treatments-xml/data/03/FD/87/03FD87DC0068FA5C5895FD42FA116A74.xml

<document id="C9C2A6E78D10A99AF975571611D92343" ID-DOI="10.11646/zootaxa.4816.1.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954133" ID-ZooBank="15A7F121-C570-4978-A83E-289CA54D69EF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339302206" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Hirabayashi, Isao" docDate="2020" docId="03FD87DC0068FA5C5895FD42FA116A74" docLanguage="en" docName="zootaxa.4816.1.5.pdf" docOrigin="Zootaxa 4816 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Naushonia kiiensis Komai &amp; Hirabayashi 2020, n. sp." docType="treatment" docUuid="A06C40A9-BEA5-463E-8B25-620F96909022" docUuidSource="ZooBank" docVersion="3" lastPageNumber="100" masterDocId="FFC4FFA40069FA545802FFD3FF916B13" masterDocTitle="A new species of the laomediid mud shrimp genus Naushonia Kingsley, 1897 (Decapoda: Gebiidea) from Japan" masterLastPageNumber="100" masterPageNumber="92" pageNumber="93" updateTime="1698782348775" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="527B1F707C69271BC70EF972D1BFF80A">A new species of the laomediid mud shrimp genus Naushonia Kingsley, 1897 (Decapoda: Gebiidea) from Japan</mods:title>
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<mods:namePart id="28F20E98275327E1734F1BBAD338451C">Komai, Tomoyuki</mods:namePart>
<mods:affiliation id="BE1BBCE852D779667629DB5938653DFC">Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan.</mods:affiliation>
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<mods:namePart id="13A688798AACBAE3119FECD9EDEF6309">Hirabayashi, Isao</mods:namePart>
<mods:affiliation id="464B23BAD8D9126680DC5C97463B4841">Kushimoto Marine Park Center Ltd, 1157 Arita, Kushimoto, Wakayama, 649 - 3514 Japan.</mods:affiliation>
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<mods:title id="8FF898EE9FFE59144FFC967DCFE83324">Zootaxa</mods:title>
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<mods:date id="8687B8448B1F62F7C51B46EBEAD17A17">2020</mods:date>
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<treatment id="03FD87DC0068FA5C5895FD42FA116A74" LSID="urn:lsid:zoobank.org:act:A06C40A9-BEA5-463E-8B25-620F96909022" httpUri="http://treatment.plazi.org/id/03FD87DC0068FA5C5895FD42FA116A74" lastPageId="8" lastPageNumber="100" pageId="1" pageNumber="93">
<subSubSection id="C34E65410068FA555895FD42FD1269DC" pageId="1" pageNumber="93" type="nomenclature">
<paragraph id="8BEB36CA0068FA555895FD42FE5869B8" blockId="1.[151,643,657,755]" box="[151,457,657,684]" pageId="1" pageNumber="93">
<heading id="D0A381A60068FA555895FD42FE5869B8" bold="true" box="[151,457,657,684]" fontSize="11" level="1" pageId="1" pageNumber="93" reason="1">
<emphasis id="B920EAD80068FA555895FD42FE5869B8" bold="true" box="[151,457,657,684]" pageId="1" pageNumber="93">
<taxonomicName id="4C544D490068FA555895FD42FEEE69BF" ID-CoL="3f1f3afe-9731-456b-8a80-5038d99805fa" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[151,383,657,684]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="93" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD80068FA555895FD42FEEE69BF" bold="true" box="[151,383,657,684]" italics="true" pageId="1" pageNumber="93">Naushonia kiiensis</emphasis>
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<taxonomicNameLabel id="A21357A30068FA555985FD42FE5869B8" box="[391,457,657,683]" pageId="1" pageNumber="93" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BEB36CA0068FA555895FD66FD1269DC" blockId="1.[151,643,657,755]" box="[151,643,692,719]" pageId="1" pageNumber="93">[New Japanese name: Kii-kagite-shako-ebi]</paragraph>
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<subSubSection id="C34E65410068FA555895FD0AFF6969E0" box="[151,248,728,755]" pageId="1" pageNumber="93" type="description">
<paragraph id="8BEB36CA0068FA555895FD0AFF6969E0" blockId="1.[151,643,657,755]" box="[151,248,728,755]" pageId="1" pageNumber="93">
<figureCitation id="136F2A4F0068FA555895FD0AFF6969E0" box="[151,248,728,755]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1778,1803]" captionStartId-1="3.[151,250,1751,1776]" captionStartId-2="4.[151,250,1626,1651]" captionStartId-3="6.[151,250,1734,1759]" captionStartId-4="7.[151,250,1947,1972]" captionTargetBox-0="[164,1424,354,1742]" captionTargetBox-1="[164,1426,496,1716]" captionTargetBox-2="[164,1425,360,1591]" captionTargetBox-3="[164,1426,192,1697]" captionTargetBox-4="[151,1436,309,1921]" captionTargetId-0="figure@2.[151,1436,345,1753]" captionTargetId-1="figure@3.[151,1436,489,1726]" captionTargetId-2="figure@4.[151,1436,345,1601]" captionTargetId-3="figure@6.[151,1436,181,1709]" captionTargetId-4="figure@7.[151,1436,309,1922]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." captionText-1="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." captionText-2="FIGURE 3. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted)." captionText-3="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." captionText-4="FIGURE 5. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. Habitus, showing coloration in life. A, dorsal view; B, right lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.3954135" figureDoi-1="http://doi.org/10.5281/zenodo.3954137" figureDoi-2="http://doi.org/10.5281/zenodo.3954139" figureDoi-3="http://doi.org/10.5281/zenodo.3954141" figureDoi-4="http://doi.org/10.5281/zenodo.3954143" httpUri-0="https://zenodo.org/record/3954135/files/figure.png" httpUri-1="https://zenodo.org/record/3954137/files/figure.png" httpUri-2="https://zenodo.org/record/3954139/files/figure.png" httpUri-3="https://zenodo.org/record/3954141/files/figure.png" httpUri-4="https://zenodo.org/record/3954143/files/figure.png" pageId="1" pageNumber="93">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E65410068FA555895FCF3FAB1684C" pageId="1" pageNumber="93" type="materials_examined">
<paragraph id="8BEB36CA0068FA555895FCF3FAB1684C" blockId="1.[151,1437,800,2015]" pageId="1" pageNumber="93">
<emphasis id="B920EAD80068FA555895FCF3FEE86828" bold="true" box="[151,377,800,827]" pageId="1" pageNumber="93">Material examined</emphasis>
. 
