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<mods:title id="D6AFF08A60BB9EB8DF629AEDE683569B">On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="21C95299B1C16AE9FD15C033A7C637B5">Chen, Jia-Heng</mods:namePart>
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<paragraph id="8BEB36BA65429661FF1FFBB72FB0548B" blockId="1.[151,661,1146,1173]" box="[151,661,1146,1173]" pageId="1" pageNumber="452">
<heading id="D0A381D665429661FF1FFBB72FB0548B" bold="true" box="[151,661,1146,1173]" fontSize="11" level="1" pageId="1" pageNumber="452" reason="1">
<emphasis id="B920EAA865429661FF1FFBB72FB0548B" bold="true" box="[151,661,1146,1173]" pageId="1" pageNumber="452">
Subgenus
<taxonomicName id="4C544D3965429661FE9FFBB72FB0548B" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[279,661,1146,1173]" class="Insecta" family="Carabidae" genus="Wraseiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAA865429661FE9FFBB72C86548A" bold="true" box="[279,419,1146,1173]" italics="true" pageId="1" pageNumber="452">Wraseiellus</emphasis>
<bibRefCitation id="EFC54B4B65429661FE22FBB72FB0548B" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[426,661,1146,1172]" pageId="1" pageNumber="452" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BEB36BA65429661FF1FFB762C4054CC" blockId="1.[151,938,1211,1266]" box="[151,357,1211,1235]" pageId="1" pageNumber="452">
<bibRefCitation id="EFC54B4B65429661FF1FFB762C4454CC" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[151,353,1211,1235]" pageId="1" pageNumber="452" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865429661FF37FB712DD654CC" box="[191,243,1211,1235]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
2013: 104
</bibRefCitation>
.
</paragraph>
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<paragraph id="8BEB36BA65429661FF1FFB172E8F54ED" blockId="1.[151,938,1211,1266]" box="[151,938,1242,1266]" pageId="1" pageNumber="452">
Type-species:
<taxonomicName id="4C544D3965429661FEA0FB172F8A54ED" authority="Jedlicka, 1931" authorityName="Jedlicka" authorityYear="1931" box="[296,687,1242,1266]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865429661FEA0FB172F3654ED" box="[296,531,1242,1266]" italics="true" pageId="1" pageNumber="452">Pterostichus andrewesi</emphasis>
<bibRefCitation id="EFC54B4B65429661FD92FB172F8A54ED" author="Jedlicka, A." box="[538,687,1242,1266]" pageId="1" pageNumber="452" pagination="21 - 30" refId="ref13081" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China- Szetschuan. Casopis Ceskoslovenske spolecnosti entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
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<paragraph id="8BEB36BA65429661FF1FFAD628AE5776" blockId="1.[151,1437,1307,2038]" pageId="1" pageNumber="452">
<emphasis id="B920EAA865429661FF1FFAD62C87552A" bold="true" box="[151,418,1307,1333]" pageId="1" pageNumber="452">Diagnostic characters.</emphasis>
The subgenus
<taxonomicName id="4C544D3965429661FDC5FAD62FF5552A" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[589,720,1307,1333]" pageId="1" pageNumber="452" rank="subGenus" subGenus="Wraseiellus">
<emphasis id="B920EAA865429661FDC5FAD62FF5552A" box="[589,720,1307,1333]" italics="true" pageId="1" pageNumber="452">Wraseiellus</emphasis>
</taxonomicName>
was defined in
<bibRefCitation id="EFC54B4B65429661FC0DFAD62964552A" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[901,1089,1307,1333]" pageId="1" pageNumber="452" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865429661FC3AFAD12EC9552A" box="[946,1004,1307,1333]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
(2013)
</bibRefCitation>
. Due to the five species newly described in the present paper, the diagnosis for the subgenus is slightly modified as below. Adults of subgenus
<taxonomicName id="4C544D3965429661FF1FFAAC2C3F5564" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[151,282,1377,1403]" pageId="1" pageNumber="452" rank="subGenus" subGenus="Wraseiellus">
<emphasis id="B920EAA865429661FF1FFAAC2C3F5564" box="[151,282,1377,1403]" italics="true" pageId="1" pageNumber="452">Wraseiellus</emphasis>
</taxonomicName>
can be recognized among Asian
<taxonomicName id="4C544D3965429661FD1CFAAC2E3A5564" authority="Bonelli, 1810" authorityName="Bonelli" authorityYear="1810" box="[660,799,1377,1403]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAA865429661FD1CFAAC2E3A5564" box="[660,799,1377,1403]" italics="true" pageId="1" pageNumber="452">Pterostichus</emphasis>
</taxonomicName>
by: medium sized pterostichine ground beetles with BL between 10.016.0 mm; dorsal surface dark brown to black, usually not metallic except males of one species that have a bluish-green metallic luster; elytra usually with granular microsculpture (formed by extraordinarily strong isodiametric meshes) in females, but microsculpture less strong than other species in females of
<taxonomicName id="4C544D3965429661FB6CFA01285655FA" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1252,1395,1483,1509]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n.">
<emphasis id="B920EAA865429661FB6CFA01285655FA" box="[1252,1395,1483,1509]" italics="true" pageId="1" pageNumber="452">P. idiopterus</emphasis>
</taxonomicName>
<emphasis id="B920EAA865429661FAF3FA062D8B5617" bold="true" pageId="1" pageNumber="452">
<taxonomicNameLabel id="A21357D365429661FAF3FA062D8B5617" pageId="1" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; penultimate labial palpomere with two setae on inner side; terminal labial palpomere at most weakly expanded; pronotum with one or more (up to five) mid-lateral setae on each side; one basal-lateral seta near posterior angle, posterior angles distinct; both parascutellar stria and basal pore present; elytral interval 3 with two setigerous pores adjacent to stria 2; umbilicate series on interval 9 discontinuous, the middle group present; metaepisterna short, with length subequal to the width of anterior margin; mesofemora with two long setae on posterior margin; male mesotibiae with a triangular projection at the apex of inner margin; metacoxae with two setae; metatrochanters without seta; metatarsomeres 13 with outer-lateral ridge distinct, metatarsomere 4 with such ridge indistinct; setae on abdominal sternite VII distant from apical margin; male abdominal sternite VII without secondary sexual modification or with a faint transverse ridge or tubercle; male genitalia relatively slender, apical lamella of aedeagus usually more or less bent dorsally or thickened on apex, but simple in two species (
<taxonomicName id="4C544D3965429661FC38F8E12967575A" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[944,1090,1835,1862]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n.">
<emphasis id="B920EAA865429661FC38F8E12EE4575A" box="[944,961,1836,1861]" italics="true" pageId="1" pageNumber="452">P</emphasis>
.
<emphasis id="B920EAA865429661FC58F8E62967575A" box="[976,1090,1835,1861]" italics="true" pageId="1" pageNumber="452">idiopterus</emphasis>
</taxonomicName>
<emphasis id="B920EAA865429661FBC1F8E629AF575A" bold="true" box="[1097,1162,1835,1861]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="A21357D365429661FBC1F8E629AF575A" box="[1097,1162,1835,1861]" pageId="1" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C544D3965429661FB10F8E12866575A" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1176,1347,1835,1862]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="hemichlorus" status="sp. n.">
<emphasis id="B920EAA865429661FB10F8E1298C575A" box="[1176,1193,1836,1861]" italics="true" pageId="1" pageNumber="452">P</emphasis>
.
<emphasis id="B920EAA865429661FB31F8E62866575A" box="[1209,1347,1835,1861]" italics="true" pageId="1" pageNumber="452">hemichlorus</emphasis>
</taxonomicName>
<emphasis id="B920EAA865429661FAC3F8E628A9575A" bold="true" box="[1355,1420,1835,1861]" pageId="1" pageNumber="452">
<taxonomicNameLabel id="A21357D365429661FAC3F8E628A9575A" box="[1355,1420,1835,1861]" pageId="1" pageNumber="452" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); endophallus simple, ventrally bent, without any chitinized pieces, often with setose or scaled area near gonopore.
