plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/FD/87/03FD87A1F11BFFEB7598FD0357C47819.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

227 lines
38 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="177E861B1B3B75511EBBF85FB5934FD9" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F11BFFEB7598FD0357C47819" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) rutrilobatus Assing 2017, spec. nov." docType="treatment" docUuid="67BAD12F-50BC-4983-A8BC-363FA67F01F1" docUuidSource="ZooBank" docVersion="10" lastPageNumber="236" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="236" updateTime="1699471357828" updateUser="ExternalLinkService">
<mods:mods id="E72839748D21ED2F89046CE11049CF36" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FDE58AA4CED73A3F08A558DCA028410">
<mods:title id="9AE6A4AB3A21A01332E06D9422D03344">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
<mods:name id="78D5638EBF6D51A28ECB2902596543E6" type="personal">
<mods:role id="852DEDD2420DB61700493DE3099DC465">
<mods:roleTerm id="CE3D59DB7A57082F85D1F2030B26589A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BFAD73CFB274F20CCF1BE30F4C7EDDFC">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="94161910B89BABB8C0203C01BABA3286">text</mods:typeOfResource>
<mods:relatedItem id="C07EE00FEA6B40C62531F02422BFD227" type="host">
<mods:titleInfo id="62B2AE3B1F2AE3A2EF42A1181CECEDAC">
<mods:title id="19BE7F2D5EFFD81B51D34A9F80593587">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="9608296F45764A20260DAC238F2CD1F3">
<mods:date id="9CD35E73A5EC58DC0908AD3009A6E936">2017</mods:date>
<mods:detail id="31247C9388A2B988C545A76599E7A41E" type="pubDate">
<mods:number id="12520B36DF9C98BCB9D295879694D7F4">2017-12-08</mods:number>
</mods:detail>
<mods:detail id="C6C1E27B628ACC0E1BA9A4C1EED65C74" type="volume">
<mods:number id="F78910F4D967F93020428A3B2E5E9B93">67</mods:number>
</mods:detail>
<mods:detail id="F8078B5EBD8013045A5C4C8EEC63181E" type="issue">
<mods:number id="F2AB935B8116703BD47EE5C8A4CD7F80">2</mods:number>
</mods:detail>
<mods:extent id="5A678B9FFDD2CA7EBEB83FFAEF025A31" unit="page">
<mods:start id="EEA5A11382ADA839D79E939641F79598">213</mods:start>
<mods:end id="D5A249E1BA180E7912831C00F996F17A">246</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0F931C652978DB935F776C221D84D222">journal article</mods:classification>
<mods:identifier id="CAF1E97D5B42D666E38969F57DDC17F2" type="CLB-Dataset">2470</mods:identifier>
<mods:identifier id="F0D856B5DD5A517362471D2E16796400" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
<mods:identifier id="964F6E411D693C3713950AA64DD1A0EB" type="GBIF-Dataset">43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924</mods:identifier>
<mods:identifier id="DDE0BBCDD562A05FA33ECB0E9085BE18" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="3AEB96882A65D3D8A1D8E911FF264AF2" type="Zenodo-Dep">5742339</mods:identifier>
<mods:identifier id="F67E29FFD8A61BD140F7D8035FD406EE" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03FD87A1F11BFFEB7598FD0357C47819" ID-DOI="http://doi.org/10.5281/zenodo.5889029" ID-GBIF-Taxon="192500456" ID-Zenodo-Dep="5889029" LSID="urn:lsid:zoobank.org:act:67BAD12F-50BC-4983-A8BC-363FA67F01F1" httpUri="http://treatment.plazi.org/id/03FD87A1F11BFFEB7598FD0357C47819" lastPageNumber="236" pageId="23" pageNumber="236">
<subSubSection id="C34E653CF11BFFEB7598FD0356B07C37" pageId="23" pageNumber="236" type="nomenclature">
<paragraph id="8BEB36B7F11BFFEB7598FD03565F7DD6" blockId="23.[189,708,723,815]" box="[189,555,723,750]" pageId="23" pageNumber="236">
<heading id="D0A381DBF11BFFEB7598FD03565F7DD6" box="[189,555,723,750]" fontSize="12" level="2" pageId="23" pageNumber="236" reason="2">
<taxonomicName id="4C544D34F11BFFEB7598FD0355CF7DD6" ID-CoL="5DDFL" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,443,723,750]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="rutrilobatus" status="sp. nov." subGenus="Zyras">
<emphasis id="B920EAA5F11BFFEB7598FD0354877DD6" box="[189,243,723,750]" italics="true" pageId="23" pageNumber="236">Zyras</emphasis>
(
<emphasis id="B920EAA5F11BFFEB7424FD0355437DD6" box="[257,311,723,750]" italics="true" pageId="23" pageNumber="236">Zyras</emphasis>
)