<materialsCitation id="3B3C3C970068FA55598AFCF2FAB1684C" collectingDate="2018-05-22" collectionCode="SCUBA" collectorName="I. Hirabayashi." country="Japan" county="Kii Peninsula" elevation="14" location="Sabiura" municipality="Kushimoto" pageId="1" pageNumber="93" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EF88680068FA55598AFCF2FE646828" box="[392,501,801,827]" pageId="1" pageNumber="93" type="holotype">Holotype</typeStatus>
: CBM-ZC 15914, male (cl 
<quantity id="4CAC9B2F0068FA555B2AFCF3FC136828" box="[808,898,800,827]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="93" unit="mm" value="3.5">3.5 mm</quantity>
), 
<location id="8E8B60110068FA555B97FCF2FC7C6828" LSID="urn:lsid:plazi:treatment:03FD87DC0068FA5C5895FD42FA116A74:8E8B60110068FA555B97FCF2FC7C6828" box="[917,1005,801,827]" country="Japan" county="Kii Peninsula" municipality="Kushimoto" name="Sabiura" pageId="1" pageNumber="93">Sabiura</location>
, 
<collectingMunicipality id="6B8FACB00068FA555BFAFCF2FBE66828" box="[1016,1143,801,827]" pageId="1" pageNumber="93">Kushimoto</collectingMunicipality>
, 
<collectingCounty id="628A4E460068FA555C81FCF2FA8F6828" box="[1155,1310,801,827]" pageId="1" pageNumber="93">Kii Peninsula</collectingCounty>
, 
<collectingCountry id="F343765A0068FA555D2BFCF2FAFD6828" box="[1321,1388,801,827]" name="Japan" pageId="1" pageNumber="93">Japan</collectingCountry>
, un- der dead coral rubble on sand bottom at 
<elevation id="0079D1F90068FA555A58FC96FD4A684C" box="[602,731,837,863]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="93" unit="m" value="14.0">
<quantity id="4CAC9B2F0068FA555A58FC96FD02684C" box="[602,659,837,863]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="93" unit="m" value="14.0">14 m</quantity>
depth
</elevation>
, 
<date id="FFEA100A0068FA555AE4FC96FCEE684C" box="[742,895,837,863]" pageId="1" pageNumber="93" value="2018-05-22">
<collectingDate id="EFAEE9E20068FA555AE4FC96FCEE684C" box="[742,895,837,863]" pageId="1" pageNumber="93" value="2018-05-22">22 May 2018</collectingDate>
</date>
, 
<collectionCode id="ED45AE0F0068FA555B88FC96FC77684C" box="[906,998,837,863]" pageId="1" pageNumber="93">SCUBA</collectionCode>
diving, coll. 
<collectorName id="26A1531C0068FA555C78FC96FAB1684C" box="[1146,1312,837,863]" pageId="1" pageNumber="93">I. Hirabayashi.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E65410068FA5558C5FCBBFEC96FF8" pageId="1" pageNumber="93" type="diagnosis">
<paragraph id="8BEB36CA0068FA5558C5FCBBFEC96FF8" blockId="1.[151,1437,800,2015]" pageId="1" pageNumber="93">
<emphasis id="B920EAD80068FA5558C5FCBBFED36890" bold="true" box="[199,322,872,899]" pageId="1" pageNumber="93">Diagnosis.</emphasis>
Rostrum subtriangular, with spinose lateral margins. Carapace anterior dorsum with 5 longitudinal carinae, all spinulose, middorsal carina extending nearly to posterodorsal margin, interrupted by W-shaped cervical groove; posterior dorsum rugose. Pleomeres 1 and 2 each with low but clearly delimited middorsal carina, pleomeres 3–5 without clearly discernible middorsal carina; pleura 2–5 all smooth on margins. Telson lateral margins with 3 pairs of spinules. Antennal scaphocerite lateral margin serrated with 7 or 8 spines, including straight terminal spine. Maxilliped 3 merus with spinose distolateral margin. Pereopod 1 ischium with spinulose lateral margin; merus lateral margin also spinulose, dorsomesial distal lobe with 3 strong spines, dorsal surface with submarginal ridge adjacent to mesial margin; palm lateral margin serrated with 14–22 small spines, mesial margin also serrated proximal to pollex, occlusal margin serrated along entire length, middle tooth prominent, stronger than pollex. Pereopods 3 and 4 dactyli bearing few minute spiniform setae on lateral surfaces. Uropodal protopod smooth on posterior margin.
</paragraph>
</subSubSection>
<subSubSection id="C34E65410068FA5158C5FB27FD316E8C" lastPageId="5" lastPageNumber="97" pageId="1" pageNumber="93" type="description">
<paragraph id="8BEB36CA0068FA5558C5FB27FB9C6E1C" blockId="1.[151,1437,800,2015]" box="[199,1037,1268,1295]" pageId="1" pageNumber="93">
<emphasis id="B920EAD80068FA5558C5FB27FEC86E1C" bold="true" box="[199,345,1268,1295]" pageId="1" pageNumber="93">Description.</emphasis>
<emphasis id="B920EAD80068FA555962FB26FD9A6E1C" box="[352,523,1269,1295]" italics="true" pageId="1" pageNumber="93">
<typeStatus id="54EF88680068FA555962FB26FE566E1C" box="[352,455,1269,1295]" pageId="1" pageNumber="93" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Body (
<figureCitation id="136F2A4F0068FA555A5FFB26FD4A6E1C" box="[605,731,1269,1295]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="1" pageNumber="93">Figs 1A, B</figureCitation>
, 
<figureCitation id="136F2A4F0068FA555AE5FB26FD656E1C" box="[743,756,1269,1295]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="1" pageNumber="93">2</figureCitation>
A–C) moderately robust.
</paragraph>
<paragraph id="8BEB36CA0068FA5558C5FACAFE766E8C" blockId="1.[151,1437,800,2015]" pageId="1" pageNumber="93">
Rostrum (
<figureCitation id="136F2A4F0068FA55593BFACAFE086E20" box="[313,409,1305,1331]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="1" pageNumber="93">Figs 1A</figureCitation>
, 
<figureCitation id="136F2A4F0068FA5559A5FACAFE5A6E20" box="[423,459,1305,1331]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="1" pageNumber="93">2A</figureCitation>
) strongly flattened dorsoventrally, subtriangular in dorsal view, slightly overreaching distal margin of article 2 of antennular peduncle, terminating in small spine overreaching subterminal spines; lateral margins slightly convex, each serrated with 6 small spines; dorsal surface with shallow median depression extending posteriorly to rostral base.
</paragraph>
<paragraph id="8BEB36CA0068FA5558C5FA7AFBD76C14" blockId="1.[151,1437,800,2015]" pageId="1" pageNumber="93">
Carapace (
<figureCitation id="136F2A4F0068FA55593FFA7AFE046ED0" box="[317,405,1449,1475]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="1" pageNumber="93">Fig. 1A</figureCitation>
, 
<figureCitation id="136F2A4F0068FA55599DFA7AFE526ED0" box="[415,451,1449,1475]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="1" pageNumber="93">2A</figureCitation>
) subcylindrical, with distinct linea thalassinica extending over entire length; surface with few very short setae. External orbital spine simple, acuminate, directed forward. Anterolateral margin spinulose, deeply notched just inferior to small branchiostegal spine; pterygostomial angle acuminate with 4small spines. Gastric region convex, sloping down to rostrum, with five longitudinal carinae, including mid-dorsal carina interrupted by cervical groove, all carinae spinulose; mid-dorsal carina low, anterior section starting from level of post-antennal notch, posterior section not reaching to posterodorsal margin of carapace; submedian carinae slightly diverging posteriorly, extending from posterior to orbital margin to about half length of carapace; lateral carinae diverging posteriorly, extending from level of orbital margin to 0.4 of carapace length. Cervical groove conspicuous, across slightly posterior to mid-length of carapace, W-shaped in dorsal view. Posterior dorsum rugose with sparse granules and faint elevations. Branchiostegite largely smooth, but with few granules dorsally.
</paragraph>
<paragraph id="8BEB36CA0068FA5658C5F8C2FD416A50" blockId="1.[151,1437,800,2015]" lastBlockId="2.[151,1437,152,323]" lastPageId="2" lastPageNumber="94" pageId="1" pageNumber="93">
Pleon (
<figureCitation id="136F2A4F0068FA555915F8C2FEE56C38" box="[279,372,1809,1835]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="1" pageNumber="93">Figs 1B</figureCitation>
, 
<figureCitation id="136F2A4F0068FA555980F8C2FE576C38" box="[386,454,1809,1835]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="1" pageNumber="93">2B, C</figureCitation>
) slightly depressed dorsoventrally; pleura 1–5 all rounded marginally. Pleomere 1 unarmed on anterolateral margin; posterior tergum with distinct median carina and weakly delimited transverse carina on either side, anteromesial part forming distinct shoulder; anterior tergum depressed below; pleuron concealed by anterior pleuron of pleomere 2. Pleomere 2 with obsolescent mid-dorsal carina on tergum; lateral surface with distinct longitudinal carina (= lateral carina) dividing tergum and pleura; pleuron slightly asymmetrical. Pleomere 3 tergum with trace of middorsal carina anteriorly; lateral carina distinct. Pleomere 4 tergum without even trace of mid-dorsal carina; lateral carina distinct. Pleomere 5 tergum with trace of mid-dorsal carina and transverse carina on either side; lateral carina short, but distinct. Pleomere 6 widened posteriorly, smooth; posterolateral process blunt. Telson (
<figureCitation id="136F2A4F006BFA56593AFF32FE016BE8" box="[312,400,225,251]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="2" pageNumber="94">Fig. 2D</figureCitation>
) 1.3 times longer than wide, slightly narrowing posteriorly; dorsal surface shallowly sulcate medially, with longitudinal row of 4 minute denticles on either side; lateral margins with 3 pairs of minute spines in posterior half; posterior margin rounded, unarmed.
</paragraph>
<caption id="DF2B6642006BFA565895F921FD746C3C" ID-DOI="http://doi.org/10.5281/zenodo.3954135" ID-Zenodo-Dep="3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="2" pageNumber="94" startId="2.[151,250,1778,1803]" targetBox="[164,1424,354,1742]" targetPageId="2">
<paragraph id="8BEB36CA006BFA565895F921FD746C3C" blockId="2.[151,1436,1778,1839]" pageId="2" pageNumber="94">
<emphasis id="B920EAD8006BFA565895F921FE9C6C18" bold="true" box="[151,269,1778,1803]" pageId="2" pageNumber="94">FIGURE 1</emphasis>
. 
<taxonomicName id="4C544D49006BFA56591BF920FE486C18" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[281,473,1779,1803]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="94" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006BFA56591BF920FE486C18" box="[281,473,1779,1803]" italics="true" pageId="2" pageNumber="94">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006BFA5659DDF920FD896C18" bold="true" box="[479,536,1779,1803]" pageId="2" pageNumber="94">
<taxonomicNameLabel id="A21357A3006BFA5659DDF920FD896C18" box="[479,536,1779,1803]" pageId="2" pageNumber="94" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view.
</paragraph>
</caption>
<paragraph id="8BEB36CA006BFA5658C5F88EFCA76C88" blockId="2.[151,1436,1885,2020]" pageId="2" pageNumber="94">
Ocular peduncles (
<figureCitation id="136F2A4F006BFA56599DF88EFD916C64" box="[415,512,1885,1911]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="2" pageNumber="94">Figs 1A</figureCitation>
, 
<figureCitation id="136F2A4F006BFA565A0DF88EFDA26C64" box="[527,563,1885,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="2" pageNumber="94">2A</figureCitation>
) short, without tubercle or spine on distomesial margin; cornea darkly pigmented, as wide as eyestalk, partially visible in dorsal view.
</paragraph>
<paragraph id="8BEB36CA006BFA5758C5F876FD656BE8" blockId="2.[151,1436,1885,2020]" lastBlockId="3.[151,1437,153,467]" lastPageId="3" lastPageNumber="95" pageId="2" pageNumber="94">
Antennular peduncle (
<figureCitation id="136F2A4F006BFA5659C0F876FDB16CAC" box="[450,544,1957,1983]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="2" pageNumber="94">Figs 1A</figureCitation>
, 
<figureCitation id="136F2A4F006BFA565A2FF876FDC06CAC" box="[557,593,1957,1983]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="2" pageNumber="94">2A</figureCitation>
) short, nearly reaching distal margin of article 5 of antennal peduncle. Article 1 not visible in dorsal view, nearly as long as distal two articles combined, armed with minute distolateral and small ventromesial distal spines; dorsal surface with statolith opening closed by stiff setae. Article 2 with minute distolateral spine. Article 3 unarmed. Lateral flagellum slightly longer than peduncle, composed of about 10 articles; mesial flagellum about 0.7 length of lateral flagellum.
</paragraph>
<paragraph id="8BEB36CA006AFA5758C5FED6FA056AC0" blockId="3.[151,1437,153,467]" pageId="3" pageNumber="95">
Antennal peduncle (
<figureCitation id="136F2A4F006AFA5759AEFED6FD9A6A0C" box="[428,523,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="3" pageNumber="95">Figs 1A</figureCitation>
, 
<figureCitation id="136F2A4F006AFA575A1AFED6FDAD6A0C" box="[536,572,261,287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="3" pageNumber="95">2A</figureCitation>
) stout. Article 1 with spinulose ventrodistal margin. Article 2 with row of 4 small spines on distolateral margin and 1 small spine on ventrodistal margin. Article 3 with small subdistal spine on ventromesial margin. Articles 4 and 5 each with distolateral spine; article 5 wider than long. Scaphocerite moderately broad, short, overreaching base of peduncular article 5; lateral margin slightly convex, serrated with 7 or 8 spines (including straight terminal spine); mesial margin strongly convex, with row of stiff setae; dorsal surface with low longitudinal carina. Antennal flagellum stout, longer than body; articles with several short setae on distal margins.
</paragraph>
<caption id="DF2B6642006AFA575895F904FDA06C2B" ID-DOI="http://doi.org/10.5281/zenodo.3954137" ID-Zenodo-Dep="3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="3" pageNumber="95" startId="3.[151,250,1751,1776]" targetBox="[164,1426,496,1716]" targetPageId="3">
<paragraph id="8BEB36CA006AFA575895F904FDA06C2B" blockId="3.[151,1437,1751,1848]" pageId="3" pageNumber="95">
<emphasis id="B920EAD8006AFA575895F904FE9E6DE3" bold="true" box="[151,271,1751,1776]" pageId="3" pageNumber="95">FIGURE 2</emphasis>
. 
<taxonomicName id="4C544D49006AFA57591CF90BFE716DE3" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[286,480,1752,1776]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="95" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006AFA57591CF90BFE716DE3" box="[286,480,1752,1776]" italics="true" pageId="3" pageNumber="95">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006AFA5759EAF90BFDB56DE3" bold="true" box="[488,548,1752,1776]" pageId="3" pageNumber="95">
<taxonomicNameLabel id="A21357A3006AFA5759EAF90BFDB56DE3" box="[488,548,1752,1776]" pageId="3" pageNumber="95" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view.