</paragraph>
<paragraph id="8BEB36BA65429661FF4FF8BC2EAA57EA" blockId="1.[151,1437,1307,2038]" pageId="1" pageNumber="452">
The infra-subgeneric taxonomy of
<taxonomicName id="4C544D3965429661FDCEF8BC2FEC5794" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[582,713,1905,1931]" pageId="1" pageNumber="472" rank="subGenus" subGenus="Wraseiellus">
<emphasis id="B920EAA865429661FDCEF8BC2FEC5794" box="[582,713,1905,1931]" italics="true" pageId="1" pageNumber="452">Wraseiellus</emphasis>
</taxonomicName>
was proposed in
<bibRefCitation id="EFC54B4B65429661FC0CF8BC291D5793" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[900,1080,1905,1932]" pageId="1" pageNumber="452" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865429661FC27F8BF2EC05794" box="[943,997,1905,1931]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
(2013)
</bibRefCitation>
. Here we present supplementary differential characters: In
<taxonomicName id="4C544D3965429661FE31F8582F9857B0" box="[441,701,1941,1967]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865429661FE31F8582F9857B0" box="[441,701,1941,1967]" italics="true" pageId="1" pageNumber="452">Pterostichus andrewesi</emphasis>
</taxonomicName>
species group, endophallus with coarse setae on the dorsal-apical surface, two lobes present: ventral-right lobe (
<emphasis id="B920EAA865429661FD16F8752F9D57CD" bold="true" box="[670,696,1976,2002]" pageId="1" pageNumber="452">vr</emphasis>
) and pre-apical lobe (
<emphasis id="B920EAA865429661FC39F8752EEA57CD" bold="true" box="[945,975,1976,2002]" pageId="1" pageNumber="452">pa</emphasis>
). In
<taxonomicName id="4C544D3965429661FB8DF87529D157CD" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[1029,1268,1976,2002]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="452" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865429661FB8DF87529D157CD" box="[1029,1268,1976,2002]" italics="true" pageId="1" pageNumber="452">Pterostichus diversus</emphasis>
</taxonomicName>
species group, endophallus without developed setae, ventral-right lobe (
<emphasis id="B920EAA865429661FC9EF8162E1557EA" bold="true" box="[790,816,2011,2037]" pageId="1" pageNumber="452">vr</emphasis>
) absent.
</paragraph>
<paragraph id="8BEB36BA65419662FF4FFF5B29DD515E" blockId="2.[151,1436,150,321]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FF4FFF5B2C8850AE" bold="true" box="[199,429,150,177]" pageId="2" pageNumber="453">Geographical notes</emphasis>
. This subgenus includes 13 species and one subspecies. For the
<taxonomicName id="4C544D3965419662FBF8FF5A29E850AE" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[1136,1229,151,177]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865419662FBF8FF5A29E850AE" box="[1136,1229,151,177]" italics="true" pageId="2" pageNumber="453">diversus</emphasis>
</taxonomicName>
-group, except one species that occurs in the Middle-Himalayas in Xizang,
<collectingCountry id="F343762A65419662FC8DFF762E6E50CA" box="[773,843,187,213]" name="China" pageId="2" pageNumber="453">China</collectingCountry>
, all other species of this species-group are distributed in the Northwest
<collectingRegion id="4990F85865419662FEE8FF122C9350E6" box="[352,438,223,249]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
or adjacent area (
<figureCitation id="136F2A3F65419662FD01FF122FFA50E6" box="[649,735,223,249]" captionStart="FIGURE 14" captionStartId="20.[151,250,1423,1447]" captionTargetBox="[151,1436,181,1399]" captionTargetId="figure-23@20.[151,1436,181,1399]" captionTargetPageId="20" captionText="FIGURE 14. Distribution maps for species of subgenus Pterostichus (Wraseiellus)." figureDoi="http://doi.org/10.5281/zenodo.11150303" httpUri="https://zenodo.org/record/11150303/files/figure.png" pageId="2" pageNumber="453">Fig. 14</figureCitation>
). While the range of the
<taxonomicName id="4C544D3965419662FB8DFF12295250E6" box="[1029,1143,223,249]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865419662FB8DFF12295250E6" box="[1029,1143,223,249]" italics="true" pageId="2" pageNumber="453">andrewesi</emphasis>
</taxonomicName>
-group is further east and broader than that of the
<taxonomicName id="4C544D3965419662FE2FFECE2F215102" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[423,516,259,285]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865419662FE2FFECE2F215102" box="[423,516,259,285]" italics="true" pageId="2" pageNumber="453">diversus</emphasis>
</taxonomicName>
-group: widely in the central-west
<collectingCountry id="F343762A65419662FC03FECE2EF45102" box="[907,977,259,285]" name="China" pageId="2" pageNumber="453">China</collectingCountry>
, recorded from
<collectingRegion id="4990F85865419662FB0BFECE29F95102" box="[1155,1244,259,285]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
,
<collectingRegion id="4990F85865419662FB60FECE28605102" box="[1256,1349,259,285]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
,
<collectingRegion id="4990F85865419662FAD9FECE28BD5102" box="[1361,1432,259,285]" country="China" name="Hubei" pageId="2" pageNumber="453">Hubei</collectingRegion>
,
<collectingRegion id="4990F85865419662FF1FFEEA2DDF515E" box="[151,250,295,321]" country="China" name="Guizhou" pageId="2" pageNumber="453">Guizhou</collectingRegion>
, and
<collectingRegion id="4990F85865419662FEBBFEEA2C5C515E" box="[307,377,295,321]" country="China" name="Anhui" pageId="2" pageNumber="453">Anhui</collectingRegion>
Provinces,
<collectingRegion id="4990F85865419662FE75FEEA2F9F515E" box="[509,698,295,321]" country="China" name="Guangxi" pageId="2" pageNumber="453">Guangxi Zhuang</collectingRegion>
Autonomous Region and
<collectingRegion id="4990F85865419662FC56FEEA297F515E" box="[990,1114,295,321]" country="China" name="Chongqing" pageId="2" pageNumber="453">Chongqing</collectingRegion>
Municipality.
</paragraph>
</subSubSection>
<subSubSection id="C34E653165419662FF1FFE5F295F53C0" pageId="2" pageNumber="453" type="taxon_list">
<paragraph id="8BEB36BA65419662FF1FFE5F2E0851B3" blockId="2.[151,813,402,429]" box="[151,813,402,429]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FF1FFE5F2E0851B3" bold="true" box="[151,813,402,429]" pageId="2" pageNumber="453">
Checklist of subgenus
<taxonomicName id="4C544D3965419662FE27FE5F2E0851B3" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[431,813,402,429]" pageId="2" pageNumber="453" rank="subGenus" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FE27FE5F2F1E51B2" bold="true" box="[431,571,402,429]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
<bibRefCitation id="EFC54B4B65419662FDCAFE5F2E0851B3" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[578,813,402,428]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFE172F6C51EA" blockId="2.[151,1437,474,991]" box="[151,585,474,501]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FF1FFE172F6C51EA" bold="true" box="[151,585,474,501]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFE172C8751EA" ID-CoL="4Q4HK" box="[151,418,474,501]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865419662FF1FFE172C8751EA" bold="true" box="[151,418,474,501]" italics="true" pageId="2" pageNumber="453">Pterostichus andrewesi</emphasis>
</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFE3128A6520C" blockId="2.[151,1437,474,991]" box="[151,1411,507,531]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFE312F64520C" ID-CoL="4Q4HK" authority="Jedlicka, 1931" authorityName="Jedlicka" authorityYear="1931" box="[151,577,507,531]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="andrewesi" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFE312D83520C" box="[151,166,508,531]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FE362C17520C" box="[188,306,507,531]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FE362C8D520C" box="[321,424,507,531]" italics="true" pageId="2" pageNumber="453">andrewesi</emphasis>
<bibRefCitation id="EFC54B4B65419662FE26FE362F64520C" author="Jedlicka, A." box="[430,577,507,531]" pageId="2" pageNumber="453" pagination="21 - 30" refId="ref13081" refString="Jedlicka, A. (1931) Novi Carabidi ze Secuanu v Cine. (Col.) Neue Carabiden aus China- Szetschuan. Casopis Ceskoslovenske spolecnosti entomologicke, 28, 21 - 30." type="journal article" year="1931">Jedlička, 1931</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FDC7FE312FC2520E" bold="true" box="[591,743,508,529]" pageId="2" pageNumber="453">ṘKaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FD7EFE362E6F520C" box="[758,842,507,531]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FCDCFE362EE0520C" box="[852,965,507,531]" country="China" name="Chongqing" pageId="2" pageNumber="453">Chongqing</collectingRegion>
(
<collectingMunicipality id="6B8FACC065419662FC5AFE362946520C" box="[978,1123,507,531]" pageId="2" pageNumber="453">Mt. Jinfoshan</collectingMunicipality>
);
<collectingRegion id="4990F85865419662FBFCFE3629E3520C" box="[1140,1222,507,531]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
(
<location id="8E8B606165419662FB5CFE362858520C" LSID="urn:lsid:plazi:treatment:03FD87AC65429663FF1FFBB728B955CA:8E8B606165419662FB5CFE362858520C" box="[1236,1405,507,531]" country="China" municipality="Mt. Jinfoshan" name="Mt. Micangshan" pageId="2" pageNumber="453" stateProvince="Chongqing">Mt. Micangshan</location>
)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFDD6298F522D" blockId="2.[151,1437,474,991]" box="[151,1194,538,562]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFDD62F3A522D" ID-CoL="4Q4HQ" authority="Jedlicka, 1934" authorityName="Jedlicka" authorityYear="1934" box="[151,543,538,562]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="meyeri" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFDD62D83522D" box="[151,166,539,562]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FDD72C17522D" box="[188,306,538,562]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FDD72CA3522E" box="[321,390,538,561]" italics="true" pageId="2" pageNumber="453">meyeri</emphasis>
<bibRefCitation id="EFC54B4B65419662FE04FDD72F3A522D" author="Jedlicka, A." box="[396,543,538,562]" pageId="2" pageNumber="453" pagination="116 - 124" refId="ref13116" refString="Jedlicka, A. (1934) Novi Carabidi z vychodni Asie (VI dil). Neue Carabiden aus Ostasien (VI. Teil). Sbornik Entomologickeho Oddeleni pri Zoologichych Sbirkach Narodniho Musea v Praze, 12, 116 - 124." type="journal article" year="1934">Jedlička, 1934</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FDA5FDD62FE0522F" bold="true" box="[557,709,539,560]" pageId="2" pageNumber="453">NJAEaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FD5CFDD72E0D522D" box="[724,808,538,562]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FCB9FDD72E5B522E" box="[817,894,538,561]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(“Soling-ho”, “Pe Yen Tsin”)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFDF429FF524F" blockId="2.[151,1437,474,991]" box="[151,1242,568,593]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFDF42F5C524E" ID-CoL="4Q4HM" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[151,633,568,593]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="crassiapex" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFDF42D83524F" box="[151,166,569,592]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FDF52C17524F" box="[188,306,568,592]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FDF42C88524F" box="[321,429,569,592]" italics="true" pageId="2" pageNumber="453">crassiapex</emphasis>
<bibRefCitation id="EFC54B4B65419662FE3BFDF52F5C524E" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[435,633,568,593]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FD0FFDF42E3A5251" bold="true" box="[647,799,569,590]" pageId="2" pageNumber="453">Ṟŵaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCA6FDF42EA7524E" box="[814,898,569,593]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC03FDF52EEC524F" box="[907,969,568,592]" country="China" name="Hubei" pageId="2" pageNumber="453">Hubei</collectingRegion>
(Badong);
<collectingRegion id="4990F85865419662FBB3FDF5295F524F" box="[1083,1146,568,592]" country="China" name="Anhui" pageId="2" pageNumber="453">Anhui</collectingRegion>
(Jinzhai)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFD9529585270" blockId="2.[151,1437,474,991]" box="[151,1149,599,623]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFD952F445270" ID-CoL="4Q4HL" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[151,609,599,623]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="comatus" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFD952D835270" box="[151,166,600,623]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FD9A2C175270" box="[188,306,599,623]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FD952CB05270" box="[321,405,600,623]" italics="true" pageId="2" pageNumber="453">comatus</emphasis>
<bibRefCitation id="EFC54B4B65419662FE14FD9A2F445270" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[412,609,599,623]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FDE7FD952E225272" bold="true" box="[623,775,600,621]" pageId="2" pageNumber="453">Øfaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FC9EFD9A2E4F5270" box="[790,874,599,623]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FCFBFD9A2EEF5270" box="[883,970,599,623]" country="China" name="Guangxi" pageId="2" pageNumber="453">Guangxi</collectingRegion>
(Mt. Maoershan)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFDBA29B45291" blockId="2.[151,1437,474,991]" box="[151,1169,630,654]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFDBA2FB15291" authority="Chen, Yin &amp; Shi" authorityName="Chen, Yin &amp; Shi" authorityYear="2024" box="[151,660,630,654]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. n." subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFDBA2D835291" box="[151,166,631,654]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FDBB2C175291" box="[188,306,630,654]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FDBB2CC65291" box="[321,483,630,654]" italics="true" pageId="2" pageNumber="453">pseudandrewesi</emphasis>
Chen, Yin &amp; Shi
</taxonomicName>
,
<emphasis id="B920EAA865419662FD16FDBB2FF25291" bold="true" box="[670,727,630,654]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A21357D365419662FD16FDBB2FF25291" box="[670,727,630,654]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
[
<emphasis id="B920EAA865419662FD6EFDBA2E5B5293" bold="true" box="[742,894,631,652]" pageId="2" pageNumber="453">fDzaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FC05FDBB2EC45291" box="[909,993,630,654]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC62FDBB29195291" box="[1002,1084,630,654]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
(Butuo)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFD5829FA52B3" blockId="2.[151,1437,474,991]" box="[151,1247,660,685]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFD582F4852B3" authority="Chen, Yin &amp; Shi" authorityName="Chen, Yin &amp; Shi" authorityYear="2024" box="[151,621,660,685]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="leishanensis" status="sp. n." subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFD582D8352B3" box="[151,166,661,684]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FD592C1752B3" box="[188,306,660,684]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FD592C9852B3" box="[321,445,660,684]" italics="true" pageId="2" pageNumber="453">leishanensis</emphasis>
Chen, Yin &amp; Shi
</taxonomicName>
,
<emphasis id="B920EAA865419662FDF0FD582F9452B2" bold="true" box="[632,689,661,685]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A21357D365419662FDF0FD582F9452B2" box="[632,689,661,685]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
[
<emphasis id="B920EAA865419662FD37FD582E7252B5" bold="true" box="[703,855,661,682]" pageId="2" pageNumber="453">DZƜaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCEEFD582E9F52B2" box="[870,954,661,685]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC4CFD59293E52B3" box="[964,1051,660,684]" country="China" name="Guizhou" pageId="2" pageNumber="453">Guizhou</collectingRegion>
(Mt. Leigongshan)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFD79298652D4" blockId="2.[151,1437,474,991]" box="[151,1187,691,715]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFD792F7052D4" authority="Chen, Yin &amp; Shi" authorityName="Chen, Yin &amp; Shi" authorityYear="2024" box="[151,597,691,715]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="strumatus" status="sp. n." subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFD792D8352D4" box="[151,166,692,715]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FD7E2C1752D4" box="[188,306,691,715]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FD792C8052D4" box="[321,421,692,715]" italics="true" pageId="2" pageNumber="453">strumatus</emphasis>
Chen, Yin &amp; Shi
</taxonomicName>
,
<emphasis id="B920EAA865419662FDE8FD7E2FBC52D4" bold="true" box="[608,665,691,715]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A21357D365419662FDE8FD7E2FBC52D4" box="[608,665,691,715]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
[
<emphasis id="B920EAA865419662FD2FFD792E6552D6" bold="true" box="[679,832,692,713]" pageId="2" pageNumber="453">OiẸaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCC6FD7E2E8752D4" box="[846,930,691,715]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC24FD7E2EDB52D4" box="[940,1022,691,715]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
(Mt. Emeishan)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFD1C2F2952F5" blockId="2.[151,1437,474,991]" box="[151,524,721,746]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FF1FFD1C2F2952F5" bold="true" box="[151,524,721,746]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFD1C2C5052F6" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[151,373,721,745]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865419662FF1FFD1C2C5052F6" bold="true" box="[151,373,721,745]" italics="true" pageId="2" pageNumber="453">Pterostichus diversus</emphasis>
</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFD3C28335317" blockId="2.[151,1437,474,991]" box="[151,1302,752,777]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFD3C2F6B5317" ID-CoL="4Q4HN" authority="(Fairmaire, 1886)" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[151,590,752,777]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="diversus" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFD3C2D835317" box="[151,166,753,776]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FD3D2C175317" box="[188,306,752,776]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FD3D2CB05317" box="[321,405,752,776]" italics="true" pageId="2" pageNumber="453">diversus</emphasis>
(
<bibRefCitation id="EFC54B4B65419662FE2AFD3D2F625317" author="Fairmaire, L." box="[418,583,752,776]" pageId="2" pageNumber="453" pagination="303 - 356" refId="ref13016" refString="Fairmaire, L. (1886) Descriptions de coleopteres de l'interieur de la Chine. Annales de la Societe Entomologique de France, Series 6, 6, 303 - 356." type="journal article" year="1886">Fairmaire, 1886</bibRefCitation>
)
</taxonomicName>
[
