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<document id="3C6DF51622373FE0F6FDF1CCBF24EDD5" ID-CLB-Dataset="25317" ID-DOI="10.11646/zootaxa.4679.3.3" ID-GBIF-Dataset="41cb435c-1867-4d73-9f58-fdd326298e5e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3471142" ID-ZooBank="5723773A-49BD-4CDB-99CD-B3A0AFC24F66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570099201260" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2019" docId="03FD87C3FFA5D512DE8EE928FE5CFD29" docLanguage="en" docName="zootaxa.4679.3.3.pdf" docOrigin="Zootaxa 4679 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kenyacus trechoides Kataev 2019, sp. n." docType="treatment" docVersion="7" lastPageNumber="484" masterDocId="FFC4FFBBFFB6D507DE19EF11FF9FFFAE" masterDocTitle="A review of the Afrotropical genus Kenyacus (Coleoptera: Carabidae: Harpalini) from the Rwenzori Mountains, with description of seven new species" masterLastPageNumber="498" masterPageNumber="463" pageNumber="482" updateTime="1698757568096" updateUser="ExternalLinkService">
<mods:mods id="4A503D60A85474008CA9CBDCACE01244" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="33F3145B3B4FFF6EF22C2D2A51D37CBF">
<mods:title id="666EA60174F8AA208F27B494130E2FD0">A review of the Afrotropical genus Kenyacus (Coleoptera: Carabidae: Harpalini) from the Rwenzori Mountains, with description of seven new species</mods:title>
</mods:titleInfo>
<mods:name id="A97B7EEF602804287A74E86B4B1ABC1A" type="personal">
<mods:role id="D912A21F2AC9ECFAC22282331B328CB7">
<mods:roleTerm id="E959776156B6F49513443ACFC0E31882">Author</mods:roleTerm>
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<mods:namePart id="132D1AF52056B442EB8A92906B910E14">Kataev, Boris M.</mods:namePart>
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<mods:typeOfResource id="B8FD8ED605298EB203E54CCBB4BBC679">text</mods:typeOfResource>
<mods:relatedItem id="2CB20E81AF9BA1BB90A7DAFCACFA08E0" type="host">
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<mods:title id="F842BFCE1CCE630EE3EA04EB097D8E4F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F9040C4D6E75914DE170411D36964B00">
<mods:date id="A033D80C18ABFC06FD4B439709D9C2CD">2019</mods:date>
<mods:detail id="35F38780076099C6A61175B4010C6E4C" type="pubDate">
<mods:number id="C9E53165A32B5A322A3A2BBBE69C309F">2019-10-03</mods:number>
</mods:detail>
<mods:detail id="31AE2004DB09BB661519A00839F56E22" type="volume">
<mods:number id="7C6A449353BA52EA622C163EA1F6472B">4679</mods:number>
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<mods:number id="478972464BD46CB92CB3D6EE7F4377E0">3</mods:number>
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<mods:classification id="E675EA430BF223F3900FACFA381CBCAE">journal article</mods:classification>
<mods:identifier id="5DCE9CB8844321ABDE2A7F5CD13DDC8A" type="CLB-Dataset">25317</mods:identifier>
<mods:identifier id="ECDD4AB1C760CC9719B2491E97DAF50B" type="DOI">10.11646/zootaxa.4679.3.3</mods:identifier>
<mods:identifier id="EA927D4E4988BC6CB0576233DA5D7DE9" type="GBIF-Dataset">41cb435c-1867-4d73-9f58-fdd326298e5e</mods:identifier>
<mods:identifier id="D15E97030F9E5F524EC6928D70EDA969" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="72737756AF6A16FD6A582DF7AAB44800" type="Zenodo-Dep">3471142</mods:identifier>
<mods:identifier id="91AA25FB9BF27A7E18516AE45B20FA8C" type="ZooBank">5723773A-49BD-4CDB-99CD-B3A0AFC24F66</mods:identifier>
</mods:mods>
<treatment id="03FD87C3FFA5D512DE8EE928FE5CFD29" ID-GBIF-Taxon="159628594" LSID="urn:lsid:plazi:treatment:03FD87C3FFA5D512DE8EE928FE5CFD29" httpUri="http://treatment.plazi.org/id/03FD87C3FFA5D512DE8EE928FE5CFD29" lastPageId="21" lastPageNumber="484" pageId="19" pageNumber="482">
<subSubSection id="C34E655EFFA5D514DE8EE928FE46F9FD" box="[151,473,1593,1620]" pageId="19" pageNumber="482" type="nomenclature">
<paragraph id="8BEB36D5FFA5D514DE8EE928FE46F9FD" blockId="19.[151,473,1593,1655]" box="[151,473,1593,1620]" pageId="19" pageNumber="482">
<heading id="D0A381B9FFA5D514DE8EE928FE46F9FD" bold="true" box="[151,473,1593,1620]" fontSize="11" level="1" pageId="19" pageNumber="482" reason="1">
<emphasis id="B920EAC7FFA5D514DE8EE928FE46F9FD" bold="true" box="[151,473,1593,1620]" pageId="19" pageNumber="482">