</paragraph>
</caption>
<paragraph id="8BEB36CA006AFA5758C5F8B5FABA6C93" blockId="3.[151,1437,1894,2029]" box="[199,1323,1894,1921]" pageId="3" pageNumber="95">Epistomial horn relatively slender, almost reaching distal margin of article 1 of antennular peduncle.</paragraph>
<paragraph id="8BEB36CA006AFA5058C5F859FB2E6A50" blockId="3.[151,1437,1894,2029]" lastBlockId="4.[151,1437,152,323]" lastPageId="4" lastPageNumber="96" pageId="3" pageNumber="95">
Mouthparts not dissected. Maxilliped 3 (
<figureCitation id="136F2A4F006AFA575A8CF859FD746CB6" box="[654,741,1930,1957]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="3" pageNumber="95">Fig. 4A</figureCitation>
) with moderately stout endopod composed of 5 articles. Coxa unarmed. Basis short, with 2 spinules on mesial surface. Ischium slightly widened distally, distodorsal angle only slightly produced; crista dentata well developed, with row of unequal, sharp teeth, distomesial angle produced (
<figureCitation id="136F2A4F006AFA505D6DF801FF2A6BA0" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" lastPageId="4" lastPageNumber="96" pageId="3" pageNumber="95">Fig. 4B</figureCitation>
). Merus shorter than ischium; distolateral to distoventral margin with 5 spines; dorsolateral margin also with 1 small spine subdistally. Carpus short, cup-shaped; extensor surface unarmed. Propodus subcylindrical. Dactylus subequal in length to carpus, tapering into blunt tip, with thick cluster of setae on flexor surface. Exopod proximal article overreaching distal margin of ischium; flagellum consisting of 5 articles. Epipod large, margins with sharp denticles; mastigobranch well developed; podobranch consisting of numerous, slender lamella.
</paragraph>
<caption id="DF2B6642006DFA505895F989FA016D84" ID-DOI="http://doi.org/10.5281/zenodo.3954139" ID-Zenodo-Dep="3954139" httpUri="https://zenodo.org/record/3954139/files/figure.png" pageId="4" pageNumber="96" startId="4.[151,250,1626,1651]" targetBox="[164,1425,360,1591]" targetPageId="4">
<paragraph id="8BEB36CA006DFA505895F989FA016D84" blockId="4.[151,1436,1626,1687]" pageId="4" pageNumber="96">
<emphasis id="B920EAD8006DFA505895F989FE9E6D60" bold="true" box="[151,271,1626,1651]" pageId="4" pageNumber="96">FIGURE 3</emphasis>
. 
<taxonomicName id="4C544D49006DFA50591FF988FE4E6D60" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[285,479,1627,1651]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="96" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006DFA50591FF988FE4E6D60" box="[285,479,1627,1651]" italics="true" pageId="4" pageNumber="96">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006DFA5059E5F988FDB36D60" bold="true" box="[487,546,1627,1651]" pageId="4" pageNumber="96">
<taxonomicNameLabel id="A21357A3006DFA5059E5F988FDB36D60" box="[487,546,1627,1651]" pageId="4" pageNumber="96" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted).
</paragraph>
</caption>
<paragraph id="8BEB36CA006DFA5158C5F913FC8C6908" blockId="4.[151,1437,1728,2042]" lastBlockId="5.[151,1437,153,2015]" lastPageId="5" lastPageNumber="97" pageId="4" pageNumber="96">
First pereopods (= chelipeds) (
<figureCitation id="136F2A4F006DFA505A22F913FDF26DC9" box="[544,611,1728,1754]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1651]" captionTargetBox="[164,1425,360,1591]" captionTargetId="figure@4.[151,1436,345,1601]" captionTargetPageId="4" captionText="FIGURE 3. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954139" httpUri="https://zenodo.org/record/3954139/files/figure.png" pageId="4" pageNumber="96">Fig. 3</figureCitation>
) large, subchelate, equal in size, symmetrical in shape, strongly flattened dorsoventrally. Ischium widened distally; mesial surface flanked by clearly delimited dorsomesial and ventromesial margins, both with row of microscopic spinules; distomesial distal angle produced into small blunt spine; lateral margin carinate, also with row of microscopic spinules. Merus widened distally, cross section nearly triangular; ventromesial margin clearly delimited only in distal half, almost smooth; dorsomesial margin distinct over entire length, microscopically spinulose, distal angle produced into 3-spined lobe; mesial surface excavate distally; dorsal surface with some small spines distally and with longitudinal ridge adjacent to ventromesial margin, merging proximally to ventromesial margin; lateral margin bluntly carinate, serrated with row of spinules increasing in size distally. Carpus short, slightly widened distally, surface microscopically granular; lateral margin sharply carinate; dorsodistal margin spinulose. Palm elongate subovate in general outline, rhomboidal in cross-section, about twice as long as wide (including pollex), longer than ischium and merus combined; lateral margin carinate, serrated with row of spinules increasing in size distally (
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right, 
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left); mesial margin sharply carinate, serrated with row of 7 small spines, and with row of sparse setae; dorsal surface also weakly elevated along midline, weakly granular along midline, with few spines adjacent to occlusal margin; ventral surface weakly elevated along midline, with scattered granules lateral to midline, nearly smooth mesial to midline; pollex directed forward, acuminate, arising at about mid-length of palm; occlusal margin strongly oblique, with prominent tooth located at about midlength of occlusal margin, distal part serrated with row of sharp teeth, proximal part also serrated with smaller spines of similar size and sparse setae. Dactylus moderately slender, gently curved, closing completely against occlusal margin, tip overlapping dorsal surface of palm when closed; extensor margin with row of thin, numerous setae, proximal part not particularly expanded; flexor margin sharply carinate.
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FDF6FDD869DC" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
Pereopod 2 (
<figureCitation id="136F2A4F006CFA515954FDF6FE3C692C" box="[342,429,549,575]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4C</figureCitation>
) stout, simple. Ischium with produced ventrodistal angle. Merus with sparse row of elongate setae on ventral margin. Carpus about 0.3 times as long as merus. Propodus slightly longer than wide, slightly shorter than carpus. Dactylus lanceolate, terminating in acute unguis; extensor margin gently arcuate in mesial view; extensor surface with dense, more or less elongate serrate setae; flexor margin minutely pectinate in distal half of its length; mesial face glabrous (
<figureCitation id="136F2A4F006CFA5159E3FD66FDAD69DC" box="[481,572,693,719]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FD0AFA8A68B4" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
Pereopods 3–5 generally similar, but decreasing in length posteriorly. Pereopod 3 (
<figureCitation id="136F2A4F006CFA515C77FD0AFB5A69E0" box="[1141,1227,729,755]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4E</figureCitation>
) with sparse short setae on merus to dactylus; ischium about 0.6 length of merus; merus slightly narrowed distally, 4.8 times longer than wide, unarmed; carpus about 0.4 times as long as merus; dactylus (
<figureCitation id="136F2A4F006CFA515BBDFCF2FB826828" box="[959,1043,801,827]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4F</figureCitation>
) about 0.5 times as long propodus, about 6.3 times longer than wide, ending in slender, basally demarcated unguis, nearly straight except for slightly curved distal part, lateral surface with 3 minute spiniform setae adjacent to extensor margin, slightly sinuous flexor margin with row of minute setae followed by comb-like, toothed lamella on proximal half of its length.
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FC62FDDC6F00" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
Pereopod 4 (
<figureCitation id="136F2A4F006CFA515957FC62FE5E68D8" box="[341,463,945,971]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4G, H</figureCitation>
) dactylus with 2 minute spiniform setae on lateral surface. Pereopod 5 (
<figureCitation id="136F2A4F006CFA515CF5FC62FACC68D8" box="[1271,1373,945,971]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4I, J</figureCitation>
) dactylus flexor margin with comb-like toothed lamella on proximal 0.6, no spiniform setae on outer surface; gonopore located at tip of papilla-like projection.
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FBCEFDCB6F48" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
Branchial formula identical with other congeneric species (cf. 
<bibRefCitation id="EFC54B3B006CFA515B79FBCEFBE16F24" author="Komai, T. &amp; Anker, A." box="[891,1136,1052,1079]" pageId="5" pageNumber="97" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
: table 1). Epipod on pereopod 4 small, without marginal denticles.
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FBB6FE666FB0" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">Pleopod 1 absent. Pleopods 2–5 biramous, rami slender, lanceolate, subequal, without appendix interna; no appendix masculina of pleopod 2.</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FB7EFDDE6E44" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
Uropod (
<figureCitation id="136F2A4F006CFA51592FFB7EFE176FD4" box="[301,390,1197,1223]" captionStart="FIGURE 4" captionStartId="6.[151,250,1734,1759]" captionTargetBox="[164,1426,192,1697]" captionTargetId="figure@6.[151,1436,181,1709]" captionTargetPageId="6" captionText="FIGURE 4. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="5" pageNumber="97">Fig. 4K</figureCitation>
) with spinulose transverse suture on both exopod and endopod. Exopod with 5 small spines on distal half of lateral margin, with spiniform seta at posterolateral angle; dorsal surface with 2 longitudinal ridges, lateral ridge with few spinules, middorsal ridge unarmed. Endopod lateral margin with 3 small spines (including posterolateral spine), no spiniform seta at posterolateral angle; dorsal surface with spinose middorsal ridge. Protopod with smooth posterodorsal margin.
</paragraph>
<paragraph id="8BEB36CA006CFA5158C5FAB3FD316E8C" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
<emphasis id="B920EAD8006CFA5158C5FAB3FEFC6E68" bold="true" box="[199,365,1376,1403]" pageId="5" pageNumber="97">Colour in life.</emphasis>
Body and appendages generally semi-translucent white, hepatopancrea inside of cephalothorax pale yellow; cornea darkly pigmented (
<figureCitation id="136F2A4F006CFA515A4CFA56FD036E8C" box="[590,658,1412,1439]" captionStart="FIGURE 5" captionStartId="7.[151,250,1947,1972]" captionTargetBox="[151,1436,309,1921]" captionTargetId="figure@7.[151,1436,309,1922]" captionTargetPageId="7" captionText="FIGURE 5. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. Habitus, showing coloration in life. A, dorsal view; B, right lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954143" httpUri="https://zenodo.org/record/3954143/files/figure.png" pageId="5" pageNumber="97">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E6541006CFA5158C5FA7BFC4B6ED0" box="[199,986,1448,1475]" pageId="5" pageNumber="97" type="distribution">
<paragraph id="8BEB36CA006CFA5158C5FA7BFC4B6ED0" blockId="5.[151,1437,153,2015]" box="[199,986,1448,1475]" pageId="5" pageNumber="97">
<emphasis id="B920EAD8006CFA5158C5FA7BFEF06ED0" bold="true" box="[199,353,1448,1475]" pageId="5" pageNumber="97">Distribution.</emphasis>
Known only from the 
<typeStatus id="54EF8868006CFA515A62FA7AFD016ED0" box="[608,656,1449,1475]" pageId="5" pageNumber="97">type</typeStatus>
locality in 
<collectingCountry id="F343765A006CFA515B0DFA7AFCC36ED0" box="[783,850,1449,1475]" name="Japan" pageId="5" pageNumber="97">Japan</collectingCountry>
; 
<quantity id="4CAC9B2F006CFA515B5CFA7AFC066ED0" box="[862,919,1449,1475]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="97" unit="m" value="14.0">14 m</quantity>
deep.
</paragraph>
</subSubSection>
<subSubSection id="C34E6541006CFA5C58C5FA1EFCF66A50" lastPageId="8" lastPageNumber="100" pageId="5" pageNumber="97" type="discussion">
<paragraph id="8BEB36CA006CFA5158C5FA1EFC796CCC" blockId="5.[151,1437,153,2015]" pageId="5" pageNumber="97">
<emphasis id="B920EAD8006CFA5158C5FA1EFEAA6EF4" bold="true" box="[199,315,1485,1511]" pageId="5" pageNumber="97">Remarks.</emphasis>
<taxonomicName id="4C544D49006CFA515940FA1EFD876EF4" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[322,534,1485,1511]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515940FA1EFD876EF4" box="[322,534,1485,1511]" italics="true" pageId="5" pageNumber="97">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515A1FFA1EFDCA6EF4" bold="true" box="[541,603,1485,1511]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515A1FFA1EFDCA6EF4" box="[541,603,1485,1511]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
most closely resembles 
<taxonomicName id="4C544D49006CFA515B6FFA1DFB8F6EF4" authorityName="Komai &amp; Anker" authorityYear="2010" box="[877,1054,1485,1511]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515B6FFA1DFB8F6EF4" box="[877,1054,1485,1511]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
in the anterior carapace ornamentation and the shape and general armature of the pereopod 1. Nevertheless, the new species is distinguished from 
<taxonomicName id="4C544D49006CFA515895F9C5FEDB6D3C" authorityName="Komai &amp; Anker" authorityYear="2010" box="[151,330,1557,1583]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515895F9C5FEDB6D3C" box="[151,330,1557,1583]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
by the following particulars: (1) the gastric median and submedian carinae are spinulose in 
<taxonomicName id="4C544D49006CFA515D5FF9C5FF466D40" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515D5FF9C5FF466D40" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA5158DCF9EAFE8C6D40" bold="true" box="[222,285,1593,1619]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA5158DCF9EAFE8C6D40" box="[222,285,1593,1619]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA51592EF9EAFE146D40" box="[300,389,1593,1619]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="5" pageNumber="97">Fig. 1A</figureCitation>
), while generally granular in 
<taxonomicName id="4C544D49006CFA515ACDF9E9FC106D40" authorityName="Komai &amp; Anker" authorityYear="2010" box="[719,897,1593,1619]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515ACDF9E9FC106D40" box="[719,897,1593,1619]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006CFA515BB6F9EAFB3D6D40" author="Komai, T. &amp; Anker, A." box="[948,1196,1593,1619]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 5B, C; 
<bibRefCitation id="EFC54B3B006CFA515D31F9EAFEB06D64" author="Komai, T. &amp; Anker, A." pageId="5" pageNumber="97" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
: fig. 7A); (2) the cervical groove is W-shaped in the dorsal view in 
<taxonomicName id="4C544D49006CFA515C1DF98DFB056D64" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[1055,1172,1629,1655]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515C1DF98DFB056D64" box="[1055,1172,1629,1655]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515C9EF98EFB4A6D64" bold="true" box="[1180,1243,1629,1655]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515C9EF98EFB4A6D64" box="[1180,1243,1629,1655]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA515CEEF98EFAD76D64" box="[1260,1350,1629,1655]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="5" pageNumber="97">Fig. 2A</figureCitation>
), rather than broadly U-shaped or transverse in 
<taxonomicName id="4C544D49006CFA515A56F951FC966D88" authorityName="Komai &amp; Anker" authorityYear="2010" box="[596,775,1665,1691]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515A56F951FC966D88" box="[596,775,1665,1691]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006CFA515B3EF952FBA46D88" author="Komai, T. &amp; Anker, A." box="[828,1077,1665,1691]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 5A; 
<bibRefCitation id="EFC54B3B006CFA515C9DF952FA096D88" author="Komai, T. &amp; Anker, A." box="[1183,1432,1664,1691]" pageId="5" pageNumber="97" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
: fig. 7A); (3) the posterior carapace dorsum is rugose in 
<taxonomicName id="4C544D49006CFA515B0BF975FCEF6DAC" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[777,894,1701,1727]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515B0BF975FCEF6DAC" box="[777,894,1701,1727]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515B87F976FC556DAC" bold="true" box="[901,964,1701,1727]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515B87F976FC556DAC" box="[901,964,1701,1727]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA515BD1F976FBA06DAC" box="[979,1073,1701,1727]" captionStart="FIGURE 1" captionStartId="2.[151,250,1778,1803]" captionTargetBox="[164,1424,354,1742]" captionTargetId="figure@2.[151,1436,345,1753]" captionTargetPageId="2" captionText="FIGURE 1. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, left lateral view; B. pleon, telson and uropod, left lateral view." figureDoi="http://doi.org/10.5281/zenodo.3954135" httpUri="https://zenodo.org/record/3954135/files/figure.png" pageId="5" pageNumber="97">Figs 1A</figureCitation>
, 
<figureCitation id="136F2A4F006CFA515C3FF976FBF36DAC" box="[1085,1122,1701,1727]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="5" pageNumber="97">2A</figureCitation>
), whereas smooth in 
<taxonomicName id="4C544D49006CFA515D52F975FE946DF0" authorityName="Komai &amp; Anker" authorityYear="2010" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515D52F975FE946DF0" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006CFA51593BF91AFDBE6DF0" author="Komai, T. &amp; Anker, A." box="[313,559,1737,1763]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 5A, B); (4) the pleomere 1 is provided with a sharply delimited middorsal carina in 
<taxonomicName id="4C544D49006CFA515900F93DFEE96C14" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[258,376,1773,1799]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515900F93DFEE96C14" box="[258,376,1773,1799]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515982F93EFE516C14" bold="true" box="[384,448,1773,1799]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515982F93EFE516C14" box="[384,448,1773,1799]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA5159D3F93EFDBB6C14" box="[465,554,1773,1799]" captionStart="FIGURE 2" captionStartId="3.[151,250,1751,1776]" captionTargetBox="[164,1426,496,1716]" captionTargetId="figure@3.[151,1436,489,1726]" captionTargetPageId="3" captionText="FIGURE 2. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914. A, carapace and cephalic appendages, dorsal view (setae on left antennae omitted); B, pleomeres 1–4, dorsal view; C, pleomere 5, dorsal view; D, telson, dorsal view; E, right antennal peduncle, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3954137" httpUri="https://zenodo.org/record/3954137/files/figure.png" pageId="5" pageNumber="97">Fig. 2B</figureCitation>
), but only a trace is discernible in 
<taxonomicName id="4C544D49006CFA515BB6F93DFBF66C14" authorityName="Komai &amp; Anker" authorityYear="2010" box="[948,1127,1773,1799]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515BB6F93DFBF66C14" box="[948,1127,1773,1799]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006CFA515C9FF93EFA096C14" author="Komai, T. &amp; Anker, A." box="[1181,1432,1773,1799]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 5E); (5) the pereopod 1 merus is provided with a longitudinal carina on the dorsal surface adjacent to the mesial margin in 
<taxonomicName id="4C544D49006CFA51590BF8E5FEEF6C5C" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[265,382,1845,1871]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA51590BF8E5FEEF6C5C" box="[265,382,1845,1871]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515986F8E6FE526C5C" bold="true" box="[388,451,1845,1871]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515986F8E6FE526C5C" box="[388,451,1845,1871]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA5159D0F8E6FDB86C5C" box="[466,553,1845,1871]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1651]" captionTargetBox="[164,1425,360,1591]" captionTargetId="figure@4.[151,1436,345,1601]" captionTargetPageId="4" captionText="FIGURE 3. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954139" httpUri="https://zenodo.org/record/3954139/files/figure.png" pageId="5" pageNumber="97">Fig. 3B</figureCitation>
), whereas such a carina is absent in 
<taxonomicName id="4C544D49006CFA515BBFF8E5FBFE6C5C" authorityName="Komai &amp; Anker" authorityYear="2010" box="[957,1135,1845,1871]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515BBFF8E5FBFE6C5C" box="[957,1135,1845,1871]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006CFA515CA0F8E6FA096C5C" author="Komai, T. &amp; Anker, A." box="[1186,1432,1845,1871]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 6A; 
<bibRefCitation id="EFC54B3B006CFA5158F4F88AFE7F6C60" author="Komai, T. &amp; Anker, A." box="[246,494,1880,1907]" pageId="5" pageNumber="97" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
: fig. 7B); (6) the distomesial lobe of the pereopod 1 merus is armed with three teeth in 
<taxonomicName id="4C544D49006CFA5158B1F8ADFEB96C84" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[179,296,1917,1943]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA5158B1F8ADFEB96C84" box="[179,296,1917,1943]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA51592CF8AEFEFD6C84" bold="true" box="[302,364,1917,1943]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA51592CF8AEFEFD6C84" box="[302,364,1917,1943]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A4F006CFA515979F8AEFE436C84" box="[379,466,1917,1943]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1651]" captionTargetBox="[164,1425,360,1591]" captionTargetId="figure@4.[151,1436,345,1601]" captionTargetPageId="4" captionText="FIGURE 3. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954139" httpUri="https://zenodo.org/record/3954139/files/figure.png" pageId="5" pageNumber="97">Fig. 3B</figureCitation>
), rather than armed with more than four smaller teeth in 
<taxonomicName id="4C544D49006CFA515C45F8ADFB686C84" authorityName="Komai &amp; Anker" authorityYear="2010" box="[1095,1273,1917,1943]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515C45F8ADFB686C84" box="[1095,1273,1917,1943]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
; (7) the lateral margin of the pereopod 1 palm bears fewer spinules in 
<taxonomicName id="4C544D49006CFA515B16F871FC1A6CA8" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[788,907,1953,1979]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006CFA515B16F871FC1A6CA8" box="[788,907,1953,1979]" italics="true" pageId="5" pageNumber="97">N. kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006CFA515B96F872FC446CA8" bold="true" box="[916,981,1953,1979]" pageId="5" pageNumber="97">
<taxonomicNameLabel id="A21357A3006CFA515B96F872FC446CA8" box="[916,981,1953,1979]" pageId="5" pageNumber="97" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than in 
<taxonomicName id="4C544D49006CFA515C34F871FB7B6CA8" authorityName="Komai &amp; Anker" authorityYear="2010" box="[1078,1258,1953,1979]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="97" phylum="Arthropoda" rank="species" species="serratipalma">
<emphasis id="B920EAD8006CFA515C34F871FB7B6CA8" box="[1078,1258,1953,1979]" italics="true" pageId="5" pageNumber="97">N. serratipalma</emphasis>
</taxonomicName>
(cf. 
<figureCitation id="136F2A4F006CFA515D20F872FA0C6CA8" box="[1314,1437,1953,1979]" captionStart="FIGURE 3" captionStartId="4.[151,250,1626,1651]" captionTargetBox="[164,1425,360,1591]" captionTargetId="figure@4.[151,1436,345,1601]" captionTargetPageId="4" captionText="FIGURE 3. Naushonia kiiensis n. sp., holotype, male (cl 3.5 mm), CBM-ZC 15914, right pereopod 1. A, chela and carpus, dorsal view; B, same, ventral view (setae omitted); C, merus and ischium, dorsal view; D, same, ventral view (setae omitted)." figureDoi="http://doi.org/10.5281/zenodo.3954139" httpUri="https://zenodo.org/record/3954139/files/figure.png" pageId="5" pageNumber="97">Fig. 3A, B</figureCitation>
versus 
<bibRefCitation id="EFC54B3B006CFA5158E4F816FE4C6CCC" author="Komai, T. &amp; Anker, A." box="[230,477,1989,2015]" pageId="5" pageNumber="97" pagination="31 - 46" refId="ref4510" refString="Komai, T. &amp; Anker, A. (2010) Two new species of the laomediid genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea) from the Indo-West Pacific. Zootaxa, 2504 (1), 31 - 46. https: // doi. org / 10.11646 / zootaxa. 2504.1.3" type="journal article" year="2010">Komai &amp; Anker 2010</bibRefCitation>
: fig. 6C, D; 
<bibRefCitation id="EFC54B3B006CFA515A65F816FCCF6CCC" author="Komai, T. &amp; Anker, A." box="[615,862,1988,2015]" pageId="5" pageNumber="97" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
: fig. 7B, C).
</paragraph>
<caption id="DF2B6642006FFA525895F915FB516C58" ID-DOI="http://doi.org/10.5281/zenodo.3954141" ID-Zenodo-Dep="3954141" httpUri="https://zenodo.org/record/3954141/files/figure.png" pageId="6" pageNumber="98" startId="6.[151,250,1734,1759]" targetBox="[164,1426,192,1697]" targetPageId="6">
<paragraph id="8BEB36CA006FFA525895F915FB516C58" blockId="6.[151,1437,1734,1867]" pageId="6" pageNumber="98">
<emphasis id="B920EAD8006FFA525895F915FE9E6DCC" bold="true" box="[151,271,1734,1759]" pageId="6" pageNumber="98">FIGURE 4</emphasis>
. 
<taxonomicName id="4C544D49006FFA52591CF914FE716DCC" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[286,480,1735,1759]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006FFA52591CF914FE716DCC" box="[286,480,1735,1759]" italics="true" pageId="6" pageNumber="98">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006FFA5259EAF914FDB26DCC" bold="true" box="[488,547,1735,1759]" pageId="6" pageNumber="98">
<taxonomicNameLabel id="A21357A3006FFA5259EAF914FDB26DCC" box="[488,547,1735,1759]" pageId="6" pageNumber="98" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm), CBM-ZC 15914. A, right maxilliped 3, lateral view; B, ischium of left maxilliped 3, ventral view; C, left pereopod 2, lateral view; D, same, dactylus, mesial view; E, left pereopod 3, lateral view; F, same, dactylus, lateral view; G, left pereopod 4, lateral view; H, same, dactylus, lateral view; I, right pereopod 5, lateral view; J, same, dactylus, lateral view; K, left uropod, dorsal view (perpendicular; setae omitted).
</paragraph>
</caption>
<paragraph id="8BEB36CA006FFA5358C5F8AAFE966A0C" blockId="6.[151,1437,1913,2012]" lastBlockId="7.[151,1437,152,287]" lastPageId="7" lastPageNumber="99" pageId="6" pageNumber="98">
Prior to this study, three species of 
<taxonomicName id="4C544D49006FFA525A54F8AAFD5E6C80" authorityName="Kingsley" authorityYear="1897" box="[598,719,1913,1939]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAD8006FFA525A54F8AAFD5E6C80" box="[598,719,1913,1939]" italics="true" pageId="6" pageNumber="98">Naushonia</emphasis>
</taxonomicName>
have been recorded from Japanese waters, viz., 
<taxonomicName id="4C544D49006FFA525CF1F8A9FAE56C80" authorityName="Dworschak, Marin &amp; Anker" authorityYear="2006" box="[1267,1396,1914,1939]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="B920EAD8006FFA525CF1F8A9FAE56C80" box="[1267,1396,1914,1939]" italics="true" pageId="6" pageNumber="98">N. carinata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C544D49006FFA525D81F8A9FF686CA4" authorityName="Komai" authorityYear="2004" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B920EAD8006FFA525D81F8A9FF686CA4" italics="true" pageId="6" pageNumber="98">N. japonica</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C544D49006FFA52592FF84DFE5C6CA4" authorityName="Berggren" authorityYear="1992" box="[301,461,1949,1975]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="B920EAD8006FFA52592FF84DFE5C6CA4" box="[301,461,1949,1975]" italics="true" pageId="6" pageNumber="98">N. lactoalbida</emphasis>
</taxonomicName>
(cf. 
<bibRefCitation id="EFC54B3B006FFA5259FCF84EFD626CA4" author="Komai, T. &amp; Anker, A." box="[510,755,1948,1975]" pageId="6" pageNumber="98" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
). Of the three, 
<taxonomicName id="4C544D49006FFA525B96F84DFB826CA4" authorityName="Dworschak, Marin &amp; Anker" authorityYear="2006" box="[916,1043,1950,1975]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="carinata">
<emphasis id="B920EAD8006FFA525B96F84DFB826CA4" box="[916,1043,1950,1975]" italics="true" pageId="6" pageNumber="98">N. carinata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C544D49006FFA525C45F84DFB776CA4" authorityName="Berggren" authorityYear="1992" box="[1095,1254,1949,1975]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="98" phylum="Arthropoda" rank="species" species="lactoalbida">
<emphasis id="B920EAD8006FFA525C45F84DFB776CA4" box="[1095,1254,1949,1975]" italics="true" pageId="6" pageNumber="98">N. lactoalbida</emphasis>
</taxonomicName>
are known from the Ryukyu Islands, and in fact, the two species are wide spread in the tropical West Pacific or Indo-West Pacific (
<bibRefCitation id="EFC54B3B006EFA53589DFF4AFEDB6BA0" author="Berggren, M." box="[159,330,153,179]" pageId="7" pageNumber="99" pagination="514 - 22" refId="ref4307" refString="Berggren, M. (1992) Naushonia lactoalbida, new species (Decapoda: Thalassinidea: Laomediidae), a mud shrimp from Inhaca Island, Mocambique. Journal of Crustacean Biology, 12, 514 - 22. https: // doi. org / 10.2307 / 1549043" type="journal article" year="1992">Berggren 1992</bibRefCitation>
; 
<bibRefCitation id="EFC54B3B006EFA535957FF4AFE726BA0" author="Komai, T." box="[341,483,153,179]" pageId="7" pageNumber="99" pagination="15 - 26" refId="ref4450" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26. https: // doi. org / 10.18353 / crustacea. 33.0 _ 15" type="journal article" year="2004">Komai 2004</bibRefCitation>
; Dworschak et al. 2006; 
<bibRefCitation id="EFC54B3B006EFA535AF7FF4AFC2B6BA0" author="Anker, A. &amp; Paratama, I. S. &amp; Firdaus, M. &amp; Rahayu, D. L." box="[757,954,152,179]" pageId="7" pageNumber="99" pagination="301 - 342" refId="ref4167" refString="Anker, A., Paratama, I. S., Firdaus, M. &amp; Rahayu, D. L. (2015) On some interesting marine decapod crustaceans (Alpheidae, Laomediidae, Strahlaxiidae) from Lombok, Indonesia. Zootaxa, 3911 (3), 301 - 342. https: // doi. org / 10.11646 / zootaxa. 3911.3.1" type="journal article" year="2015">
Anker 
<emphasis id="B920EAD8006EFA535B43FF49FCE36BA0" box="[833,882,153,179]" italics="true" pageId="7" pageNumber="99">et al</emphasis>
. 2015
</bibRefCitation>
; 
<bibRefCitation id="EFC54B3B006EFA535BC7FF4AFB2D6BA0" author="Komai, T. &amp; Anker, A." box="[965,1212,152,179]" pageId="7" pageNumber="99" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
). On the other hand, 
<taxonomicName id="4C544D49006EFA535895FF6DFE8A6BC4" authorityName="Komai" authorityYear="2004" box="[151,283,190,215]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="99" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B920EAD8006EFA535895FF6DFE8A6BC4" box="[151,283,190,215]" italics="true" pageId="7" pageNumber="99">N. japonica</emphasis>
</taxonomicName>
is so far restricted to the warm temperate Japanese mainland (Honshu) (
<bibRefCitation id="EFC54B3B006EFA535C51FF6EFAC06BC4" author="Komai, T. &amp; Anker, A." box="[1107,1361,188,215]" pageId="7" pageNumber="99" pagination="341 - 360" refId="ref4568" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
). Thus the present new species is the second occurring in the Japanese mainland, although its actual distribution remains unknown.
</paragraph>
<caption id="DF2B6642006EFA535895F848FE7B6CCB" ID-DOI="http://doi.org/10.5281/zenodo.3954143" ID-Zenodo-Dep="3954143" httpUri="https://zenodo.org/record/3954143/files/figure.png" pageId="7" pageNumber="99" startId="7.[151,250,1947,1972]" targetBox="[151,1436,309,1921]" targetPageId="7">
<paragraph id="8BEB36CA006EFA535895F848FE7B6CCB" blockId="7.[151,1436,1947,2008]" pageId="7" pageNumber="99">
<emphasis id="B920EAD8006EFA535895F848FE846CA7" bold="true" box="[151,277,1947,1972]" pageId="7" pageNumber="99">FIGURE 5.</emphasis>
<taxonomicName id="4C544D49006EFA53591FF84FFE4E6CA7" authority="Komai &amp; Hirabayashi, 2020" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[285,479,1948,1972]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="99" phylum="Arthropoda" rank="species" species="kiiensis" status="sp. nov.">
<emphasis id="B920EAD8006EFA53591FF84FFE4E6CA7" box="[285,479,1948,1972]" italics="true" pageId="7" pageNumber="99">Naushonia kiiensis</emphasis>
</taxonomicName>
<emphasis id="B920EAD8006EFA5359E5F84FFDB06CA7" bold="true" box="[487,545,1948,1972]" pageId="7" pageNumber="99">
<taxonomicNameLabel id="A21357A3006EFA5359E5F84FFDB06CA7" box="[487,545,1948,1972]" pageId="7" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm), CBM-ZC 15914. Habitus, showing coloration in life. A, dorsal view; B, right lateral view.
</paragraph>
</caption>
<paragraph id="8BEB36CA0061FA5C58C5FF4AFCF66A50" blockId="8.[151,1436,153,359]" pageId="8" pageNumber="100">
<bibRefCitation id="EFC54B3B0061FA5C58C5FF4AFEF76BA0" author="Komai, T." box="[199,358,153,179]" pageId="8" pageNumber="100" pagination="15 - 26" refId="ref4450" refString="Komai, T. (2004) Rare mud shrimp genus Naushonia Kingsley (Decapoda: Thalassinidea: Laomediidae) from Japan: Description of a new species and new record of N. lactoalbida Berggren. Crustacean Research, 33, 15 - 26. https: // doi. org / 10.18353 / crustacea. 33.0 _ 15" type="journal article" year="2004">Komai (2004)</bibRefCitation>
suggested that plankotonic zoeas collected in Gokasho Bay, 
<collectingRegion id="4990F8280061FA5C5C18FF4AFB546BA0" box="[1050,1221,153,179]" country="Japan" name="Mie" pageId="8" pageNumber="100">Mie Prefecture</collectingRegion>
, central 
<collectingCountry id="F343765A0061FA5C5D27FF4AFAF96BA0" box="[1317,1384,153,179]" name="Japan" pageId="8" pageNumber="100">Japan</collectingCountry>
, and attributed to an unidentified 
<taxonomicName id="4C544D490061FA5C59D6FF6EFDDC6BC4" authorityName="Kingsley" authorityYear="1897" box="[468,589,189,215]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="100" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAD80061FA5C59D6FF6EFDDC6BC4" box="[468,589,189,215]" italics="true" pageId="8" pageNumber="100">Naushonia</emphasis>
</taxonomicName>
sp. by 
<bibRefCitation id="EFC54B3B0061FA5C5A9EFF6EFCD96BC4" author="Konishi, K." box="[668,840,189,215]" pageId="8" pageNumber="100" pagination="611 - 617" refId="ref4628" refString="Konishi, K. (2001) First record of larvae of the rare mud shrimp Naushonia Kingsley (Crustacea: Decapoda: Laomediidae) from Asian waters. Proceedings of the Biological Society of Washington, 114, 611 - 617." type="journal article" year="2001">Konishi (2001)</bibRefCitation>
, might actually represent 
<taxonomicName id="4C544D490061FA5C5C65FF6DFB7B6BC4" authorityName="Komai" authorityYear="2004" box="[1127,1258,190,215]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="100" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B920EAD80061FA5C5C65FF6DFB7B6BC4" box="[1127,1258,190,215]" italics="true" pageId="8" pageNumber="100">N. japonica</emphasis>
</taxonomicName>
, but the discovery of the present new species from closer locality may obscure the identity of the larvae of 
<bibRefCitation id="EFC54B3B0061FA5C5CA5FF32FAC46BE8" author="Konishi, K." box="[1191,1365,225,251]" pageId="8" pageNumber="100" pagination="611 - 617" refId="ref4628" refString="Konishi, K. (2001) First record of larvae of the rare mud shrimp Naushonia Kingsley (Crustacea: Decapoda: Laomediidae) from Asian waters. Proceedings of the Biological Society of Washington, 114, 611 - 617." type="journal article" year="2001">Konishi (2001)</bibRefCitation>
. DNA barcoding identification will be useful to determine the real specific identity of larvae, when sequence data based on adult specimens of certain specific identity will be accumulated.
</paragraph>
</subSubSection>
<subSubSection id="C34E65410061FA5C58C5FE9EFA116A74" box="[199,1408,333,359]" pageId="8" pageNumber="100" type="etymology">
<paragraph id="8BEB36CA0061FA5C58C5FE9EFA116A74" blockId="8.[151,1436,153,359]" box="[199,1408,333,359]" pageId="8" pageNumber="100">
<emphasis id="B920EAD80061FA5C58C5FE9EFEDF6A74" bold="true" box="[199,334,333,359]" pageId="8" pageNumber="100">Etymology.</emphasis>
The specific epithet “ 
<taxonomicName id="4C544D490061FA5C5A47FE9EFD086A74" authorityName="Komai &amp; Hirabayashi" authorityYear="2020" box="[581,665,333,359]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="100" phylum="Arthropoda" rank="species" species="kiiensis">
<emphasis id="B920EAD80061FA5C5A47FE9EFD086A74" box="[581,665,333,359]" italics="true" pageId="8" pageNumber="100">kiiensis</emphasis>
</taxonomicName>
” is derived from Kii District, where the 
<typeStatus id="54EF88680061FA5C5C62FE9EFB016A74" box="[1120,1168,333,359]" pageId="8" pageNumber="100">type</typeStatus>
locality is embraced.
</paragraph>
</subSubSection>
</treatment>
</document>