<emphasis id="B920EAA865419662FDD5FD3C2FD05319" bold="true" box="[605,757,753,774]" pageId="2" pageNumber="453">ƂNjaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FC8CFD3C2E7D5316" box="[772,856,753,777]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FCE8FD3C2E885317" box="[864,941,753,776]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(“Ta-pin-tze”, Mt. Yulongxueshan)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFCDD2CEE5359" blockId="2.[151,1437,474,991]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFCDD2F975338" ID-CoL="4Q4HR" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[151,690,783,807]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="pseudodiversus" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFCDD2D835338" box="[151,166,784,807]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF35FCC22C165338" box="[189,307,783,807]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FECCFCC22CC55338" box="[324,480,783,807]" italics="true" pageId="2" pageNumber="453">pseudodiversus</emphasis>
<bibRefCitation id="EFC54B4B65419662FE60FCC22F975338" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[488,690,783,807]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FD4AFCDD2E7F533A" bold="true" box="[706,858,784,805]" pageId="2" pageNumber="453">ṃdzaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCE2FCC22E9B5338" box="[874,958,783,807]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC40FCDD29305338" box="[968,1045,784,807]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(Deqin, Weixi, Ninglang, Zhongdian, Lanping);
<collectingRegion id="4990F85865419662FEB9FCE32CA65359" box="[305,387,814,838]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
(Muli)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFC802845537B" blockId="2.[151,1437,474,991]" box="[151,1376,844,869]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFC802E23537A" ID-CoL="8M83F" authority="(Fairmaire, 1888)" baseAuthorityName="Fairmaire" baseAuthorityYear="1888" box="[151,774,844,869]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subGenus="Wraseiellus" subSpecies="stictopleurus">
<emphasis id="B920EAA865419662FF1FFC802D83537B" box="[151,166,845,868]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FC812C17537B" box="[188,306,844,868]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FC812F68537B" box="[321,589,844,868]" italics="true" pageId="2" pageNumber="453">stictopleurus stictopleurus</emphasis>
(
<bibRefCitation id="EFC54B4B65419662FDD2FC812FDA537A" author="Fairmaire, L." box="[602,767,844,869]" pageId="2" pageNumber="453" pagination="7 - 46" refId="ref13049" refString="Fairmaire, L. (1888) Coleopteres de l'interieur de la Chine. (suite). Annales de la Societe Entomologique de Belgique, 32, 7 - 46." type="journal article" year="1888">Fairmaire, 1888</bibRefCitation>
)
</taxonomicName>
[
<emphasis id="B920EAA865419662FC9DFC802937537D" bold="true" box="[789,1042,845,866]" pageId="2" pageNumber="453">RUaeữfflmDzadzȅ</emphasis>
]
<collectingCountry id="F343762A65419662FBA9FC802950537A" box="[1057,1141,845,869]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FBF5FC8029EF537B" box="[1149,1226,845,868]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(“Ta-pin-tze”)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFCA12815539C" blockId="2.[151,1437,474,991]" box="[151,1328,875,899]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFCA12E0C539C" ID-CoL="8MFXD" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[151,809,875,899]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subGenus="Wraseiellus" subSpecies="cangshanensis">
<emphasis id="B920EAA865419662FF1FFCA12D83539C" box="[151,166,876,899]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FCA62C17539C" box="[188,306,875,899]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FCA62F7B539C" box="[321,606,875,899]" italics="true" pageId="2" pageNumber="453">stictopleurus cangshanensis</emphasis>
<bibRefCitation id="EFC54B4B65419662FDECFCA62E0C539C" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[612,809,875,899]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
[
<emphasis id="B920EAA865419662FCB0FCA12910539E" bold="true" box="[824,1077,876,897]" pageId="2" pageNumber="453">RUaeữfflmẼƜdzȅ</emphasis>
]
<collectingCountry id="F343762A65419662FBCCFCA629BD539C" box="[1092,1176,875,899]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FB28FCA129C8539C" box="[1184,1261,876,899]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(Dali)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFC46293053BD" blockId="2.[151,1437,474,991]" box="[151,1045,906,930]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFC462F7353BD" ID-CoL="4Q4HP" authority="(Andrewes, 1947)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[151,598,906,930]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="kambaiti" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFC462D8353BD" box="[151,166,907,930]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FC472C1753BD" box="[188,306,906,930]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FC472CBF53BD" box="[321,410,906,930]" italics="true" pageId="2" pageNumber="453">kambaiti</emphasis>
(
<bibRefCitation id="EFC54B4B65419662FE2FFC472F7553BD" author="Andrewes, H. E." box="[423,592,906,930]" pageId="2" pageNumber="453" pagination="1946 - 1947" refId="ref12909" refString="Andrewes, H. E. (1947) Entomological results from the Swedish expedition 1934 to Burma and British India. Coleoptera: Carabidae. Collected by Rene Malaise. Arkiv for Zoologi, 38 A (Nr. 20), 1 - 49. [1946 - 1947]" type="book chapter" year="1947">Andrewes, 1947</bibRefCitation>
)
</taxonomicName>
[
<emphasis id="B920EAA865419662FDEDFC462E3253BF" bold="true" box="[613,791,907,928]" pageId="2" pageNumber="453">ffpďaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCAEFC472E8B53BD" box="[806,942,906,930]" name="Myanmar" pageId="2" pageNumber="453">MYANMAR</collectingCountry>
: Kambaiti
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFC64296B53DF" blockId="2.[151,1437,474,991]" box="[151,1102,936,961]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFC642F7D53DF" authority="Chen, Yin &amp; Shi" authorityName="Chen, Yin &amp; Shi" authorityYear="2024" box="[151,600,936,961]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n." subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFC642D8353DF" box="[151,166,937,960]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FC652C1753DF" box="[188,306,936,960]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FC652C8D53DF" box="[321,424,936,960]" italics="true" pageId="2" pageNumber="453">idiopterus</emphasis>
Chen, Yin &amp; Shi
</taxonomicName>
,
<emphasis id="B920EAA865419662FDEAFC642FBE53DE" bold="true" box="[610,667,937,961]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A21357D365419662FDEAFC642FBE53DE" box="[610,667,937,961]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
[
<emphasis id="B920EAA865419662FD22FC642E6753A1" bold="true" box="[682,834,937,958]" pageId="2" pageNumber="453">Ƃaeaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCD9FC642E8053DE" box="[849,933,937,961]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC26FC642EDE53DF" box="[942,1019,937,960]" country="China" name="Yunnan" pageId="2" pageNumber="453">Yunnan</collectingRegion>
(Weixi)
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFC05295F53C0" blockId="2.[151,1437,474,991]" box="[151,1146,967,991]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FF1FFC052F4A53C0" authority="Chen, Yin &amp; Shi" authorityName="Chen, Yin &amp; Shi" authorityYear="2024" box="[151,623,967,991]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="hemichlorus" status="sp. n." subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FF1FFC052D8353C0" box="[151,166,968,991]" italics="true" pageId="2" pageNumber="453">P</emphasis>
. (
<emphasis id="B920EAA865419662FF34FC0A2C1753C0" box="[188,306,967,991]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
)
<emphasis id="B920EAA865419662FEC9FC0A2C9B53C0" box="[321,446,967,991]" italics="true" pageId="2" pageNumber="453">hemichlorus</emphasis>
Chen, Yin &amp; Shi
</taxonomicName>
,
<emphasis id="B920EAA865419662FDF1FC0A2F9753C0" bold="true" box="[633,690,967,991]" pageId="2" pageNumber="453">
<taxonomicNameLabel id="A21357D365419662FDF1FC0A2F9753C0" box="[633,690,967,991]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
[
<emphasis id="B920EAA865419662FD49FC052E7D53C2" bold="true" box="[705,856,968,989]" pageId="2" pageNumber="453">fiffiaeữfflm</emphasis>
]
<collectingCountry id="F343762A65419662FCE0FC0A2E9953C0" box="[872,956,967,991]" name="China" pageId="2" pageNumber="453">CHINA</collectingCountry>
:
<collectingRegion id="4990F85865419662FC4DFC0A292B53C0" box="[965,1038,967,991]" country="China" name="Xizang" pageId="2" pageNumber="453">Xizang</collectingRegion>
(Gyirong)
</paragraph>
</subSubSection>
<subSubSection id="C34E653165419663FF1FFBFC28B955CA" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="key">
<paragraph id="8BEB36BA65419662FF1FFBFC29775454" blockId="2.[151,1106,1073,1100]" box="[151,1106,1073,1100]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FF1FFBFC29775454" bold="true" box="[151,1106,1073,1100]" pageId="2" pageNumber="453">
An improved key to species and subspecies of
<taxonomicName id="4C544D3965419662FD5CFBFC29775454" authority="Shi &amp; Sciaky, 2013" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[724,1106,1073,1100]" pageId="2" pageNumber="453" rank="subGenus" subGenus="Wraseiellus">
<emphasis id="B920EAA865419662FD5CFBFC2E455453" bold="true" box="[724,864,1073,1100]" italics="true" pageId="2" pageNumber="453">Wraseiellus</emphasis>
<bibRefCitation id="EFC54B4B65419662FCEFFBFC29775454" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[871,1106,1073,1099]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">Shi &amp; Sciaky, 2013</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
<key id="FFC5618B65419663FF1FFBBB28B955CA" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453">
<keyStep id="30A02E1F65419662FF1FFBBB28B95501" pageId="2" pageNumber="453">
<paragraph id="8BEB36BA65419662FF1FFBBB28B954D9" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFBBB28B954D9" pageId="2" pageNumber="453">
