNumber="119" masterDocId="FF936870565BFFE9C14FFF95FFA9FF9E" masterDocTitle="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions" masterLastPageNumber="135" masterPageNumber="101" pageNumber="118" updateTime="1698330049886" updateUser="plazi">
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<mods:titleInfo id="5CC6145CA054142ED1A0243A622D79CA">
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<mods:title id="43D41A68AA4D3C78C9CB80D6A7666AA3">A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions</mods:title>
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</mods:titleInfo>
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<mods:name id="4020E5C85D22328107452BFC57F53E07" type="personal">
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<mods:roleTerm id="99BE1DAD9AB14BCDB71C9CD167A85383">Author</mods:roleTerm>
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<mods:namePart id="9F8BC922F4F353D0604935B4BBB40ECF">Shi, Hongliang</mods:namePart>
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<mods:roleTerm id="7A1C0CBBF572431A456CAB36D8DF0099">Author</mods:roleTerm>
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<mods:namePart id="60502DBD2F440A1722CC04A26DBF2DA8">Sciaky, Riccardo</mods:namePart>
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<mods:roleTerm id="5780102FB962CFCE4A8AD83865498D78">Author</mods:roleTerm>
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<mods:namePart id="591649B540086115BAFA73E4FF82BBE9">Liang, Hongbin</mods:namePart>
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<mods:namePart id="C5F7D66E0C3453AE4235DA1A8C22399C">Zhou, Hongzhang</mods:namePart>
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<mods:title id="1F448345108FFF45EC5D2B725239CF98">Zootaxa</mods:title>
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<mods:part id="70704D2353CC681399A7B3B8DA924B8A">
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<mods:date id="7DA83500F0DD55410B1955365002905E">2013</mods:date>
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<mods:number id="81ABA4A418622620978447D7EB205600">3664</mods:number>
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<mods:classification id="9BEEC5FCC76FE662C9190A590FA5D4C4">journal article</mods:classification>
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<mods:identifier id="EF9C9B34C784C15ADBBBE4C66A632BE8" type="DOI">10.11646/zootaxa.3664.2.1</mods:identifier>
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<treatment id="03AA1008564AFFFBC1D8FDD2FDC1FB91" ID-DOI="http://doi.org/10.5281/zenodo.6154517" ID-GBIF-Taxon="119568524" ID-Zenodo-Dep="6154517" LSID="urn:lsid:plazi:treatment:03AA1008564AFFFBC1D8FDD2FDC1FB91" httpUri="http://treatment.plazi.org/id/03AA1008564AFFFBC1D8FDD2FDC1FB91" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118">
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<subSubSection id="C319F295564AFFF8C1D8FDD2FD01FD1C" pageId="17" pageNumber="118" type="nomenclature">
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<paragraph id="8BBCA11E564AFFF8C1D8FDD2FB08FDFF" blockId="17.[151,1185,583,642]" box="[151,1185,583,609]" pageId="17" pageNumber="118">
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<heading id="D0F41672564AFFF8C1D8FDD2FB08FDFF" bold="true" box="[151,1185,583,609]" fontSize="11" level="1" pageId="17" pageNumber="118" reason="1">
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<emphasis id="B9777D0C564AFFF8C1D8FDD2FB08FDFF" bold="true" box="[151,1185,583,609]" pageId="17" pageNumber="118">
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<taxonomicName id="4C03DA9D564AFFF8C1D8FDD2FC71FDFF" ID-CoL="8MFXD" authority="Shi & Sciaky" authorityName="Shi & Sciaky" box="[151,984,583,609]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" status="subsp. nov." subGenus="Wraseiellus" subSpecies="cangshanensis">
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<emphasis id="B9777D0C564AFFF8C1D8FDD2FE84FDFF" bold="true" box="[151,301,583,609]" italics="true" pageId="17" pageNumber="118">Pterostichus</emphasis>
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(
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<emphasis id="B9777D0C564AFFF8C072FDD2FE61FDFF" bold="true" box="[317,456,583,609]" italics="true" pageId="17" pageNumber="118">Wraseiellus</emphasis>
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)
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<emphasis id="B9777D0C564AFFF8C095FDD2FC87FDFF" bold="true" box="[474,814,583,609]" italics="true" pageId="17" pageNumber="118">stictopleurus cangshanensis</emphasis>
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Shi & Sciaky
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</taxonomicName>