<taxonomicName id="4C544D56FFA5D514DE8EE928FE0FF9FA" ID-CoL="8M648" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[151,400,1593,1620]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="482" phylum="Arthropoda" rank="species" species="trechoides" status="sp. nov.">
<emphasis id="B920EAC7FFA5D514DE8EE928FE0FF9FA" bold="true" box="[151,400,1593,1620]" italics="true" pageId="19" pageNumber="482">Kenyacus trechoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357BCFFA5D514DF8EE928FE46F9FD" box="[407,473,1593,1619]" pageId="19" pageNumber="482" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFA5D514DE8EE94CFE18F9D9" box="[151,391,1628,1655]" pageId="19" pageNumber="482" type="description">
<paragraph id="8BEB36D5FFA5D514DE8EE94CFE18F9D9" blockId="19.[151,473,1593,1655]" box="[151,391,1628,1655]" pageId="19" pageNumber="482">
(
<figureCitation id="136F2A50FFA5D514DE86E94CFF69F9D9" box="[159,246,1629,1655]" captionStart="FIGURES 30, 31" captionStartId="15.[151,264,1809,1834]" captionTargetBox="[159,1426,744,1783]" captionTargetId="figure@15.[151,1436,733,1789]" captionTargetPageId="15" captionText="FIGURES 30, 31. Kenyacus, habitus. 30, K. berndi sp. n. (holotype); 31, K. trechoides sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471156" httpUri="https://zenodo.org/record/3471156/files/figure.png" pageId="19" pageNumber="482">Figs 31</figureCitation>
,
<figureCitation id="136F2A50FFA5D514DF1BE94CFED4F9D9" box="[258,331,1629,1655]" captionStart="FIGURES 32–49" captionStartId="17.[151,264,1856,1881]" captionTargetBox="[165,1419,181,1824]" captionTargetId="figure@17.[151,1436,174,1837]" captionTargetPageId="17" captionText="FIGURES 32–49. Kenyacus. 32–39, K. parvus sp. n. (32–34, holotype; 35–37, paratype, loc. U160; 38, 39, paratype, loc. U121); 40–46, K. berndi sp. n. (40–42, holotype; 43, 44, paratype, loc. U121; 45, 46, same, loc. U123); 47–49, K. trechoides sp. n. (holotype). 32, 35, 40, 47, median lobe, dorsal view; 34, 37, 42, 49, same, lateral view; 33, 36, 41, 48, apex of median lobe, dorso-caudal view; 38, 43, 45, stylus, lateral view; 39, 44, 46, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471158" httpUri="https://zenodo.org/record/3471158/files/figure.png" pageId="19" pageNumber="482">47–49</figureCitation>
,
<figureCitation id="136F2A50FFA5D514DF4FE94DFEE0F9D9" box="[342,383,1628,1655]" captionStart="FIGURE 51" captionStartId="21.[151,250,1926,1951]" captionTargetBox="[151,1436,661,1907]" captionTargetId="figure@21.[151,1436,661,1907]" captionTargetPageId="21" captionText="FIGURE 51. Kenyacus, distribution. a, K. trechoides sp. n.; b, K. berndi sp. n.; c, Kenyacus sp." figureDoi="http://doi.org/10.5281/zenodo.3471162" httpUri="https://zenodo.org/record/3471162/files/figure.png" pageId="19" pageNumber="482">51a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFA5D514DE8EE9B4FC6AF885" pageId="19" pageNumber="482" type="materials_examined">
<paragraph id="8BEB36D5FFA5D514DE8EE9B4FC6AF885" blockId="19.[151,1437,1700,2015]" pageId="19" pageNumber="482">
<emphasis id="B920EAC7FFA5D514DE8EE9B4FEA1F911" bold="true" box="[151,318,1700,1727]" pageId="19" pageNumber="482">Type material</emphasis>
.
<materialsCitation id="3B3C3C88FFA5D514DF52E9B4FC6AF885" ID-GBIF-Occurrence="2423812434" collectingDate="2008-08-17" collectionCode="ZMUN" collectorName="V. I. Gusarov" country="Uganda" county="Western Region" elevation="15" latitude="0.36784998" location="John Matte Hut" longLatPrecision="1" longitude="29.9563" municipality="Rwenzori Mts. National Park" pageId="19" pageNumber="482" specimenCount="1" specimenCount-male="1" stateProvince="Kasese" typeStatus="holotype">
<typeStatus id="54EF8877FFA5D514DF52E9B4FE27F911" box="[331,440,1701,1727]" pageId="19" pageNumber="482" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D52FD5CFFA5D514DFDBE9B4FE46F911" box="[450,473,1701,1727]" pageId="19" pageNumber="482" type="male">♂</specimenCount>
, “
<collectingCountry id="F3437645FFA5D514DFF7E9B4FDF2F911" box="[494,621,1701,1727]" name="Uganda" pageId="19" pageNumber="482">UGANDA</collectingCountry>
,
<collectingCounty id="628A4E59FFA5D514DC61E9B4FCB2F911" box="[632,813,1701,1727]" pageId="19" pageNumber="482">Western Region</collectingCounty>
,
<collectingRegion id="4990F837FFA5D514DD21E9B4FC7AF911" box="[824,997,1701,1727]" country="Uganda" name="Kasese" pageId="19" pageNumber="482">Kasese District</collectingRegion>
,
<locationDeviation id="9A592FF7FFA5D514DDE9E9B4FABBF911" box="[1008,1316,1701,1727]" location="Nyakalengija" pageId="19" pageNumber="482">
<quantity id="4CAC9B30FFA5D514DDE9E9B4FBA8F911" box="[1008,1079,1701,1727]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="19" pageNumber="482" unit="km" value="10.0">10 km</quantity>
NWW Nyakalengija
</locationDeviation>
,
<collectingMunicipality id="6B8FACAFFFA5D514DB36E9B4FEEDF94D" pageId="19" pageNumber="482">Rwenzori Mts. National Park</collectingMunicipality>
, main trail from
<location id="8E8B600EFFA5D514DC2CE9D8FD79F94D" LSID="urn:lsid:plazi:treatment:03FD87C3FFA5D512DE8EE928FE5CFD29:8E8B600EFFA5D514DC2CE9D8FD79F94D" box="[565,742,1737,1763]" country="Uganda" county="Western Region" latitude="0.36784998" longLatPrecision="1" longitude="29.9563" municipality="Rwenzori Mts. National Park" name="John Matte Hut" pageId="19" pageNumber="482" stateProvince="Kasese">John Matte Hut</location>
to
<location id="8E8B600EFFA5D514DD17E9D8FC28F94D" LSID="urn:lsid:plazi:treatment:03FD87C3FFA5D512DE8EE928FE5CFD29:8E8B600EFFA5D514DD17E9D8FC28F94D" box="[782,951,1737,1763]" country="Uganda" county="Western Region" latitude="0.36784998" longLatPrecision="1" longitude="29.9563" municipality="Rwenzori Mts. National Park" name="Nyabitaba Hut" pageId="19" pageNumber="482" stateProvince="Kasese">Nyabitaba Hut</location>
,
<geoCoordinate id="EE605012FFA5D514DDDAE9D8FBD8F94D" box="[963,1095,1737,1763]" degrees="0" direction="north" minutes="22.071" orientation="latitude" pageId="19" pageNumber="482" precision="1" value="0.36784998">0°22.071’N</geoCoordinate>
<geoCoordinate id="EE605012FFA5D514DA49E9D9FB7EF94D" box="[1104,1249,1736,1763]" degrees="29" direction="east" minutes="57.378" orientation="longitude" pageId="19" pageNumber="482" precision="1" value="29.9563">29°57.378’E</geoCoordinate>
, accuracy
<quantity id="4CAC9B30FFA5D514DB42E9D9FA08F94D" box="[1371,1431,1736,1763]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="482" unit="m" value="15.0">
<elevation id="0079D1E6FFA5D514DB42E9D9FA08F94D" box="[1371,1431,1736,1763]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="482" unit="m" value="15.0">15 m</elevation>
</quantity>
, extent
<quantity id="4CAC9B30FFA5D514DEFAE9FDFE82F8A9" box="[227,285,1772,1799]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="482" unit="m" value="15.0">
<elevation id="0079D1E6FFA5D514DEFAE9FDFE82F8A9" box="[227,285,1772,1799]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="482" unit="m" value="15.0">15 m</elevation>
</quantity>
, h =
<quantity id="4CAC9B30FFA5D514DF4CE9FCFE33F8A9" box="[341,428,1773,1799]" metricMagnitude="3" metricUnit="m" metricValue="2.9989999999999997" pageId="19" pageNumber="482" unit="m" value="2999.0">
<elevation id="0079D1E6FFA5D514DF4CE9FCFE33F8A9" box="[341,428,1773,1799]" metricMagnitude="3" metricUnit="m" metricValue="2.9989999999999997" pageId="19" pageNumber="482" unit="m" value="2999.0">2999 m</elevation>
</quantity>
,
<taxonomicName id="4C544D56FFA5D514DFAEE9FFFD88F8A9" box="[439,535,1774,1799]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="19" pageNumber="482" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA5D514DFAEE9FFFD88F8A9" box="[439,535,1774,1799]" italics="true" pageId="19" pageNumber="482">Hagenia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D56FFA5D514DC3FE9FFFD3DF8A9" box="[550,674,1774,1799]" class="Magnoliopsida" family="Hypericaceae" genus="Hypericum" kingdom="Plantae" order="Malpighiales" pageId="19" pageNumber="482" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA5D514DC3FE9FFFD3DF8A9" box="[550,674,1774,1799]" italics="true" pageId="19" pageNumber="482">Hypericum</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D56FFA5D514DCA9E9FCFD69F8A9" box="[688,758,1773,1799]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="19" pageNumber="482" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA5D514DCA9E9FCFD69F8A9" box="[688,758,1773,1799]" italics="true" pageId="19" pageNumber="482">Rubus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C544D56FFA5D514DD1DE9FFFCEDF8A9" box="[772,882,1774,1799]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="19" pageNumber="482" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA5D514DD1DE9FFFCEDF8A9" box="[772,882,1774,1799]" italics="true" pageId="19" pageNumber="482">Impatiens</emphasis>
</taxonomicName>
, big gaps with bushes and grass, sifting leaf litter [3595] [Garmin 60CSx; WGS84]
<collectorName id="26A15303FFA5D514DC0AE800FD3EF885" box="[531,673,1809,1835]" pageId="19" pageNumber="482">V.I. Gusarov</collectorName>
<date id="FFEA1015FFA5D514DCB1E800FCB0F885" box="[680,815,1809,1835]" pageId="19" pageNumber="482" value="2008-08-17">
<collectingDate id="EFAEE9FDFFA5D514DCB1E800FCB0F885" box="[680,815,1809,1835]" pageId="19" pageNumber="482" value="2008-08-17">17.viii.2008</collectingDate>
</date>
U161” (
<collectionCode id="ED45AE10FFA5D514DD89E800FC76F885" box="[912,1001,1809,1835]" pageId="19" pageNumber="482">ZMUN</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFA5D513DEDEE825FDAFF885" lastPageId="20" lastPageNumber="483" pageId="19" pageNumber="482" type="description">
<paragraph id="8BEB36D5FFA5D514DEDEE825FBA1F8E1" blockId="19.[151,1437,1700,2015]" box="[199,1086,1844,1872]" pageId="19" pageNumber="482">
<emphasis id="B920EAC7FFA5D514DEDEE825FECEF8E1" bold="true" box="[199,337,1844,1871]" pageId="19" pageNumber="482">Description</emphasis>
(male). Habitus as in
<figureCitation id="136F2A50FFA5D514DC50E824FD04F8E1" box="[585,667,1845,1871]" captionStart="FIGURES 30, 31" captionStartId="15.[151,264,1809,1834]" captionTargetBox="[159,1426,744,1783]" captionTargetId="figure@15.[151,1436,733,1789]" captionTargetPageId="15" captionText="FIGURES 30, 31. Kenyacus, habitus. 30, K. berndi sp. n. (holotype); 31, K. trechoides sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471156" httpUri="https://zenodo.org/record/3471156/files/figure.png" pageId="19" pageNumber="482">Fig. 31</figureCitation>
. Body length 4.0 mm, width
<quantity id="4CAC9B30FFA5D514DDF8E824FBA5F8E1" box="[993,1082,1845,1871]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="19" pageNumber="482" unit="mm" value="1.7">1.7 mm</quantity>
.
</paragraph>
<paragraph id="8BEB36D5FFA5D514DEDEE848FC54F815" blockId="19.[151,1437,1700,2015]" pageId="19" pageNumber="482">Colour: Body brown, dorsal surface slightly shiny, four basal abdominal sternites clearly infuscate; base of mandibles, labrum, clypeus, lateral margins of pronotum and elytra, as well as elytral suture slightly paler; palpi, antennae and legs brownish yellow; antennomeres 3–5 slightly infuscate.</paragraph>
<paragraph id="8BEB36D5FFA5D513DEDEE8D4FDD2FE7D" blockId="19.[151,1437,1700,2015]" lastBlockId="20.[151,1437,153,2015]" lastPageId="20" lastPageNumber="483" pageId="19" pageNumber="482">
Head: Comparatively large, HWmax/PWmax 0.81, HWmin/PWmax 0.67. Eyes with large facets, slightly con- vex, HWmax/HWmin 1.20, separated from buccal fissure at distance of width of antennomere 1. Tempora short, about quarter as long as eye, slightly convex. Frons and vertex convex. Fronto-clypeal suture distinct, superficial. Supraorbital setigerous pore located just behind level of posterior margin of eye, removed from supraorbital furrow at a distance of length of antennomere 2. Labrum almost straight anteriorly. Clypeus with slightly prominent apical angles and almost straight apical margin between them. Mentum and submentum fused, submentum with two setae on each side. Mandibles medium-sized; left mandible evenly bent in apical portion, with apex acute, only slightly blunted just at tip. Dorsal microsculpture in both sexes distinct throughout, similar to that of
<taxonomicName id="4C544D56FFA2D513DA86EE63FA97FE25" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1183,1288,369,395]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DA86EE63FA97FE25" box="[1183,1288,369,395]" italics="true" pageId="20" pageNumber="483">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DB17EE60FAD2FE25" bold="true" box="[1294,1357,369,395]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DB17EE60FAD2FE25" box="[1294,1357,369,395]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C544D56FFA2D513DB9AEE63FF7CFE01" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DB9AEE63FF7CFE01" italics="true" pageId="20" pageNumber="483">K. parvus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DEF3EE84FEB5FE01" bold="true" box="[234,298,405,431]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DEF3EE84FEB5FE01" box="[234,298,405,431]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, consisting of isodiametric meshes. Antennae slightly surpassing basal edge of pronotum, with antennomeres 7–9 slightly longer than wide.
</paragraph>
<paragraph id="8BEB36D5FFA2D513DEDEEECCFDC2FC41" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">
Pronotum: Slightly transverse, PWmax/PL 1.32, widest at the end of anterior third, markedly narrowed posteriorly, PWmax/PWmin 1.32; lateral seta inserted slightly before widest point. Sides rounded anteriorly, almost straightly converging from widest point to basal angles, without distinct border. Apical margin very shallowly emarginate; apical angles slightly projecting ahead, acute, blunted at tip. Basal margin slightly narrower than apical margin and noticeably narrower than elytral base, very shallowly concave medially and rounded laterally; both apical and basal margins not bordered. Basal angles obtuse, narrowly rounded at tip, not projecting posteriorly. Disc moderately convex, more strongly sloped to apical angles, slightly depressed basally and flattened at basal angles, forming latero-basal depressions not distinctly delimited from other parts of disc. Lateral depressions beginning from apical angles as narrow furrows, markedly widened in basal half, reaching basal margin and fused with laterobasal depressions. Basal fovea shallow, rather wide, reaching basal pronotal edge, fused with lateral depression at pronotal base and separated from it by small and narrow convexity anteriorly. Surface of pronotum impunctate. Anterior transverse depression distinct and long. Median line thin, superficial, situated between anterior transverse depression and basal margins of pronotum. Microsculpture distinct throughout, similar to that of
<taxonomicName id="4C544D56FFA2D513DAF1EC9FFACCFC09" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1256,1363,909,935]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DAF1EC9FFACCFC09" box="[1256,1363,909,935]" italics="true" pageId="20" pageNumber="483">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DB45EC9CFA02FC09" bold="true" box="[1372,1437,909,935]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DB45EC9CFA02FC09" box="[1372,1437,909,935]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C544D56FFA2D513DEDCECA3FEAEFC65" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[197,305,946,971]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DEDCECA3FEAEFC65" box="[197,305,946,971]" italics="true" pageId="20" pageNumber="483">K. parvus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DF2EECA0FEEAFC65" bold="true" box="[311,373,945,971]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DF2EECA0FEEAFC65" box="[311,373,945,971]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A50FFA2D513DF9AECA0FE4BFC62" box="[387,468,945,972]" captionStart="FIGURES 14–21" captionStartId="8.[151,264,1512,1537]" captionTargetBox="[157,1415,739,1485]" captionTargetId="figure@8.[151,1423,733,1492]" captionTargetPageId="8" captionText="FIGURES 14–21. Kenyacus, microsculpture (14–17, male pronotal; 18–21, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="20" pageNumber="483">Fig. 16</figureCitation>
), consisting of large isodiametric meshes along basal and apical margins and of slightly transverse meshes on remaining surface.
</paragraph>
<paragraph id="8BEB36D5FFA2D513DEDEECE8FEF5FAF9" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">
Elytra: Oval, slightly widened posteriorly, widest just behind middle, EL/EW 1.38, EL/PL 2.57, and EW/ PWmax 1.41. Humeri prominent, evenly rounded at tip. Sides in apical fifth rather sharply curved inside and then almost straightly converging to apex. Sutural angle very narrowly rounded at tip. Basal edge oblique, widely arcuately curving inside humerus up to lateral margin. Lateral gutter distinct, narrow along entire length. Elytral disc convex in lateral and caudal views. Striae thin, impunctate, slightly crenulate, impressed along entire length, shallowly basally and slightly more deeply apically, their bases removed from basal elytral border. Parascutellar setigerous pore present, large. Parascutellar striole present, very short. Intervals almost flat, only very slightly convex, almost not narrowed posteriorly; intervals 2 and 3 almost parallel-sided at apex; interval 3 without discal pore. Microsculpture distinct throughout, similar to that of
<taxonomicName id="4C544D56FFA2D513DCEBEA0BFCC2FA9D" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[754,861,1305,1331]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DCEBEA0BFCC2FA9D" box="[754,861,1305,1331]" italics="true" pageId="20" pageNumber="483">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DD7CEA08FC3AFA9D" bold="true" box="[869,933,1305,1331]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DD7CEA08FC3AFA9D" box="[869,933,1305,1331]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C544D56FFA2D513DDC4EA0BFBD5FA9D" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[989,1098,1306,1331]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DDC4EA0BFBD5FA9D" box="[989,1098,1306,1331]" italics="true" pageId="20" pageNumber="483">K. parvus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DA4BEA08FB0DFA9D" bold="true" box="[1106,1170,1305,1331]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DA4BEA08FB0DFA9D" box="[1106,1170,1305,1331]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136F2A50FFA2D513DABBEA08FB69FA9D" box="[1186,1270,1305,1331]" captionStart="FIGURES 14–21" captionStartId="8.[151,264,1512,1537]" captionTargetBox="[157,1415,739,1485]" captionTargetId="figure@8.[151,1423,733,1492]" captionTargetPageId="8" captionText="FIGURES 14–21. Kenyacus, microsculpture (14–17, male pronotal; 18–21, male elytral). 14, 18, K. ruwenzorii (Alluaud, 1917); 15, 19, K. gusarovi sp. n.; 16, 20, K. parvus sp. n.; 17, 21, K. angustatus sp. n. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3471148" httpUri="https://zenodo.org/record/3471148/files/figure.png" pageId="20" pageNumber="483">Fig. 20</figureCitation>
), consisting of transverse meshes.
</paragraph>
<paragraph id="8BEB36D5FFA2D513DEDEEA70FEDEFA49" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">Ventral surface of thorax and abdomen: Prosternum with several moderately long erect setae medio-apically. Metepisternum slightly wider than long, narrowed posteriorly. Basal abdominal sternites glabrous, three last visible (V, VI and VII) sternites very sparsely pubescent; apex of last visible sternite evenly rounded, with two pairs of marginal setae.</paragraph>
<paragraph id="8BEB36D5FFA2D513DEDEEAE0FBC4F9D9" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">Legs: Metacoxae without any additional setae. Tarsi glabrous dorsally. Tarsomere 5 without setae ventrally. Meso- and metatarsomeres 1 about as long as meso- and metatarsomeres 2+3, respectively. Male protarsomeres 2–4 only scarcely dilated, with biseriately arranged long adhesive scales ventrally; mesotarsomeres not dilated, with a pair of scales present only on tarsomere 4; mesotarsomere 4 rather deeply emarginate.</paragraph>
<paragraph id="8BEB36D5FFA2D513DEDEE990FDAFF885" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">
Male genitalia: Median lobe (
<figureCitation id="136F2A50FFA2D513DC09E990FD10F935" box="[528,655,1665,1691]" captionStart="FIGURES 32–49" captionStartId="17.[151,264,1856,1881]" captionTargetBox="[165,1419,181,1824]" captionTargetId="figure@17.[151,1436,174,1837]" captionTargetPageId="17" captionText="FIGURES 32–49. Kenyacus. 32–39, K. parvus sp. n. (32–34, holotype; 35–37, paratype, loc. U160; 38, 39, paratype, loc. U121); 40–46, K. berndi sp. n. (40–42, holotype; 43, 44, paratype, loc. U121; 45, 46, same, loc. U123); 47–49, K. trechoides sp. n. (holotype). 32, 35, 40, 47, median lobe, dorsal view; 34, 37, 42, 49, same, lateral view; 33, 36, 41, 48, apex of median lobe, dorso-caudal view; 38, 43, 45, stylus, lateral view; 39, 44, 46, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471158" httpUri="https://zenodo.org/record/3471158/files/figure.png" pageId="20" pageNumber="483">Figs 47, 49</figureCitation>
) stout, strongly arcuate in lateral view, with large basal bulb; in dorsal view rather evenly narrowed distally in apical third. Terminal lamella in lateral view short, straight, without apical capitulum; in dorso-caudal view comparatively narrow, narrowly rounded at apex and with evenly rounded sides (
<figureCitation id="136F2A50FFA2D513DEB9E9FCFF6AF8A9" box="[160,245,1773,1799]" captionStart="FIGURES 32–49" captionStartId="17.[151,264,1856,1881]" captionTargetBox="[165,1419,181,1824]" captionTargetId="figure@17.[151,1436,174,1837]" captionTargetPageId="17" captionText="FIGURES 32–49. Kenyacus. 32–39, K. parvus sp. n. (32–34, holotype; 35–37, paratype, loc. U160; 38, 39, paratype, loc. U121); 40–46, K. berndi sp. n. (40–42, holotype; 43, 44, paratype, loc. U121; 45, 46, same, loc. U123); 47–49, K. trechoides sp. n. (holotype). 32, 35, 40, 47, median lobe, dorsal view; 34, 37, 42, 49, same, lateral view; 33, 36, 41, 48, apex of median lobe, dorso-caudal view; 38, 43, 45, stylus, lateral view; 39, 44, 46, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471158" httpUri="https://zenodo.org/record/3471158/files/figure.png" pageId="20" pageNumber="483">Fig. 48</figureCitation>
). Internal sac without any sclerotic elements; apical folding spiny patch, as in
<taxonomicName id="4C544D56FFA2D513DA69E9FFFB44F8A9" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1136,1243,1773,1799]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DA69E9FFFB44F8A9" box="[1136,1243,1773,1799]" italics="true" pageId="20" pageNumber="483">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DAFAE9FCFABBF8A9" bold="true" box="[1251,1316,1773,1799]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DAFAE9FCFABBF8A9" box="[1251,1316,1773,1799]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, comparatively short, not reaching basal bulb.
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFA2D513DEDEE824FB84F8DD" pageId="20" pageNumber="483" type="etymology">
<paragraph id="8BEB36D5FFA2D513DEDEE824FB84F8DD" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">
<emphasis id="B920EAC7FFA2D513DEDEE824FED7F8E1" bold="true" box="[199,328,1845,1871]" pageId="20" pageNumber="483">Etymology</emphasis>
. This specific name is derived from the name of the carabid genus
<taxonomicName id="4C544D56FFA2D513DA25E824FB0BF8E1" authorityName="Clairville" authorityYear="1806" box="[1084,1172,1845,1871]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFA2D513DA25E824FB0BF8E1" box="[1084,1172,1845,1871]" italics="true" pageId="20" pageNumber="483">Trechus</emphasis>
</taxonomicName>
, the members of which are somewhat similar in appearance to this and many other species of
<taxonomicName id="4C544D56FFA2D513DDBEE84BFB8BF8DD" authorityName="Alluaud" authorityYear="1917" box="[935,1044,1882,1907]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAC7FFA2D513DDBEE84BFB8BF8DD" box="[935,1044,1882,1907]" italics="true" pageId="20" pageNumber="483">Kenyacus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E655EFFA2D512DEDEE86DFE5CFD29" lastPageId="21" lastPageNumber="484" pageId="20" pageNumber="483" type="distribution">
<paragraph id="8BEB36D5FFA2D513DEDEE86DFED8F871" blockId="20.[151,1437,153,2015]" pageId="20" pageNumber="483">
<emphasis id="B920EAC7FFA2D513DEDEE86DFEC6F839" bold="true" box="[199,345,1916,1943]" pageId="20" pageNumber="483">Distribution</emphasis>
(
<figureCitation id="136F2A50FFA2D513DF70E86CFE55F839" box="[361,458,1916,1943]" captionStart="FIGURE 51" captionStartId="21.[151,250,1926,1951]" captionTargetBox="[151,1436,661,1907]" captionTargetId="figure@21.[151,1436,661,1907]" captionTargetPageId="21" captionText="FIGURE 51. Kenyacus, distribution. a, K. trechoides sp. n.; b, K. berndi sp. n.; c, Kenyacus sp." figureDoi="http://doi.org/10.5281/zenodo.3471162" httpUri="https://zenodo.org/record/3471162/files/figure.png" pageId="20" pageNumber="483">Fig. 51a</figureCitation>
). This new species is known from one locality in the Bujuku Valley at altitude of
<quantity id="4CAC9B30FFA2D513DB7DE86CFF32F815" metricMagnitude="3" metricUnit="m" metricValue="2.9989999999999997" pageId="20" pageNumber="483" unit="m" value="2999.0">2999 m</quantity>
in the south-eastern part of the Rwenzori Mountains, where it was found together with
<taxonomicName id="4C544D56FFA2D513DA92E8B3FABAF815" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1163,1317,1954,1979]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="angustatus" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DA92E8B3FABAF815" box="[1163,1317,1954,1979]" italics="true" pageId="20" pageNumber="483">K. angustatus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DB34E8B0FAF2F815" bold="true" box="[1325,1389,1953,1979]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DB34E8B0FAF2F815" box="[1325,1389,1953,1979]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C544D56FFA2D513DE8EE8D7FE9EF871" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[151,257,1989,2015]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="483" phylum="Arthropoda" rank="species" species="similis" status="sp. nov.">
<emphasis id="B920EAC7FFA2D513DE8EE8D7FE9EF871" box="[151,257,1989,2015]" italics="true" pageId="20" pageNumber="483">K. similis</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA2D513DF11E8D4FED8F871" bold="true" box="[264,327,1989,2015]" pageId="20" pageNumber="483">
<taxonomicNameLabel id="A21357BCFFA2D513DF11E8D4FED8F871" box="[264,327,1989,2015]" pageId="20" pageNumber="483" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BEB36D5FFA3D512DEDEEF89FE9FFF55" blockId="21.[151,1437,152,647]" pageId="21" pageNumber="484">
<emphasis id="B920EAC7FFA3D512DEDEEF89FEDCFF1D" bold="true" box="[199,323,152,179]" pageId="21" pageNumber="484">Bionomics</emphasis>
. According to the label data, the specimen was collected by sifting leaf litter in forest with
<taxonomicName id="4C544D56FFA3D512DB25EF88FA03FF1C" box="[1340,1436,153,178]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="484" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA3D512DB25EF88FA03FF1C" box="[1340,1436,153,178]" italics="true" pageId="21" pageNumber="484">Hagenia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D56FFA3D512DEDCEFACFEDEFF78" box="[197,321,189,214]" class="Magnoliopsida" family="Hypericaceae" genus="Hypericum" kingdom="Plantae" order="Malpighiales" pageId="21" pageNumber="484" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA3D512DEDCEFACFEDEFF78" box="[197,321,189,214]" italics="true" pageId="21" pageNumber="484">Hypericum</emphasis>
</taxonomicName>
, with large gaps between trees and pronounced bush and herbaceous layer with
<taxonomicName id="4C544D56FFA3D512DAADEFADFB65FF78" box="[1204,1274,188,214]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="21" pageNumber="484" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA3D512DAADEFADFB65FF78" box="[1204,1274,188,214]" italics="true" pageId="21" pageNumber="484">Rubus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D56FFA3D512DB37EFACFA03FF78" box="[1326,1436,189,214]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="21" pageNumber="484" phylum="Tracheophyta" rank="genus">
<emphasis id="B920EAC7FFA3D512DB37EFACFA03FF78" box="[1326,1436,189,214]" italics="true" pageId="21" pageNumber="484">Impatiens</emphasis>
</taxonomicName>
(
<figureCitation id="136F2A50FFA3D512DEB9EFF1FF6BFF55" box="[160,244,224,251]" captionStart="FIGURE 92" captionStartId="32.[151,250,1944,1969]" captionTargetBox="[168,1420,1117,1925]" captionTargetId="figure@32.[168,1420,1117,1925]" captionTargetPageId="32" captionText="FIGURE 92. Bujuku valley between Nyabitaba Hut and John Matte Hut (ca. 2800–2900 m) near the type localities of K. trechoides sp. n., K. angustatus sp. n. and K. pusillus sp. n. Forest patches with Podocarpus, Hagenia and Hypericum separated by large gaps with bushes and herbaceous vegetation. Photo: Vladimir Gusarov." figureDoi="http://doi.org/10.5281/zenodo.3471176" httpUri="https://zenodo.org/record/3471176/files/figure.png" pageId="21" pageNumber="484">Fig. 92</figureCitation>
).
</paragraph>
<paragraph id="8BEB36D5FFA3D512DEDEEE15FE5CFD29" blockId="21.[151,1437,152,647]" pageId="21" pageNumber="484">
<emphasis id="B920EAC7FFA3D512DEDEEE15FEC4FEB1" bold="true" box="[199,347,260,287]" pageId="21" pageNumber="484">Comparison</emphasis>
. In size and general appearance with rather short oval elytra,
<taxonomicName id="4C544D56FFA3D512DA0AEE14FB37FEB0" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1043,1192,260,286]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="trechoides" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DA0AEE14FB37FEB0" box="[1043,1192,260,286]" italics="true" pageId="21" pageNumber="484">K. trechoides</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DAB6EE14FB70FEB1" bold="true" box="[1199,1263,261,287]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DAB6EE14FB70FEB1" box="[1199,1263,261,287]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C544D56FFA3D512DEAAEE38FE83FEEC" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[179,284,296,322]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DEAAEE38FE83FEEC" box="[179,284,296,322]" italics="true" pageId="21" pageNumber="484">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DF3BEE38FEFEFEED" bold="true" box="[290,353,297,323]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DF3BEE38FEFEFEED" box="[290,353,297,323]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but distinctly differs from it in the shape of the pronotum with more widely rounded basal angles and more shallowly emarginate apical and basal margins (
<figureCitation id="136F2A50FFA3D512DD22EE5DFC0EFEC9" box="[827,913,332,359]" captionStart="FIGURES 30, 31" captionStartId="15.[151,264,1809,1834]" captionTargetBox="[159,1426,744,1783]" captionTargetId="figure@15.[151,1436,733,1789]" captionTargetPageId="15" captionText="FIGURES 30, 31. Kenyacus, habitus. 30, K. berndi sp. n. (holotype); 31, K. trechoides sp. n. (holotype). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.3471156" httpUri="https://zenodo.org/record/3471156/files/figure.png" pageId="21" pageNumber="484">Fig. 31</figureCitation>
). Additionally,
<taxonomicName id="4C544D56FFA3D512DA5DEE5CFB43FEC8" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[1092,1244,332,358]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="trechoides" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DA5DEE5CFB43FEC8" box="[1092,1244,332,358]" italics="true" pageId="21" pageNumber="484">K. trechoides</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DAFFEE5CFAB6FEC9" bold="true" box="[1254,1321,333,359]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DAFFEE5CFAB6FEC9" box="[1254,1321,333,359]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
markedly differs from
<taxonomicName id="4C544D56FFA3D512DF3CEE60FE0FFE24" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[293,400,368,394]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="berndi" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DF3CEE60FE0FFE24" box="[293,400,368,394]" italics="true" pageId="21" pageNumber="484">K. berndi</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DF8EEE60FE48FE25" bold="true" box="[407,471,369,395]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DF8EEE60FE48FE25" box="[407,471,369,395]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in having supraorbital setigerous pore located more posteriorly; abdominal sternite V sparsely pubescent; male protarsus less dilated; mesotarsus almost not dilated, with adhesive scales present only on tarsomere 4; and median lobe of aedeagus (
<figureCitation id="136F2A50FFA3D512DC61EEA9FD68FE7D" box="[632,759,440,467]" captionStart="FIGURES 32–49" captionStartId="17.[151,264,1856,1881]" captionTargetBox="[165,1419,181,1824]" captionTargetId="figure@17.[151,1436,174,1837]" captionTargetPageId="17" captionText="FIGURES 32–49. Kenyacus. 32–39, K. parvus sp. n. (32–34, holotype; 35–37, paratype, loc. U160; 38, 39, paratype, loc. U121); 40–46, K. berndi sp. n. (40–42, holotype; 43, 44, paratype, loc. U121; 45, 46, same, loc. U123); 47–49, K. trechoides sp. n. (holotype). 32, 35, 40, 47, median lobe, dorsal view; 34, 37, 42, 49, same, lateral view; 33, 36, 41, 48, apex of median lobe, dorso-caudal view; 38, 43, 45, stylus, lateral view; 39, 44, 46, laterotergite and stylus, ventral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3471158" httpUri="https://zenodo.org/record/3471158/files/figure.png" pageId="21" pageNumber="484">Figs 47, 49</figureCitation>
) more strongly curved in lateral view and with narrower apical portion in dorsal view. In shape of pronotum and many other characters,
<taxonomicName id="4C544D56FFA3D512DDF8EECCFBEBFE58" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[993,1140,476,502]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="trechoides" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DDF8EECCFBEBFE58" box="[993,1140,476,502]" italics="true" pageId="21" pageNumber="484">K. trechoides</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DA63EECCFB26FE59" bold="true" box="[1146,1209,477,503]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DA63EECCFB26FE59" box="[1146,1209,477,503]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is somewhat similar to
<taxonomicName id="4C544D56FFA3D512DEACED10FEBCFDB4" authority="Kataev, 2019" authorityName="Kataev" authorityYear="2019" box="[181,291,513,538]" class="Insecta" family="Carabidae" genus="Kenyacus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="484" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B920EAC7FFA3D512DEACED10FEBCFDB4" box="[181,291,513,538]" italics="true" pageId="21" pageNumber="484">K. parvus</emphasis>
</taxonomicName>
<emphasis id="B920EAC7FFA3D512DF35ED10FEF2FDB5" bold="true" box="[300,365,513,539]" pageId="21" pageNumber="484">
<taxonomicNameLabel id="A21357BCFFA3D512DF35ED10FEF2FDB5" box="[300,365,513,539]" pageId="21" pageNumber="484" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but smaller (body length 4.0 mm versus
<quantity id="4CAC9B30FFA3D512DD52ED10FC48FDB5" box="[843,983,513,539]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.7" metricValueMin="4.1" pageId="21" pageNumber="484" unit="mm" value="4.4" valueMax="4.7" valueMin="4.1">4.1–4.7 mm</quantity>
), its elytra are relatively shorter (EL/PL 2.57 versus 2.59–2.87), abdominal sternite V is sparsely pubescent, male pro- and mesotarsi are less dilated, and median lobe of aedeagus is more strongly curved, with narrower terminal lamella and with shorter apical folding spiny patch in internal sac.
</paragraph>
</subSubSection>
</treatment>
</document>
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