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<mods:title id="6D241C090F101E23B784B410C83BE4DD">Two new genera and species of the family Scathophagidae (Diptera) from the Palaearctic and Oriental Regions with additional faunistic records</mods:title>
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<paragraph id="8BEB36F6FF92FFB2F9DA090CFD3BAED6" blockId="1.[464,739,891,915]" box="[464,739,891,915]" pageId="1" pageNumber="610">
<heading id="D0A3819AFF92FFB2F9DA090CFD3BAED6" bold="true" box="[464,739,891,915]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="610" reason="3">
<emphasis id="B920EAE4FF92FFB2F9DA090CFD3BAED6" bold="true" box="[464,739,891,915]" pageId="1" pageNumber="610">
Genus
<taxonomicName id="4C544D75FF92FFB2FA15090CFDAEAED6" ID-CoL="89QQP" authority="Šifner, 2010" authorityName="Šifner" authorityYear="2010" box="[543,630,891,915]" class="Insecta" family="Scathophagidae" genus="Milania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="610" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EAE4FF92FFB2FA15090CFDAEAED6" bold="true" box="[543,630,891,915]" italics="true" pageId="1" pageNumber="610">Milania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213579FFF92FFB2FA77090CFD3BAED6" box="[637,739,891,915]" pageId="1" pageNumber="610" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E657DFF92FFB2F87B09DEFD00AEFE" box="[113,728,936,956]" pageId="1" pageNumber="610" type="type_taxon">
<paragraph id="8BEB36F6FF92FFB2F87B09DEFD00AEFE" blockId="1.[113,728,936,956]" box="[113,728,936,956]" pageId="1" pageNumber="610">
<emphasis id="B920EAE4FF92FFB2F87B09DEFF31AEF9" bold="true" box="[113,233,936,956]" pageId="1" pageNumber="610">
<typeStatus id="54EF8854FF92FFB2F87B09DEFF47AEF9" box="[113,159,937,956]" pageId="1" pageNumber="610">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C544D75FF92FFB2F8E409DEFDD0AEF9" ID-CoL="89KSH" authority="Seguy, 1948" authorityName="♀, Cotyp., E. Seguy" authorityYear="1948" box="[238,520,936,956]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="610" phylum="Arthropoda" rank="species" species="agrion">
<emphasis id="B920EAE4FF92FFB2F8E409DEFE4EAEFE" box="[238,406,937,955]" italics="true" pageId="1" pageNumber="610">Norellisoma agrion</emphasis>
<bibRefCitation id="EFC54B07FF92FFB2F99109DFFDD0AEF9" author="SEGUY E." box="[411,520,936,956]" pageId="1" pageNumber="610" refId="ref5326" refString="SEGUY E. 1948: Dipteres nouveaux ou peu connus d'Extreme-Orient. Notes d'Entomologie Chinoise 12: 153 - 142." type="journal volume" year="1948">Séguy, 1948</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C34E657DFF92FFB2F87B09A3FD33A847" pageId="1" pageNumber="610" type="diagnosis">
<paragraph id="8BEB36F6FF92FFB2F87B09A3FC4BAEA9" blockId="1.[113,1074,980,1204]" box="[113,915,980,1004]" pageId="1" pageNumber="610">
<emphasis id="B920EAE4FF92FFB2F87B09A3FF3EAEA9" bold="true" box="[113,230,980,1004]" pageId="1" pageNumber="610">Diagnosis.</emphasis>
Principal diagnostic characters of the new genus are as follows:
</paragraph>
<paragraph id="8BEB36F6FF92FFB2F89A098CFBEAA9F6" blockId="1.[113,1074,980,1204]" pageId="1" pageNumber="610">
i) ratio of the length of thorax to the length of abdomen equal to 1: 22.2; ii) double rows of bristles on anterior femora and tibiae very weakly developed; iii) all legs covered with long hair-like bristles; iv) epiphallus straight and dilated apically; v) all abdominal sternites in both sexes weakly sclerotized; vi) sternites
<quantity id="4CAC9B13FF92FFB2F9190EEBFEE4A9F6" box="[275,316,1179,1204]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="1" pageNumber="610" unit="in" value="8.0">8 in</quantity>
female weakly sclerotized, without thorns and only with short bristles.
</paragraph>
<paragraph id="8BEB36F6FF92FFB2F87B0EBDFD33A847" blockId="1.[113,1089,1226,1282]" pageId="1" pageNumber="610">
<emphasis id="B920EAE4FF92FFB2F87B0EBDFEBFA9A7" bold="true" box="[113,359,1226,1250]" pageId="1" pageNumber="610">Differential diagnosis</emphasis>
.
<taxonomicName id="4C544D75FF92FFB2F97C0EBBFE12A9A6" authority="Šifner, 2010" authorityName="Šifner" authorityYear="2010" box="[374,458,1228,1251]" class="Insecta" family="Scathophagidae" genus="Milania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="610" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EAE4FF92FFB2F97C0EBBFE12A9A6" box="[374,458,1228,1251]" italics="true" pageId="1" pageNumber="610">Milania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213579FFF92FFB2F9D90EBCFDE0A9A7" box="[467,568,1227,1251]" pageId="1" pageNumber="610" rank="genus">gen. nov.</taxonomicNameLabel>
can be separated from other genera of the tribe
<taxonomicName id="4C544D75FF92FFB2F87B0E9DFF30A847" box="[113,232,1258,1282]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="610" phylum="Arthropoda" rank="tribe" tribe="Cordilurini">Cordilurini</taxonomicName>
by characters summarized in the following key.
</paragraph>
</subSubSection>
<subSubSection id="C34E657DFF92FFB1F87B0F6DFBBEACB1" lastPageId="2" lastPageNumber="611" pageId="1" pageNumber="610" type="key">
<key id="FFC561C7FF92FFB1F87B0F6DFBBEACB1" lastPageId="2" lastPageNumber="611" pageId="1" pageNumber="610">
<keyStep id="30A02E53FF92FFB2F87B0F6DFB99AB17" pageId="1" pageNumber="610">
<paragraph id="8BEB36F6FF92FFB2F87B0F6DFB98A837" blockId="1.[113,1089,1305,1618]" pageId="1" pageNumber="610">
<keyLead id="30A595C3FF92FFB2F87B0F6DFB98A837" pageId="1" pageNumber="610">1 All bristles on thorax distinctly developed; ratio of length of thorax to length of abdomen equal to 1: 1.51.7; all abdominal sclerites in both sexes distinctly sclerotized; epiphallus slightly arched and pointed. ........................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8BEB36F6FF92FFB2F87B0F0DFB99AB17" blockId="1.[113,1089,1305,1618]" pageId="1" pageNumber="610">
<keyLead id="30A595C3FF92FFB2F87B0F0DFB99AB17" pageId="1" pageNumber="610">
Bristles on thorax weakly developed; ratio of length of thorax to length of abdomen equal to 1: 22.2; all abdominal sclerites in both sexes weakly sclerotized; epiphallus almost straight and dilated apically. Double row of bristles at inner side of anterior femur and tibia weakly developed, all femora and tibiae with long and fine hair-like bristles; palpi narrow with 12 weak apical or subapical bristles; sternite 7 of female long, sclerotized in lateral part and membranous only medially; sternites 8 distinctly prolonged, without thorns and only apically with short fine bristles. ...............................
<emphasis id="B920EAE4FF92FFB2FB770C4EFB99AB17" bold="true" box="[893,1089,1593,1618]" pageId="1" pageNumber="610">
<taxonomicName id="4C544D75FF92FFB2FB770C4EFC0CAB14" ID-CoL="89QQP" authority="Šifner, 2010" authorityName="Šifner" authorityYear="2010" box="[893,980,1593,1617]" class="Insecta" family="Scathophagidae" genus="Milania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="610" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EAE4FF92FFB2FB770C4EFC0CAB14" bold="true" box="[893,980,1593,1617]" italics="true" pageId="1" pageNumber="610">Milania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213579FFF92FFB2FBD10C4DFB99AB17" box="[987,1089,1594,1618]" pageId="1" pageNumber="610" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E53FF91FFB1F89D0AABFBBEACB1" pageId="2" pageNumber="611">
<paragraph id="8BEB36F6FF91FFB1F89D0AABFBBEAC11" blockId="2.[151,1127,219,500]" pageId="2" pageNumber="611">
<keyLead id="30A595C3FF91FFB1F89D0AABFBBEAC11" pageId="2" pageNumber="611">
2 Double row of bristles at inner side of anterior femur and tibia strong; palpi uniformly narrow with 12 apical or subapical bristles; sternite 7 of female partly or entirely divided in two sclerites; sternites 8 of female with varying number of thorns. ............................. ............................................................................................
<taxonomicName id="4C544D75FF91FFB1FB270B4CFBBEAC11" ID-CoL="63GDT" authority="Wahlgren, 1917" authorityName="Wahlgren" authorityYear="1917" box="[813,1126,315,340]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1FB270B4CFC6CAC16" bold="true" box="[813,948,315,339]" italics="true" pageId="2" pageNumber="611">Norellisoma</emphasis>
Wahlgren, 1917
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36F6FF91FFB1F89D0B2BFBBEACB1" blockId="2.[151,1127,219,500]" pageId="2" pageNumber="611">
<keyLead id="30A595C3FF91FFB1F89D0B2BFBBEACB1" pageId="2" pageNumber="611">
Double row of bristles at inner side of anterior femur and tibia not developed; palpi narrow and dilated in their median portion with 12 strong apical or subapical bristles; sternite 7 of female divided in three separate sclerites; sternites 8 of female with short and weak bristles. ............................................................................................................................... ...
<taxonomicName id="4C544D75FF91FFB1F8D60BACFE0FACB1" ID-CoL="3TZY" authority="Fallen, 1810" authorityName="Fallen" authorityYear="1810" box="[220,471,475,500]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1F8D60BACFE90ACB6" bold="true" box="[220,328,475,499]" italics="true" pageId="2" pageNumber="611">Cordilura</emphasis>
Fallén, 1810
</taxonomicName>
,
<taxonomicName id="4C544D75FF91FFB1F9E80BACFCCAACB1" ID-CoL="6FBJ" authority="Becker, 1894" authorityName="Becker" authorityYear="1894" box="[482,786,475,500]" class="Insecta" family="Scathophagidae" genus="Parallelomma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1F9E80BACFDA6ACB6" bold="true" box="[482,638,475,499]" italics="true" pageId="2" pageNumber="611">Parallelomma</emphasis>
Becker, 1894
</taxonomicName>
and
<taxonomicName id="4C544D75FF91FFB1FB4C0BACFBBEACB1" authority="Becker, 1894" authorityName="Becker" authorityYear="1894" box="[838,1126,475,500]" class="Insecta" family="Scathophagidae" genus="Scoliaphleps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1FB4C0BACFC09ACB6" bold="true" box="[838,977,475,499]" italics="true" pageId="2" pageNumber="611">Scoliaphleps</emphasis>
Becker, 1894
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C34E657DFF91FFB1F89D087CFF15AF06" pageId="2" pageNumber="611" type="etymology">
<paragraph id="8BEB36F6FF91FFB1F89D087CFF15AF06" blockId="2.[151,1127,523,579]" pageId="2" pageNumber="611">
<emphasis id="B920EAE4FF91FFB1F89D087CFEC1AF66" bold="true" box="[151,281,523,547]" pageId="2" pageNumber="611">Etymology.</emphasis>
I dedicate this genus to the memory of my late son Milan. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C34E657DFF91FFB1F89D082EFB92AEF4" pageId="2" pageNumber="611" type="discussion">
<paragraph id="8BEB36F6FF91FFB1F89D082EFB92AEF4" blockId="2.[151,1127,601,946]" pageId="2" pageNumber="611">
<emphasis id="B920EAE4FF91FFB1F89D082EFEC2AF34" bold="true" box="[151,282,601,625]" pageId="2" pageNumber="611">Comments.</emphasis>
<bibRefCitation id="EFC54B07FF91FFB1F92A082DFE0DAF37" author="HACKMAN W." box="[288,469,602,626]" pageId="2" pageNumber="611" refId="ref5054" refString="HACKMAN W. 1956: The Scathophagidae (Diptera) of Eastern Fennoscandia. Fauna Fennica. Vol. 2. Societas pro Fauna et Flora Fennica, Helsingforsiae, 67 pp." type="book" year="1956">HACKMAN (1956)</bibRefCitation>
was the first author to notice the differential characters of the Japanese species of
<taxonomicName id="4C544D75FF91FFB1F960080CFE37AFD7" authorityName="Wahlgren" authorityYear="1917" box="[362,495,635,658]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1F960080CFE37AFD7" box="[362,495,635,658]" italics="true" pageId="2" pageNumber="611">Norellisoma</emphasis>
</taxonomicName>
(or the subfamily
<taxonomicName id="4C544D75FF91FFB1FAA5080DFCFEAFD7" authorityName="Ager" authorityYear="1959" box="[687,806,634,658]" class="Rhynchonellata" family="Norellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rhynchonellida" pageId="2" pageNumber="611" phylum="Brachiopoda" rank="subFamily" subFamily="Norellinae">Norellinae</taxonomicName>
).
<bibRefCitation id="EFC54B07FF91FFB1FB3F080EFC1FAFD7" author="SIFNER F." box="[821,967,633,658]" pageId="2" pageNumber="611" pagination="1 - 90" refId="ref5452" refString="SIFNER F. 2003: The family Scathophagidae (Diptera) of the Czech and Slovak Republics (with notes on selected Palaearctic taxa). Acta Musei Nationalis Pragae, Series B - Historia Naturalis 59: 1 - 90." type="journal article" year="2003">ŠIFNER (2003)</bibRefCitation>
considered the Japanese species as intermediate between the genera
<taxonomicName id="4C544D75FF91FFB1FADE08ECFC81AFF7" authorityName="Wahlgren" authorityYear="1917" box="[724,857,667,690]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1FADE08ECFC81AFF7" box="[724,857,667,690]" italics="true" pageId="2" pageNumber="611">Norellisoma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D75FF91FFB1FB8708ECFC21AFF7" authorityName="Fallen" authorityYear="1810" box="[909,1017,667,690]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1FB8708ECFC21AFF7" box="[909,1017,667,690]" italics="true" pageId="2" pageNumber="611">Cordilura</emphasis>
</taxonomicName>
, probably representing a new genus.
<bibRefCitation id="EFC54B07FF91FFB1F9B208CDFD82AF97" author="OZEROV A. L." box="[440,602,698,722]" pageId="2" pageNumber="611" pagination="419 - 427" refId="ref5124" refString="OZEROV A. L. 2008: New species of Scathophagidae (Diptera). Russian Entomological Journal 17: 419 - 427." type="journal article" year="2008">OZEROV (2008)</bibRefCitation>
placed the Japanese and Chinese species
<taxonomicName id="4C544D75FF91FFB1FC2A08CCFDFAAFB7" authority="(Sun, 1992)" baseAuthorityName="Sun" baseAuthorityYear="1992" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="species" species="longiabdominum">
<emphasis id="B920EAE4FF91FFB1FC2A08CCFE44AFB7" italics="true" pageId="2" pageNumber="611">Norellisoma longiabdominum</emphasis>
(
<bibRefCitation id="EFC54B07FF91FFB1F9A108ADFDC2AFB7" author="SUN X." box="[427,538,730,754]" pageId="2" pageNumber="611" pagination="335 - 338" refId="ref5387" refString="SUN X. 1992: (Notes on the genus Norellia from China (Diptera, Scathophagidae )). Sinozoologia 4: 335 - 338 (in Chinese, with English summary)." type="journal article" year="1992">Sun, 1992</bibRefCitation>
)
</taxonomicName>
in the genus
<taxonomicName id="4C544D75FF91FFB1FABA08ACFCC4AFB7" authorityName="Fallen" authorityYear="1810" box="[688,796,731,754]" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAE4FF91FFB1FABA08ACFCC4AFB7" box="[688,796,731,754]" italics="true" pageId="2" pageNumber="611">Cordilura</emphasis>
</taxonomicName>
(subgenus
<taxonomicName id="4C544D75FF91FFB1FB9D08ACFF08AE57" authority="James, 1955" authorityName="James" authorityYear="1955" class="Insecta" family="Scathophagidae" genus="Cordilura" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="subGenus" subGenus="Cordilurina">
<emphasis id="B920EAE4FF91FFB1FB9D08ACFBC0AFB7" box="[919,1048,731,754]" italics="true" pageId="2" pageNumber="611">Cordilurina</emphasis>
James, 1955
</taxonomicName>
), citing the presence of rows of long anteroventral and posterovetral bristles on the fore femora and tibiae (
<emphasis id="B920EAE4FF91FFB1F968096CFE58AE77" box="[354,384,795,818]" italics="true" pageId="2" pageNumber="611">sic</emphasis>
!), a long apical seta on palpus, two pairs of setae on scutum and long hairs on arista. Based on the characters mentioned above under the generic diagnosis, I instead propose for
<taxonomicName id="4C544D75FF91FFB1F91D092CFD80AE37" baseAuthorityName="Sun" baseAuthorityYear="1992" box="[279,600,859,882]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="species" species="longiabdominum">
<emphasis id="B920EAE4FF91FFB1F91D092CFD80AE37" box="[279,600,859,882]" italics="true" pageId="2" pageNumber="611">Norellisoma longiabdominum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D75FF91FFB1FA81092CFC56AE34" authority="Seguy, 1948 a" authorityName="♀, Cotyp., E. Seguy" authorityYear="1948" box="[651,910,858,882]" class="Insecta" family="Scathophagidae" genus="Norellisoma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="species" species="agrion" status="gen. nov.">
<emphasis id="B920EAE4FF91FFB1FA81092CFD29AE37" box="[651,753,859,882]" italics="true" pageId="2" pageNumber="611">N. agrion</emphasis>
<bibRefCitation id="EFC54B07FF91FFB1FAFD092DFCA4AE37" author="SEGUY E." box="[759,892,858,882]" pageId="2" pageNumber="611" refId="ref5326" refString="SEGUY E. 1948: Dipteres nouveaux ou peu connus d'Extreme-Orient. Notes d'Entomologie Chinoise 12: 153 - 142." type="journal volume" year="1948">Séguy, 1948</bibRefCitation>
a
</taxonomicName>
<taxonomicNameLabel id="A213579FFF91FFB1FB99092DFBDFAE37" box="[915,1031,858,882]" pageId="2" pageNumber="611" rank="genus">new genus</taxonomicNameLabel>
,
<taxonomicName id="4C544D75FF91FFB1FC1B092CFBBEAE37" authority="Šifner, 2010" authorityName="Šifner" authorityYear="2010" box="[1041,1126,859,882]" class="Insecta" family="Scathophagidae" genus="Milania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EAE4FF91FFB1FC1B092CFBBEAE37" box="[1041,1126,859,882]" italics="true" pageId="2" pageNumber="611">Milania</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A213579FFF91FFB1F89D090DFF27AED4" box="[151,255,890,914]" pageId="2" pageNumber="611" rank="genus">gen. nov.</taxonomicNameLabel>
, belonging to the tribe
<taxonomicName id="4C544D75FF91FFB1FA08090DFDA2AED7" box="[514,634,890,914]" class="Insecta" family="Scathophagidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="611" phylum="Arthropoda" rank="tribe" tribe="Cordilurini">Cordilurini</taxonomicName>
(cf.
<bibRefCitation id="EFC54B07FF91FFB1FABA090EFCE0AED7" author="SIFNER F." box="[688,824,889,914]" pageId="2" pageNumber="611" pagination="1 - 90" refId="ref5452" refString="SIFNER F. 2003: The family Scathophagidae (Diptera) of the Czech and Slovak Republics (with notes on selected Palaearctic taxa). Acta Musei Nationalis Pragae, Series B - Historia Naturalis 59: 1 - 90." type="journal article" year="2003">ŠIFNER 2003</bibRefCitation>
). Both included species are redescribed below with emphasis on the previously undescribed genitalia of both sexes.
</paragraph>
</subSubSection>
</treatment>
</document>