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,
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<taxonomicNameLabel id="A244C077564AFFF8C2AAFDD2FB08FDFF" box="[997,1185,583,609]" pageId="17" pageNumber="118" rank="subSpecies">new subspecies</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBCA11E564AFFF8C1D8FDFFFD01FD1C" blockId="17.[151,1185,583,642]" box="[151,680,618,642]" pageId="17" pageNumber="118">
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(
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<figureCitation id="1338BD9B564AFFF8C1D1FDFFFEB0FD1C" box="[158,281,618,642]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="17" pageNumber="118">Figures 12</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C06AFDFFFEEAFD1C" box="[293,323,618,642]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="17" pageNumber="118">25</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C000FDFFFEC4FD1C" box="[335,365,618,642]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="17" pageNumber="118">36</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C036FDFFFE31FD1C" box="[377,408,618,642]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 43, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="17" pageNumber="118">44</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C0ECFDFFFE42FD1C" box="[419,491,618,642]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="17" pageNumber="118">65, 66</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C0B8FDFFFDBCFD1C" box="[503,533,618,642]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from " Yunnan "; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 77, Pt. diversus (Fairmaire), a male from " Yunnan "; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="17" pageNumber="118">73</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C36EFDFFFD96FD1C" box="[545,575,618,642]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="17" pageNumber="118">88</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C304FDFFFDC0FD1C" box="[587,617,618,642]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="17" pageNumber="118">91</figureCitation>
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,
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<figureCitation id="1338BD9B564AFFF8C33AFDFFFD09FD1C" box="[629,672,618,642]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from " Yunnan "; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="17" pageNumber="118">105</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C319F295564AFFF8C1D8FD26FDF2FB50" pageId="17" pageNumber="118" type="materials_examined">
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<paragraph id="8BBCA11E564AFFF8C1D8FD26FC4FFD54" blockId="17.[151,1436,690,2022]" box="[151,998,690,715]" pageId="17" pageNumber="118">
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<emphasis id="B9777D0C564AFFF8C1D8FD26FE84FD55" bold="true" box="[151,301,690,715]" pageId="17" pageNumber="118">
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<typeStatus id="54B81FBC564AFFF8C1D8FD26FF7BFD55" box="[151,210,691,715]" pageId="17" pageNumber="118">Type</typeStatus>
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locality
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</emphasis>
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. Yunnan: Dali, E. slope of Cangshan Mt. (
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<geoCoordinate id="EE37C7D9564AFFF8C245FD27FCC3FD54" box="[778,874,690,714]" direction="north" orientation="latitude" pageId="17" pageNumber="118" precision="555" value="25.63">N25.63º</geoCoordinate>
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,
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<geoCoordinate id="EE37C7D9564AFFF8C23AFD27FC77FD54" box="[885,990,690,714]" direction="east" orientation="longitude" pageId="17" pageNumber="118" precision="555" value="100.14">E100.14º</geoCoordinate>
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)
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</paragraph>
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<paragraph id="8BBCA11E564AFFF8C188FD42FDF2FB50" blockId="17.[151,1436,690,2022]" pageId="17" pageNumber="118">
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<emphasis id="B9777D0C564AFFF8C188FD42FED4FD71" bold="true" box="[199,381,726,751]" pageId="17" pageNumber="118">
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<typeStatus id="54B81FBC564AFFF8C188FD42FEADFD71" box="[199,260,727,751]" pageId="17" pageNumber="118">Type</typeStatus>
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materials
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</emphasis>
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||
.
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<emphasis id="B9777D0C564AFFF8C0C1FD42FE52FD71" bold="true" box="[398,507,727,751]" pageId="17" pageNumber="118">
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<typeStatus id="54B81FBC564AFFF8C0C1FD42FE52FD71" box="[398,507,727,751]" pageId="17" pageNumber="118" type="holotype">Holotype</typeStatus>
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</emphasis>
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(IZAS), male, body length =
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<quantity id="4CFB0CFB564AFFF8C217FD43FC69FD70" box="[856,960,726,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="17" pageNumber="118" unit="mm" value="12.8">12.8 mm</quantity>
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, pinned, genitalia dissected and glued on plastic film pinned under specimen, "
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<collectingCountry id="F314E18E564AFFF8C378FD6FFD3CFC8C" box="[567,661,762,786]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
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, Yunnan prov., / Dali, E. slope of Cang- / shan Mt., 2008.
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<date id="FFBD87DE564AFFF8C46DFD6FFAF1FC8C" box="[1314,1368,762,786]" pageId="17" pageNumber="118" value="1918-10">X.18</date>
|
||
, / leg. HUANG Rong etc., / NO.2008101801"; " / 2008.
|
||
<date id="FFBD87DE564AFFF8C2BDFC8BFB8EFCA8" box="[1010,1063,798,822]" pageId="17" pageNumber="118" value="1918-10">X.18</date>
|
||
, / NO.2008101801"; "IOZ(E)1891916"; "
|
||
<typeStatus id="54B81FBC564AFFF8C034FCD7FDBBFCC4" box="[379,530,834,858]" pageId="17" pageNumber="118" type="holotype">HOLOTYPE</typeStatus>
|
||
3 /
|
||
<taxonomicName id="4C03DA9D564AFFF8C306FCD7FCDBFCC4" authority="Wraseiellus" authorityName="Wraseiellus" box="[585,882,834,858]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="genus">Pterostichus (Wraseiellus)</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C03DA9D564AFFF8C2C4FCD7FB6AFCC4" box="[907,1219,834,858]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="118" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="cangshanensis">stictopleurus cangshanensis</taxonomicName>
|
||
/ new subspecies / Des. SHI H.L. 2012" [red label] [
|
||
<figureCitation id="1338BD9B564AFFF8C340FCF3FDEDFCEA" box="[527,580,870,884]" captionStart="FIGURES 7 – 12" captionStartId="22.[151,264,1765,1788]" captionTargetBox="[167,1429,198,1733]" captionTargetId="figure@22.[151,1436,183,1744]" captionTargetPageId="22" captionText="FIGURES 7 – 12. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 7, Pt. crassiapex n. sp. holotype; 8, Pt. near crassiapex n. sp., a female from Liziping; 9, Pt. diversus (Fairmaire), a female syntype; 10, Pt. diversus (Fairmaire), a male from Yulongshan; 11, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 12, Pt. stictopleurus cangshanensis n. ssp., holotype." httpUri="https://zenodo.org/record/220505/files/figure.png" pageId="17" pageNumber="118">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C304FCF3FDF5FCEA" box="[587,604,870,884]" captionStart="FIGURES 19 – 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 – 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="17" pageNumber="118">25</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C32CFCF3FDDDFCEA" box="[611,628,870,884]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="17" pageNumber="118">36</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C335FCF3FD22FCEA" box="[634,651,870,884]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 43, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="17" pageNumber="118">44</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C3DDFCF3FD0AFCEA" box="[658,675,870,884]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="17" pageNumber="118">65</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C3E5FCF3FD12FCEA" box="[682,699,870,884]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from " Yunnan "; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 77, Pt. diversus (Fairmaire), a male from " Yunnan "; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="17" pageNumber="118">73</figureCitation>
|
||
,
|
||
<figureCitation id="1338BD9B564AFFF8C38DFCF3FD73FCEA" box="[706,730,870,884]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from " Yunnan "; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="17" pageNumber="118">105</figureCitation>
|
||
].