<emphasis id="B920EAA5F11BFFEB7460FD0355CF7DD6" box="[325,443,723,750]" italics="true" pageId="23" pageNumber="236">rutrilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11BFFEB74E4FD03565F7DD6" box="[449,555,723,750]" pageId="23" pageNumber="236" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BEB36B7F11BFFEB7598FD2B56B07C37" blockId="23.[189,708,723,815]" box="[189,708,763,783]" pageId="23" pageNumber="236">
<uri id="FFC53AB5F11BFFEB7598FD2B56B07C37" box="[189,708,763,783]" pageId="23" pageNumber="236">
urn:lsid: zoobank.org:act:
<uuid id="FFF20C62F11BFFEB744BFD2B56B07C37" box="[366,708,763,783]" pageId="23" pageNumber="236">67BAD12F-50BC-4983-A8BC-363FA67F01F1</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFEB7598FCCB55117C17" box="[189,357,795,815]" pageId="23" pageNumber="236" type="description">
<paragraph id="8BEB36B7F11BFFEB7598FCCB55117C17" blockId="23.[189,708,723,815]" box="[189,357,795,815]" pageId="23" pageNumber="236">
(
<figureCitation id="136F2A32F11BFFEB75E4FCCB54847C17" box="[193,240,795,815]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="23" pageNumber="236">Figs 9</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB75DDFCCB55797C17" box="[248,269,795,815]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">31</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB7431FCCB555D7C17" box="[276,297,795,815]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="23" pageNumber="236">45</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB740AFCCB55147C17" box="[303,352,795,815]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">81–86</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFEB75B2FC9955B77CF9" pageId="23" pageNumber="236" type="materials_examined">
<paragraph id="8BEB36B7F11BFFEB75B2FC9955B77CF9" blockId="23.[151,756,841,961]" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB75B2FC9955D77C59" bold="true" box="[151,419,841,865]" pageId="23" pageNumber="236">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11BFFEB7494FC9955B77CF9" ID-GBIF-Occurrence="3450187310" collectingDate="2004-05-01" collectingDateMax="2004-05-15" collectingDateMin="2004-05-01" collectorName="Lao" country="Laos" determinerName="V. Assing" elevation="1400" location="Phongsali env." pageId="23" pageNumber="236" specimenCount="1" stateProvince="Phongsali Prov." typeStatus="holotype">
<typeStatus id="54EF8815F11BFFEB7494FC9956607C59" box="[433,532,841,865]" pageId="23" pageNumber="236" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF11BFFEB7739FC9A56457C59" box="[540,561,842,865]" pageId="23" pageNumber="236" type="generic">♂</specimenCount>
: “N
<collectingCountry id="F3437627F11BFFEB7741FC9956D37C59" box="[612,679,841,865]" name="Laos" pageId="23" pageNumber="236">LAOS</collectingCountry>
,
<quantity id="4CAC9B52F11BFFEB7796FC99549C7CB9" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="236" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<elevation id="0079D184F11BFFEB7796FC99549C7CB9" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="236" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300– 1500 m</elevation>
</quantity>
,
<collectingRegion id="4990F855F11BFFEB75D7FCB955EE7CB9" box="[242,410,873,897]" country="Laos" name="Phongsali" pageId="23" pageNumber="236">Phongsali Prov.</collectingRegion>
,
<location id="8E8B606CF11BFFEB7486FCB9564A7CB9" LSID="urn:lsid:plazi:treatment:03FD87A1F11BFFEB7598FD0357C47819:8E8B606CF11BFFEB7486FCB9564A7CB9" box="[419,574,873,897]" country="Laos" name="Phongsali env." pageId="23" pageNumber="236" stateProvince="Phongsali Prov.">Phongsali env.</location>
,
<date id="FFEA1077F11BFFEB776DFCB956847CBA" box="[584,752,873,898]" pageId="23" pageNumber="236" value="2004-05-01" valueMax="2004-05-15" valueMin="2004-05-01">
<collectingDate id="EFAEE99FF11BFFEB776DFCB956847CBA" box="[584,752,873,898]" pageId="23" pageNumber="236" value="2004-05-01" valueMax="2004-05-15" valueMin="2004-05-01">2004, 1–15 May</collectingDate>
</date>
, Lao collector leg. / Holotypus ♂
<taxonomicName id="4C544D34F11BFFEB74D6FC5A56C47C99" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[499,688,905,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="rutrilobatus" status="sp. nov.">
<emphasis id="B920EAA5F11BFFEB74D6FC5A56C47C99" box="[499,688,905,929]" italics="true" pageId="23" pageNumber="236">Zyras rutrilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11BFFEB7793FC5A569B7C99" box="[694,751,906,929]" pageId="23" pageNumber="236" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E37DA73F11BFFEB75E0FC7A55537CF9" box="[197,295,937,961]" pageId="23" pageNumber="236">
V.
