treatments-xml/data/03/FD/87/03FD87C3FFB2D50EDE8EE97CFA8CFE00.xml
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<document id="6282F740CFD46BF51EA93BB53FF64F7C" ID-CLB-Dataset="25317" ID-DOI="10.11646/zootaxa.4679.3.3" ID-GBIF-Dataset="41cb435c-1867-4d73-9f58-fdd326298e5e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471142" ID-ZooBank="5723773A-49BD-4CDB-99CD-B3A0AFC24F66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099201260" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2019" docId="03FD87C3FFB2D50EDE8EE97CFA8CFE00" docLanguage="en" docName="zootaxa.4679.3.3.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kenyacus ruwenzorii" docType="treatment" docVersion="8" lastPageNumber="472" masterDocId="FFC4FFBBFFB6D507DE19EF11FF9FFFAE" masterDocTitle="A review of the Afrotropical genus Kenyacus (Coleoptera: Carabidae: Harpalini) from the Rwenzori Mountains, with description of seven new species" masterLastPageNumber="498" masterPageNumber="463" pageNumber="467" updateTime="1698757568096" updateUser="ExternalLinkService">
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<mods:title id="F3C02EC442119AC5620451C7AF235CD4">A review of the Afrotropical genus Kenyacus (Coleoptera: Carabidae: Harpalini) from the Rwenzori Mountains, with description of seven new species</mods:title>
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<mods:namePart id="06108DB38851CE8CB73C59F5D9A942F0">Kataev, Boris M.</mods:namePart>
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<mods:date id="DE200AA309A1CC2C8A3675CA314CF8E1">2019</mods:date>
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<treatment id="03FD87C3FFB2D50EDE8EE97CFA8CFE00" ID-GBIF-Taxon="159628588" LSID="urn:lsid:plazi:treatment:03FD87C3FFB2D50EDE8EE97CFA8CFE00" httpUri="http://treatment.plazi.org/id/03FD87C3FFB2D50EDE8EE97CFA8CFE00" lastPageId="9" lastPageNumber="472" pageId="4" pageNumber="467">
<subSubSection id="C34E655EFFB2D503DE8EE97CFDC0F929" box="[151,607,1645,1671]" pageId="4" pageNumber="467" type="nomenclature">
<paragraph id="8BEB36D5FFB2D503DE8EE97CFDC0F929" blockId="4.[151,607,1645,1707]" box="[151,607,1645,1671]" pageId="4" pageNumber="467">
<heading id="D0A381B9FFB2D503DE8EE97CFDC0F929" bold="true" box="[151,607,1645,1671]" fontSize="11" level="1" pageId="4" pageNumber="467" reason="1">
<taxonomicName id="4C544D56FFB2D503DE8EE97CFDC0F929" ID-CoL="8MJ5X" authority="(Alluaud, 1917)" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[151,607,1645,1671]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB2D503DE8EE97CFDC0F929" bold="true" box="[151,607,1645,1671]" pageId="4" pageNumber="467">
<emphasis id="B920EAC7FFB2D503DE8EE97CFE09F929" bold="true" box="[151,406,1645,1671]" italics="true" pageId="4" pageNumber="467">Kenyacus ruwenzorii</emphasis>
(
<bibRefCitation id="EFC54B24FFB2D503DFBFE97CFDC8F929" author="Alluaud, C." box="[422,599,1645,1671]" pageId="4" pageNumber="467" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFB2D503DE8EE980FE2EF905" box="[151,433,1680,1707]" pageId="4" pageNumber="467" type="description">
<paragraph id="8BEB36D5FFB2D503DE8EE980FE2EF905" blockId="4.[151,607,1645,1707]" box="[151,433,1680,1707]" pageId="4" pageNumber="467">
(
<figureCitation id="136F2A50FFB2D503DE86E980FE9BF905" box="[159,260,1681,1707]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURES 3, 4" captionStartId-0="5.[151,264,1505,1530]" captionStartId-1="6.[151,264,1902,1927]" captionTargetBox-0="[152,1297,301,1482]" captionTargetBox-1="[152,1375,777,1870]" captionTargetId-0="figure@5.[151,1382,301,1485]" captionTargetId-1="figure@6.[151,1379,769,1882]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1, 2. Kenyacus ruwenzorii (Alluaud, 1917), holotype. 1, habitus; 2, labels. Scale bar = 1.0 mm." captionText-1="FIGURES 3, 4. Kenyacus, habitus (males). 3, K. ruwenzorii (Alluaud, 1917) (loc. U166); 4, K. gusarovi sp. n. (holotype). Scale bar = 1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3471144" figureDoi-1="http://doi.org/10.5281/zenodo.3471146" httpUri-0="https://zenodo.org/record/3471144/files/figure.png" httpUri-1="https://zenodo.org/record/3471146/files/figure.png" pageId="4" pageNumber="467">Figs 13</figureCitation>
,
<figureCitation id="136F2A50FFB2D503DF09E981FEEBF905" box="[272,372,1680,1707]" captionStart-0="FIGURES 513" captionStart-1="FIGURES 1421" captionStartId-0="7.[151,264,1619,1644]" captionStartId-1="8.[151,264,1512,1537]" captionTargetBox-0="[168,1411,184,1582]" captionTargetBox-1="[157,1415,739,1485]" captionTargetId-0="figure@7.[151,1426,174,1600]" captionTargetId-1="figure@8.[151,1423,733,1492]" captionTargetPageId-1="8" captionText-0="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." captionText-1="FIGURES 1421. Kenyacus, microsculpture (1417, male pronotal; 1821, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3471324" httpUri-0="https://zenodo.org/record/3471324/files/figure.png" httpUri-1="https://zenodo.org/record/3471148/files/figure.png" pageId="4" pageNumber="467">514, 18</figureCitation>
, 27a)
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFB2D503DE8EE9C7FDD4F943" box="[151,587,1748,1773]" pageId="4" pageNumber="467" type="reference_group">
<paragraph id="8BEB36D5FFB2D503DE8EE9C7FDD4F943" blockId="4.[151,587,1748,1773]" box="[151,587,1748,1773]" pageId="4" pageNumber="467">
<treatmentCitationGroup id="AB4411FBFFB2D503DE8EE9C7FDD4F943" box="[151,587,1748,1773]" pageId="4" pageNumber="467">
<taxonomicName id="4C544D56FFB2D503DE8EE9C7FDD8F943" ID-CoL="8LPH7" authority="Alluaud, 1917: 95" authorityName="Alluaud" authorityPageNumber="95" authorityYear="1917" box="[151,583,1748,1773]" class="Insecta" family="Carabidae" genus="Tropicoritus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB2D503DE8EE9C7FE17F943" box="[151,392,1750,1773]" italics="true" pageId="4" pageNumber="467">Tropicoritus ruwenzorii</emphasis>
<treatmentCitation id="0AF510C4FFB2D503DF94E9C5FDD8F943" author="Alluaud, C." box="[397,583,1748,1773]" page="95" pageId="4" pageNumber="467" year="1917">
<bibRefCitation id="EFC54B24FFB2D503DF94E9C5FDD8F943" author="Alluaud, C." box="[397,583,1748,1773]" pageId="4" pageNumber="467" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917: 95</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFB2D50FDE8EE80DFB58F953" lastPageId="8" lastPageNumber="471" pageId="4" pageNumber="467" type="materials_examined">
<paragraph id="8BEB36D5FFB2D503DE8EE80DFCBEF8D0" blockId="4.[151,1437,1819,2026]" pageId="4" pageNumber="467">
<emphasis id="B920EAC7FFB2D503DE8EE80DFE59F898" bold="true" box="[151,454,1819,1846]" pageId="4" pageNumber="467">Type material examined.</emphasis>
<materialsCitation id="3B3C3C88FFB2D503DFD7E80AFCBEF8D0" ID-GBIF-Occurrence="2423812432" collectionCode="MNHN" collectorName="Ch. Alluaud" determinerName="Alluaud" elevation="2650" pageId="4" pageNumber="467" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54EF8877FFB2D503DFD7E80AFDA2F89B" box="[462,573,1819,1845]" pageId="4" pageNumber="467" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D52FD5CFFB2D503DC50E80DFDFFF898" box="[585,608,1820,1846]" pageId="4" pageNumber="467" type="male"></specimenCount>
, labelled “Museum Paris, monts Rouwenzori, zone de forets, Makitawa [! sic] (
<quantity id="4CAC9B30FFB2D503DEF2E82EFED6F8F4" box="[235,329,1855,1882]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="4" pageNumber="467" unit="m" value="2650.0">
<elevation id="0079D1E6FFB2D503DEF2E82EFED6F8F4" box="[235,329,1855,1882]" metricMagnitude="3" metricUnit="m" metricValue="2.65" pageId="4" pageNumber="467" unit="m" value="2650.0">2650 m</elevation>
</quantity>
.),
<collectorName id="26A15303FFB2D503DF7AE82EFE72F8F7" box="[355,493,1855,1881]" pageId="4" pageNumber="467">Ch. Alluaud</collectorName>
1909”, “Type” [white, with red letters]”, “Type” [on red paper], “
<taxonomicName id="4C544D56FFB2D503DAFCE851FEFFF8D3" authority="Alluaud" authorityName="Alluaud" authorityYear="1917" class="Insecta" family="Carabidae" genus="Tropicoritus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB2D503DAFCE851FF65F8D3" italics="true" pageId="4" pageNumber="467">Tropicoritus ruwenzorii</emphasis>
Alluaud
</taxonomicName>
, type,
<determinerName id="7E37DA11FFB2D503DFB3E872FD96F8D3" box="[426,521,1891,1917]" pageId="4" pageNumber="467">Alluaud</determinerName>
, det.” (
<collectionCode id="ED45AE10FFB2D503DC42E875FD29F8D0" box="[603,694,1892,1918]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="467">MNHN</collectionCode>
) (
<figureCitation id="136F2A50FFB2D503DCD4E872FC8BF8D0" box="[717,788,1891,1918]" captionStart="FIGURES 1, 2" captionStartId="5.[151,264,1505,1530]" captionTargetBox="[152,1297,301,1482]" captionTargetId="figure@5.[151,1382,301,1485]" captionTargetPageId="5" captionText="FIGURES 1, 2. Kenyacus ruwenzorii (Alluaud, 1917), holotype. 1, habitus; 2, labels. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471144" httpUri="https://zenodo.org/record/3471144/files/figure.png" pageId="4" pageNumber="467">Fig. 2</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEB36D5FFB2D502DEDEE896FC7FFEB1" blockId="4.[151,1437,1819,2026]" lastBlockId="5.[151,1437,152,287]" lastPageId="5" lastPageNumber="468" pageId="4" pageNumber="467">
<emphasis id="B920EAC7FFB2D503DEDEE896FD0EF80C" bold="true" box="[199,657,1927,1954]" pageId="4" pageNumber="467">Additional material examined. Uganda</emphasis>
:
<materialsCitation id="3B3C3C88FFB2D503DC84E899FA02F847" ID-GBIF-Occurrence="2423812429" country="Uganda" county="Western Region" elevation="7" latitude="0.35059997" location="Nyabitaba Hut" longLatPrecision="1" longitude="29.968334" municipality="Rwenzori Mts. National Park" pageId="4" pageNumber="467" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Kasese">
<specimenCount id="9D52FD5CFFB2D503DC84E899FD43F80C" box="[669,732,1928,1954]" pageId="4" pageNumber="467" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D52FD5CFFB2D503DCFCE899FC92F80C" box="[741,781,1928,1954]" pageId="4" pageNumber="467" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F3437645FFB2D503DD3BE899FC3EF80C" box="[802,929,1928,1954]" name="Uganda" pageId="4" pageNumber="467">UGANDA</collectingCountry>
,
<collectingCounty id="628A4E59FFB2D503DDB0E899FBC2F80F" box="[937,1117,1927,1954]" pageId="4" pageNumber="467">Western Region</collectingCounty>
,
<collectingRegion id="4990F837FFB2D503DA7CE899FA8FF80F" box="[1125,1296,1927,1954]" country="Uganda" name="Kasese" pageId="4" pageNumber="467">Kasese District</collectingRegion>
,
<locationDeviation id="9A592FF7FFB2D503DB00E899FEB0F86B" location="Nyakalengija" pageId="4" pageNumber="467">
<quantity id="4CAC9B30FFB2D503DB00E899FAD0F80F" box="[1305,1359,1927,1954]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="467" unit="km" value="8.0">8 km</quantity>
NWW Nyakalengija
</locationDeviation>
,
<collectingMunicipality id="6B8FACAFFFB2D503DF22E8BAFD19F86B" box="[315,646,1963,1990]" pageId="4" pageNumber="467">Rwenzori Mts. National Park</collectingMunicipality>
, trail from
<location id="8E8B600EFFB2D503DD1AE8BAFC38F868" LSID="urn:lsid:plazi:treatment:03FD87C3FFB2D50EDE8EE97CFA8CFE00:8E8B600EFFB2D503DD1AE8BAFC38F868" box="[771,935,1963,1990]" country="Uganda" county="Western Region" latitude="0.35059997" longLatPrecision="1" longitude="29.968334" municipality="Rwenzori Mts. National Park" name="Nyabitaba Hut" pageId="4" pageNumber="467" stateProvince="Kasese">Nyabitaba Hut</location>
to
<location id="8E8B600EFFB2D503DDD2E8BAFBEEF86B" LSID="urn:lsid:plazi:treatment:03FD87C3FFB2D50EDE8EE97CFA8CFE00:8E8B600EFFB2D503DDD2E8BAFBEEF86B" box="[971,1137,1963,1989]" country="Uganda" county="Western Region" latitude="0.35059997" longLatPrecision="1" longitude="29.968334" municipality="Rwenzori Mts. National Park" name="Mahoma Lake" pageId="4" pageNumber="467" stateProvince="Kasese">Mahoma Lake</location>
,
<geoCoordinate id="EE605012FFB2D503DA65E8BAFA9FF86B" box="[1148,1280,1963,1989]" degrees="0" direction="north" minutes="21.036" orientation="latitude" pageId="4" pageNumber="467" precision="1" value="0.35059997">0°21.036N</geoCoordinate>
<geoCoordinate id="EE605012FFB2D503DB1EE8BAFA07F868" box="[1287,1432,1963,1990]" degrees="29" direction="east" minutes="58.100" orientation="longitude" pageId="4" pageNumber="467" precision="1" value="29.968334">29°58.100E</geoCoordinate>
, accuracy
<quantity id="4CAC9B30FFB2D503DF18E8C1FEB1F844" box="[257,302,2000,2026]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="467" unit="m" value="7.0">
<elevation id="0079D1E6FFB2D503DF18E8C1FEB1F844" box="[257,302,2000,2026]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="467" unit="m" value="7.0">7 m</elevation>
</quantity>
, extent
<quantity id="4CAC9B30FFB2D503DF9DE8C1FE20F844" box="[388,447,2000,2026]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="467" unit="m" value="10.0">
<elevation id="0079D1E6FFB2D503DF9DE8C1FE20F844" box="[388,447,2000,2026]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="467" unit="m" value="10.0">10 m</elevation>
</quantity>
, h =
<quantity id="4CAC9B30FFB2D503DFECE8DEFDD3F844" box="[501,588,1999,2026]" metricMagnitude="3" metricUnit="m" metricValue="2.957" pageId="4" pageNumber="467" unit="m" value="2957.0">
<elevation id="0079D1E6FFB2D503DFECE8DEFDD3F844" box="[501,588,1999,2026]" metricMagnitude="3" metricUnit="m" metricValue="2.957" pageId="4" pageNumber="467" unit="m" value="2957.0">2957 m</elevation>
</quantity>
,
<taxonomicName id="4C544D56FFB2D503DC4EE8DEFD41F847" box="[599,734,1999,2025]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="467" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFB2D503DC4EE8DEFD41F847" box="[599,734,1999,2025]" italics="true" pageId="4" pageNumber="467">Podocarpus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D56FFB2D503DCF5E8C1FCB5F847" box="[748,810,2000,2025]" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFB2D503DCF5E8C1FCB5F847" box="[748,810,2000,2025]" italics="true" pageId="4" pageNumber="467">Erica</emphasis>
</taxonomicName>
, bamboo, on the ridge, sifting leaf litter [3601] [Garmin
</materialsCitation>
60CSx; WGS84] V.I. Gusarov
<date id="FFEA1015FFB3D502DFF7EF88FDEAFF1D" box="[494,629,153,179]" pageId="5" pageNumber="468" value="2008-08-18">18.viii.2008</date>
U166” (
<collectionCode id="ED45AE10FFB3D502DCCDEF88FCB2FF1D" box="[724,813,153,179]" pageId="5" pageNumber="468">ZMUN</collectionCode>
);
<specimenCount id="9D52FD5CFFB3D502DD59EF89FC1FFF1D" box="[832,896,152,179]" pageId="5" pageNumber="468" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="9D52FD5CFFB3D502DD92EF88FC2AFF1D" box="[907,949,153,179]" pageId="5" pageNumber="468" type="female">2 ♀</specimenCount>
(
<specimenCount id="9D52FD5CFFB3D502DDDEEF88FC77FF1D" box="[967,1000,153,179]" pageId="5" pageNumber="468" type="female">1♀</specimenCount>
teneral), “
<collectingCountry id="F3437645FFB3D502DA43EF88FB46FF1D" box="[1114,1241,153,179]" name="Uganda" pageId="5" pageNumber="468">UGANDA</collectingCountry>
, Western Region,
<collectingRegion id="4990F837FFB3D502DE8EEFACFEDAFF79" box="[151,325,189,215]" country="Uganda" name="Kasese" pageId="5" pageNumber="468">Kasese District</collectingRegion>
,
<quantity id="4CAC9B30FFB3D502DF48EFACFE13FF79" box="[337,396,189,215]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="468" unit="km" value="8.0">8 km</quantity>
W Nyakalengija, Rwenzori Mts. National Park, trail from Nyabitaba Hut to Mahoma Lake,
<geoCoordinate id="EE605012FFB3D502DE8EEFF0FE84FF55" box="[151,283,225,251]" degrees="0" direction="north" minutes="20.748" orientation="latitude" pageId="5" pageNumber="468" precision="1" value="0.34579998">0°20.748N</geoCoordinate>
<geoCoordinate id="EE605012FFB3D502DF3AEFF1FE2BFF55" box="[291,436,224,251]" degrees="29" direction="east" minutes="58.085" orientation="longitude" pageId="5" pageNumber="468" precision="1" value="29.968082">29°58.085E</geoCoordinate>
, accuracy
<quantity id="4CAC9B30FFB3D502DC34EFF0FDC4FF55" box="[557,603,225,251]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="468" unit="m" value="6.0">6 m</quantity>
, extent
<quantity id="4CAC9B30FFB3D502DCACEFF1FD6EFF55" box="[693,753,224,251]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="468" unit="m" value="15.0">15 m</quantity>
, h =
<quantity id="4CAC9B30FFB3D502DD34EFF0FC1AFF55" box="[813,901,225,251]" metricMagnitude="3" metricUnit="m" metricValue="2.988" pageId="5" pageNumber="468" unit="m" value="2988.0">2988 m</quantity>
, bamboo,
<taxonomicName id="4C544D56FFB3D502DDE5EFF0FB1CFF55" box="[1020,1155,225,251]" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="468" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFB3D502DDE5EFF0FB1CFF55" box="[1020,1155,225,251]" italics="true" pageId="5" pageNumber="468">Podocarpus</emphasis>
</taxonomicName>
, sifting leaf litter [3600] [Garmin 60CSx; WGS84] V.I. Gusarov
<date id="FFEA1015FFB3D502DC4FEE14FD41FEB1" box="[598,734,261,287]" pageId="5" pageNumber="468" value="2008-08-18">18.viii.2008</date>
U165” (
<collectionCode id="ED45AE10FFB3D502DD27EE14FC0AFEB1" box="[830,917,261,287]" pageId="5" pageNumber="468">ZMUN</collectionCode>
,
<collectionCode id="ED45AE10FFB3D502DDB9EE14FC4CFEB1" box="[928,979,261,287]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="5" pageNumber="468" type="Museum">ZIN</collectionCode>
).
</paragraph>
<caption id="DF2B665DFFB3D502DE8EEAF0FB4FFA57" ID-DOI="http://doi.org/10.5281/zenodo.3471144" ID-Zenodo-Dep="3471144" box="[151,1232,1505,1530]" httpUri="https://zenodo.org/record/3471144/files/figure.png" pageId="5" pageNumber="468" startId="5.[151,264,1505,1530]" targetBox="[152,1297,301,1482]" targetPageId="5">
<paragraph id="8BEB36D5FFB3D502DE8EEAF0FB4FFA57" blockId="5.[151,1232,1505,1530]" box="[151,1232,1505,1530]" pageId="5" pageNumber="468">
<emphasis id="B920EAC7FFB3D502DE8EEAF0FEA5FA54" bold="true" box="[151,314,1505,1530]" pageId="5" pageNumber="468">FIGURES 1, 2.</emphasis>
<taxonomicName id="4C544D56FFB3D502DF58EAF3FD5DFA57" authority="(Alluaud, 1917)" authorityName="Alluaud" authorityYear="1917" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[321,706,1505,1530]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB3D502DF58EAF3FD87FA57" box="[321,536,1506,1529]" italics="true" pageId="5" pageNumber="468">Kenyacus ruwenzorii</emphasis>
(
<bibRefCitation id="EFC54B24FFB3D502DC3CEAF0FD25FA57" author="Alluaud, C." box="[549,698,1505,1530]" pageId="5" pageNumber="468" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
)
</taxonomicName>
, holotype. 1, habitus; 2, labels. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BEB36D5FFB3D502DEDEE936FD9EF9C8" blockId="5.[151,1437,1575,2034]" pageId="5" pageNumber="468">
<emphasis id="B920EAC7FFB3D502DEDEE936FEE8F9EC" bold="true" box="[199,375,1575,1602]" pageId="5" pageNumber="468">Re-description</emphasis>
(
<specimenCount id="9D52FD5CFFB3D502DF9FE936FE43F9EC" box="[390,476,1575,1602]" pageId="5" pageNumber="468" type="male">5 males</specimenCount>
and
<specimenCount id="9D52FD5CFFB3D502DC08E939FDE2F9EC" box="[529,637,1576,1602]" pageId="5" pageNumber="468" type="female">3 females</specimenCount>
measured). Habitus as in
<figureCitation id="136F2A50FFB3D502DD85E939FC7EF9EC" box="[924,993,1576,1602]" captionStart="FIGURES 1, 2" captionStartId="5.[151,264,1505,1530]" captionTargetBox="[152,1297,301,1482]" captionTargetId="figure@5.[151,1382,301,1485]" captionTargetPageId="5" captionText="FIGURES 1, 2. Kenyacus ruwenzorii (Alluaud, 1917), holotype. 1, habitus; 2, labels. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471144" httpUri="https://zenodo.org/record/3471144/files/figure.png" pageId="5" pageNumber="468">Figs 1</figureCitation>
and
<figureCitation id="136F2A50FFB3D502DA0FE939FBB9F9EC" box="[1046,1062,1576,1602]" captionStart="FIGURES 3, 4" captionStartId="6.[151,264,1902,1927]" captionTargetBox="[152,1375,777,1870]" captionTargetId="figure@6.[151,1379,769,1882]" captionTargetPageId="6" captionText="FIGURES 3, 4. Kenyacus, habitus (males). 3, K. ruwenzorii (Alluaud, 1917) (loc. U166); 4, K. gusarovi sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471146" httpUri="https://zenodo.org/record/3471146/files/figure.png" pageId="5" pageNumber="468">3</figureCitation>
. Large for the genus: body length 5.36.0 mm, width
<quantity id="4CAC9B30FFB3D502DF6AE95DFE62F9C8" box="[371,509,1612,1638]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="5" pageNumber="468" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 mm</quantity>
.
</paragraph>
<paragraph id="8BEB36D5FFB3D502DEDEE961FC10F97C" blockId="5.[151,1437,1575,2034]" pageId="5" pageNumber="468">Colour: Body brown to blackish brown, dorsal surface somewhat dull, darker than ventral surface; base of mandibles, labrum, margins of pronotum and elytra, as well as elytral suture usually paler; palpi, antennae and legs light brown or brownish yellow; antennae infuscate from antennomere 2.</paragraph>
<paragraph id="8BEB36D5FFB3D501DEDEE9CDFF61FEB1" blockId="5.[151,1437,1575,2034]" lastBlockId="6.[151,1437,153,755]" lastPageId="6" lastPageNumber="469" pageId="5" pageNumber="468">
Head: Comparatively large, in males HWmax/PWmax 0.750.78 (
<emphasis id="B920EAC7FFB3D502DDC0E9CDFC72F95B" box="[985,1005,1756,1781]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 0.77), HWmin/PWmax 0.640.67 (
<emphasis id="B920EAC7FFB3D502DEB8E811FF2AF8B7" box="[161,181,1792,1817]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 0.66); in females these indices respectively 0.730.80 (
<emphasis id="B920EAC7FFB3D502DD7FE811FCE5F8B7" box="[870,890,1792,1817]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 0.77), and 0.630.70 (
<emphasis id="B920EAC7FFB3D502DABDE811FB27F8B7" box="[1188,1208,1792,1817]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 0.67). Eyes with large facets, only slightly convex, separated from buccal fissure at distance of width of antennomere 1, HWmax/HWmin 1.141.20 (
<emphasis id="B920EAC7FFB3D502DFA4E859FE4EF8CF" box="[445,465,1864,1889]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 1.17) in males and 1.141.16 (
<emphasis id="B920EAC7FFB3D502DD42E859FCF0F8CF" box="[859,879,1864,1889]" italics="true" pageId="5" pageNumber="468">m</emphasis>
= 1.15) in females. Tempora very short and flat. Frons and vertex convex. Fronto-clypeal suture distinct, slightly impressed. Supraorbital setigerous pore located at level of posterior margin of eye, removed from supraorbital furrow at a distance of length of antennomere 2. Labrum almost straight anteriorly. Clypeus with prominent apical angles and with slightly convex apical margin between them. Mentum and submentum (
<figureCitation id="136F2A50FFB3D502DC1BE8C9FDCCF85C" box="[514,595,2008,2034]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="5" pageNumber="468">Fig. 10</figureCitation>
) completely fused, submentum with two setae on each side. Mandibles me- dium-sized; left mandible evenly bent in apical portion, with acute apex. Dorsal microsculpture in both sexes very distinct throughout, consisting of large isodiametric meshes. Antennae in male extending to basal edge of pronotum, with antennomeres 79 only slightly longer than wide; in female antennae noticeably shorter, with shorter antennomeres.
</paragraph>
<paragraph id="8BEB36D5FFB0D501DEDEEE38FC3CFD5D" blockId="6.[151,1437,153,755]" pageId="6" pageNumber="469">
Pronotum: Slightly transverse, PWmax/PL 1.251.33 (
<emphasis id="B920EAC7FFB0D501DD2DEE3BFCD7FEED" box="[820,840,298,323]" italics="true" pageId="6" pageNumber="469">m</emphasis>
= 1.29), widest at the end of anterior third, strongly narrowed posteriorly, PWmax/PWmin 1.281.33 (
<emphasis id="B920EAC7FFB0D501DCD3EE5FFD41FEC9" box="[714,734,334,359]" italics="true" pageId="6" pageNumber="469">m</emphasis>
= 1.33); lateral seta inserted slightly before widest point. Sides rounded anteriorly, almost straightly converging from widest point to basal angles, without distinct border. Apical margin almost straight, with slightly projecting, acute or blunted, apical angles. Basal margin slightly narrower than apical margin and noticeably narrower than elytral base, deeply emarginate, almost straight medially and oblique laterally; both apical and basal margins not bordered. Basal angles large, almost right, at most only slightly blunted at tip, markedly projecting posteriorly and covering elytral base. Disc moderately convex, strongly sloped to apical angles, depressed basally and flattened at basal angles, forming latero-basal impressions distinctly delimited from other parts of disc. Lateral depressions beginning from apical angles, narrow in apical third, evenly widened after lateral seta, reaching basal margin. Basal foveae somewhat deep, reaching basal pronotal edge, usually fused basally with lateral depressions. Surface of pronotum impunctate. Anterior transverse depression distinct and long, usually deepened. Median line thin, superficial, usually reaching apical and basal margins of pronotum. Microsculpture distinct throughout, consisting of large isodiametric meshes (
<figureCitation id="136F2A50FFB0D501DD5BEDC8FC0AFD5D" box="[834,917,729,755]" captionStart="FIGURES 1421" captionStartId="8.[151,264,1512,1537]" captionTargetBox="[157,1415,739,1485]" captionTargetId="figure@8.[151,1423,733,1492]" captionTargetPageId="8" captionText="FIGURES 1421. Kenyacus, microsculpture (1417, male pronotal; 1821, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="6" pageNumber="469">Fig. 14</figureCitation>
).
</paragraph>
<caption id="DF2B665DFFB0D501DE8EE87FFEFBF804" ID-DOI="http://doi.org/10.5281/zenodo.3471146" ID-Zenodo-Dep="3471146" httpUri="https://zenodo.org/record/3471146/files/figure.png" pageId="6" pageNumber="469" startId="6.[151,264,1902,1927]" targetBox="[152,1375,777,1870]" targetPageId="6">
<paragraph id="8BEB36D5FFB0D501DE8EE87FFEFBF804" blockId="6.[151,1437,1902,1963]" pageId="6" pageNumber="469">
<emphasis id="B920EAC7FFB0D501DE8EE87FFEA0F829" bold="true" box="[151,319,1902,1927]" pageId="6" pageNumber="469">FIGURES 3, 4.</emphasis>
<taxonomicName id="4C544D56FFB0D501DF51E861FE35F829" authorityName="Alluaud" authorityYear="1917" box="[328,426,1904,1927]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFB0D501DF51E861FE35F829" box="[328,426,1904,1927]" italics="true" pageId="6" pageNumber="469">Kenyacus</emphasis>
</taxonomicName>
, habitus (males). 3,
<taxonomicName id="4C544D56FFB0D501DC9BE861FC23F829" authority="(Alluaud, 1917)" authorityName="Alluaud" authorityYear="1917" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[642,956,1902,1927]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB0D501DC9BE861FC8FF829" box="[642,784,1904,1927]" italics="true" pageId="6" pageNumber="469">K. ruwenzorii</emphasis>
(
<bibRefCitation id="EFC54B24FFB0D501DD06E87FFC2AF829" author="Alluaud, C." box="[799,949,1902,1927]" pageId="6" pageNumber="469" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
)
</taxonomicName>
(loc. U166); 4,
<taxonomicName id="4C544D56FFB0D501DA71E861FB7FF829" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1128,1248,1904,1927]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="gusarovi" status="sp. nov.">
<emphasis id="B920EAC7FFB0D501DA71E861FB7FF829" box="[1128,1248,1904,1927]" italics="true" pageId="6" pageNumber="469">K. gusarovi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFB0D501DAF1E87EFABBF829" bold="true" box="[1256,1316,1903,1927]" pageId="6" pageNumber="469">
<taxonomicNameLabel id="A21357BCFFB0D501DAF1E87EFABBF829" box="[1256,1316,1903,1927]" pageId="6" pageNumber="469" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype). Scale bar = 1.0 mm.
</paragraph>
</caption>
<caption id="DF2B665DFFB1D500DE8EE942FC14F91A" ID-DOI="http://doi.org/10.5281/zenodo.3471324" ID-Zenodo-Dep="3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="7" pageNumber="470" startId="7.[151,264,1619,1644]" targetBox="[168,1411,184,1582]" targetPageId="7">
<paragraph id="8BEB36D5FFB1D500DE8EE942FC14F91A" blockId="7.[151,1437,1619,1716]" pageId="7" pageNumber="470">
<emphasis id="B920EAC7FFB1D500DE8EE942FEDEF9C2" bold="true" box="[151,321,1619,1644]" pageId="7" pageNumber="470">FIGURES 513</emphasis>
.
<taxonomicName id="4C544D56FFB1D500DF57E944FD52F9C2" authority="(Alluaud, 1917)" authorityName="Alluaud" authorityYear="1917" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[334,717,1619,1644]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFB1D500DF57E944FDBAF9C2" box="[334,549,1621,1644]" italics="true" pageId="7" pageNumber="470">Kenyacus ruwenzorii</emphasis>
(
<bibRefCitation id="EFC54B24FFB1D500DC2BE942FD59F9C2" author="Alluaud, C." box="[562,710,1619,1644]" pageId="7" pageNumber="470" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
)
</taxonomicName>
(5, 6, 9,
<typeStatus id="54EF8877FFB1D500DD30E942FC1AF9C2" box="[809,901,1619,1644]" pageId="7" pageNumber="470" type="holotype">holotype</typeStatus>
; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar =
<quantity id="4CAC9B30FFB1D500DD2FE98AFC18F91A" box="[822,903,1691,1716]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="470" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<paragraph id="8BEB36D5FFB1D50FDEDEE817FD5FFEED" blockId="7.[151,1437,1798,2041]" lastBlockId="8.[151,1437,153,719]" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470">
Elytra: Oval, widest at middle, in males EL/EW 1.401.45 (
<emphasis id="B920EAC7FFB1D500DD74E816FC1EF88E" box="[877,897,1799,1824]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 1.42), EL/PL 2.362.45 (
<emphasis id="B920EAC7FFB1D500DAA2E816FB50F88E" box="[1211,1231,1799,1824]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 2.39), EW/PWmax 1.281.34 (
<emphasis id="B920EAC7FFB1D500DF48E83AFEFAF8EA" box="[337,357,1835,1860]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 1.31); in females these indices respectively 1.431.46 (
<emphasis id="B920EAC7FFB1D500DDEEE83AFB94F8EA" box="[1015,1035,1835,1860]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 1.44), 2.362.46 (
<emphasis id="B920EAC7FFB1D500DAEFE83AFA95F8EA" box="[1270,1290,1835,1860]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 2.40), and 1.261.32 (
<emphasis id="B920EAC7FFB1D500DF00E85EFEB2F8C6" box="[281,301,1871,1896]" italics="true" pageId="7" pageNumber="470">m</emphasis>
= 1.28). Humeri prominent, angularly rounded at tip. Preapical sinuation absent, but sides in apical fifth rather sharply curved and then almost straightly converging to apex (elytra obliquely truncate apically). Sutural angle in both sexes acute and somewhat sharp at tip. Basal edge markedly sinuate laterally, arcuately curving inside humerus up to lateral margin. Lateral gutter distinct, slightly widened at humerus and prolonged on elytral base to stria 5. Elytral disc convex in lateral and caudal views, depressed basally on each side between humerus and scutellum. Striae thin, impunctate, faintly crenulate, slightly impressed along entire length, their bases removed from basal elytral border. Parascutellar setigerous pore present, large. Parascutellar striole present, with apex usually connected with stria 1. Intervals slightly convex, slightly narrowed posteriorly; interval 2 almost parallel-sided or slightly widened at apex; interval 3 without discal pore. Microsculpture very distinct throughout, almost granulate, consisting of large isodiametric meshes (
<figureCitation id="136F2A50FFBED50FDC79EE38FD2CFEED" box="[608,691,297,323]" captionStart="FIGURES 1421" captionStartId="8.[151,264,1512,1537]" captionTargetBox="[157,1415,739,1485]" captionTargetId="figure@8.[151,1423,733,1492]" captionTargetPageId="8" captionText="FIGURES 1421. Kenyacus, microsculpture (1417, male pronotal; 1821, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="8" pageNumber="471">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BEB36D5FFBED50FDEDEEE5CFB92FE7D" blockId="8.[151,1437,153,719]" pageId="8" pageNumber="471">
Ventral surface of thorax and abdomen: Prosternum with 15 short or moderately long erect setae medio-apically. Metepisternum slightly wider than long, narrowed posteriorly. Abdominal sternites glabrous, only last visible (
<collectionCode id="ED45AE10FFBED50FDE86EE84FF56FE01" box="[159,201,405,431]" pageId="8" pageNumber="471">VII</collectionCode>
) sternite sparsely pubescent apically in some specimens; its apical margin in both sexes with two pairs of setae, subtruncate at tip, occasionally with very shallow, indistinct incision at middle.
</paragraph>
<paragraph id="8BEB36D5FFBED50FDEDEEECCFAB9FDCD" blockId="8.[151,1437,153,719]" pageId="8" pageNumber="471">Legs: Metacoxae without any additional setae. Tarsi glabrous dorsally. Tarsomere 5 without setae ventrally. Meso- and metatarsomeres 1 about as long as meso- and metatarsomeres 2+3, respectively. Pro- and mesotarsomeres 24 of male slightly dilated (mesotarsi very faintly) and with biseriately arranged long adhesive scales ventrally; mesotarsomere 1 lacking adhesive vestiture ventrally; mesotarsomere 4 rather deeply emarginate.</paragraph>
<paragraph id="8BEB36D5FFBED50FDEDEED7CFCFCFD61" blockId="8.[151,1437,153,719]" pageId="8" pageNumber="471">
Male genitalia: Median lobe (
<figureCitation id="136F2A50FFBED50FDC00ED7CFDE1FD29" box="[537,638,620,647]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="8" pageNumber="471">Figs 58</figureCitation>
) stout, arcuate in lateral view, with large basal bulb. Terminal lamella short, slightly curved dorsally, without apical capitulum; in dorso-caudal view wide, triangular, narrowly rounded at apex (
<figureCitation id="136F2A50FFBED50FDEEFEDA4FEF7FD61" box="[246,360,693,719]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="8" pageNumber="471">Figs 9, 11</figureCitation>
). Internal sac without any sclerotic elements.
</paragraph>
<caption id="DF2B665DFFBED50FDE8EEAF9FB4CF98A" ID-DOI="http://doi.org/10.5281/zenodo.3471148" ID-Zenodo-Dep="3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="8" pageNumber="471" startId="8.[151,264,1512,1537]" targetBox="[157,1415,739,1485]" targetPageId="8">
<paragraph id="8BEB36D5FFBED50FDE8EEAF9FB4CF98A" blockId="8.[151,1436,1512,1573]" pageId="8" pageNumber="471">
<emphasis id="B920EAC7FFBED50FDE8EEAF9FEC8F9AF" bold="true" box="[151,343,1512,1537]" pageId="8" pageNumber="471">FIGURES 1421.</emphasis>
<taxonomicName id="4C544D56FFBED50FDF78EAF8FE5CF9AE" authorityName="Alluaud" authorityYear="1917" box="[353,451,1513,1536]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFBED50FDF78EAF8FE5CF9AE" box="[353,451,1513,1536]" italics="true" pageId="8" pageNumber="471">Kenyacus</emphasis>
</taxonomicName>
, microsculpture (1417, male pronotal; 1821, male elytral). 14, 18,
<taxonomicName id="4C544D56FFBED50FDABAEAF8FF49F98A" authority="(Alluaud, 1917)" authorityName="Alluaud" authorityYear="1917" baseAuthorityName="Alluaud" baseAuthorityYear="1917" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFBED50FDABAEAF8FAADF9AE" box="[1187,1330,1513,1536]" italics="true" pageId="8" pageNumber="471">K. ruwenzorii</emphasis>
(
<bibRefCitation id="EFC54B24FFBED50FDB5AEAF9FF52F98A" author="Alluaud, C." pageId="8" pageNumber="471" pagination="73 - 116" refId="ref24649" refString="Alluaud, C. (1917) Les carabiques de la faune alpine des hautes montagnes de l'Afrique orientale. Annales de la Societe Entomologique de France, 86, 73 - 116." type="journal article" year="1917">Alluaud, 1917</bibRefCitation>
)
</taxonomicName>
; 15, 19,
<taxonomicName id="4C544D56FFBED50FDF35E91CFE3EF98A" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[300,417,1549,1572]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="gusarovi" status="sp. nov.">
<emphasis id="B920EAC7FFBED50FDF35E91CFE3EF98A" box="[300,417,1549,1572]" italics="true" pageId="8" pageNumber="471">K. gusarovi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFBED50FDFBEE91DFE7FF98A" bold="true" box="[423,480,1548,1572]" pageId="8" pageNumber="471">
<taxonomicNameLabel id="A21357BCFFBED50FDFBEE91DFE7FF98A" box="[423,480,1548,1572]" pageId="8" pageNumber="471" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 16, 20,
<taxonomicName id="4C544D56FFBED50FDC23E91CFD03F98A" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[570,668,1549,1572]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B920EAC7FFBED50FDC23E91CFD03F98A" box="[570,668,1549,1572]" italics="true" pageId="8" pageNumber="471">K. parvus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFBED50FDCBAE91DFD43F98A" bold="true" box="[675,732,1548,1572]" pageId="8" pageNumber="471">
<taxonomicNameLabel id="A21357BCFFBED50FDCBAE91DFD43F98A" box="[675,732,1548,1572]" pageId="8" pageNumber="471" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; 17, 21,
<taxonomicName id="4C544D56FFBED50FDD2CE91CFC5EF98A" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[821,961,1549,1572]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="angustatus" status="sp. nov.">
<emphasis id="B920EAC7FFBED50FDD2CE91CFC5EF98A" box="[821,961,1549,1572]" italics="true" pageId="8" pageNumber="471">K. angustatus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFBED50FDDDEE91DFB9FF98A" bold="true" box="[967,1024,1548,1572]" pageId="8" pageNumber="471">
<taxonomicNameLabel id="A21357BCFFBED50FDDDEE91DFB9FF98A" box="[967,1024,1548,1572]" pageId="8" pageNumber="471" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BEB36D5FFBED50FDEDEE966FB58F953" blockId="8.[151,1437,1655,2041]" pageId="8" pageNumber="471">
Female genitalia: Very small (
<figureCitation id="136F2A50FFBED50FDC38E966FD39F93F" box="[545,678,1655,1681]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="8" pageNumber="471">Figs 12, 13</figureCitation>
); laterotergite (hemisternite) somewhat symmetrical, membranous apically, without setae and spines. Basal stylomere elongate, setae and spines not recognized, probably absent. Apical stylomere moderately curved, slightly shorter than basal, with short thick seta at ventral edge of outer margin, in ventral view with somewhat wide base; in lateral view (
<figureCitation id="136F2A50FFBED50FDD11E9F2FCC5F953" box="[776,858,1763,1789]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="8" pageNumber="471">Fig. 12</figureCitation>
) evenly narrowed to acute apex.
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFBED50EDEDEE817FE74FEB1" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471" type="distribution">
<paragraph id="8BEB36D5FFBED50FDEDEE817FABAF823" blockId="8.[151,1437,1655,2041]" pageId="8" pageNumber="471">
<emphasis id="B920EAC7FFBED50FDEDEE817FEC9F88F" bold="true" box="[199,342,1798,1825]" pageId="8" pageNumber="471">Distribution</emphasis>
(
<figureCitation id="136F2A50FFBED50FDF7FE816FE59F88F" box="[358,454,1799,1825]" captionStart="FIGURE 27" captionStartId="11.[151,250,1783,1808]" captionTargetBox="[151,1436,515,1761]" captionTargetId="figure@11.[151,1436,515,1761]" captionTargetPageId="11" captionText="FIGURE 27. Kenyacus, distribution. a, K. ruwenzorii (Alluaud, 1917); b, K. gusarovi sp. n." figureDoi="http://doi.org/10.5281/zenodo.3471152" httpUri="https://zenodo.org/record/3471152/files/figure.png" pageId="8" pageNumber="471">Fig. 27a</figureCitation>
). Originally described from one male collected in the Mubuku (Mobuko) Valley slightly above Nakitawa till
<quantity id="4CAC9B30FFBED50FDF6BE83AFE5AF8EB" box="[370,453,1835,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="471" unit="m" value="2800.0">2800 m</quantity>
(
<quantity id="4CAC9B30FFBED50FDFCBE83BFDB6F8EB" box="[466,553,1834,1861]" metricMagnitude="3" metricUnit="m" metricValue="2.659" pageId="8" pageNumber="471" unit="m" value="2659.0">2659 m</quantity>
, according to the label data) in the south-eastern part of the Rwenzori Mountains. Additional known specimens were collected in the same area on the trail from Nyabitaba Hut to the Mahoma Lake at altitudes of
<quantity id="4CAC9B30FFBED50FDF01E863FE2CF823" box="[280,435,1906,1933]" metricMagnitude="3" metricUnit="m" metricValue="2.9725" metricValueMax="2.988" metricValueMin="2.957" pageId="8" pageNumber="471" unit="m" value="2972.5" valueMax="2988.0" valueMin="2957.0">29572988 m</quantity>
. This species occurs sympatrically with
<taxonomicName id="4C544D56FFBED50FDD73E862FB9EF822" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[874,1025,1907,1932]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="angustatus" status="sp. nov.">
<emphasis id="B920EAC7FFBED50FDD73E862FB9EF822" box="[874,1025,1907,1932]" italics="true" pageId="8" pageNumber="471">K. angustatus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFBED50FDA1EE862FBDAF823" bold="true" box="[1031,1093,1907,1933]" pageId="8" pageNumber="471">
<taxonomicNameLabel id="A21357BCFFBED50FDA1EE862FBDAF823" box="[1031,1093,1907,1933]" pageId="8" pageNumber="471" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C544D56FFBED50FDA63E862FB7EF823" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1146,1249,1907,1933]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="471" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B920EAC7FFBED50FDA63E862FB7EF823" box="[1146,1249,1907,1933]" italics="true" pageId="8" pageNumber="471">K. similis</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFBED50FDAF1E862FABAF823" bold="true" box="[1256,1317,1907,1933]" pageId="8" pageNumber="471">
<taxonomicNameLabel id="A21357BCFFBED50FDAF1E862FABAF823" box="[1256,1317,1907,1933]" pageId="8" pageNumber="471" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEB36D5FFBED50FDEDEE887FD94F87B" blockId="8.[151,1437,1655,2041]" pageId="8" pageNumber="471">
<emphasis id="B920EAC7FFBED50FDEDEE887FEDCF81F" bold="true" box="[199,323,1942,1969]" pageId="8" pageNumber="471">Bionomics</emphasis>
. According to the label data, specimens were collected by sifting leaf litter in forest with
<taxonomicName id="4C544D56FFBED50FDB2AE886FF21F87A" class="Pinopsida" family="Podocarpaceae" genus="Podocarpus" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="471" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFBED50FDB2AE886FF21F87A" italics="true" pageId="8" pageNumber="471">Podocarpus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D56FFBED50FDED5E8AAFE95F87A" box="[204,266,1979,2004]" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="471" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFBED50FDED5E8AAFE95F87A" box="[204,266,1979,2004]" italics="true" pageId="8" pageNumber="471">Erica</emphasis>
</taxonomicName>
and bamboo (
<figureCitation id="136F2A50FFBED50FDFB3E8AAFE61F87B" box="[426,510,1979,2005]" captionStart="FIGURE 91" captionStartId="32.[151,250,969,994]" captionTargetBox="[176,1411,174,950]" captionTargetId="figure@32.[176,1411,174,950]" captionTargetPageId="32" captionText="FIGURE 91. Mahoma lake (ca. 2960 m) near the type locality of K. similis sp. n. Patches of Erica, Hagenia, Podocarpus and bamboo forest, separated by large gaps with bushes and herbaceous vegetation. Habitats of K. ruwenzorii, K. angustatus sp. n. and K. similis sp. n. Photo: Vladimir Gusarov." figureDoi="http://doi.org/10.5281/zenodo.3471174" httpUri="https://zenodo.org/record/3471174/files/figure.png" pageId="8" pageNumber="471">Fig. 91</figureCitation>
).
</paragraph>
<paragraph id="8BEB36D5FFBED50EDEDEE8CFFE74FEB1" blockId="8.[151,1437,1655,2041]" lastBlockId="9.[151,1436,152,431]" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471">
<emphasis id="B920EAC7FFBED50FDEDEE8CFFEC4F857" bold="true" box="[199,347,2014,2041]" pageId="8" pageNumber="471">Comparison</emphasis>
. Distinctly differs from other congeners in larger size (body length 5.36.0 mm) and in pronotum with basal angles large, almost right, at most only slightly blunted at tip, and markedly projecting posteriorly above elytral base (
<figureCitation id="136F2A50FFBFD50EDF3FEFADFEF2FF79" box="[294,365,188,215]" captionStart="FIGURES 1, 2" captionStartId="5.[151,264,1505,1530]" captionTargetBox="[152,1297,301,1482]" captionTargetId="figure@5.[151,1382,301,1485]" captionTargetPageId="5" captionText="FIGURES 1, 2. Kenyacus ruwenzorii (Alluaud, 1917), holotype. 1, habitus; 2, labels. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471144" httpUri="https://zenodo.org/record/3471144/files/figure.png" pageId="9" pageNumber="472">Figs 1</figureCitation>
,
<figureCitation id="136F2A50FFBFD50EDF63EFACFE15FF79" box="[378,394,189,215]" captionStart="FIGURES 3, 4" captionStartId="6.[151,264,1902,1927]" captionTargetBox="[152,1375,777,1870]" captionTargetId="figure@6.[151,1379,769,1882]" captionTargetPageId="6" captionText="FIGURES 3, 4. Kenyacus, habitus (males). 3, K. ruwenzorii (Alluaud, 1917) (loc. U166); 4, K. gusarovi sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471146" httpUri="https://zenodo.org/record/3471146/files/figure.png" pageId="9" pageNumber="472">3</figureCitation>
). Unlike other species from the Rwenzori Mountains having all slightly transverse microsculpture on elytra, the elytral microsculpture of
<taxonomicName id="4C544D56FFBFD50EDC87EFF0FCA6FF54" baseAuthorityName="Alluaud" baseAuthorityYear="1917" box="[670,825,225,250]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="ruwenzorii">
<emphasis id="B920EAC7FFBFD50EDC87EFF0FCA6FF54" box="[670,825,225,250]" italics="true" pageId="9" pageNumber="472">K. ruwenzorii</emphasis>
</taxonomicName>
is more distinct, almost granulate, consisting of large isodiametric meshes (
<figureCitation id="136F2A50FFBFD50EDF93EE15FE41FEB1" box="[394,478,260,287]" captionStart="FIGURES 1421" captionStartId="8.[151,264,1512,1537]" captionTargetBox="[157,1415,739,1485]" captionTargetId="figure@8.[151,1423,733,1492]" captionTargetPageId="8" captionText="FIGURES 1421. Kenyacus, microsculpture (1417, male pronotal; 1821, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="9" pageNumber="472">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFBFD50EDEDEEE38FA8CFE00" pageId="9" pageNumber="472" type="discussion">
<paragraph id="8BEB36D5FFBFD50EDEDEEE38FA8CFE00" blockId="9.[151,1436,152,431]" pageId="9" pageNumber="472">
<emphasis id="B920EAC7FFBFD50EDEDEEE38FEABFEED" bold="true" box="[199,308,297,323]" pageId="9" pageNumber="472">Remarks</emphasis>
. In the
<typeStatus id="54EF8877FFBFD50EDF97EE39FE6CFEEC" box="[398,499,296,322]" pageId="9" pageNumber="472" type="holotype">holotype</typeStatus>
, the ventral margin of the apical half of the median lobe of the aedeagus in lateral view is more straight (
<figureCitation id="136F2A50FFBFD50EDF8BEE5DFE4AFEC9" box="[402,469,332,359]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="9" pageNumber="472">Fig. 6</figureCitation>
) and its apex in dorsal view is slightly narrower (
<figureCitation id="136F2A50FFBFD50EDDE6EE5DFBDDFEC9" box="[1023,1090,332,359]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="9" pageNumber="472">Fig. 9</figureCitation>
) than in other examined specimens (
<figureCitation id="136F2A50FFBFD50EDEFAEE61FE1DFE25" box="[227,386,368,395]" captionStart="FIGURES 513" captionStartId="7.[151,264,1619,1644]" captionTargetBox="[168,1411,184,1582]" captionTargetId="figure@7.[151,1426,174,1600]" captionText="FIGURES 513. Kenyacus ruwenzorii (Alluaud, 1917) (5, 6, 9, holotype; 7, 8, 1113, loc. U166; 10, loc. U165). 5, 7, median lobe, dorsal view; 6, 8, same, lateral view; 9, 11, apex of median lobe, dorso-caudal view; 10, labium, ventral view; 12, stylus, lateral view; 13, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471324" httpUri="https://zenodo.org/record/3471324/files/figure.png" pageId="9" pageNumber="472">Figs 8 and 11</figureCitation>
, respectively); there are also some differences in configuration of the internal sacs. Because only few males are available, additional collecting is needed to determine the nature of this variability.
</paragraph>
</subSubSection>
</treatment>
</document>