1. Area between outer basal foveal groove and lateral margin of pronotum strongly convex, forming a distinct ridge and tarsomere 5 setose beneath; males with a faint transverse ridge or tubercle on sternite VII (
<figureCitation id="136F2A3F65419662FC62FB59291754B6" box="[1002,1074,1172,1194]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="2" pageNumber="453">Fig. 10</figureCitation>
); median lobe of male genitalia more or less thickened on extreme apex (
<emphasis id="B920EAA865419662FDBEFB7D2FFA54D9" bold="true" box="[566,735,1200,1222]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FDBEFB7D2F8754D9" bold="true" box="[566,674,1200,1222]" italics="true" pageId="2" pageNumber="453">andrewesi-</emphasis>
group
</emphasis>
)........................................................ 2
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFB0228B95501" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFB0228B95501" pageId="2" pageNumber="453">
Area between outer basal foveal groove and lateral margin of pronotum slightly convex, not forming a distinct ridge or tarsomere 5 glabrous beneath; males without sexual structure on sternite VII; median lobe of male genitalia not thickened on extreme apex, sometimes laterally widened or bent upward (
<emphasis id="B920EAA865419662FD5BFAC52E4C5501" bold="true" box="[723,873,1288,1310]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FD5BFAC52E015501" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[723,804,1288,1310]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865419662FD5BFAC52E015501" bold="true" box="[723,804,1288,1310]" italics="true" pageId="2" pageNumber="453">diversus</emphasis>
</taxonomicName>
-group
</emphasis>
)............................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65419662FF1FFAEB28B95546" pageId="2" pageNumber="453">
<paragraph id="8BEB36BA65419662FF1FFAEB28B95524" blockId="2.[151,1437,1142,2015]" box="[151,1436,1318,1340]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFAEB28B95524" box="[151,1436,1318,1340]" pageId="2" pageNumber="453">2. Pronotum with 35 mid-lateral setae on each side............................................................ 3</keyLead>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFA8928B95546" blockId="2.[151,1437,1142,2015]" box="[151,1436,1348,1370]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFA8928B95546" box="[151,1436,1348,1370]" pageId="2" pageNumber="453"> Pronotum with only one mid-lateral seta on each side......................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65419662FF1FFAAC28B9565B" pageId="2" pageNumber="453">
<paragraph id="8BEB36BA65419662FF1FFAAC28B955D1" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFAAC28B955D1" pageId="2" pageNumber="453">
3. Tempora more developed (TL/EYL = 0.590.63); frontal furrows shorter, just reaching level of anterior margin of eyes; two supraorbital setae placed respectively near anterior and posterior margins of eyes; pronotal posterior angles rectangular, forming a prominent denticle (
<figureCitation id="136F2A3F65419662FE32FA512F2C55AE" box="[442,521,1436,1458]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="2" pageNumber="453">Fig. 9A</figureCitation>
); apical lamella of aedeagus short and wide (AL/AW=1.09), weakly bent to the left;
<collectingRegion id="4990F85865419662FAC7FA5128B955AD" box="[1359,1436,1436,1458]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
(Butuo)..........................................................................
<emphasis id="B920EAA865419662FB23FA7428B955D1" bold="true" box="[1195,1436,1464,1486]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FB23FA74284555D1" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1195,1376,1464,1486]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="pseudandrewesi" status="sp. n.">
<emphasis id="B920EAA865419662FB23FA74284555D1" bold="true" box="[1195,1376,1464,1486]" italics="true" pageId="2" pageNumber="453">P. pseudandrewesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365419662FAEEFA7428B955D1" box="[1382,1436,1465,1486]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FFA1A28B9565B" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FFA1A28B9565B" pageId="2" pageNumber="453">
Tempora less developed (TL/EYL = 0.310.38); frontal furrows longer, reaching level of middle of eyes; two supraorbital setae placed near middle of eyes and behind posterior margin of eyes; pronotal posterior angles a little obtuse angled, not forming prominent denticle; apical lamella of aedeagus narrow and long (AL/AW
<emphasis id="B920EAA865419662FB88F9DE293D5639" bold="true" box="[1024,1048,1555,1574]" pageId="2" pageNumber="453">&gt;</emphasis>
2), strongly twisted to the left;
<collectingRegion id="4990F85865419662FAC2F9DC28B85638" box="[1354,1437,1553,1575]" country="China" name="Guangxi" pageId="2" pageNumber="453">Guangxi</collectingRegion>
(Maoershan)......................................................................
<taxonomicName id="4C544D3965419662FB2FF9E228B9565B" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[1191,1436,1582,1604]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="comatus">
<emphasis id="B920EAA865419662FB2FF9E228B9565B" bold="true" box="[1191,1436,1582,1604]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FB2FF9E22834565B" bold="true" box="[1191,1297,1583,1604]" italics="true" pageId="2" pageNumber="453">P. comatus</emphasis>
Shi &amp; Sciaky
</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="30A02E1F65419662FF1FF98128B9570D" pageId="2" pageNumber="453">
<paragraph id="8BEB36BA65419662FF1FF98128B95683" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
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4. Pronotum with posterior margin wider than anterior margin; area between inner and outer basal foveal grooves more or less convex (
<figureCitation id="136F2A3F65419662FEBFF9A42CA35661" box="[311,390,1641,1663]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="2" pageNumber="453">Fig. 9B</figureCitation>
); median lobe of aedeagus with ventral margin distinctly curved ventrally at apical fourth; apical lamella weakly bent dorsally when in lateral view, slightly inclined to the right when in dorsal view (
<figureCitation id="136F2A3F65419662FBD5F94B298D5684" box="[1117,1192,1670,1692]" captionStart="FIGURE 3" captionStartId="7.[151,250,1064,1088]" captionTargetBox="[159,1428,190,1031]" captionTargetId="figure-17@7.[151,1436,181,1040]" captionTargetPageId="7" captionText="FIGURE 3. Male genitalia of Pterostichus leishanensis sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: vr ventral-right lobe; pa pre-apical lobe; gp gonopore; gpl gonopore lobe." figureDoi="http://doi.org/10.5281/zenodo.11150259" httpUri="https://zenodo.org/record/11150259/files/figure.png" pageId="2" pageNumber="453">Fig. 3B</figureCitation>
);
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,
<collectingRegion id="4990F85865419662FA87F94B28475683" box="[1295,1378,1670,1692]" country="China" name="Guizhou" pageId="2" pageNumber="453">Guizhou</collectingRegion>
... 5
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FF96928B9570D" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FF96928B9570D" pageId="2" pageNumber="453">
Pronotum with posterior margin narrower than anterior margin; area between inner and outer basal foveal grooves hardly convex (
<figureCitation id="136F2A3F65419662FEBDF90C2CA456C9" box="[309,385,1729,1751]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="2" pageNumber="453">Fig. 9C</figureCitation>
); median lobe of aedeagus with ventral margin nearly straight, sometimes slightly sinuate near apical fourth; apical lamella strongly bent dorsally when in lateral view, more or less inclined to the left when in dorsal view (
<figureCitation id="136F2A3F65419662FAB0F91328B556EC" box="[1336,1424,1758,1780]" captionStart="FIGURE 11" captionStartId="17.[151,250,1938,1962]" captionTargetBox="[172,1415,649,1892]" captionTargetId="figure-97@17.[151,1436,628,1915]" captionTargetPageId="17" captionText="FIGURE 11. Pterostichus crassiapex Shi &amp; Sciaky from Tianma, Anhui. (A) habitus, male; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD." figureDoi="http://doi.org/10.5281/zenodo.11150293" httpUri="https://zenodo.org/record/11150293/files/figure.png" pageId="2" pageNumber="453">Fig. 11B</figureCitation>
);
<collectingRegion id="4990F85865419662FF6AF9312C17570D" box="[226,306,1788,1810]" country="China" name="Sichuan" pageId="2" pageNumber="453">Sichuan</collectingRegion>
,
<collectingRegion id="4990F85865419662FEB3F9312C8D570D" box="[315,424,1788,1810]" country="China" name="Chongqing" pageId="2" pageNumber="453">Chongqing</collectingRegion>
,
<collectingRegion id="4990F85865419662FE3AF9312CCA570D" box="[434,495,1788,1810]" country="China" name="Hubei" pageId="2" pageNumber="453">Hubei</collectingRegion>
,
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....................................................................... 6
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<paragraph id="8BEB36BA65419662FF1FF8D428B95798" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FF8D428B95798" pageId="2" pageNumber="453">
5. Pronotum with dense punctures on mid-basal area, basal fovea strongly convex between inner and outer grooves (fig.
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<bibRefCitation id="EFC54B4B65419662FAF4F8D42C6F5754" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
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2013
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); lateral margins of elytra subparallel; apical lamella of aedeagus rounded-truncated on apex, hardly bent to the right; right paramere obtuse apically (fig.