<bibRefCitation id="EFC54B46F11BFFEB75C5FC7955537CF9" author="ASSING, V." box="[224,295,937,961]" pageId="23" pageNumber="236" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (cMar).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFEB75B2FC3956F17B79" pageId="23" pageNumber="236" type="etymology">
<paragraph id="8BEB36B7F11BFFEB75B2FC3956F17B79" blockId="23.[151,756,1001,1089]" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB75B2FC39557A7B39" bold="true" box="[151,270,1001,1025]" pageId="23" pageNumber="236">Etymology</emphasis>
: The specific epithet (derived from the Latin noun rutrum: shovel) is an adjective and alludes to the shovel-shaped ventral process of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFEB75B2FBB957967999" pageId="23" pageNumber="236" type="description">
<paragraph id="8BEB36B7F11BFFEB75B2FBB956667AB9" blockId="23.[151,756,1129,2081]" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB75B2FBB9556D7BB9" bold="true" box="[151,281,1129,1153]" pageId="23" pageNumber="236">Description</emphasis>
: Body length 6.0 mm; length of forebody
<quantity id="4CAC9B52F11BFFEB75B2FB59549F7B99" box="[151,235,1161,1185]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="23" pageNumber="236" unit="mm" value="2.8">2.8 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F11BFFEB74B7FB5955A07B99" box="[402,468,1161,1185]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="23" pageNumber="236">Figs 9</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB74FAFB59558E7B99" box="[479,506,1161,1185]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">31</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB7720FB5956567B99" box="[517,546,1161,1185]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="23" pageNumber="236">45</figureCitation>
): forebody black; abdomen black with the paratergites and the posterior margins of the posterior tergites paler brown; legs yellowish with the pro- and mesofemora, and the apical halves of the metafemora dark-brown; antennae blackishbrown with antennomeres I–II reddish-brown, X brown, and XI reddish-brown; maxillary palpi pale-brown with the apical palpomere yellowish-red.
</paragraph>
<paragraph id="8BEB36B7F11BFFEB75B2FA59551779B9" blockId="23.[151,756,1129,2081]" pageId="23" pageNumber="236">
Head (
<figureCitation id="136F2A32F11BFFEB75C4FA5955587A99" box="[225,300,1417,1441]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">Fig. 31</figureCitation>
) distinctly transverse; punctation moderately coarse and very sparse; median dorsal portion extensively impunctate. Eyes large, much longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F11BFFEB777CFA3956E27939" box="[601,662,1513,1537]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="23" pageNumber="236">Fig. 9</figureCitation>
)
<quantity id="4CAC9B52F11BFFEB7787FA3956807939" box="[674,756,1513,1537]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="23" pageNumber="236" unit="mm" value="2.3">2.3 mm</quantity>
long and slender; antennomeres IV–VII distinctly oblong, VIII weakly oblong, IX approximately as long as broad, X weakly transverse, and XI nearly as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F11BFFEB75B2F95A55D67859" blockId="23.[151,756,1129,2081]" pageId="23" pageNumber="236">
Pronotum (
<figureCitation id="136F2A32F11BFFEB743EF95955117999" box="[283,357,1673,1697]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">Fig. 31</figureCitation>
) strongly transverse, 1.21 times as broad as long and 1.30 times as broad as head, broadest in anterior half; posterior angles obtusely marked; lateral margins not sinuate in posterior half in dorsal view; punctation moderately coarse and very irregularly distributed, laterally with extensive impunctate patches; pubescence long, pale, and sub-erect.
</paragraph>
<paragraph id="8BEB36B7F11BFFEB75B2F8B9554F7719" blockId="23.[151,756,1129,2081]" pageId="23" pageNumber="236">
Elytra (
<figureCitation id="136F2A32F11BFFEB75CEF8B9554178B9" box="[235,309,1897,1921]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">Fig. 31</figureCitation>
) nearly 0.9 times as long as pronotum, distinctly dilated posteriad; punctation rather sparse and moderately coarse, somewhat sparser posteriorly than anteriorly; pubescence similar to that of pronotum. Hind wings present. Metatarsomere I as long as the combined length of II–IV.
</paragraph>
<paragraph id="8BEB36B7F11BFFEB763FFF3957877C59" blockId="23.[794,1399,233,1089]" pageId="23" pageNumber="236">
Abdomen (
<figureCitation id="136F2A32F11BFFEB76B1FF3957AA7E39" box="[916,990,233,257]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="23" pageNumber="236">Fig. 45</figureCitation>
) distinctly narrower than elytra, with deep anterior impressions on tergites III–V; tergites III–V each with a transverse row of moderately coarse and rather weakly defined puncture-like impressions in anterior impressions; tergite III with 1–2 lateral setiferous punctures on either side and with numerous setiferous punctures at or near posterior margin; tergites IV–V with a lateral setiferous puncture on either side, with a median pair of setiferous punctures, and with numerous setiferous punctures at posterior margins; tergite VI with a narrow transverse band of sparse non-setiferous punctures anteriorly, with few lateral setiferous punctures on either side, with a median pair of setiferous punctures in posterior portion, and with numerous setiferous punctures at posterior margin; tergite VII with a narrow transverse band of sparse non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; posterior margin of tergite VIII weakly concave in the middle (
<figureCitation id="136F2A32F11BFFEB76BEFC9957937C59" box="[923,999,841,865]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">Fig. 84</figureCitation>
).
</paragraph>
<paragraph id="8BEB36B7F11BFFEB763FFCBA57817B19" blockId="23.[794,1399,233,1089]" pageId="23" pageNumber="236">
♂: sternite VIII (
<figureCitation id="136F2A32F11BFFEB76E9FCB950637CB9" box="[972,1047,873,897]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">Fig. 85</figureCitation>
) longer than tergite VIII, wegdeshaped, and with truncate posterior margin (
<figureCitation id="136F2A32F11BFFEB703BFC59511F7C99" box="[1310,1387,905,929]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">Fig. 86</figureCitation>
); median lobe of aedeagus (
<figureCitation id="136F2A32F11BFFEB710DFC7950EA7CF9" box="[1064,1182,937,961]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">Figs 81–82</figureCitation>
) nearly
<quantity id="4CAC9B52F11BFFEB71D4FC7951377CF9" box="[1265,1347,937,961]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="236" unit="mm" value="0.9">0.9 mm</quantity>
long and with ventral process of distinctive shape; paramere (
<figureCitation id="136F2A32F11BFFEB7604FC3957187B39" box="[801,876,1001,1025]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="23" pageNumber="236">Fig. 83</figureCitation>
)
<quantity id="4CAC9B52F11BFFEB765AFC3957A97B39" box="[895,989,1001,1025]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="23" pageNumber="236" unit="mm" value="0.95">0.95 mm</quantity>
long and with very long, somewhat flattened apical lobe.
</paragraph>
<paragraph id="8BEB36B7F11BFFEB763FFBFA57DC7B79" blockId="23.[794,1399,233,1089]" box="[794,936,1065,1089]" pageId="23" pageNumber="236">♀: unknown.</paragraph>
<paragraph id="8BEB36B7F11BFFEB763FFBB957967999" blockId="23.[794,1399,1129,1697]" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB763FFBB957847BB9" bold="true" box="[794,1008,1129,1153]" pageId="23" pageNumber="236">Comparative notes</emphasis>
: Based on the morphology of the median lobe of the aedeagus, the derived shape of the paramere (very long and flattened apical lobe), the modifications of the male sternite VIII, and external characters (long and slender antennae; body glossy with sparse punctation and long pubescence),
<taxonomicName id="4C544D34F11BFFEB71FBFADA51027A19" authorityName="Assing" authorityYear="2017" box="[1246,1398,1289,1313]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F11BFFEB71FBFADA51027A19" box="[1246,1398,1289,1313]" italics="true" pageId="23" pageNumber="236">Z. rutrilobatus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C544D34F11BFFEB7124FAFA51067A79" authority="SCHEERPELTZ, 1965" authorityName="SCHEERPELTZ" authorityYear="1965" box="[1025,1394,1321,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="glabricollis">
<emphasis id="B920EAA5F11BFFEB7124FAFA50F97A79" box="[1025,1165,1321,1345]" italics="true" pageId="23" pageNumber="236">Z. glabricollis</emphasis>
<bibRefCitation id="EFC54B46F11BFFEB71BCFAF951067A79" author="SCHEERPELTZ, O." box="[1177,1394,1321,1345]" pageId="23" pageNumber="236" pagination="93 - 371" refId="ref27516" refString="SCHEERPELTZ, O. 1965: Wissenschaftliche Ergebnisse der Schwedischen Expedition 1934 nach Indien und Burma. Coleoptera Staphylinidae (except. Megalopsidiinae et Steninae). - Arkiv for Zoologie 17 (2): 93 - 371." type="journal article" year="1965">SCHEERPELTZ, 1965</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C544D34F11BFFEB763FFA9A504D7A59" authority="CAMERON, 1944" authorityName="CAMERON" authorityYear="1944" box="[794,1081,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F11BFFEB763FFA9A57007A59" box="[794,884,1354,1377]" italics="true" pageId="23" pageNumber="236">Z. nitens</emphasis>
<bibRefCitation id="EFC54B46F11BFFEB76A6FA99504D7A59" author="CAMERON, M." box="[899,1081,1353,1377]" pageId="23" pageNumber="236" pagination="104 - 108" refId="ref26241" refString="CAMERON, M. 1944: Descriptions of new Staphylinidae (Coleoptera). - Proceedings of the Royal Entomological Society of London (B) 13: 11 - 15, 49 - 52, 104 - 108." type="journal article" year="1944">CAMERON, 1944</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C544D34F11BFFEB7169FA9A51067A59" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[1100,1394,1353,1377]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B920EAA5F11BFFEB7169FA9A50B87A59" box="[1100,1228,1354,1377]" italics="true" pageId="23" pageNumber="236">Z. truncatus</emphasis>
<bibRefCitation id="EFC54B46F11BFFEB71FFFA9951067A59" author="ASSING, V." box="[1242,1394,1353,1377]" pageId="23" pageNumber="236" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING, 2017</bibRefCitation>
</taxonomicName>
, and allied species. The geographically closest species of this group is
<taxonomicName id="4C544D34F11BFFEB76FCFA5A50477A99" authorityName="CAMERON" authorityYear="1944" box="[985,1075,1418,1441]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F11BFFEB76FCFA5A50477A99" box="[985,1075,1418,1441]" italics="true" pageId="23" pageNumber="236">Z. nitens</emphasis>
</taxonomicName>
(Peninsular
<collectingCountry id="F3437627F11BFFEB71E2FA59515D7A99" box="[1223,1321,1417,1441]" name="Malaysia" pageId="23" pageNumber="236">Malaysia</collectingCountry>
), from which
<taxonomicName id="4C544D34F11BFFEB7640FA7A57897AF9" authorityName="Assing" authorityYear="2017" box="[869,1021,1449,1473]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F11BFFEB7640FA7A57897AF9" box="[869,1021,1449,1473]" italics="true" pageId="23" pageNumber="236">Z. rutrilobatus</emphasis>
</taxonomicName>
differs by a much broader, larger, and more robust body, a much more transverse pronotum, darker coloration of the body, the femora, and the antennae, more massive antennae, more finely punctate elytra, a larger median lobe of the aedeagus with a ventral process of completely different shape, and by the longer apical lobe of the paramere. For illustrations of
<taxonomicName id="4C544D34F11BFFEB7039F9BA510279B9" authorityName="CAMERON" authorityYear="1944" box="[1308,1398,1642,1665]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B920EAA5F11BFFEB7039F9BA510279B9" box="[1308,1398,1642,1665]" italics="true" pageId="23" pageNumber="236">Z. nitens</emphasis>
</taxonomicName>
see
<bibRefCitation id="EFC54B46F11BFFEB7664F95957AA7999" author="ASSING, V." box="[833,990,1673,1697]" pageId="23" pageNumber="236" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFEB763FF91957C47819" pageId="23" pageNumber="236" type="distribution">
<paragraph id="8BEB36B7F11BFFEB763FF91957C47819" blockId="23.[794,1399,1737,1825]" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB763FF91950F879D9" bold="true" box="[794,1164,1737,1761]" pageId="23" pageNumber="236">Distribution and natural history</emphasis>
: The
<typeStatus id="54EF8815F11BFFEB71F6F91A517579DA" box="[1235,1281,1738,1762]" pageId="23" pageNumber="236">type</typeStatus>
locality is situated near
<collectingRegion id="4990F855F11BFFEB768AF939506C7839" box="[943,1048,1769,1793]" country="Laos" name="Phongsali" pageId="23" pageNumber="236">Phongsali</collectingRegion>
in North
<collectingCountry id="F3437627F11BFFEB71AEF93A50C97839" box="[1163,1213,1770,1793]" name="Laos" pageId="23" pageNumber="236">Laos</collectingCountry>
at an altitude of
<quantity id="4CAC9B52F11BFFEB763FF8D957D87819" box="[794,940,1801,1825]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="236" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink