240 lines
37 KiB
XML
240 lines
37 KiB
XML
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<mods:title id="59D7130D781681523D2F4D406F3E990D">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="8AB14B8B2014CF2143718CE4CAFE29E9">Assing, Volker</mods:namePart>
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<mods:title id="E83E535277FAD86EBBDE1F0DB724285A">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="16F383F77A28611BA928B06A2F1C987A">2017</mods:date>
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<mods:number id="BB7FAF18525E9C4D73B9464025541428">2017-12-08</mods:number>
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<treatment id="03FD87A1F119FFEA761AFA23508B7C97" ID-DOI="http://doi.org/10.5281/zenodo.5888806" ID-GBIF-Taxon="192500460" ID-Zenodo-Dep="5888806" LSID="urn:lsid:plazi:treatment:03FD87A1F119FFEA761AFA23508B7C97" httpUri="http://treatment.plazi.org/id/03FD87A1F119FFEA761AFA23508B7C97" lastPageId="22" lastPageNumber="235" pageId="21" pageNumber="234">
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<subSubSection id="C34E653CF119FFE9761AFA2350AC7936" box="[831,1240,1523,1550]" pageId="21" pageNumber="234" type="nomenclature">
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<paragraph id="8BEB36B7F119FFE9761AFA2350AC7936" blockId="21.[831,1240,1523,1583]" box="[831,1240,1523,1550]" pageId="21" pageNumber="234">
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<heading id="D0A381DBF119FFE9761AFA2350AC7936" box="[831,1240,1523,1550]" centered="true" fontSize="12" level="2" pageId="21" pageNumber="234" reason="2">
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<taxonomicName id="4C544D34F119FFE9761AFA2350AC7936" ID-CoL="7GHYP" authority="CAMERON, 1939" authorityName="Cameron" authorityYear="1939" box="[831,1240,1523,1550]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="flavorufus" subGenus="Zyras">
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<emphasis id="B920EAA5F119FFE9761AFA2357017936" box="[831,885,1523,1550]" italics="true" pageId="21" pageNumber="234">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F119FFE976A6FA2357CD7936" box="[899,953,1523,1550]" italics="true" pageId="21" pageNumber="234">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F119FFE976E2FA2350597936" box="[967,1069,1523,1550]" italics="true" pageId="21" pageNumber="234">flavorufus</emphasis>
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CAMERON, 1939
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF119FFE9761AF9CB57937917" box="[831,999,1563,1583]" pageId="21" pageNumber="234" type="description">
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<paragraph id="8BEB36B7F119FFE9761AF9CB57937917" blockId="21.[831,1240,1523,1583]" box="[831,999,1563,1583]" pageId="21" pageNumber="234">
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(
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<figureCitation id="136F2A32F119FFE97666F9CB57077917" box="[835,883,1563,1583]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="21" pageNumber="234">Figs 8</figureCitation>
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,
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<figureCitation id="136F2A32F119FFE9765FF9CB57FB7917" box="[890,911,1563,1583]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="21" pageNumber="234">27</figureCitation>
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,
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<figureCitation id="136F2A32F119FFE976B3F9CB57DF7917" box="[918,939,1563,1583]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="21" pageNumber="234">42</figureCitation>
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,
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<figureCitation id="136F2A32F119FFE97697F9CB57967917" box="[946,994,1563,1583]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="21" pageNumber="234">73–75</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF119FFE9763FF99C50A67958" box="[794,1234,1611,1633]" pageId="21" pageNumber="234" type="reference_group">
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<paragraph id="8BEB36B7F119FFE9763FF99C50A67958" blockId="21.[794,1234,1611,1633]" box="[794,1234,1611,1633]" pageId="21" pageNumber="234">
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<treatmentCitationGroup id="AB441199F119FFE9763FF99C50A67958" box="[794,1234,1611,1633]" pageId="21" pageNumber="234">
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<treatmentCitation id="0AF510A6F119FFE9763FF99C50BA7958" author="CAMERON, M." box="[794,1230,1611,1633]" page="18" pageId="21" pageNumber="234" year="1939">
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<taxonomicName id="4C544D34F119FFE9763FF99C50BA7958" ID-CoL="7GHYP" authority="CAMERON, 1939 b: 18" authorityName="Cameron" authorityPageNumber="18" authorityYear="1939" box="[794,1230,1611,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="flavorufus" subGenus="Zyras">
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<emphasis id="B920EAA5F119FFE9763FF99C573A7959" box="[794,846,1612,1633]" italics="true" pageId="21" pageNumber="234">Zyras</emphasis>
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||
(
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<emphasis id="B920EAA5F119FFE97679F99C57E47959" box="[860,912,1612,1633]" italics="true" pageId="21" pageNumber="234">Zyras</emphasis>
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||
)
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<emphasis id="B920EAA5F119FFE976B8F99B57887958" box="[925,1020,1611,1632]" italics="true" pageId="21" pageNumber="234">flavorufus</emphasis>
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||
<bibRefCitation id="EFC54B46F119FFE97127F99B50BA7958" author="CAMERON, M." box="[1026,1230,1611,1633]" pageId="21" pageNumber="234" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON, 1939b: 18</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF119FFE9763FF95957F77879" pageId="21" pageNumber="234" type="materials_examined">
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||
<paragraph id="8BEB36B7F119FFE9763FF95957F77879" blockId="21.[794,1399,1673,1857]" pageId="21" pageNumber="234">
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||
<emphasis id="B920EAA5F119FFE9763FF95950517999" bold="true" box="[794,1061,1673,1697]" pageId="21" pageNumber="234">Type material examined</emphasis>
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||
:
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<materialsCitation id="3B3C3CEAF119FFE97116F95A57F77879" ID-GBIF-Occurrence="3450187335" collectingDate="1933-09" collectionCode="BMNH" determinerName="V. Assing" location="Preanger" municipality="Voet" pageId="21" pageNumber="234" specimenCount="3" stateProvince="Java" typeStatus="lectotype">
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<typeStatus id="54EF8815F119FFE97116F95A50E8799A" box="[1075,1180,1674,1698]" pageId="21" pageNumber="234" type="lectotype">Lectotype</typeStatus>
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♂, present designation: “
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<collectorName id="26A15361F119FFE97679F979578179F9" box="[860,1013,1705,1729]" pageId="21" pageNumber="234">F. C. Drescher</collectorName>
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,
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<collectorName id="26A15361F119FFE97125F979508379F9" box="[1024,1271,1705,1729]" pageId="21" pageNumber="234">G. Tangkoeban Prahoe</collectorName>
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, 4000–5000
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<collectingMunicipality id="6B8FACCDF119FFE9763FF91A573A79D9" box="[794,846,1738,1761]" pageId="21" pageNumber="234">Voet</collectingMunicipality>
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,
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<location id="8E8B606CF119FFE97644F91A57B579DA" LSID="urn:lsid:plazi:treatment:03FD87A1F119FFEA761AFA23508B7C97:8E8B606CF119FFE97644F91A57B579DA" box="[865,961,1738,1762]" municipality="Voet" name="Preanger" pageId="21" pageNumber="234" stateProvince="Java">Preanger</location>
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,
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<collectingRegion id="4990F855F119FFE976F1F91A507679D9" box="[980,1026,1738,1761]" country="Indonesia" name="Jawa Timur" pageId="21" pageNumber="234">Java</collectingRegion>
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,
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<date id="FFEA1077F119FFE97131F919501C79D9" box="[1044,1128,1737,1761]" pageId="21" pageNumber="234" value="1933-09">
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<collectingDate id="EFAEE99FF119FFE97131F919501C79D9" box="[1044,1128,1737,1761]" pageId="21" pageNumber="234" value="1933-09">IX.1933</collectingDate>
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</date>
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/
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<taxonomicName id="4C544D34F119FFE971B5F91A512B79D9" authority="Cam." authorityName="Cameron" authorityYear="1939" box="[1168,1375,1737,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="flavorufus">
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<emphasis id="B920EAA5F119FFE971B5F91A516279D9" box="[1168,1302,1737,1761]" italics="true" pageId="21" pageNumber="234">Z. flavorufus</emphasis>
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Cam.
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</taxonomicName>
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/
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<typeStatus id="54EF8815F119FFE9763FF939571B7839" box="[794,879,1769,1793]" pageId="21" pageNumber="234" type="syntype">Syntype</typeStatus>
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/
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<collectorName id="26A15361F119FFE976A3F93A50027839" box="[902,1142,1769,1793]" pageId="21" pageNumber="234">M. Cameron. Bequest.</collectorName>
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B.M. 1955-147 /
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<typeStatus id="54EF8815F119FFE97017F93A5720781A" pageId="21" pageNumber="234" type="lectotype">Lectotypus</typeStatus>
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♂
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<taxonomicName id="4C544D34F119FFE97657F8DA50F27819" authority="Cameron" authorityName="Cameron" authorityYear="1939" box="[882,1158,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="flavorufus">
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||
<location id="8E8B606CF119FFE97657F8DA50F27819" LSID="urn:lsid:plazi:treatment:03FD87A1F119FFEA761AFA23508B7C97:8E8B606CF119FFE97657F8DA50F27819" box="[882,1158,1801,1825]" municipality="Voet" name="Zyras flavorufus Cameron" pageId="21" pageNumber="234" stateProvince="Java">
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<emphasis id="B920EAA5F119FFE97657F8DA506F7819" box="[882,1051,1801,1825]" italics="true" pageId="21" pageNumber="234">Zyras flavorufus</emphasis>
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Cameron
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</location>
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</taxonomicName>
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, desig.
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<determinerName id="7E37DA73F119FFE971F4F8DA51477819" box="[1233,1331,1801,1825]" pageId="21" pageNumber="234">
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V.
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<bibRefCitation id="EFC54B46F119FFE971C9F8D951477819" author="ASSING, V." box="[1260,1331,1801,1825]" pageId="21" pageNumber="234" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
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</determinerName>
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2017” (
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<collectionCode id="ED45AE72F119FFE97604F8F957037879" box="[801,887,1833,1857]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="21" pageNumber="234">BMNH</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF119FFEA763FF8B956A77EB9" lastPageId="22" lastPageNumber="235" pageId="21" pageNumber="234" type="discussion">
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||
<paragraph id="8BEB36B7F119FFEA763FF8B956A77EB9" blockId="21.[794,1399,1897,2081]" lastBlockId="22.[189,794,233,385]" lastPageId="22" lastPageNumber="235" pageId="21" pageNumber="234">
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||
<emphasis id="B920EAA5F119FFE9763FF8B957FD78B9" bold="true" box="[794,905,1897,1921]" pageId="21" pageNumber="234">Comment</emphasis>
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: The original description is based on an unspecified number of
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<typeStatus id="54EF8815F119FFE97106F85A500B789A" box="[1059,1151,1930,1954]" pageId="21" pageNumber="234" type="syntype">syntypes</typeStatus>
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from “G. Tangkoeban Prahoe ... 20.VII.30” (
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<bibRefCitation id="EFC54B46F119FFE9712EF87950B278F9" author="CAMERON, M." box="[1035,1222,1961,1985]" pageId="21" pageNumber="234" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON 1939b</bibRefCitation>
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). The date indicated on the label of the sole specimen found in the Cameron collection differs from that indicated in the description, but it can be assumed that it is one of the specimens that
|
||
<bibRefCitation id="EFC54B46F11AFFEA7440FF39565A7E39" author="CAMERON, M." box="[357,558,233,257]" pageId="22" pageNumber="235" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON (1939b)</bibRefCitation>
|
||
based his description on and that it has type status (identification label with Cameron´s handwriting; collection date six years prior to publication year; characters matching original description). The specimen is designated as the
|
||
<typeStatus id="54EF8815F11AFFEA774EFEB956BB7EB9" box="[619,719,361,385]" pageId="22" pageNumber="235" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF11AFFEA7598FE7951567DD9" pageId="22" pageNumber="235" type="description">
|
||
<paragraph id="8BEB36B7F11AFFEA7598FE79564C7D59" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
<emphasis id="B920EAA5F11AFFEA7598FE7955227EF9" bold="true" box="[189,342,425,449]" pageId="22" pageNumber="235">Redescription</emphasis>
|
||
: Body length
|
||
<quantity id="4CAC9B52F11AFFEA74D4FE7956317EF9" box="[497,581,425,449]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="22" pageNumber="235" unit="mm" value="6.7">6.7 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CAC9B52F11AFFEA7598FE1955657ED9" box="[189,273,457,481]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="22" pageNumber="235" unit="mm" value="3.1">3.1 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="136F2A32F11AFFEA7486FE1955917ED9" box="[419,485,457,481]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="22" pageNumber="235">Figs 8</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F11AFFEA74D1FE1956647ED9" box="[500,528,457,481]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="22" pageNumber="235">27</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F11AFFEA7738FE19564F7ED9" box="[541,571,457,481]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="22" pageNumber="235">42</figureCitation>
|
||
): body reddish with the head and the postero-lateral portions slightly darker; legs dark-yellowish; antennae dark-brown with the basal three antennomeres reddish; maxillary palpi reddish with the apical palpomere yellowish-red.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598FDB956DE7CB9" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
Head (
|
||
<figureCitation id="136F2A32F11AFFEA7426FDB9553A7DB9" box="[259,334,617,641]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="22" pageNumber="235">Fig. 27</figureCitation>
|
||
) moderately transverse; punctation moderately dense and moderately coarse; median dorsal portion extensively impunctate. Eyes moderately large, slightly longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="136F2A32F11AFFEA75E0FD3955757C39" box="[197,257,745,769]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="22" pageNumber="235">Fig. 8</figureCitation>
|
||
)
|
||
<quantity id="4CAC9B52F11AFFEA7431FD3955127C39" box="[276,358,745,769]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="22" pageNumber="235" unit="mm" value="2.1">2.1 mm</quantity>
|
||
long and rather slender; antennomeres IV indistinctly oblong, V as long as broad, VI–X increasingly transverse and of gradually increasing width, X less than 1.5 times as broad as long, and XI of conical shape, shorter than the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598FC5A554F7B79" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
Pronotum (
|
||
<figureCitation id="136F2A32F11AFFEA7467FC5955FB7C99" box="[322,399,905,929]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="22" pageNumber="235">Fig. 27</figureCitation>
|
||
) 1.11 times as broad as long and 1.21 times as broad as head, broadest in anterior half; posterior angles marked; lateral margins distinctly sinuate in posterior half in dorsal view; punctation moderately coarse, rather dense, and slightly irregularly distributed.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598FB9956917BD9" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
Elytra (
|
||
<figureCitation id="136F2A32F11AFFEA742AFB99552E7B59" box="[271,346,1097,1121]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="22" pageNumber="235">Fig. 27</figureCitation>
|
||
) 0.94 times as long as pronotum; punctation moderately dense and moderately coarse, nearly regularly distributed; pubescence dense, not particularly long, and sub-depressed. Hind wings present. Metatarsomere I shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598FB3955E178B9" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
Abdomen (
|
||
<figureCitation id="136F2A32F11AFFEA741EFB3955FC7A39" box="[315,392,1257,1281]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="22" pageNumber="235">Fig. 42</figureCitation>
|
||
) narrower than elytra, with moderately deep anterior impressions on tergites III–V; tergites III–V each with a transverse row of fine, sparse, and weakly defined puncture-like impressions in anterior impressions, with few lateral setiferous punctures on either side, and minute setiferous punctures at posterior margin, disc with scattered micropunctation; tergite VI with a broad transverse band of fine nonsetiferous punctures in anterior portion, with scattered micropunctation on disc, with few lateral setiferous punctures bearing long dark setae on either side and with numerous fine setiferous punctures near posterior margin; tergite VII with a broad band of fine non-setiferous punctation in anterior portion, with non-setiferous micropunctation and larger punctures on remainder of disc, with some lateral setiferous punctures bearing long dark setae on either side, and with fine setiferous punctures near posterior margin, posterior margin with palisade fringe; tergite VIII with numerous setiferous punctures in posterior half, posterior margin convex.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598F85A55D37739" blockId="22.[189,794,425,2081]" pageId="22" pageNumber="235">
|
||
♂: sternite VIII with strongly convex posterior margin; median lobe of aedeagus
|
||
<quantity id="4CAC9B52F11AFFEA74E9F879565F78F9" box="[460,555,1961,1985]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="22" pageNumber="235" unit="mm" value="0.83">0.83 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="136F2A32F11AFFEA7598F819554678D9" box="[189,306,1993,2017]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="22" pageNumber="235">Figs 73–74</figureCitation>
|
||
; paramere
|
||
<quantity id="4CAC9B52F11AFFEA7495F819567B78D9" box="[432,527,1993,2017]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="22" pageNumber="235" unit="mm" value="0.75">0.75 mm</quantity>
|
||
(
|
||
<figureCitation id="136F2A32F11AFFEA7704F819561F78D9" box="[545,619,1993,2017]" captionStart="Figs 72–86" captionStartId="12.[189,229,1971,1993]" captionTargetBox="[198,1434,242,1959]" captionTargetId="figure-132@12.[192,1434,232,1963]" captionTargetPageId="12" captionText="Figs 72–86: Zyras latibasalis (72), Z. flavorufus, lectotype (73–75), Z. wei from Laos (76–77), Z. notaticornis from Laos (78–80), and Z. rutrilobatus (81–86): paramere (72, 75, 80, 83); median lobe of aedeagus in lateral and in ventral view (73–74, 76–79, 81–82); male tergite VIII (84); male sternite VIII (85); postero-median portion of male sternite VIII (86). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="22" pageNumber="235">Fig. 75</figureCitation>
|
||
) long and with very short apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7598F7DA553F7719" blockId="22.[189,794,425,2081]" box="[189,331,2057,2081]" pageId="22" pageNumber="235">♀: unknown.</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7665FF3951567DD9" blockId="22.[832,1437,233,737]" pageId="22" pageNumber="235">
|
||
<emphasis id="B920EAA5F11AFFEA7665FF39507A7E39" bold="true" box="[832,1038,233,257]" pageId="22" pageNumber="235">Comparative notes</emphasis>
|
||
: Based on the very short apical lobe of the paramere, on the shapes of the ventral process and of the internal structures of the aedeagus, and on the punctation pattern of the abdomen,
|
||
<taxonomicName id="4C544D34F11AFFEA71B9FE9A51567E59" authorityName="Cameron" authorityYear="1939" box="[1180,1314,329,353]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="235" phylum="Arthropoda" rank="species" species="flavorufus">
|
||
<emphasis id="B920EAA5F11AFFEA71B9FE9A51567E59" box="[1180,1314,329,353]" italics="true" pageId="22" pageNumber="235">Z. flavorufus</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C544D34F11AFFEA764DFEBA57B67EB9" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[872,962,361,385]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="235" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B920EAA5F11AFFEA764DFEBA57B67EB9" box="[872,962,361,385]" italics="true" pageId="22" pageNumber="235">Z. hirtus</emphasis>
|
||
</taxonomicName>
|
||
group. It is distinguished from most species of this group by the shorter and less dense pubescence of the forebody and of the abdomen. In habitus, coloration, and other external characters,
|
||
<taxonomicName id="4C544D34F11AFFEA7192FE1A51497ED9" authorityName="Cameron" authorityYear="1939" box="[1207,1341,457,481]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="235" phylum="Arthropoda" rank="species" species="flavorufus">
|
||
<emphasis id="B920EAA5F11AFFEA7192FE1A51497ED9" box="[1207,1341,457,481]" italics="true" pageId="22" pageNumber="235">Z. flavorufus</emphasis>
|
||
</taxonomicName>
|
||
is highly similar to
|
||
<taxonomicName id="4C544D34F11AFFEA768FFE3A508C7D39" authority="CAMERON, 1943" authorityName="Cameron" authorityYear="1943" box="[938,1272,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="235" phylum="Arthropoda" rank="species" species="matangensis">
|
||
<emphasis id="B920EAA5F11AFFEA768FFE3A50337D39" box="[938,1095,490,513]" italics="true" pageId="22" pageNumber="235">Z. matangensis</emphasis>
|
||
<bibRefCitation id="EFC54B46F11AFFEA7169FE39508C7D39" author="CAMERON, M." box="[1100,1272,489,513]" pageId="22" pageNumber="235" pagination="139 - 143" refId="ref26208" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D34F11AFFEA700FFE3A578D7D19" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="235" phylum="Arthropoda" rank="species" species="parahirtus">
|
||
<emphasis id="B920EAA5F11AFFEA700FFE3A572B7D19" italics="true" pageId="22" pageNumber="235">Z. parahirtus</emphasis>
|
||
<bibRefCitation id="EFC54B46F11AFFEA7642FDD9578D7D19" author="ASSING, V." box="[871,1017,521,545]" pageId="22" pageNumber="235" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
, both from Borneo. It is distinguished from these species by more distinctly sinuate lateral margins of the pronotum, somewhat more slender antennae with less transverse preapical antennomeres, longer elytra with more distinct punctation, finer and sparser non-setiferous punctation on tergites VI and VII, and by the shape of the median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF11AFFEA7665FCD9508B7C97" pageId="22" pageNumber="235" type="distribution">
|
||
<paragraph id="8BEB36B7F11AFFEA7665FCD950497C79" blockId="22.[832,1436,777,834]" pageId="22" pageNumber="235">
|
||
<emphasis id="B920EAA5F11AFFEA7665FCD957BD7C19" bold="true" box="[832,969,777,801]" pageId="22" pageNumber="235">Distribution</emphasis>
|
||
: This species has been recorded only from the
|
||
<typeStatus id="54EF8815F11AFFEA7642FCFA57E17C7A" box="[871,917,810,834]" pageId="22" pageNumber="235">type</typeStatus>
|
||
locality in
|
||
<collectingRegion id="4990F855F11AFFEA712EFCFA504D7C79" box="[1035,1081,810,833]" country="Indonesia" name="Jawa Timur" pageId="22" pageNumber="235">Java</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11AFFEA7640FC41508B7C97" blockId="22.[869,1279,913,943]" box="[869,1279,913,943]" pageId="22" pageNumber="235">3.3.2 Southeastern continental Asia</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |