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<document id="9334FA7B382DC3C87DB5A149B9FC5D98" ID-CLB-Dataset="101165" ID-DOI="10.17109/AZH.61.1.3.2015" ID-GBIF-Dataset="779ae168-3c95-49a5-a890-342248548e92" ID-ISSN="2064-2474" ID-Zenodo-Dep="5734508" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638187783047" checkinUser="felipe" docAuthor="Tóth, Balázs &amp; Ronkay, László" docDate="2015" docId="03FD87F45907FFD190381F14A9C6FB1C" docLanguage="en" docName="ActaZoolAcadSciH.61.1.3-23.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 61 (1)" docSource="http://dx.doi.org/10.17109/azh.61.1.3.2015" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.4:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="4" docTitle="Naarda secreta Tóth &amp; Ronkay 2015, sp. n." docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FFC4FF8C5908FFC192391B4BA863FF96" masterDocTitle="Revision Of The Palaearctic And Oriental Species Of The Genus Naarda (Lepidoptera: Erebidae, Hypeninae) Part 2. Description Of Ten New Species From Asia" masterLastPageNumber="23" masterPageNumber="3" pageNumber="18" updateTime="1699470903606" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="6BF8C40F7FC89DFAAD9581CFEE2E700C">Revision Of The Palaearctic And Oriental Species Of The Genus Naarda (Lepidoptera: Erebidae, Hypeninae) Part 2. Description Of Ten New Species From Asia</mods:title>
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<mods:namePart id="6AF54288859E2BC52335AE037868E2CF">Tóth, Balázs</mods:namePart>
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<treatment id="03FD87F45907FFD190381F14A9C6FB1C" ID-DOI="http://doi.org/10.5281/zenodo.6870814" ID-GBIF-Taxon="197829067" ID-Zenodo-Dep="6870814" LSID="urn:lsid:plazi:treatment:03FD87F45907FFD190381F14A9C6FB1C" httpUri="http://treatment.plazi.org/id/03FD87F45907FFD190381F14A9C6FB1C" lastPageId="16" lastPageNumber="19" pageId="15" pageNumber="18">
<subSubSection id="C34E65695907FFCE90381F14AB71FBEF" box="[513,786,1119,1145]" pageId="15" pageNumber="18" type="nomenclature">
<paragraph id="8BEB36E25907FFCE90381F14AB71FBEF" blockId="15.[513,786,1119,1145]" box="[513,786,1119,1145]" pageId="15" pageNumber="18">
<heading id="D0A3818E5907FFCE90381F14AB71FBEF" box="[513,786,1119,1145]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="18" reason="2">
<taxonomicName id="4C544D615907FFCE90381F14AAA6FBEF" authority="Tóth &amp; Ronkay, 2015" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[513,709,1119,1145]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="18" phylum="Arthropoda" rank="species" species="secreta" status="sp. nov.">
<emphasis id="B920EAF05907FFCE90381F14AAA6FBEF" bold="true" box="[513,709,1119,1145]" pageId="15" pageNumber="18">Naarda secreta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213578B5907FFCE90F41F14AB71FBEF" box="[717,786,1119,1145]" pageId="15" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E65695907FFD1907D1FC9A9C6FB1C" lastPageId="16" lastPageNumber="19" pageId="15" pageNumber="18" type="description">
<paragraph id="8BEB36E25907FFCE907D1FC9AAACFB0A" blockId="15.[580,719,1154,1180]" box="[580,719,1154,1180]" pageId="15" pageNumber="18">
<heading id="D0A3818E5907FFCE907D1FC9AAACFB0A" box="[580,719,1154,1180]" centered="true" fontSize="11" level="3" pageId="15" pageNumber="18" reason="3">
(
<figureCitation id="136F2A675907FFCE90751FC9AAF8FB0A" box="[588,667,1154,1180]" captionStart="Figs 7–12" captionStartId="4.[136,181,1464,1486]" captionTargetBox="[147,1080,193,1438]" captionTargetId="figure-21@4.[144,1080,189,1438]" captionTargetPageId="4" captionText="Figs 7–12. Male genitalia of newly and formerly described Naarda species: 7 = N. blepharota Strand (RL10745m), 8 = N. secreta sp. n. (RL155TFRIm), 9 = N. huettleri sp. n. (TB571m), 10 = N. palawana sp. n. (RL7992m), 11 = N. glauculalis Hampson (RL7681m), 12 = N. melinau Hol- loway (RL7977m)." figureDoi="http://doi.org/10.5281/zenodo.5734515" httpUri="https://zenodo.org/record/5734515/files/figure.png" pageId="15" pageNumber="18">Figs 8</figureCitation>
,
<figureCitation id="136F2A675907FFCE90911FC9AAA5FB0A" box="[680,710,1154,1180]" captionStart="Figs 25–39" captionStartId="13.[173,218,1427,1449]" captionTargetBox="[177,1125,387,1402]" captionTargetId="figure-178@13.[173,1125,373,1405]" captionTargetPageId="13" captionText="Figs 25–39. Adults of newly described species: 25 = N. octogesima sp. n. male, 26 = N. laoana sp. n. male, 27 = N. mirabilis sp. n. male, 28 = N. octogesima sp. n. female, 29 = N. laoana sp. n. female, 30 = N. vicina sp. n. male, 31 = N. atrata sp. n. male, 32 = N. furcatella sp. n. male, 33 = N. vicina sp. n. male (photo: ZMUC), 34 = N. atrata sp. n. female, 35 = N. furcatella sp. n. female, 36 = N. plumbea sp. n. female, 37 = N. palawana sp. n. male, 38 = N. secreta sp. n. male," figureDoi="http://doi.org/10.5281/zenodo.5734519" httpUri="https://zenodo.org/record/5734519/files/figure.png" pageId="15" pageNumber="18">38</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BEB36E25907FFCE92D31FF1A949FB7A" blockId="15.[173,1125,1210,1260]" pageId="15" pageNumber="18">
<materialsCitation id="3B3C3CBF5907FFCE92D31FF1A945FB7A" ID-GBIF-Occurrence="3864438325" collectingDate="1993-12-13" collectorName="Y. B. Fan" country="Taiwan" location="Chaofeng" pageId="15" pageNumber="18" specimenCount="1" stateProvince="Hualien" typeStatus="holotype">
<typeStatus id="54EF88405907FFCE92D31FF1A933FB59" box="[234,336,1210,1232]" pageId="15" pageNumber="18" type="holotype">Holotype</typeStatus>
. m,
<collectingCountry id="F34376725907FFCE934F1FF1A9A6FB59" box="[374,453,1210,1232]" name="Taiwan" pageId="15" pageNumber="18">Taiwan</collectingCountry>
:
<location id="8E8B60395907FFCE93F71FF1AA54FB46" LSID="urn:lsid:plazi:treatment:03FD87F45907FFD190381F14A9C6FB1C:8E8B60395907FFCE93F71FF1AA54FB46" box="[462,567,1210,1232]" country="Taiwan" name="Chaofeng" pageId="15" pageNumber="18" stateProvince="Hualien">Chaofeng</location>
,
<collectingRegion id="4990F8005907FFCE90061FF1AAF4FB46" box="[575,663,1210,1232]" country="Taiwan" name="Hualien" pageId="15" pageNumber="18">Hualien</collectingRegion>
,
<date id="FFEA10225907FFCE90991FF1AB75FB59" box="[672,790,1210,1231]" pageId="15" pageNumber="18" value="1993-12-13">
<collectingDate id="EFAEE9CA5907FFCE90991FF1AB75FB59" box="[672,790,1210,1231]" pageId="15" pageNumber="18" value="1993-12-13">13.XII.1993</collectingDate>
</date>
, leg.
<collectorName id="26A153345907FFCE91731FF1ABC2FB59" box="[842,929,1210,1231]" pageId="15" pageNumber="18">Y.B. Fan</collectorName>
; slideNo. 155TFRI (coll. TFRI)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB36E25907FFCE92D31E59A91AFAF9" blockId="15.[173,1126,1297,1391]" pageId="15" pageNumber="18">
Taxonomy. Thesize, thedarkgroundcolour, thestrongpostmedialline andthestructureofthemalegenitaliaimplycloserelationshipwith
<taxonomicName id="4C544D615907FFCE91C61E7FAC05FADB" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[1023,1126,1332,1357]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="18" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B920EAF05907FFCE91C61E7FAC05FADB" box="[1023,1126,1332,1357]" italics="true" pageId="15" pageNumber="18">N. atrata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D615907FFCE92DD1E1EA911FAF8" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[228,370,1364,1390]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="18" phylum="Arthropoda" rank="species" species="furcatella">
<emphasis id="B920EAF05907FFCE92DD1E1EA911FAF8" box="[228,370,1364,1390]" italics="true" pageId="15" pageNumber="18">N. furcatella</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEB36E25907FFD192D31ED9AC5CFE17" blockId="15.[173,1126,1426,1563]" lastBlockId="16.[136,1088,190,616]" lastPageId="16" lastPageNumber="19" pageId="15" pageNumber="18">
Description. Wingspan
<quantity id="4CAC9B075907FFCE93DD1ED9AA4DFA31" box="[484,558,1426,1447]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="18" unit="mm" value="13.0">13 mm</quantity>
, lengthofforewing
<quantity id="4CAC9B075907FFCE913A1ED9AB37FA31" box="[771,852,1426,1448]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="18" unit="mm" value="6.5">6.5 mm</quantity>
. Antennaefiliform, ciliate withsetae. Twosetaeoneachsegment; longestonestwicelongerthediameterofflagellum. Ciliationdense; ciliaaslongasthediameterofflagellum. Labialpalpslong, fivetimes longerthediameterofeye; 3rdsegmentrelativelylong, itstiplight; dorsalscalesof 2nd segmentrelativelyshort, dorsaledgeconvex. Scale-hoodofvertexbroad-based, medium long, rounded. Characteristicwingpatternfeatures: costaofforewingnotconcave; shape offorewingunusuallybroad; groundcolourdarkgreywithslightbrownishshade (the specimenisworn); terminallineverystrong, blackish; transverselineshardlyvisibleexceptforthebroad, blackishpostmediallinebeingslightlysinuousandbentinwardsbelowcell; reniformstigmalightyellow, narrow, angular, withnarrowdarkedgeandsome blackishscalesatbottomsection; orbicularstigmatiny, hardlyvisible, composedbyonlya fewlightyellowscales. Colourationandpatternofhindwingsimilartothoseofforewing.
</paragraph>
<paragraph id="8BEB36E25918FFD192FD1AC2A9D8FDF1" blockId="16.[136,1088,190,616]" pageId="16" pageNumber="19">
Malegenitalia (
<figureCitation id="136F2A675918FFD193531AC3A9C5FE08" box="[362,422,392,414]" captionStart="Figs 7–12" captionStartId="4.[136,181,1464,1486]" captionTargetBox="[147,1080,193,1438]" captionTargetId="figure-21@4.[144,1080,189,1438]" captionTargetPageId="4" captionText="Figs 7–12. Male genitalia of newly and formerly described Naarda species: 7 = N. blepharota Strand (RL10745m), 8 = N. secreta sp. n. (RL155TFRIm), 9 = N. huettleri sp. n. (TB571m), 10 = N. palawana sp. n. (RL7992m), 11 = N. glauculalis Hampson (RL7681m), 12 = N. melinau Hol- loway (RL7977m)." figureDoi="http://doi.org/10.5281/zenodo.5734515" httpUri="https://zenodo.org/record/5734515/files/figure.png" pageId="16" pageNumber="19">Fig. 8</figureCitation>
). Uncusrelativelylong, straight, withconvexdorsaledgeand hookedtip. Tegumenaslongasvinculum. Saccusbroad-based, long, graduallytapering, itstiprounded. Juxtalarge, rhomboidal, broaderthanlong. Valvalong, mediumwide, slightlytapering; asmallcostallobepresentintheapicalquarterofvalva. Sacculusnarrow, elongate, halfaslongasvalva. Distalthirdofvalvacomposedbythestronglysclerotised fusionofcosta, harpeandcucullus; thissectionbeingtapering, veryslightlycurveddorsally, itstiprounded. Aedeaguselongate, evenlywide, slightlycurved; vesica (uneverted) withlarge, straightcornutus.
</paragraph>
<paragraph id="8BEB36E25918FFD192FD19C7AAA9FC04" blockId="16.[136,1088,652,1049]" pageId="16" pageNumber="19">
Diagnosis. Theunusuallybroadforewingofthisnewspeciesisshared by
<taxonomicName id="4C544D615918FFD1928919E4AA66FD5E" authority="(Strand, 1920)" baseAuthorityName="Strand" baseAuthorityYear="1920" box="[176,517,686,713]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="blepharota">
<emphasis id="B920EAF05918FFD1928919E4A928FD5E" box="[176,331,686,712]" italics="true" pageId="16" pageNumber="19">N. blepharota</emphasis>
(
<bibRefCitation id="EFC54B135918FFD1936419E5A99FFD5E" author="STrAnd, E." box="[349,508,686,713]" pageId="16" pageNumber="19" pagination="102 - 197" refId="ref5882" refString="STrAnd, E. (1920) H. Sauter'sFormosa- Ausbeite: NoctuidaeII., nebstNachtragezuden FamilienArctiidae, Lymantriidae, Notodontidae, Geometridae, Thyrididae, Pyrali- dae, Tortricidae, GelechiidaeundOecophoridae. ArchivfurNaturgeschichte 84 A 12: 102 - 197." type="journal article" year="1920">Strand, 1920</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B920EAF05918FFD1902B19E5AADFFD5E" box="[530,700,686,712]" italics="true" pageId="16" pageNumber="19">Naardasecreta</emphasis>
isslightlysmallerinsizethan
<taxonomicName id="4C544D615918FFD192B1199BA941FD7C" baseAuthorityName="Strand" baseAuthorityYear="1920" box="[136,290,720,746]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="blepharota">
<emphasis id="B920EAF05918FFD192B1199BA941FD7C" box="[136,290,720,746]" italics="true" pageId="16" pageNumber="19">N. blepharota</emphasis>
</taxonomicName>
, thegroundcolourisdarker, thereniformstigmaislighterand smaller, andthestreaksoftheterminallinearemuchmoreelongate. Themale genitaliashowmoreconspicuousdifferencesbetweenthesetwospecies: the ventraledgeofuncusin
<taxonomicName id="4C544D615918FFD19384187EAA4CFCD8" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[445,559,821,846]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="secreta">
<emphasis id="B920EAF05918FFD19384187EAA4CFCD8" box="[445,559,821,846]" italics="true" pageId="16" pageNumber="19">N. secreta</emphasis>
</taxonomicName>
isstraight, whilethatof
<taxonomicName id="4C544D615918FFD1915E187EAC63FCD8" baseAuthorityName="Strand" baseAuthorityYear="1920" box="[871,1024,820,846]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="blepharota">
<emphasis id="B920EAF05918FFD1915E187EAC63FCD8" box="[871,1024,820,846]" italics="true" pageId="16" pageNumber="19">N. blepharota</emphasis>
</taxonomicName>
(
<figureCitation id="136F2A675918FFD19628187FA8F4FCE7" captionStart="Figs 7–12" captionStartId="4.[136,181,1464,1486]" captionTargetBox="[147,1080,193,1438]" captionTargetId="figure-21@4.[144,1080,189,1438]" captionTargetPageId="4" captionText="Figs 7–12. Male genitalia of newly and formerly described Naarda species: 7 = N. blepharota Strand (RL10745m), 8 = N. secreta sp. n. (RL155TFRIm), 9 = N. huettleri sp. n. (TB571m), 10 = N. palawana sp. n. (RL7992m), 11 = N. glauculalis Hampson (RL7681m), 12 = N. melinau Hol- loway (RL7977m)." figureDoi="http://doi.org/10.5281/zenodo.5734515" httpUri="https://zenodo.org/record/5734515/files/figure.png" pageId="16" pageNumber="19">Fig. 7</figureCitation>
) has an emergence, the juxta is broader, the valva has no harpe, but the costa hassmalllobe, whichisabsentin
<taxonomicName id="4C544D615918FFD190131833AAA0FC04" baseAuthorityName="Strand" baseAuthorityYear="1920" box="[554,707,888,914]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="blepharota">
<emphasis id="B920EAF05918FFD190131833AAA0FC04" box="[554,707,888,914]" italics="true" pageId="16" pageNumber="19">N. blepharota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BEB36E25918FFD192FD18D1AC23FB8E" blockId="16.[136,1088,652,1049]" pageId="16" pageNumber="19">
Themalegenitaliaofthisnewspeciesaresimilartothoseof
<taxonomicName id="4C544D615918FFD191E918D1AC5AFC25" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[976,1081,922,947]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B920EAF05918FFD191E918D1AC5AFC25" box="[976,1081,922,947]" italics="true" pageId="16" pageNumber="19">N. atrata</emphasis>
</taxonomicName>
. However, thecostallobeofthevalvaof
<taxonomicName id="4C544D615918FFD190A818F7AB66FC43" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[657,773,956,981]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="secreta">
<emphasis id="B920EAF05918FFD190A818F7AB66FC43" box="[657,773,956,981]" italics="true" pageId="16" pageNumber="19">N. secreta</emphasis>
</taxonomicName>
isabsentfrom
<taxonomicName id="4C544D615918FFD191F618F7AC5AFC43" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[975,1081,956,981]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B920EAF05918FFD191F618F7AC5AFC43" box="[975,1081,956,981]" italics="true" pageId="16" pageNumber="19">N. atrata</emphasis>
</taxonomicName>
, moreover, thevalvaislesstapering, itstiproundedinsteadofbeingpointedas in
<taxonomicName id="4C544D615918FFD1929F18B4A969FB8E" authorityName="Tóth &amp; Ronkay" authorityYear="2015" box="[166,266,1023,1048]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="atrata">
<emphasis id="B920EAF05918FFD1929F18B4A969FB8E" box="[166,266,1023,1048]" italics="true" pageId="16" pageNumber="19">N. atrata</emphasis>
</taxonomicName>
, thesaccusisbroaderandthecornutusinthevesicaisalsobroader.
</paragraph>
<paragraph id="8BEB36E25918FFD192FD1F77AB74FBF8" blockId="16.[136,1088,1083,1163]" pageId="16" pageNumber="19">
Etymology. Thespecieshasremainedlongbeenhiddenduetotheexternalsimilarity to
<taxonomicName id="4C544D615918FFD1929B1F12A946FBFB" baseAuthorityName="Strand" baseAuthorityYear="1920" box="[162,293,1112,1134]" class="Insecta" family="Noctuidae" genus="Naarda" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="19" phylum="Arthropoda" rank="species" species="blepharota">
<emphasis id="B920EAF05918FFD1929B1F12A946FBFB" box="[162,293,1112,1134]" italics="true" pageId="16" pageNumber="19">N. blepharota</emphasis>
</taxonomicName>
. Thespecificnameisrememberingtothisfact.
</paragraph>
<paragraph id="8BEB36E25918FFD192FD1F3EA9C6FB1C" blockId="16.[136,1088,1083,1163]" box="[196,421,1141,1163]" pageId="16" pageNumber="19">
Distribution.
<collectingCountry id="F34376725918FFD1936B1F3EA9C2FB1C" box="[338,417,1141,1163]" name="Taiwan" pageId="16" pageNumber="19">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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