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<document id="9B29AF0AE807D648261BBBDF4452FF50" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F11BFFE4761AF8A350677DD9" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) parvilobatus Assing 2017, spec. nov." docType="treatment" docUuid="6DBB36A1-C12B-4FA1-B28E-55560D6645C6" docUuidSource="ZooBank" docVersion="10" lastPageNumber="237" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="236" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="B4D534C3BD756B2AC91C7FD5E8A208A3">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="1264269C8E91E25BD8A2FFBB805ECC2C">Assing, Volker</mods:namePart>
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<mods:date id="CC41FBB4F9C612A44B385E5A212242CC">2017</mods:date>
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<treatment id="03FD87A1F11BFFE4761AF8A350677DD9" ID-DOI="http://doi.org/10.5281/zenodo.5888810" ID-GBIF-Taxon="192500457" ID-Zenodo-Dep="5888810" LSID="urn:lsid:zoobank.org:act:6DBB36A1-C12B-4FA1-B28E-55560D6645C6" httpUri="http://treatment.plazi.org/id/03FD87A1F11BFFE4761AF8A350677DD9" lastPageId="24" lastPageNumber="237" pageId="23" pageNumber="236">
<subSubSection id="C34E653CF11BFFEB761AF8A3513F7897" pageId="23" pageNumber="236" type="nomenclature">
<paragraph id="8BEB36B7F11BFFEB761AF8A350C178B6" blockId="23.[831,1355,1907,1999]" box="[831,1205,1907,1934]" pageId="23" pageNumber="236">
<heading id="D0A381DBF11BFFEB761AF8A350C178B6" box="[831,1205,1907,1934]" fontSize="12" level="2" pageId="23" pageNumber="236" reason="2">
<taxonomicName id="4C544D34F11BFFEB761AF8A3503178B6" ID-CoL="7GHXC" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[831,1093,1907,1934]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="236" phylum="Arthropoda" rank="species" species="parvilobatus" status="sp. nov." subGenus="Zyras">
<emphasis id="B920EAA5F11BFFEB761AF8A3570178B6" box="[831,885,1907,1934]" italics="true" pageId="23" pageNumber="236">Zyras</emphasis>
(
<emphasis id="B920EAA5F11BFFEB76A6F8A357CD78B6" box="[899,953,1907,1934]" italics="true" pageId="23" pageNumber="236">Zyras</emphasis>
)
<emphasis id="B920EAA5F11BFFEB76E2F8A3503178B6" box="[967,1093,1907,1934]" italics="true" pageId="23" pageNumber="236">parvilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11BFFEB7169F8A350C178B6" box="[1100,1205,1907,1934]" pageId="23" pageNumber="236" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C34E653CF11BFFEB761AF86B578478F7" box="[831,1008,1979,1999]" pageId="23" pageNumber="236" type="description">
<paragraph id="8BEB36B7F11BFFEB761AF86B578478F7" blockId="23.[831,1355,1907,1999]" box="[831,1008,1979,1999]" pageId="23" pageNumber="236">
(
<figureCitation id="136F2A32F11BFFEB7666F86B570978F7" box="[835,893,1979,1999]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="23" pageNumber="236">Figs 10</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB76A6F86B57EC78F7" box="[899,920,1979,1999]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="23" pageNumber="236">28</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB76BAF86B57C078F7" box="[927,948,1979,1999]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="23" pageNumber="236">43</figureCitation>
,
<figureCitation id="136F2A32F11BFFEB769EF86B579878F7" box="[955,1004,1979,1999]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="23" pageNumber="236">8789</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11BFFE4763FF839551F7E79" lastPageId="24" lastPageNumber="237" pageId="23" pageNumber="236" type="materials_examined">
<paragraph id="8BEB36B7F11BFFE4763FF839551F7E79" blockId="23.[794,1399,2025,2082]" lastBlockId="24.[189,794,233,321]" lastPageId="24" lastPageNumber="237" pageId="23" pageNumber="236">
<emphasis id="B920EAA5F11BFFEB763FF83950517739" bold="true" box="[794,1061,2025,2049]" pageId="23" pageNumber="236">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11BFFE47116F83955137E79" ID-GBIF-Occurrence="3450187308" collectingDate="2004-05-01" collectingDateMax="2004-05-15" collectingDateMin="2004-05-01" collectorName="Lao" country="Laos" elevation="1400" lastPageId="24" lastPageNumber="237" location="Phongsali env." pageId="23" pageNumber="236" specimenCount="1" stateProvince="Phongsali Prov." typeStatus="holotype">
<typeStatus id="54EF8815F11BFFEB7116F83950E27739" box="[1075,1174,2025,2049]" pageId="23" pageNumber="236" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF11BFFEB71BBF83A50C77739" box="[1182,1203,2026,2049]" pageId="23" pageNumber="236"></specimenCount>
: “N
<collectingCountry id="F3437627F11BFFEB71C2F839515E7739" box="[1255,1322,2025,2049]" name="Laos" pageId="23" pageNumber="236">LAOS</collectingCountry>
,
<quantity id="4CAC9B52F11BFFEB7010F839571F7719" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="236" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
<elevation id="0079D184F11BFFEB7010F839571F7719" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="236" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300 1500 m</elevation>
</quantity>
,
<collectingRegion id="4990F855F11BFFEB7650F7D950687719" box="[885,1052,2057,2081]" country="Laos" name="Phongsali" pageId="23" pageNumber="236">Phongsali Prov.</collectingRegion>
,
<location id="8E8B606CF11BFFEB7103F7D950B57719" LSID="urn:lsid:plazi:treatment:03FD87A1F11BFFE4761AF8A350677DD9:8E8B606CF11BFFEB7103F7D950B57719" box="[1062,1217,2057,2081]" country="Laos" name="Phongsali env." pageId="23" pageNumber="236" stateProvince="Phongsali Prov.">Phongsali env.</location>
,
<date id="FFEA1077F11BFFEB71EFF7D95107771A" box="[1226,1395,2057,2082]" pageId="23" pageNumber="236" value="2004-05-01" valueMax="2004-05-15" valueMin="2004-05-01">
<collectingDate id="EFAEE99FF11BFFEB71EFF7D95107771A" box="[1226,1395,2057,2082]" pageId="23" pageNumber="236" value="2004-05-01" valueMax="2004-05-15" valueMin="2004-05-01">2004, 115 May</collectingDate>
</date>
, Lao collector leg. / Holotypus ♂
<taxonomicName id="4C544D34F114FFE47731FF3A56A37E39" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[532,727,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="parvilobatus" status="sp. nov.">
<emphasis id="B920EAA5F114FFE47731FF3A56A37E39" box="[532,727,233,257]" italics="true" pageId="24" pageNumber="237">Zyras parvilobatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF114FFE477F9FF3A57607E39" box="[732,788,234,257]" pageId="24" pageNumber="237" rank="species">sp. n.</taxonomicNameLabel>
, det. V.
<bibRefCitation id="EFC54B46F114FFE47422FED955FF7E19" author="ASSING, V." box="[263,395,265,289]" pageId="24" pageNumber="237" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing 2017</bibRefCitation>
” (cMar). Paratype: 1 ♀: same data as holotype (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE47598FEB956827E99" pageId="24" pageNumber="237" type="etymology">
<paragraph id="8BEB36B7F114FFE47598FEB956827E99" blockId="24.[189,793,361,417]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47598FEB955407EB9" bold="true" box="[189,308,361,385]" pageId="24" pageNumber="237">Etymology</emphasis>
: The specific epithet is an adjective alluding to the relatively small ventral process of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE47598FE1950567D79" pageId="24" pageNumber="237" type="description">
<paragraph id="8BEB36B7F114FFE47598FE1955207DD9" blockId="24.[189,795,457,2081]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47598FE19554B7ED9" bold="true" box="[189,319,457,481]" pageId="24" pageNumber="237">Description</emphasis>
: Body length
<quantity id="4CAC9B52F114FFE474C0FE1956127ED9" box="[485,614,457,481]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" metricValueMax="6.5" metricValueMin="5.8" pageId="24" pageNumber="237" unit="mm" value="6.15" valueMax="6.5" valueMin="5.8">5.86.5 mm</quantity>
; length of forebody
<quantity id="4CAC9B52F114FFE475DFFE39550F7D39" box="[250,379,489,513]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="24" pageNumber="237" unit="mm" value="2.8" valueMax="2.9" valueMin="2.7">2.72.9 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F114FFE4772CFE3956237D39" box="[521,599,489,513]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="24" pageNumber="237">Figs 10</figureCitation>
,
<figureCitation id="136F2A32F114FFE47746FE39560B7D39" box="[611,639,489,513]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">28</figureCitation>
,
<figureCitation id="136F2A32F114FFE477AFFE3956D37D39" box="[650,679,489,513]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="24" pageNumber="237">43</figureCitation>
): forebody blackish with the humeral and anterior portions of the elytra reddish-yellow; abdomen black with the posterior margins of the tergites more or less distinctly paler reddish; legs yellowish; antennae blackish with the apical portion of antennomere XI sometimes paler brown; maxillary palpi brown with the apical palpomere yellowish-red.
</paragraph>
<paragraph id="8BEB36B7F114FFE47598FD39558D7B39" blockId="24.[189,795,457,2081]" pageId="24" pageNumber="237">
Head (
<figureCitation id="136F2A32F114FFE47422FD3955207C39" box="[263,340,745,769]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">Fig. 28</figureCitation>
) distinctly transverse; punctation moderately coarse and moderately sparse; along middle without punctation. Eyes large, much longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F114FFE4772BFC99562D7C59" box="[526,601,841,865]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="24" pageNumber="237">Fig. 10</figureCitation>
)
<quantity id="4CAC9B52F114FFE47743FC9956927C59" box="[614,742,841,865]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="24" pageNumber="237" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.92.1 mm</quantity>
long and moderately massive; antennomeres IVV weakly oblong to weakly transverse, VI as long as broad or weakly transverse, VIIX weakly transverse, X less than 1.5 times as broad as long, and XI slightly longer than the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F114FFE47598FBDA55527A39" blockId="24.[189,795,457,2081]" pageId="24" pageNumber="237">
Pronotum (
<figureCitation id="136F2A32F114FFE4741EFBD955F27B19" box="[315,390,1033,1057]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">Fig. 28</figureCitation>
) strongly transverse, 1.231.30 times as broad as long and 1.341.35 times as broad as head, broadest near anterior angles; posterior angles weakly, obtusely marked; lateral margins not sinuate in posterior half in dorsal view; punctation moderately coarse and moderately dense, regularly distributed; midline with or without narrow and somewhat irregular narrow impunctate band.
</paragraph>
<paragraph id="8BEB36B7F114FFE47598FAD955157A99" blockId="24.[189,795,457,2081]" pageId="24" pageNumber="237">
Elytra (
<figureCitation id="136F2A32F114FFE47437FAD955287A19" box="[274,348,1289,1313]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="24" pageNumber="237">Fig. 28</figureCitation>
) 0.820.85 times as long as pronotum; punctation rather dense and coarse, denser near scutellum than in posterior portion. Hind wings present. Metatarsomere I approximately as long as the combined length of IIIV.
</paragraph>
<paragraph id="8BEB36B7F114FFE47598FA79564378F9" blockId="24.[189,795,457,2081]" pageId="24" pageNumber="237">
Abdomen (
<figureCitation id="136F2A32F114FFE47462FA7955E07AF9" box="[327,404,1449,1473]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="24" pageNumber="237">Fig. 43</figureCitation>
) slightly narrower than elytra, with moderately deep anterior impressions on tergites IIIV; tergites IIIV each with a somewhat irregular transverse row or band of moderately coarse non-setiferous punctures in anterior impressions, with a lateral setiferous puncture on either side and with 46 setiferous punctures at posterior margins; tergite VI with a transverse band of rather sparse non-setiferous punctures anteriorly, with a transverse row of sparse setiferous punctures in posterior portion, and with six setiferous punctures at posterior margin; tergite VII with a transverse band of sparse non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures only in posterior fourth, posterior margin with sharply delimited concavity in the middle.
</paragraph>
<paragraph id="8BEB36B7F114FFE47598F81A503B7E19" blockId="24.[189,795,457,2081]" lastBlockId="24.[832,1437,233,353]" pageId="24" pageNumber="237">
♂: sternite VIII with strongly convex posterior margin; median lobe of aedeagus (
<figureCitation id="136F2A32F114FFE474D0F839561F7739" box="[501,619,2025,2049]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="24" pageNumber="237">Figs 8788</figureCitation>
) approximately
<quantity id="4CAC9B52F114FFE47598F7D9557B7719" box="[189,271,2057,2081]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="24" pageNumber="237" unit="mm" value="0.7">0.7 mm</quantity>
long; ventral process short in relation to capsule; paramere (
<figureCitation id="136F2A32F114FFE47692FF3950767E39" box="[951,1026,233,257]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="24" pageNumber="237">Fig. 89</figureCitation>
)
<quantity id="4CAC9B52F114FFE47131FF3950127E39" box="[1044,1126,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="24" pageNumber="237" unit="mm" value="0.8">0.8 mm</quantity>
long, apical lobe short and with four long dark setae.
</paragraph>
<paragraph id="8BEB36B7F114FFE47665FEFA57CD7E59" blockId="24.[832,1437,233,353]" pageId="24" pageNumber="237">♀: posterior margin of sternite VIII shallowly concave in the middle.</paragraph>
<paragraph id="8BEB36B7F114FFE47665FE5950567D79" blockId="24.[832,1437,393,577]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47665FE5950637E99" bold="true" box="[832,1047,393,417]" pageId="24" pageNumber="237">Comparative notes</emphasis>
:
<taxonomicName id="4C544D34F114FFE4710EFE5A50827E99" authorityName="Assing" authorityYear="2017" box="[1067,1270,393,417]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F114FFE4710EFE5A50827E99" box="[1067,1270,393,417]" italics="true" pageId="24" pageNumber="237">Zyras parvilobatus</emphasis>
</taxonomicName>
is difficult to distinguish from the extremely similar, syntopic
<taxonomicName id="4C544D34F114FFE47078FE7A51E97EF9" authorityName="PACE" authorityYear="1993" box="[1373,1437,426,449]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F114FFE47078FE7A51E97EF9" box="[1373,1437,426,449]" italics="true" pageId="24" pageNumber="237">Z. wei</emphasis>
</taxonomicName>
based on external characters alone. The only reliable diagnostic characters are the distinctive shape of the median lobe of the aedeagus and the much shorter apical lobe of the paramere.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF114FFE47665FDB950677DD9" pageId="24" pageNumber="237" type="distribution">
<paragraph id="8BEB36B7F114FFE47665FDB950677DD9" blockId="24.[832,1437,617,737]" pageId="24" pageNumber="237">
<emphasis id="B920EAA5F114FFE47665FDB950C67DB9" bold="true" box="[832,1202,617,641]" pageId="24" pageNumber="237">Distribution and natural history</emphasis>
: The
<typeStatus id="54EF8815F114FFE471DDFDBA51527DBA" box="[1272,1318,618,642]" pageId="24" pageNumber="237">type</typeStatus>
locality is situated near
<collectingRegion id="4990F855F114FFE476F0FD59504A7D99" box="[981,1086,649,673]" country="Laos" name="Phongsali" pageId="24" pageNumber="237">Phongsali</collectingRegion>
in North
<collectingCountry id="F3437627F114FFE47194FD5A50977D99" box="[1201,1251,650,673]" name="Laos" pageId="24" pageNumber="237">Laos</collectingCountry>
at an altitude of
<quantity id="4CAC9B52F114FFE47665FD7957A67DF9" box="[832,978,681,705]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="24" pageNumber="237" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">13001500 m</quantity>
.
<taxonomicName id="4C544D34F114FFE476FAFD7A50EA7DF9" authorityName="Assing" authorityYear="2017" box="[991,1182,681,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F114FFE476FAFD7A50EA7DF9" box="[991,1182,681,705]" italics="true" pageId="24" pageNumber="237">Zyras rutrilobatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C544D34F114FFE471FDFD7A516C7DF9" authorityName="PACE" authorityYear="1993" box="[1240,1304,682,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="237" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F114FFE471FDFD7A516C7DF9" box="[1240,1304,682,705]" italics="true" pageId="24" pageNumber="237">Z. wei</emphasis>
</taxonomicName>
were found in the same locality.
</paragraph>
</subSubSection>
</treatment>
</document>