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<bibRefCitation id="EFC54B4B65419662FDF8F8992E255776" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[624,768,1876,1898]" pageId="2" pageNumber="453" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865419662FD1EF8992FE25776" box="[662,711,1876,1897]" italics="true" pageId="2" pageNumber="453">et al.</emphasis>
2013
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);
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(Dayao, Yuanmou).....................................................................................................................
<taxonomicName id="4C544D3965419662FB6FF8BC28B95798" authority="Jedlicka" authorityName="Jedlicka" authorityYear="1934" box="[1255,1436,1905,1927]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="meyeri">
<emphasis id="B920EAA865419662FB6FF8BC28B95798" bold="true" box="[1255,1436,1905,1927]" pageId="2" pageNumber="453">
<emphasis id="B920EAA865419662FB6FF8BC28645799" bold="true" box="[1255,1345,1905,1926]" italics="true" pageId="2" pageNumber="453">P. meyeri</emphasis>
Jedlička
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65419662FF1FF84228B957C1" blockId="2.[151,1437,1142,2015]" pageId="2" pageNumber="453">
<keyLead id="30A5958F65419662FF1FF84228B957C1" pageId="2" pageNumber="453">
Pronotum with sparse punctures on mid-basal area, basal fovea slightly convex between inner and outer grooves (
<figureCitation id="136F2A3F65419662FACBF84328B457BC" box="[1347,1425,1934,1956]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="2" pageNumber="453">Fig. 9B</figureCitation>
); lateral margins of elytra distinctly arched near middle; apical lamella of aedeagus tapered to apex, more distinctly bent to the right; right paramere pointed apically (
<figureCitation id="136F2A3F65419662FDDEF8042F8457C1" box="[598,673,1993,2015]" captionStart="FIGURE 3" captionStartId="7.[151,250,1064,1088]" captionTargetBox="[159,1428,190,1031]" captionTargetId="figure-17@7.[151,1436,181,1040]" captionTargetPageId="7" captionText="FIGURE 3. Male genitalia of Pterostichus leishanensis sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: vr ventral-right lobe; pa pre-apical lobe; gp gonopore; gpl gonopore lobe." figureDoi="http://doi.org/10.5281/zenodo.11150259" httpUri="https://zenodo.org/record/11150259/files/figure.png" pageId="2" pageNumber="453">Fig. 3C</figureCitation>
);
<collectingRegion id="4990F85865419662FD3BF8042E2357C0" box="[691,774,1993,2015]" country="China" name="Guizhou" pageId="2" pageNumber="453">Guizhou</collectingRegion>
(Leigongshan).........................
<emphasis id="B920EAA865419662FB47F80428B957C1" bold="true" box="[1231,1436,1992,2014]" pageId="2" pageNumber="453">
<taxonomicName id="4C544D3965419662FB47F804284557C1" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1231,1376,1992,2014]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="453" phylum="Arthropoda" rank="species" species="leishanensis" status="sp. n.">
<emphasis id="B920EAA865419662FB47F804284557C1" bold="true" box="[1231,1376,1992,2014]" italics="true" pageId="2" pageNumber="453">P. leishanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365419662FAEEF80428B957C1" box="[1382,1436,1993,2014]" pageId="2" pageNumber="453" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFF5A28B9513D" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFF5A28B950F7" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFF5A28B950F7" pageId="3" pageNumber="454">
6. Metaepisterna finely punctate; elytron with humeral teeth less prominent; apical lamella of aedeagus short (length a little less than basal width) (
<figureCitation id="136F2A3F65409663FE1CFF782CC850D5" box="[404,493,181,203]" captionStart="FIGURE 12" captionStartId="18.[151,250,1859,1883]" captionTargetBox="[173,1417,603,1813]" captionTargetId="figure-135@18.[151,1436,581,1835]" captionTargetPageId="18" captionText="FIGURE 12. Pterostichus andrewesi Jedlička from Mt. Micangshan, Sichuan. (A) habitus, male; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD." figureDoi="http://doi.org/10.5281/zenodo.11150297" httpUri="https://zenodo.org/record/11150297/files/figure.png" pageId="3" pageNumber="454">Fig. 12C</figureCitation>
); right paramere stout, apex rounded (
<figureCitation id="136F2A3F65409663FCD2FF782E9650D5" box="[858,947,181,203]" captionStart="FIGURE 12" captionStartId="18.[151,250,1859,1883]" captionTargetBox="[173,1417,603,1813]" captionTargetId="figure-135@18.[151,1436,581,1835]" captionTargetPageId="18" captionText="FIGURE 12. Pterostichus andrewesi Jedlička from Mt. Micangshan, Sichuan. (A) habitus, male; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD." figureDoi="http://doi.org/10.5281/zenodo.11150297" httpUri="https://zenodo.org/record/11150297/files/figure.png" pageId="3" pageNumber="454">Fig. 12D</figureCitation>
);
<collectingRegion id="4990F85865409663FC4DFF78291550D4" box="[965,1072,181,203]" country="China" name="Chongqing" pageId="3" pageNumber="454">Chongqing</collectingRegion>
(Jinfoshan),
<collectingRegion id="4990F85865409663FB26FF7829DE50D4" box="[1198,1275,181,203]" country="China" name="Sichuan" pageId="3" pageNumber="454">Sichuan</collectingRegion>
(Micangshan)...................................................................................
<taxonomicName id="4C544D3965409663FB4DFF1F28B950F7" authority="Jedlicka" authorityName="Jedlicka" box="[1221,1436,210,232]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="andrewesi">
<emphasis id="B920EAA865409663FB4DFF1F28B950F7" bold="true" box="[1221,1436,210,232]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FB4DFF1F286450F7" bold="true" box="[1221,1345,210,232]" italics="true" pageId="3" pageNumber="454">P. andrewesi</emphasis>
Jedlička
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</taxonomicName>
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</paragraph>
<paragraph id="8BEB36BA65409663FF1FFF3D28B9513D" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFF3D28B9513D" pageId="3" pageNumber="454">
Metaepisterna coarsely punctate; elytron with humeral teeth a little more prominent; apical lamella of aedeagus slender (length much greater than basal width) (
<figureCitation id="136F2A3F65409663FD90FEC02F43513D" box="[536,614,269,291]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="3" pageNumber="454">Figs 4C</figureCitation>
,
<figureCitation id="136F2A3F65409663FDF8FEC02FBF513D" box="[624,666,269,290]" captionStart="FIGURE 11" captionStartId="17.[151,250,1938,1962]" captionTargetBox="[172,1415,649,1892]" captionTargetId="figure-97@17.[151,1436,628,1915]" captionTargetPageId="17" captionText="FIGURE 11. Pterostichus crassiapex Shi &amp; Sciaky from Tianma, Anhui. (A) habitus, male; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD." figureDoi="http://doi.org/10.5281/zenodo.11150293" httpUri="https://zenodo.org/record/11150293/files/figure.png" pageId="3" pageNumber="454">11C</figureCitation>
); right paramere slender, acinaciform (
<figureCitation id="136F2A3F65409663FB86FEC0297A513D" box="[1038,1119,269,291]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="3" pageNumber="454">Figs 4D</figureCitation>
,
<figureCitation id="136F2A3F65409663FBE0FEC029B7513D" box="[1128,1170,269,290]" captionStart="FIGURE 11" captionStartId="17.[151,250,1938,1962]" captionTargetBox="[172,1415,649,1892]" captionTargetId="figure-97@17.[151,1436,628,1915]" captionTargetPageId="17" captionText="FIGURE 11. Pterostichus crassiapex Shi &amp; Sciaky from Tianma, Anhui. (A) habitus, male; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD." figureDoi="http://doi.org/10.5281/zenodo.11150293" httpUri="https://zenodo.org/record/11150293/files/figure.png" pageId="3" pageNumber="454">11D</figureCitation>
).................... 7
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<keyLead id="30A5958F65409663FF1FFEE628B95187" pageId="3" pageNumber="454">
7. Pronotum with posterior angle distinctly prominent, forming a small denticle; proepisterna densely and coarsely punctate; median lobe of aedeagus without ridge or tubercle on ventral margin; ventral surface of apical lamella shallowly concaved, without ridged border; in dorsal view, apical lamella more constricted at base;
<collectingRegion id="4990F85865409663FC5BFEA829355164" box="[979,1040,357,379]" country="China" name="Hubei" pageId="3" pageNumber="454">Hubei</collectingRegion>
,
<collectingRegion id="4990F85865409663FB91FEA829705164" box="[1049,1109,357,379]" country="China" name="Anhui" pageId="3" pageNumber="454">Anhui</collectingRegion>
.........................................................................................................
<taxonomicName id="4C544D3965409663FB19FE4F28B95187" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[1169,1436,386,408]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="crassiapex">
<emphasis id="B920EAA865409663FB19FE4F28B95187" bold="true" box="[1169,1436,386,408]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FB19FE4F28345188" bold="true" box="[1169,1297,386,407]" italics="true" pageId="3" pageNumber="454">P. crassiapex</emphasis>
Shi &amp; Sciaky
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFE6D28B95212" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFE6D28B95212" pageId="3" pageNumber="454">
Pronotum with posterior angle not prominent (
<figureCitation id="136F2A3F65409663FD23FE522FDD51AA" box="[683,760,415,437]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="3" pageNumber="454">Fig. 9C</figureCitation>
); proepisterna finely and sparsely punctate; median lobe of aedeagus with a tubercle near base of ventral margin (
<figureCitation id="136F2A3F65409663FD1EFE702FC651CD" box="[662,739,445,467]" captionStart="FIGURE 4" captionStartId="9.[151,250,1557,1581]" captionTargetBox="[159,1427,282,1525]" captionTargetId="figure-38@9.[151,1436,273,1534]" captionTargetPageId="9" captionText="FIGURE 4. Holotype of Pterostichus strumatus sp. nov. (A) habitus; (B) median lobe of aedeagus, left lateral view; (C) median lobe of aedeagus, dorsal view; (D) right paramere. Scale bars = 5.0 mm for A; = 1.0 mm for BD. Abbreviations: t tubercle on ventral margin of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.11150263" httpUri="https://zenodo.org/record/11150263/files/figure.png" pageId="3" pageNumber="454">Fig. 4B</figureCitation>
); ventral surface of apical lamella deeply concaved, the concavity with ridged apical border (
<figureCitation id="136F2A3F65409663FE38FE172F2D51F0" box="[432,520,474,496]" captionStart="FIGURE 10" captionStartId="16.[151,250,1743,1767]" captionTargetBox="[153,1432,692,1720]" captionTargetId="figure-228@16.[151,1436,669,1720]" captionTargetPageId="16" captionText="FIGURE 10. Apical portion of aedeagus of Pterostichus (Wraseiellus) spp., left lateral view and ventral view. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. andrewesi Jedlička." figureDoi="http://doi.org/10.5281/zenodo.11150289" httpUri="https://zenodo.org/record/11150289/files/figure.png" pageId="3" pageNumber="454">Fig. 10C</figureCitation>
); in dorsal view, apical lamella less constricted at base;
<collectingRegion id="4990F85865409663FBA8FE17294851EF" box="[1056,1133,474,496]" country="China" name="Sichuan" pageId="3" pageNumber="454">Sichuan</collectingRegion>
(Emeishan)....................................................................................................
<emphasis id="B920EAA865409663FB6EFE3528B95212" bold="true" box="[1254,1436,504,525]" pageId="3" pageNumber="454">
<taxonomicName id="4C544D3965409663FB6EFE3528455212" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1254,1376,504,525]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="strumatus" status="sp. n.">
<emphasis id="B920EAA865409663FB6EFE3528455212" bold="true" box="[1254,1376,504,525]" italics="true" pageId="3" pageNumber="454">P. strumatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FAEEFE3528B95212" box="[1382,1436,504,525]" pageId="3" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFDD828B952E8" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFDD828B95280" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFDD828B95280" pageId="3" pageNumber="454">
8. Pronotum with outer basal foveal grooves obsolete (
<figureCitation id="136F2A3F65409663FD61FDD82E135235" box="[745,822,533,555]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="3" pageNumber="454">Fig. 9E</figureCitation>
); elytra with green metallic luster in males (
<figureCitation id="136F2A3F65409663FB7AFDD828645235" box="[1266,1345,533,555]" captionStart="FIGURE 7" captionStartId="13.[151,250,1977,2001]" captionTargetBox="[162,1428,750,1944]" captionTargetId="figure-215@13.[151,1436,741,1953]" captionTargetPageId="13" captionText="FIGURE 7. Habitus of Pterostichus hemichlorus sp. nov. (A) holotype; (B) Paratype, female. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11150277" httpUri="https://zenodo.org/record/11150277/files/figure.png" pageId="3" pageNumber="454">Fig. 7A</figureCitation>
); median lobe of aedeagus distinctly curved ventrally before apex, apical lamella not bent dorsally; apical portion (area distal to the articulation) of right paramere strongly curved, not narrowed to apex (
<figureCitation id="136F2A3F65409663FC0CFD822EF5527A" box="[900,976,591,613]" captionStart="FIGURE 8" captionStartId="14.[151,250,1151,1175]" captionTargetBox="[160,1427,190,1118]" captionTargetId="figure-23@14.[151,1436,181,1127]" captionTargetPageId="14" captionText="FIGURE 8. Male genitalia of Pterostichus hemichlorus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi="http://doi.org/10.5281/zenodo.11150283" httpUri="https://zenodo.org/record/11150283/files/figure.png" pageId="3" pageNumber="454">Fig. 8C</figureCitation>
);
<collectingRegion id="4990F85865409663FC6AFD822902527A" box="[994,1063,591,613]" country="China" name="Xizang" pageId="3" pageNumber="454">Xizang</collectingRegion>
(Gyirong).............................................................................................................................................................................................................
<emphasis id="B920EAA865409663FB45FD4728B95280" bold="true" box="[1229,1436,650,672]" pageId="3" pageNumber="454">
<taxonomicName id="4C544D3965409663FB45FD47284552BF" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1229,1376,650,672]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="hemichlorus" status="sp. n.">
<emphasis id="B920EAA865409663FB45FD47284552BF" bold="true" box="[1229,1376,650,672]" italics="true" pageId="3" pageNumber="454">P. hemichlorus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FAEEFD4728B95280" box="[1382,1436,650,671]" pageId="3" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFD6528B952E8" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFD6528B952E8" pageId="3" pageNumber="454">
Pronotum with outer basal foveal grooves well-defined (
<figureCitation id="136F2A3F65409663FC9EFD6A2E4252A2" box="[790,871,679,701]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="3" pageNumber="454">Fig. 9D</figureCitation>
); elytra without metallic luster in males; median lobe of aedeagus with ventral margin straight before apex, apical lamella usually slightly bent dorsally; apical portion of right paramere straight and strongly narrowed to apex.................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFCCD28B95372" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFCCD28B9532D" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFCCD28B9532D" pageId="3" pageNumber="454">9. Area between outer basal foveal groove and lateral margin of pronotum convex, forming a distinct ridge; lateral margins of pronotum faintly sinuate before posterior angles............................................................ 10</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFCF628B95372" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFCF628B95372" pageId="3" pageNumber="454"> Area between outer basal foveal groove and lateral margin of pronotum hardly convex, without a distinct ridge; lateral margins of pronotum straight before posterior angles............................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFCB828B953FD" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFCB828B953B8" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFCB828B953B8" pageId="3" pageNumber="454">
10. Elytral odd intervals (, III, V and VII) much wider than (about twice width as) even ones (
<figureCitation id="136F2A3F65409663FBC1FCB829B35394" box="[1097,1174,885,907]" captionStart="FIGURE 5" captionStartId="11.[151,250,1418,1442]" captionTargetBox="[159,1427,189,1386]" captionTargetId="figure-17@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 5. Habitus of Pterostichus idiopterus sp. nov. (A) holotype; (B) Paratype, female. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11150273" httpUri="https://zenodo.org/record/11150273/files/figure.png" pageId="3" pageNumber="454">Fig. 5A</figureCitation>
); apical portion (area distal to the articulation) of right paramere very short, length subequal to greatest width;
<collectingRegion id="4990F85865409663FC73FC5E296153B7" box="[1019,1092,915,936]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(Weixi).......
<emphasis id="B920EAA865409663FB6EFC5F28B953B8" bold="true" box="[1254,1436,914,936]" pageId="3" pageNumber="454">
<taxonomicName id="4C544D3965409663FB6EFC5F284453B7" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[1254,1377,914,936]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n.">
<emphasis id="B920EAA865409663FB6EFC5F284453B7" bold="true" box="[1254,1377,914,936]" italics="true" pageId="3" pageNumber="454">P. idiopterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FAEEFC5F28B953B8" box="[1382,1436,914,935]" pageId="3" pageNumber="454" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFC7D28B953FD" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFC7D28B953FD" pageId="3" pageNumber="454"> Elytral intervals normal and regular; apical portion of right paramere relatively elongated, length at least one and a half time as greatest width....................................................................................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFC2728B954D2" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFC2728B95447" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFC2728B95447" pageId="3" pageNumber="454">
11. Area between inner and outer basal foveal groove usually more convex; antennomere 3 usually glabrous except the apical ring; median lobe of aedeagus with apex of apical lamella distinctly widened (figs
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Shi
<emphasis id="B920EAA865409663FBC4FBC529585403" box="[1100,1149,1031,1053]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
); right paramere longer, apex truncate (figs 74B77B in
<bibRefCitation id="EFC54B4B65409663FD9DFBE82F805425" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[533,677,1061,1083]" pageId="3" pageNumber="454" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865409663FDB3FBE82F495425" box="[571,620,1061,1082]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
);
<collectingRegion id="4990F85865409663FD3DFBE82FDB5425" box="[693,766,1061,1082]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(Dali, Lijiang)............................................................................................................................
<taxonomicName id="4C544D3965409663FB3EFB8F28B95447" authority="(Fairmaire)" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[1206,1436,1090,1112]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA865409663FB3EFB8F28B95447" bold="true" box="[1206,1436,1090,1112]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FB3EFB8F283B5447" bold="true" box="[1206,1310,1090,1112]" italics="true" pageId="3" pageNumber="454">P. diversus</emphasis>
(Fairmaire)
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFBAD28B954D2" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFBAD28B954D2" pageId="3" pageNumber="454">
Area between inner and outer basal foveal groove usually less convex; antennomere 3 usually with few additional setae; median lobe of aedeagus with apex of apical lamella not widened, narrow or slightly truncate (figs
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<bibRefCitation id="EFC54B4B65409663FA88FBB028B7548D" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[1280,1426,1149,1171]" pageId="3" pageNumber="454" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865409663FAAEFBB0287D548D" box="[1318,1368,1149,1170]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
); right paramere shorter, apex round or slightly pointed (figs 79B84B in
<bibRefCitation id="EFC54B4B65409663FC25FB57296654B0" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[941,1091,1178,1200]" pageId="3" pageNumber="454" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865409663FC5DFB56292D54B0" box="[981,1032,1178,1200]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
);
<collectingRegion id="4990F85865409663FBDDFB5629BB54AF" box="[1109,1182,1179,1200]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(Deqin, Weixi, Ninglang, Zhongdian, Lanping),
<collectingRegion id="4990F85865409663FE32FB7A2F2254D2" box="[442,519,1207,1229]" country="China" name="Sichuan" pageId="3" pageNumber="454">Sichuan</collectingRegion>
(Muli)............................................
<taxonomicName id="4C544D3965409663FBEBFB7528B954D2" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[1123,1436,1207,1229]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="pseudodiversus">
<emphasis id="B920EAA865409663FBEBFB7528B954D2" bold="true" box="[1123,1436,1207,1229]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FBEBFB75283454D2" bold="true" box="[1123,1297,1207,1229]" italics="true" pageId="3" pageNumber="454">P. pseudodiversus</emphasis>
Shi &amp; Sciaky
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BEB36BA65409663FF1FFB1828B95517" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFB1828B95517" pageId="3" pageNumber="454">
12. Proepisterna and metaepisterna impunctate; right paramere with apex wide and obtuse (fig. 78B in
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Shi
<emphasis id="B920EAA865409663FB4BFB1829D054F5" box="[1219,1269,1237,1258]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
);
<collectingCountry id="F343762A65409663FAB7FB1828B954F5" box="[1343,1436,1237,1258]" name="Myanmar" pageId="3" pageNumber="454">Myanmar</collectingCountry>
(Kambaiti).........................................................................
<taxonomicName id="4C544D3965409663FB3FFB3F28B95517" authority="(Andrewes)" baseAuthorityName="Andrewes" baseAuthorityYear="1947" box="[1207,1436,1266,1288]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="kambaiti">
<emphasis id="B920EAA865409663FB3FFB3F28005517" bold="true" box="[1207,1317,1266,1288]" italics="true" pageId="3" pageNumber="454">P. kambaiti</emphasis>
(Andrewes)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFADD28B9555D" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFADD28B9555D" pageId="3" pageNumber="454">
Proepisterna and metaepisterna distinctly punctate; right paramere with apex narrow and sharp (fig. 72B in
<bibRefCitation id="EFC54B4B65409663FB76FAC228B7553A" author="Shi, H. L. &amp; Sciaky, R. &amp; Liang, H. B. &amp; Zhou, H. Z." box="[1278,1426,1295,1317]" pageId="3" pageNumber="454" pagination="101 - 135" refId="ref13337" refString="Shi, H. L., Sciaky, R., Liang, H. B. &amp; Zhou, H. Z. (2013) A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions. Zootaxa, 3664 (2), 101 - 135. https: // doi. org / 10.11646 / zootaxa. 3664.2.1" type="journal article" year="2013">
Shi
<emphasis id="B920EAA865409663FAADFADD2872553B" box="[1317,1367,1295,1317]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2013
</bibRefCitation>
);
<collectingRegion id="4990F85865409663FF6AFAE02C0E555D" box="[226,299,1325,1346]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(
<taxonomicName id="4C544D3965409663FEB1FAE02CF5555D" baseAuthorityName="Fairmaire" baseAuthorityYear="1889" box="[313,464,1324,1346]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="stictopleurus">
<emphasis id="B920EAA865409663FEB1FAE02CF5555D" bold="true" box="[313,464,1324,1346]" italics="true" pageId="3" pageNumber="454">P. stictopleurus</emphasis>
</taxonomicName>
).............................................................................. 13
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A02E1F65409663FF1FFA8728B955CA" pageId="3" pageNumber="454">
<paragraph id="8BEB36BA65409663FF1FFA8728B95562" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFA8728B95562" pageId="3" pageNumber="454">
13. Punctures on proepisterna less fine and sparse; tarsomere 5 glabrous beneath; in lateral view, median lobe of aedeagus slightly sinuate before apical lamella and then strongly bent dorsally at apex;
<collectingRegion id="4990F85865409663FCF3FAA52EE15562" box="[891,964,1384,1405]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(Jizushan)..
<emphasis id="B920EAA865409663FBC5FAA528B95562" bold="true" box="[1101,1436,1383,1405]" pageId="3" pageNumber="454">
<taxonomicName id="4C544D3965409663FBC5FAA529C15562" baseAuthorityName="Fairmaire" baseAuthorityYear="1889" box="[1101,1252,1383,1405]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="stictopleurus">
<emphasis id="B920EAA865409663FBC5FAA529C15562" bold="true" box="[1101,1252,1383,1405]" italics="true" pageId="3" pageNumber="454">P. stictopleurus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D365409663FB62FAA5283A5562" box="[1258,1311,1384,1405]" pageId="3" pageNumber="454" sensu="stricto">s. str.</taxonomicNameLabel>
(Fairmaire)
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEB36BA65409663FF1FFA4828B955CA" blockId="3.[151,1437,151,1493]" pageId="3" pageNumber="454">
<keyLead id="30A5958F65409663FF1FFA4828B955CA" pageId="3" pageNumber="454">
Punctures on proepisterna much finer and sparser; tarsomere 5 distinctly setose beneath; in lateral view, median lobe of aedeagus straight before apical lamella, and slightly bent dorsally at apex;
<collectingRegion id="4990F85865409663FCC7FA6E2EBD55A7" box="[847,920,1443,1464]" country="China" name="Yunnan" pageId="3" pageNumber="454">Yunnan</collectingRegion>
(Cangshan)................................................................................................
<taxonomicName id="4C544D3965409663FC6BFA0D28B955CA" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[995,1436,1471,1493]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="subSpecies" species="stictopleurus" subSpecies="cangshanensis">
<emphasis id="B920EAA865409663FC6BFA0D28B955CA" bold="true" box="[995,1436,1471,1493]" pageId="3" pageNumber="454">
<emphasis id="B920EAA865409663FC6BFA0D283555CA" bold="true" box="[995,1296,1471,1493]" italics="true" pageId="3" pageNumber="454">P. stictopleurus cangshanensis</emphasis>
Shi &amp; Sciaky
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>