|
||
<emphasis id="B9777D0C564AFFF8C3A2FCFFFCCCFC1C" bold="true" box="[749,869,874,898]" pageId="17" pageNumber="118">
|
||
<typeStatus id="54B81FBC564AFFF8C3A2FCFFFCCCFC1C" box="[749,869,874,898]" pageId="17" pageNumber="118" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(total
|
||
<specimenCount id="9D056A97564AFFF8C2FFFCFCFBE3FC1F" box="[944,1098,873,897]" pageId="17" pageNumber="118" type="generic">18 specimens</specimenCount>
|
||
all from Dali):
|
||
<specimenCount id="9D056A97564AFFF8C5B1FCFCFAFCFC1F" box="[1278,1365,873,897]" pageId="17" pageNumber="118" type="male">3 males</specimenCount>
|
||
and
|
||
<specimenCount id="9D056A97564AFFF8C4C1FCFCFF47FC3B" pageId="17" pageNumber="118" type="female">2 females</specimenCount>
|
||
(IZAS), the same label data as
|
||
<typeStatus id="54B81FBC564AFFF8C313FC18FD68FC3B" box="[604,705,909,933]" pageId="17" pageNumber="118" type="holotype">holotype</typeStatus>
|
||
, 2 of them (
|
||
<specimenCount id="9D056A97564AFFF8C21EFC18FC36FC3B" box="[849,927,909,933]" pageId="17" pageNumber="118" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D056A97564AFFF8C2E3FC18FBB8FC3B" box="[940,1041,909,933]" pageId="17" pageNumber="118" type="female">1 female</specimenCount>
|
||
) with different IOZ database label (1891914, 1891915), others without database label.
|
||
<specimenCount id="9D056A97564AFFF8C3B9FC24FCECFC57" box="[758,837,945,969]" pageId="17" pageNumber="118" type="male">1 male</specimenCount>
|
||
(IZAS), "
|
||
<collectingCountry id="F314E18E564AFFF8C2F3FC24FBB3FC57" box="[956,1050,945,969]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov., Dali, S. slope of Cangshan Mt., 2008.
|
||
<date id="FFBD87DE564AFFF8C0CAFC40FE11FC73" box="[389,440,981,1005]" pageId="17" pageNumber="118">IX.6</date>
|
||
, leg. LI Yunchun etc., NO.2008090608".
|
||
<specimenCount id="9D056A97564AFFF8C2DCFC40FC76FC73" box="[915,991,981,1005]" pageId="17" pageNumber="118" type="male">1 male</specimenCount>
|
||
(IZAS), "
|
||
<collectingCountry id="F314E18E564AFFF8C51FFC40FB07FC73" box="[1104,1198,981,1005]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov., Dali, E. slope of Cangshan Mt., 2008.
|
||
<date id="FFBD87DE564AFFF8C345FC6CFD95FB8F" box="[522,572,1017,1041]" pageId="17" pageNumber="118">IX.6</date>
|
||
, leg. LI Yunchun etc., NO.2008090608".
|
||
<specimenCount id="9D056A97564AFFF8C555FC6CFBCFFB8F" box="[1050,1126,1017,1041]" pageId="17" pageNumber="118" type="male">1 male</specimenCount>
|
||
(IZAS), "
|
||
<collectingCountry id="F314E18E564AFFF8C597FC6CFA9EFB8F" box="[1240,1335,1017,1041]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov., Dali, E. slope of Cangshan Mt., 2008.
|
||
<date id="FFBD87DE564AFFF8C3D6FBB5FD70FBA6" box="[665,729,1056,1080]" pageId="17" pageNumber="118" value="1913-09">IX.13</date>
|
||
, leg. ZHANG Youliang etc., NO.2008091301" [
|
||
<figureCitation id="1338BD9B564AFFF8C44FFB88FA86FBB5" box="[1280,1327,1053,1067]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="17" pageNumber="118">Fig. 66</figureCitation>
|
||
].
|
||
<specimenCount id="9D056A97564AFFF8C40CFBB5FA35FBA6" box="[1347,1436,1056,1080]" pageId="17" pageNumber="118" type="male">2 males</specimenCount>
|
||
(CRS), "
|
||
<collectingCountry id="F314E18E564AFFF8C1B5FBD1FEF1FBC2" box="[250,344,1092,1116]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov. Dali,
|
||
<date id="FFBD87DE564AFFF8C31AFBD1FD13FBC2" box="[597,698,1092,1116]" pageId="17" pageNumber="118" value="1993-07-06">6.7.1993</date>
|
||
, lgt.S.Becvar".
|
||
<specimenCount id="9D056A97564AFFF8C23DFBD1FC79FBC2" box="[882,976,1092,1116]" pageId="17" pageNumber="118" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D056A97564AFFF8C2AFFBD1FBF8FBC2" box="[992,1105,1092,1116]" pageId="17" pageNumber="118" type="female">3 females</specimenCount>
|
||
(SCAU), "
|
||
<collectingCountry id="F314E18E564AFFF8C59AFBD1FA9AFBC2" box="[1237,1331,1092,1116]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov., Dali, Cangshan Mt., 2009.
|
||
<date id="FFBD87DE564AFFF8C345FBFEFD39FB1D" box="[522,656,1131,1155]" pageId="17" pageNumber="118" value="2600-06-25">
|
||
VI.25,
|
||
<quantity id="4CFB0CFB564AFFF8C319FBFEFD39FB1D" box="[598,656,1131,1155]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="118" unit="m" value="2600.0">2600</quantity>
|
||
</date>
|
||
m, leg. YANG Zhenxing, ZHANG Youliang etc., S1986WDm" [
|
||
<figureCitation id="1338BD9B564AFFF8C42CFBFDFA39FBE8" box="[1379,1424,1128,1142]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="17" pageNumber="118">Fig. 91</figureCitation>
|
||
].
|
||
<specimenCount id="9D056A97564AFFF8C1D8FB1AFF4EFB39" box="[151,231,1167,1191]" pageId="17" pageNumber="118" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D056A97564AFFF8C1BAFB1AFEF0FB39" box="[245,345,1167,1191]" pageId="17" pageNumber="118" type="female">1 female</specimenCount>
|
||
(SCAU), "
|
||
<collectingCountry id="F314E18E564AFFF8C096FB1AFD9EFB39" box="[473,567,1167,1191]" name="China" pageId="17" pageNumber="118">CHINA</collectingCountry>
|
||
, Yunnan prov., Dali, Cangshan Mt., 2009.
|
||
<date id="FFBD87DE564AFFF8C56AFB1AFB00FB39" box="[1061,1193,1167,1191]" pageId="17" pageNumber="118" value="2500-07-08">
|
||
VII.8,
|
||
<quantity id="4CFB0CFB564AFFF8C520FB1AFB00FB39" box="[1135,1193,1167,1191]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="118" unit="m" value="2500.0">2500</quantity>
|
||
</date>
|
||
m, leg. Ma Yanhong, WANG Mingtao etc., S1991EDo" [
|
||
<figureCitation id="1338BD9B564AFFF8C36DFB26FDF9FB5F" box="[546,592,1203,1217]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="17" pageNumber="118">Fig. 88</figureCitation>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319F295564AFFF8C188FB4CFB6BF9A8" pageId="17" pageNumber="118" type="diagnosis">
|
||
<paragraph id="8BBCA11E564AFFF8C188FB4CFC98FAA7" blockId="17.[151,1436,690,2022]" pageId="17" pageNumber="118">
|
||
<emphasis id="B9777D0C564AFFF8C188FB4CFEE8FB6C" bold="true" box="[199,321,1241,1266]" pageId="17" pageNumber="118">Diagnosis.</emphasis>
|
||
The new subspecies can be distinguished from all the other members of this subgenus by the combination of the following characters: (1) 5th tarsomere setose beneath; (2) males with elytral microsculpture isodiametric; (3) pronotum hind angles obtuse, not pointed.
|
||
</paragraph>
|
||
<paragraph id="8BBCA11E564AFFF8C188FAD3FB6BF9A8" blockId="17.[151,1436,690,2022]" pageId="17" pageNumber="118">
|
||
Comparing with the nominotypical subspecies, the new subspecies is different in: (1) 5th tarsomere setose beneath (glabrous in the nominotypical subspecies); (2) ridge between outer groove of basal fovea and lateral margin of pronotum stronger than in the nominotypical subspecies (
|
||
<figureCitation id="1338BD9B564AFFF8C2DAFA1BFB92FA38" box="[917,1083,1422,1446]" captionStart="FIGURES 98 – 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 – 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from " Yunnan "; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="17" pageNumber="118">Figs. 104, 105</figureCitation>
|
||
); (3) punctures on proepisterna much finer and sparser than in the nominotypical subspecies; (4) outer margin of right paramere slightly curved (straight in the nominotypical subspecies) (
|
||
<figureCitation id="1338BD9B564AFFF8C338FA43FD7CFA70" box="[631,725,1494,1518]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from " Yunnan "; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 77, Pt. diversus (Fairmaire), a male from " Yunnan "; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="17" pageNumber="118">Figs. 72</figureCitation>
|
||
B, 73B); (5) median lobe of male genitalia straight before apical lamella (in lateral view), and slightly bent to the dorsal side at apex (
|
||
<figureCitation id="1338BD9B564AFFF8C299FA6CFBC8F98F" box="[982,1121,1529,1553]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="17" pageNumber="118">Figs 65, 66</figureCitation>
|
||
), but in the nominotypical subspecies, median lobe slightly sinuate and then strongly bent to the dorsal side (
|
||
<figureCitation id="1338BD9B564AFFF8C563F98BFB1FF9A8" box="[1068,1206,1566,1590]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="17" pageNumber="118">Figs. 63, 64</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319F295564AFFFBC188F9D4FC9DFD39" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118" type="description">
|
||
<paragraph id="8BBCA11E564AFFFBC188F9D4FC9DFD39" blockId="17.[151,1436,690,2022]" lastBlockId="18.[151,1437,151,1040]" lastPageId="18" lastPageNumber="119" pageId="17" pageNumber="118">
|
||
<emphasis id="B9777D0C564AFFF8C188F9D4FEF8F9C4" bold="true" box="[199,337,1601,1626]" pageId="17" pageNumber="118">Description</emphasis>
|
||
. Body length
|
||
<quantity id="4CFB0CFB564AFFF8C0BCF9D4FD33F9C7" box="[499,666,1601,1625]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.34" metricValueMin="1.16" pageId="17" pageNumber="118" unit="mm" value="12.5" valueMax="13.4" valueMin="11.6">11.6–13.4 mm</quantity>
|
||
; dorsal side black, shining in males; tibiae dark brown, tarsi brown; ventral side black or dark brown. Males with elytral microsculpture isodiametric.
|
||
<emphasis id="B9777D0C564AFFF8C533F9F3FB15F9E0" bold="true" box="[1148,1212,1638,1662]" pageId="17" pageNumber="118">Head</emphasis>
|
||
. Frons and vertex impunctate, few punctures present in frontal furrows; antennae reaching elytra basal eighth; 3rd antennomere glabrous except apical setae; males with the terminal labial palpomere similar as in females, slightly expanded and truncate.
|
||
<emphasis id="B9777D0C564AFFF8C04FF947FED3F974" bold="true" box="[256,378,1746,1770]" pageId="17" pageNumber="118">Pronotum</emphasis>
|
||
rounded, widest at about middle, PW/PL = 1.32–1.39; lateral margins fully rounded near the middle and then gradually curved posteriorly, almost straight before hind angles; hind angles obtuse, not pointed; one mid-lateral seta present at about anterior third; basal fovea shallow, faintly defined; inner groove of basal fovea weak, subparallel to median line, gradually disappeared posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about two-third length as inner one, somewhat more distinct than the inner one, reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct, weakly convex; area between inner and outer basal foveal grooves slightly convex and rugose; basal foveal area densely punctate, some sparse punctures usually present in the area between two basal foveae.
|
||
<emphasis id="B9777D0C564AFFF8C533F85BFB61F878" bold="true" box="[1148,1224,1998,2022]" pageId="17" pageNumber="118">Elytra</emphasis>
|
||
with base slightly depressed between 3rd and 5th intervals; elytral shoulders slightly narrowed, humeral teeth small but distinct; intervals feebly convex; striae moderately deep, with very faint punctures; scutellar striae complete or incomplete, between 1st stria and elytral suture or between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 15–17 pores (5–6, 2–3, 7–8).
|
||
<emphasis id="B9777D0C5649FFFBC3ABFE91FCD1FE82" bold="true" box="[740,888,259,284]" pageId="18" pageNumber="119">Ventral side</emphasis>
|
||
. Proepisterna finely rugose, usually with sparse and fine punctures at anterior half; metaepisterna finely punctate; terminal sternum of males without special structure.
|
||
<emphasis id="B9777D0C5649FFFBC044FED9FEEAFEFA" bold="true" box="[267,323,332,356]" pageId="18" pageNumber="119">Legs</emphasis>
|
||
. 5th tarsomere with one to three pairs of setae beneath.
|
||
<emphasis id="B9777D0C5649FFFBC2AAFED9FB3DFEFA" bold="true" box="[997,1172,331,356]" pageId="18" pageNumber="119">Male genitalia</emphasis>
|
||
. Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifth, ventral margin almost straight in the middle (
|
||
<figureCitation id="1338BD9B5649FFFBC421FEFAFF1CFE32" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="18" pageNumber="119">Fig. 36</figureCitation>
|
||
A); apical part straight and then the apical lamella slightly bent to the dorsal side (
|
||
<figureCitation id="1338BD9B5649FFFBC52BFE01FB58FE32" box="[1124,1265,404,428]" captionStart="FIGURES 57 – 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 – 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from " Yunnan "; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 64, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="18" pageNumber="119">Figs. 65, 66</figureCitation>
|
||
); apical orifice large, slightly turned to the left side, apical lamella twisted to the left (
|
||
<figureCitation id="1338BD9B5649FFFBC287FE22FBB7FE51" box="[968,1054,439,463]" captionStart="FIGURES 35 – 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 – 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="18" pageNumber="119">Fig. 36</figureCitation>
|
||
B); apical lamella laminar, length about twice as its basal width, apex slightly capitate and rounded, hardly bent to the left (
|
||
<figureCitation id="1338BD9B5649FFFBC5CAFE49FB73FE6A" box="[1157,1242,476,500]" captionStart="FIGURES 41 – 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 – 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 43, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 – 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 – 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="18" pageNumber="119">Fig. 44</figureCitation>
|
||
). Right paramere dirk-form, elongate and curved, right margin slightly curved, narrowed apically, apex sharp (
|
||
<figureCitation id="1338BD9B5649FFFBC58AFE6AFAB2FD89" box="[1221,1307,511,535]" captionStart="FIGURES 67 – 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 – 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from " Yunnan "; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from " Ta-pin-tze "; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 – 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from " Ta-pin-tze "; 77, Pt. diversus (Fairmaire), a male from " Yunnan "; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="18" pageNumber="119">Fig. 73</figureCitation>
|
||
B).
|
||
<emphasis id="B9777D0C5649FFFBC40AFD95FF57FDA2" bold="true" pageId="18" pageNumber="119">Female genitalia</emphasis>
|
||
(
|
||
<figureCitation id="1338BD9B5649FFFBC041FDB1FEC8FDA2" box="[270,353,548,572]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="18" pageNumber="119">Fig. 88</figureCitation>
|
||
) very similar to those of
|
||
<taxonomicName id="4C03DA9D5649FFFBC3CAFDB0FB8BFDA2" authority="Fairmaire. Spermatheca" authorityName="Fairmaire. Spermatheca" box="[645,1058,548,572]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="species" species="diversus">
|
||
<emphasis id="B9777D0C5649FFFBC3CAFDB0FCA2FDA2" box="[645,779,548,572]" italics="true" pageId="18" pageNumber="119">Pt. diversus</emphasis>
|
||
Fairmaire. Spermatheca
|
||
</taxonomicName>
|
||
with the seminal canal about ten times long as the receptaculum; spermathecal gland strongly expanded; the seminal canal inserted at the base of common oviduct, base of seminal canal sclerotized (as in
|
||
<figureCitation id="1338BD9B5649FFFBC26FFDF9FCDBFD1A" box="[800,882,620,644]" captionStart="FIGURES 85 – 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 – 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="18" pageNumber="119">Fig. 85</figureCitation>
|
||
). Stylomere 2 with two or three ensiform setae at the outer margin and three or four at inner margin (
|
||
<figureCitation id="1338BD9B5649FFFBC39CFD1AFC8EFD39" box="[723,807,655,679]" captionStart="FIGURES 90 – 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 – 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="18" pageNumber="119">Fig. 91</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319F2955649FFFBC188FD26FF78FD71" pageId="18" pageNumber="119" type="distribution">
|
||
<paragraph id="8BBCA11E5649FFFBC188FD26FF78FD71" blockId="18.[151,1437,151,1040]" pageId="18" pageNumber="119">
|
||
<emphasis id="B9777D0C5649FFFBC188FD26FEF0FD52" bold="true" box="[199,345,691,716]" pageId="18" pageNumber="119">Distribution</emphasis>
|
||
. This subspecies is only known from Cangshan Mt., near Dali (Xiaguan Township) (
|
||
<figureCitation id="1338BD9B5649FFFBC465FD21FF6CFD71" captionStart="FIGURES 110 – 111" captionStartId="33.[151,264,1885,1908]" captionTargetBox="[224,1312,207,1827]" captionTargetId="figure@33.[224,1343,193,1844]" captionTargetPageId="33" captionText="FIGURES 110 – 111. Distribution maps for species of subgenus Pterostichus (Wraseiellus): - Pt. andrewesi Jedlička; « Pt. meyeri Jedlička; ˑ Pt. crassiapex n. sp.; ¿ Pt. comatus n. sp.; ● Pt. diversus (Fairmaire); Ο Pt. pseudodiversus n. sp.; ˔ Pt. stictopleurus stictopleurus (Fairmaire); r Pt. stictopleurus cangshanensis n. ssp.; ․ Pt. kambaiti (Andrewes). 111. shows an enlarged scale of the area in figure 110 marked by the dotted line square." httpUri="https://zenodo.org/record/220516/files/figure.png" pageId="18" pageNumber="119">Figs. 110, 111</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319F2955649FFFBC188FD69FC98FCA6" pageId="18" pageNumber="119" type="etymology">
|
||
<paragraph id="8BBCA11E5649FFFBC188FD69FC98FCA6" blockId="18.[151,1437,151,1040]" pageId="18" pageNumber="119">
|
||
<emphasis id="B9777D0C5649FFFBC188FD69FEE1FC8A" bold="true" box="[199,328,764,788]" pageId="18" pageNumber="119">Etymology</emphasis>
|
||
. The Latinlized name "
|
||
<emphasis id="B9777D0C5649FFFBC31DFD69FD5CFC8A" box="[594,757,764,788]" italics="true" pageId="18" pageNumber="119">cangshanensis</emphasis>
|
||
" refers to the
|
||
<typeStatus id="54B81FBC5649FFFBC2D9FD68FC6FFC8B" box="[918,966,765,789]" pageId="18" pageNumber="119">type</typeStatus>
|
||
locality of the new subspecies, Cangshan Mt., a mountain range located on the west of Dali, Yunnan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C319F2955649FFFBC188FCD1FDC1FB91" pageId="18" pageNumber="119" type="discussion">
|
||
<paragraph id="8BBCA11E5649FFFBC188FCD1FDC1FB91" blockId="18.[151,1437,151,1040]" pageId="18" pageNumber="119">
|
||
<emphasis id="B9777D0C5649FFFBC188FCD1FE9DFCC2" bold="true" box="[199,308,836,860]" pageId="18" pageNumber="119">Remarks</emphasis>
|
||
. We propose this new subspecies mainly for the presence of setae on 5th tarsomeres, which is an important character in
|
||
<taxonomicName id="4C03DA9D5649FFFBC0DAFCFDFD88FC1E" box="[405,545,872,896]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9777D0C5649FFFBC0DAFCFDFD88FC1E" box="[405,545,872,896]" italics="true" pageId="18" pageNumber="119">Pterostichus</emphasis>
|
||
</taxonomicName>
|
||
, even on subgenera level. Besides this, the external and genital differences (see diagnosis section) are constant though slight, and its geographical range is isolated from the nominotypical subspecies (Cangshan and Dapingdi belong to mountain ranges isolated by Erhai basin), which support the validity of the new subspecies. But the differences between their male genitalia are so small that we prefer to regard it as a subspecies rather than a different species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |