119 lines
19 KiB
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119 lines
19 KiB
XML
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<mods:title id="D63C396DF225AA96F901F4A241F737F1">The Early Cretaceous Mesofossil Flora Of Catefica, Portugal: Angiosperms</mods:title>
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<mods:namePart id="BB6C5C314EDEFD14B8A3596459BE3163">Friis, Else Marie</mods:namePart>
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<mods:namePart id="D996E014F55BDA32775103964AC210FD">Crane, Peter R.</mods:namePart>
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<mods:namePart id="782C77FE3EFD2EB9730436EE1B7887FC">Pedersen, Kaj Raunsgaard</mods:namePart>
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<mods:namePart id="889B62C59C2AECA5245AB754BC105B5A">Mendes, Mário Miguel</mods:namePart>
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<mods:namePart id="356FE8A09D4146285DD663FC6C820576">Kvaček, Jiří</mods:namePart>
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<treatment id="03FD87F2FFD1FFC2FF7DF9D6C62AF994" ID-DOI="http://doi.org/10.5281/zenodo.7535341" ID-GBIF-Taxon="208830437" ID-Zenodo-Dep="7535341" LSID="urn:lsid:plazi:treatment:03FD87F2FFD1FFC2FF7DF9D6C62AF994" httpUri="http://treatment.plazi.org/id/03FD87F2FFD1FFC2FF7DF9D6C62AF994" lastPageNumber="385" pageId="44" pageNumber="385">
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<taxonomicName id="4C544D67FFD1FFC2FF7DF9D6C148F9C7" authority="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN, 2019" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN, Mugideiriflora" authorityYear="2019" class="Magnoliopsida" family="Ceratopogonidae" genus="Kempia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="44" pageNumber="385" phylum="Tracheophyta" rank="species" species="longicolpites">
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<emphasis id="B920EAF6FFD1FFC2FF7DF9D6C2D0F9E6" bold="true" box="[194,424,1576,1601]" italics="true" pageId="44" pageNumber="385">Kempia longicolpites</emphasis>
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E.M.FRIIS, P.R.CRANE et K.R.PEDERSEN, 2019
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<paragraph id="8BEB36E4FFD1FFC2FECEF996C177F927" blockId="44.[369,527,1641,1665]" box="[369,527,1641,1665]" pageId="44" pageNumber="385">
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<figureCitation id="136F2A61FFD1FFC2FECEF996C177F927" box="[369,527,1641,1665]" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30a–f</figureCitation>
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</paragraph>
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<subSubSection id="C34E656FFFD1FFC2FF19F966C62AF994" pageId="44" pageNumber="385" type="description">
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<paragraph id="8BEB36E4FFD1FFC2FF19F966C644FA05" blockId="44.[129,768,1688,1840]" lastBlockId="44.[843,1483,1035,1443]" pageId="44" pageNumber="385">
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Description and remarks. The material comprises two adhering stamens (only one cut stamen illustrated) with dithecate, tetrasporangiate anthers (
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<figureCitation id="136F2A61FFD1FFC2FD1AF927C3D0F8B6" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30a</figureCitation>
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) and in situ pollen. The stamens are about
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<quantity id="4CAC9B01FFD1FFC2FD39F907C1BFF8B6" box="[646,711,1784,1808]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="44" pageNumber="385" unit="mm" value="1.0">1 mm</quantity>
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long and
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<quantity id="4CAC9B01FFD1FFC2FF13F8E7C270F896" box="[172,264,1816,1840]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="44" pageNumber="385" unit="mm" value="0.25">0.25 mm</quantity>
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broad with a distinct, triangular apical extension of the connective. One stamen was cut transversely into two pieces to expose the pollen for SEM. The other stamen was removed for TEM. Stamens and anthers are closely similar to
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<taxonomicName id="4C544D67FFD1FFC2FCD4FB94C737FB25" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN, Mugideiriflora" authorityYear="2019" box="[875,1103,1131,1155]" class="Magnoliopsida" family="Ceratopogonidae" genus="Kempia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="44" pageNumber="385" phylum="Tracheophyta" rank="species" species="longicolpites">
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<emphasis id="B920EAF6FFD1FFC2FCD4FB94C737FB25" box="[875,1103,1131,1155]" italics="true" pageId="44" pageNumber="385">Kempia longicolpites</emphasis>
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described from the Torres Vedras locality (
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<bibRefCitation id="EFC54B15FFD1FFC2FC13FB74C708FB05" author="Friis, E. M. & Crane, P. R. & Pedersen, K. R." box="[940,1136,1163,1187]" pageId="44" pageNumber="385" pagination="153 - 257" refId="ref55433" refString="Friis, E. M., Crane, P. R., Pedersen, K. R. (2019 a): The Early Cretaceous mesofossil flora of Torres Vedras (NE of Forte da Forca), Portugal: a palaeofloristic analysis of an early angiosperm community. - Fossil Imprint, 75: 153 - 257. https: // doi. org / 10.2478 / if- 2019 - 0013" type="journal article" year="2019">Friis et al. 2019a</bibRefCitation>
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) and the specimen is assigned here to the same species. Pollen grains are small, about 11– 12 µm long, monoaperturate and with the colpus extending beyond the full length of the grains. The exine is semitectatereticulate, columellate (
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<figureCitation id="136F2A61FFD1FFC2FB84FAF4C7A1FA85" box="[1083,1241,1291,1315]" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30b–f</figureCitation>
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) with the reticulum and columellae only loosely attached to the foot layer (
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<figureCitation id="136F2A61FFD1FFC2FAD0FAD4C00AFAC5" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30f</figureCitation>
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). The muri have a rounded profile and a smooth surface (
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<figureCitation id="136F2A61FFD1FFC2FCEDFA94C0A7FA25" box="[850,991,1387,1411]" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30c</figureCitation>
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). The foot layer is thick, and the endexine is restricted to the apertural region (
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<figureCitation id="136F2A61FFD1FFC2FB1AFA74C657FA05" box="[1189,1327,1419,1443]" captionStart="Text-fig" captionStartId="48.[147,226,976,997]" captionTargetBox="[148,1462,151,960]" captionTargetId="figure-518@48.[145,1466,148,963]" captionTargetPageId="48" captionText="Text-fig. 30. Scanning electron microscope (SEM) images of anther and pollen of Kempia longicolpites; Catefica locality, Portugal. a) Transverse section through an anther showing the four pollen sacs with in situ pollen; b) Pollen in situ from anther in (a) showing very long colpus and loosely attached reticulum; c) Detail of reticulumshowing smooth muri supported by short, scattered columellae; d) Pollen grain from anther in (a) enlarged showing colpus extended beyond the equator and loosely attached reticulum; e) Detail of pollen wall showing thick, homogeneous foot layer, columellae and reticulate tectum; note remains of granular endexine (arrows) in the apertural region of the grain; f) Internal view of reticulum showing the short columellae adhering to the muri detached from the foot layer. Specimen, Catefica 49-S101208 (a–f). Scale bars = 100 Μm (a), 20 Μm (b), 6 Μm (d), 3 Μm (c, e, f)." figureDoi="http://doi.org/10.5281/zenodo.7522867" httpUri="https://zenodo.org/record/7522867/files/figure.png" pageId="44" pageNumber="385">Text-fig. 30e</figureCitation>
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).
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</paragraph>
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<paragraph id="8BEB36E4FFD1FFC2FCCEFA45C62AF994" blockId="44.[843,1482,1466,1586]" pageId="44" pageNumber="385">
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Affinity and other occurrences.
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<taxonomicName id="4C544D67FFD1FFC2FAC8FA45C0ADFA54" authorityName="E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN, Mugideiriflora" authorityYear="2019" class="Magnoliopsida" family="Ceratopogonidae" genus="Kempia" higherTaxonomySource="CoL" kingdom="Plantae" order="Diptera" pageId="44" pageNumber="385" phylum="Tracheophyta" rank="species" species="longicolpites">
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<emphasis id="B920EAF6FFD1FFC2FAC8FA45C0ADFA54" italics="true" pageId="44" pageNumber="385">Kempia longicolpites</emphasis>
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</taxonomicName>
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was first described from the Torres Vedras locality (
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<bibRefCitation id="EFC54B15FFD1FFC2FC17FA05C724F9B4" author="Friis, E. M. & Crane, P. R. & Pedersen, K. R." box="[936,1116,1530,1554]" pageId="44" pageNumber="385" pagination="153 - 257" refId="ref55433" refString="Friis, E. M., Crane, P. R., Pedersen, K. R. (2019 a): The Early Cretaceous mesofossil flora of Torres Vedras (NE of Forte da Forca), Portugal: a palaeofloristic analysis of an early angiosperm community. - Fossil Imprint, 75: 153 - 257. https: // doi. org / 10.2478 / if- 2019 - 0013" type="journal article" year="2019">Friis et al. 2019a</bibRefCitation>
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) and is currently known from only the Torres Vedras and Catefica mesofossil floras.
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</paragraph>
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</treatment>
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