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<document id="8BA127CEE9419692260B5D247834D8E1" ID-CLB-Dataset="106789" ID-DOI="10.11646/zootaxa.5169.1.1" ID-GBIF-Dataset="b76236d0-0336-4d17-804a-80f0344b097e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911151" ID-ZooBank="3CB49021-90C5-46F0-AAAF-619EE8068F42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658900973851" checkinUser="plazi" docAuthor="Grall, Elena &amp; Jäger, Peter" docDate="2022" docId="03FD8792583DFFD1FF56F938838F46B9" docLanguage="en" docName="zootaxa.5169.1.1.pdf" docOrigin="Zootaxa 5169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tiomaniella ladam Grall &amp; Jäger 2022, spec. nov." docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="FFC4FFEA5832FFC5FFC1FFA480644008" masterDocTitle="Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="16" updateTime="1699362128362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C9D0F49A3D76601F5667A440A622CB7C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CE6EB4A984260157EC2D1969F8AB2630">
<mods:title id="DC0814CE16B2186F25216D56F0C0B9FD">Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)</mods:title>
</mods:titleInfo>
<mods:name id="F9A187170BCF95D7781762F372CFEF1C" type="personal">
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<mods:roleTerm id="B905B709460DAA402EAE74408470A268">Author</mods:roleTerm>
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<mods:namePart id="1C1D47F2F28BAB9DD9741B038C0548AD">Grall, Elena</mods:namePart>
<mods:nameIdentifier id="7BABEF782A81CF33314E24935F468457" type="ORCID">0000-0002-8938-4105</mods:nameIdentifier>
<mods:affiliation id="E84DCC1A1D7BD71875F540AD796BA8C1">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; Institute of Ecology, Diversity and Evolution, Campus Riedberg, Goethe University, 60438 Frankfurt am Main, Germany; elena. grall @ senckenberg. de, https: // orcid. org / 0000 - 0002 - 8938 - 4105</mods:affiliation>
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<mods:namePart id="0A0AD05CAAFC5F6B3DFA8A8FA0A28207">Jäger, Peter</mods:namePart>
<mods:nameIdentifier id="7CB3F02D8628F001D96BA8464F98A336" type="ORCID">0000-0003-1563-0147</mods:nameIdentifier>
<mods:affiliation id="C2C6A57B9486541D03C0E5B94C501F6B">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; peter. jaeger @ senckenberg. de, https: // orcid. org / 0000 - 0003 - 1563 - 0147</mods:affiliation>
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<mods:title id="573A452308F5495A8604E74CEF5F9358">Zootaxa</mods:title>
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<mods:part id="A9E020A244109B4427F29FB92D323FB5">
<mods:date id="729901254C2D6797206D52F7C56DDED1">2022</mods:date>
<mods:detail id="B22A2DCD964510AA36A72AFFA6EB7D2D" type="pubDate">
<mods:number id="337BF3A3BE29BB2BD859477B526CFE2E">2022-07-27</mods:number>
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<mods:number id="721C085FBE7F54596C82F75896230778">5169</mods:number>
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<mods:start id="2B28D9972E77FF16A5CB6E7694CBC831">1</mods:start>
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<mods:classification id="1EABAC19BA815CB9F641142701DBC5EB">journal article</mods:classification>
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<mods:identifier id="52185FFE355EEDE3593FF9276EFAA574" type="DOI">10.11646/zootaxa.5169.1.1</mods:identifier>
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<mods:identifier id="E48C04B48B4D7D1158A21879AF53BF75" type="Zenodo-Dep">6911151</mods:identifier>
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<treatment id="03FD8792583DFFD1FF56F938838F46B9" ID-DOI="http://doi.org/10.5281/zenodo.6914322" ID-GBIF-Taxon="197859752" ID-Zenodo-Dep="6914322" LSID="urn:lsid:plazi:treatment:03FD8792583DFFD1FF56F938838F46B9" httpUri="http://treatment.plazi.org/id/03FD8792583DFFD1FF56F938838F46B9" lastPageId="20" lastPageNumber="21" pageId="15" pageNumber="16">
<subSubSection id="C34E650F583DFFCAFF56F938819E46BF" box="[151,506,1692,1719]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BEB3684583DFFCAFF56F938819E46BF" blockId="15.[151,506,1692,1755]" box="[151,506,1692,1719]" pageId="15" pageNumber="16">
<heading id="D0A381E8583DFFCAFF56F938819E46BF" bold="true" box="[151,506,1692,1719]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B920EA96583DFFCAFF56F938819E46BF" bold="true" box="[151,506,1692,1719]" pageId="15" pageNumber="16">
<taxonomicName id="4C544D07583DFFCAFF56F938811F46BF" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[151,379,1692,1719]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ladam" status="sp. nov.">
<emphasis id="B920EA96583DFFCAFF56F938811F46BF" bold="true" box="[151,379,1692,1719]" italics="true" pageId="15" pageNumber="16">Tiomaniella ladam</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357ED583DFFCAFE43F939819E46BF" box="[386,506,1693,1719]" pageId="15" pageNumber="16" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF56F965810C46D3" box="[151,360,1729,1755]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEB3684583DFFCAFF56F965810C46D3" blockId="15.[151,506,1692,1755]" box="[151,360,1729,1755]" pageId="15" pageNumber="16">
<figureCitation id="136F2A01583DFFCAFF56F965817346D3" box="[151,279,1729,1755]" captionStart-0="FIGURES 66–69" captionStart-1="FIGURES 70–72" captionStart-2="FIGURES 73–81" captionStartId-0="16.[151,264,1679,1703]" captionStartId-1="17.[151,264,1831,1855]" captionStartId-2="18.[151,264,1842,1866]" captionTargetBox-0="[151,1420,464,1644]" captionTargetBox-1="[151,1436,309,1808]" captionTargetBox-2="[151,1436,536,1819]" captionTargetId-0="figure-135@16.[151,1436,452,1655]" captionTargetId-1="figure-101@17.[151,1436,309,1808]" captionTargetId-2="figure-169@18.[151,1436,525,1819]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURES 66–69. Tiomaniella ladam gen. &amp; spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis." captionText-1="FIGURES 70–72. Tiomaniella ladam gen. &amp; spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca." captionText-2="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi-0="http://doi.org/10.5281/zenodo.6911193" figureDoi-1="http://doi.org/10.5281/zenodo.6911197" figureDoi-2="http://doi.org/10.5281/zenodo.6911201" httpUri-0="https://zenodo.org/record/6911193/files/figure.png" httpUri-1="https://zenodo.org/record/6911197/files/figure.png" httpUri-2="https://zenodo.org/record/6911201/files/figure.png" pageId="15" pageNumber="16">Figs 66–81</figureCitation>
,
<figureCitation id="136F2A01583DFFCAFEE3F965810C46D3" box="[290,360,1729,1755]" captionStart-0="FIGURES 82–97" captionStart-1="FIGURE 98" captionStartId-0="19.[151,264,1899,1923]" captionStartId-1="20.[151,250,1436,1460]" captionTargetBox-0="[152,1435,182,1873]" captionTargetBox-1="[151,1436,956,1413]" captionTargetId-0="figure-19@19.[151,1436,181,1875]" captionTargetId-1="figure-372@20.[151,1436,956,1413]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." captionText-1="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi-0="http://doi.org/10.5281/zenodo.6911203" figureDoi-1="http://doi.org/10.5281/zenodo.6911205" httpUri-0="https://zenodo.org/record/6911203/files/figure.png" httpUri-1="https://zenodo.org/record/6911205/files/figure.png" pageId="15" pageNumber="16">94–98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF56F8AD808D47F3" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BEB3684583DFFCAFF56F8AD808D47F3" blockId="15.[151,1436,1800,2043]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF56F8AD812C472B" bold="true" box="[151,328,1801,1827]" pageId="15" pageNumber="16">Type material.</emphasis>
<materialsCitation id="3B3C3CD9583DFFCAFE90F8AD83FE472B" ID-GBIF-Occurrence="3866517344" box="[337,922,1800,1827]" collectionCode="SD" country="Malaysia" location="Pahang" pageId="15" pageNumber="16" specimenCode="SD 1471" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<emphasis id="B920EA96583DFFCAFE90F8AD82B0472B" bold="true" box="[337,724,1800,1827]" pageId="15" pageNumber="16">
<collectingCountry id="F3437614583DFFCAFE90F8AD8183472B" box="[337,487,1801,1827]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
:
<emphasis id="B920EA96583DFFCAFE3BF8AC8230472B" bold="true" box="[506,596,1800,1827]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="4990F866583DFFCAFE3BF8AC8230472B" box="[506,596,1800,1827]" country="Malaysia" name="Pahang" pageId="15" pageNumber="16">Pahang</collectingRegion>
</emphasis>
:
<typeStatus id="54EF8826583DFFCAFDA6F8AD82B0472B" box="[615,724,1801,1827]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<specimenCode id="DBF29EFF583DFFCAFCE4F8AD83F5472B" box="[805,913,1801,1827]" collectionCode="SD" pageId="15" pageNumber="16">SD 1471</specimenCode>
)
</materialsCitation>
:
<materialsCitation id="3B3C3CD9583DFFCAFC66F8AD8491474F" ID-GBIF-Occurrence="3866517304" collectingDate="2001-10-02" collectionCode="MHNG" collectorName="L. Monod" country="Malaysia" elevation="160" latitude="2.78635" location="Gunung" longLatPrecision="1" longitude="104.13153" municipality="Pulau Tioman" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Pahang" typeStatus="holotype">
<collectingMunicipality id="6B8FACFE583DFFCAFC66F8AD842D472B" box="[935,1097,1801,1827]" pageId="15" pageNumber="16">Pulau Tioman</collectingMunicipality>
, pied du
<location id="8E8B605F583DFFCAFB77F8AD8577472B" LSID="urn:lsid:plazi:treatment:03FD8792583DFFD1FF56F938838F46B9:8E8B605F583DFFCAFB77F8AD8577472B" box="[1206,1299,1801,1827]" country="Malaysia" latitude="2.78635" longLatPrecision="1" longitude="104.13153" municipality="Pulau Tioman" name="Gunung" pageId="15" pageNumber="16" stateProvince="Pahang">Gunung</location>
, rain forest,
<geoCoordinate id="EE605043583DFFCAFF56F889814F474F" box="[151,299,1837,1863]" degrees="02" direction="north" minutes="47.181" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="2.78635">N02°47.181’</geoCoordinate>
,
<geoCoordinate id="EE605043583DFFCAFEFAF88981BD474F" box="[315,473,1837,1863]" degrees="104" direction="east" minutes="07.892" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="104.13153">E104°07.892’</geoCoordinate>
,
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</quantity>
,
<date id="FFEA1044583DFFCAFD84F8898299474F" box="[581,765,1837,1863]" pageId="15" pageNumber="16" value="2001-10-02">
<collectingDate id="EFAEE9AC583DFFCAFD84F8898299474F" box="[581,765,1837,1863]" pageId="15" pageNumber="16" value="2001-10-02">2 October 2001</collectingDate>
</date>
,
<collectorName id="26A15352583DFFCAFCCDF88983E7474F" box="[780,899,1837,1863]" pageId="15" pageNumber="16">L. Monod</collectorName>
, field number TM-17 (
<collectionCode id="ED45AE41583DFFCAFB51F8898488474F" box="[1168,1260,1837,1863]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CD9583DFFCAFAC2F88982344763" ID-GBIF-Occurrence="3866517324" collectionCode="SD" country="Malaysia" pageId="15" pageNumber="16" specimenCode="SD 1472" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Pahang" typeStatus="paratype">
<emphasis id="B920EA96583DFFCAFAC2F88981074762" bold="true" pageId="15" pageNumber="16">
<typeStatus id="54EF8826583DFFCAFAC2F889851F474F" box="[1283,1403,1837,1863]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D52FD0D583DFFCAFA4FF88880B64763" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<specimenCount id="9D52FD0D583DFFCAFF21F8F581334762" box="[224,343,1872,1899]" pageId="15" pageNumber="16" type="female">3 females</specimenCount>
):
</emphasis>
<specimenCount id="9D52FD0D583DFFCAFEADF8F581AA4763" box="[364,462,1873,1899]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
(
<specimenCode id="DBF29EFF583DFFCAFE1EF8F5822D4763" box="[479,585,1873,1899]" collectionCode="SD" pageId="15" pageNumber="16">SD 1472</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3C3CD9583DFFCAFD9CF8F584544763" ID-GBIF-Occurrence="3866517311" box="[605,1072,1873,1899]" collectingDate="2001-10-02" collectionCode="MHNG" collectorName="L. Monod" country="Malaysia" elevation="160" latitude="2.78635" location="Gunung" longLatPrecision="1" longitude="104.13153" municipality="Pulau Tioman" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
with same data as for holotype (
<collectionCode id="ED45AE41583DFFCAFC0DF8F5844C4763" box="[972,1064,1873,1899]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CD9583DFFCAFBFDF8F5816A4787" ID-GBIF-Occurrence="3866517308" collectionCode="SD" country="Malaysia" location="Pahang" pageId="15" pageNumber="16" specimenCode="SD 1474" specimenCount="1" specimenCount-male="1" stateProvince="Pahang" typeStatus="paratype">
<emphasis id="B920EA96583DFFCAFBFDF8F585234763" bold="true" box="[1084,1351,1872,1899]" pageId="15" pageNumber="16">
<collectingCountry id="F3437614583DFFCAFBFDF8F584B64763" box="[1084,1234,1873,1899]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
:
<emphasis id="B920EA96583DFFCAFB22F8F485594763" bold="true" box="[1251,1341,1872,1899]" italics="true" pageId="15" pageNumber="16">
<collectingRegion id="4990F866583DFFCAFB22F8F485594763" box="[1251,1341,1872,1899]" country="Malaysia" name="Pahang" pageId="15" pageNumber="16">Pahang</collectingRegion>
</emphasis>
:
</emphasis>
<specimenCount id="9D52FD0D583DFFCAFA8EF8F585F84763" box="[1359,1436,1873,1899]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
(
<specimenCode id="DBF29EFF583DFFCAFF5EF8D181634787" box="[159,263,1909,1935]" collectionCode="SD" pageId="15" pageNumber="16">SD 1474</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3C3CD9583DFFCAFED8F8D182F647BB" ID-GBIF-Occurrence="3866517338" collectingDate="2001-06-28" collectionCode="SMF" collectorName="A. Schulz &amp; K. Vock" country="Malaysia" elevation="100" latitude="2.7833333" location="Westside" longLatPrecision="1307" longitude="104.13333" municipality="Tioman Island" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
<collectingMunicipality id="6B8FACFE583DFFCAFED8F8D181D94787" box="[281,445,1909,1935]" pageId="15" pageNumber="16">Tioman Island</collectingMunicipality>
,
<location id="8E8B605F583DFFCAFE06F8D1824A4787" LSID="urn:lsid:plazi:treatment:03FD8792583DFFD1FF56F938838F46B9:8E8B605F583DFFCAFE06F8D1824A4787" box="[455,558,1909,1935]" country="Malaysia" latitude="2.7833333" longLatPrecision="1307" longitude="104.13333" municipality="Tioman Island" name="Westside" pageId="15" pageNumber="16" stateProvince="Pahang">Westside</location>
,
<locationDeviation id="9A592FA6583DFFCAFDF8F8D1832D4787" box="[569,841,1909,1935]" location="Kg. Genting" pageId="15" pageNumber="16">
<quantity id="4CAC9B61583DFFCAFDF8F8D182154787" box="[569,625,1909,1935]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="15" pageNumber="16" unit="km" value="2.0">2 km</quantity>
SE of Kg. Genting
</locationDeviation>
,
<geoCoordinate id="EE605043583DFFCAFC92F8D183D34787" box="[851,951,1909,1935]" degrees="02" direction="north" minutes="47" orientation="latitude" pageId="15" pageNumber="16" precision="925" value="2.7833333">N02°47’</geoCoordinate>
,
<geoCoordinate id="EE605043583DFFCAFC03F8D184544787" box="[962,1072,1909,1935]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="15" pageNumber="16" precision="925" value="104.13333">E104°08’</geoCoordinate>
,
<quantity id="4CAC9B61583DFFCAFBFAF8D184E74787" box="[1083,1155,1909,1935]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="100.0">
<elevation id="0079D1B7583DFFCAFBFAF8D184E74787" box="[1083,1155,1909,1935]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="16" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FFEA1044583DFFCAFB4FF8D185414787" box="[1166,1317,1909,1935]" pageId="15" pageNumber="16" value="2001-06-28">
<collectingDate id="EFAEE9AC583DFFCAFB4FF8D185414787" box="[1166,1317,1909,1935]" pageId="15" pageNumber="16" value="2001-06-28">28 June 2001</collectingDate>
</date>
,
<collectorName id="26A15352583DFFCAFAEFF8D185F84787" box="[1326,1436,1909,1935]" pageId="15" pageNumber="16">A. Schulz</collectorName>
&amp;
<collectorName id="26A15352583DFFCAFF75F83D816A47BB" box="[180,270,1945,1971]" pageId="15" pageNumber="16">K. Vock</collectorName>
leg., field number M01-71 (
<collectionCode id="ED45AE41583DFFCAFD8DF83D82EE47BB" box="[588,650,1945,1971]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="15" pageNumber="16">SMF</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD9583DFFCAFD5CF83D82BE47DF" ID-GBIF-Occurrence="3866517305" collectingDate="2001-06-26" collectionCode="MHNG" collectorName="A. Schulz &amp; K. Vock" country="Malaysia" elevation="100" latitude="2.7833333" location="Westside" longLatPrecision="1307" longitude="104.13333" municipality="Tioman Island" pageId="15" pageNumber="16" specimenCount="3" specimenCount-female="3" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="9D52FD0D583DFFCAFD5CF83D836D47BB" box="[669,777,1945,1971]" pageId="15" pageNumber="16" type="female">2 females</specimenCount>
with same data as for previous specimen, except:
<specimenCount id="9D52FD0D583DFFCAFAFAF83D85F847BB" box="[1339,1436,1945,1971]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
<date id="FFEA1044583DFFCAFF56F819814A47DF" box="[151,302,1981,2007]" pageId="15" pageNumber="16" value="2001-06-26">
<collectingDate id="EFAEE9AC583DFFCAFF56F819814A47DF" box="[151,302,1981,2007]" pageId="15" pageNumber="16" value="2001-06-26">26 June 2001</collectingDate>
</date>
with field number M01–39 (
<collectionCode id="ED45AE41583DFFCAFDB7F81982B647DF" box="[630,722,1981,2007]" country="Switzerland" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="16" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD9583DFFCAFD27F81983A147DF" ID-GBIF-Occurrence="3866517340" box="[742,965,1981,2007]" collectionCode="SD" country="Malaysia" pageId="15" pageNumber="16" specimenCode="SD 1473" specimenCount="1" specimenCount-female="1" stateProvince="Pahang" typeStatus="paratype">
<specimenCount id="9D52FD0D583DFFCAFD27F819832347DF" box="[742,839,1981,2007]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
(
<specimenCode id="DBF29EFF583DFFCAFC96F81983D947DF" box="[855,957,1981,2007]" collectionCode="SD" pageId="15" pageNumber="16">SD 1473</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B3C3CD9583DFFCAFC0CF819808147F3" ID-GBIF-Occurrence="3866517316" collectingDate="2001-06-26" collectionCode="SMF" country="Malaysia" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Pahang" typeStatus="paratype">
<date id="FFEA1044583DFFCAFC0CF819840047DF" box="[973,1124,1981,2007]" pageId="15" pageNumber="16" value="2001-06-26">
<collectingDate id="EFAEE9AC583DFFCAFC0CF819840047DF" box="[973,1124,1981,2007]" pageId="15" pageNumber="16" value="2001-06-26">26 June 2001</collectingDate>
</date>
with field number M01–40 (
<collectionCode id="ED45AE41583DFFCAFF5EF84580B947F3" box="[159,221,2017,2043]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="15" pageNumber="16">SMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5822FFD5FF06FF32833F40DC" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BEB36845822FFD5FF06FF32833F40DC" blockId="16.[151,1437,150,429]" pageId="16" pageNumber="17">
<emphasis id="B920EA965822FFD5FF06FF32812A40B8" bold="true" box="[199,334,150,176]" pageId="16" pageNumber="17">Etymology.</emphasis>
The species name refers to the Malayan word “ladam” meaning “horse shoe” and refers to the brown U-shaped patterning of the prosoma; term in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5822FFD5FF06FF7A81CA41A5" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BEB36845822FFD5FF06FF7A81CA41A5" blockId="16.[151,1437,150,429]" pageId="16" pageNumber="17">
<emphasis id="B920EA965822FFD5FF06FF7A812640F0" bold="true" box="[199,322,222,248]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
<taxonomicName id="4C544D075822FFD5FE86FF7A827D40F0" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[327,537,222,248]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ladam" status="sp. nov.">
<emphasis id="B920EA965822FFD5FE86FF7A827D40F0" box="[327,537,222,248]" italics="true" pageId="16" pageNumber="17">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA965822FFD5FDDEFF7A82F440F0" bold="true" box="[543,656,222,248]" pageId="16" pageNumber="17">
<taxonomicNameLabel id="A21357ED5822FFD5FDDEFF7A82F440F0" box="[543,656,222,248]" pageId="16" pageNumber="17" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
may be recognised by the following combination of characters: Males (
<figureCitation id="136F2A015822FFD5FF5EFEA681404114" box="[159,292,258,284]" captionStart="FIGURES 66–69" captionStartId="16.[151,264,1679,1703]" captionTargetBox="[151,1420,464,1644]" captionTargetId="figure-135@16.[151,1436,452,1655]" captionTargetPageId="16" captionText="FIGURES 66–69. Tiomaniella ladam gen. &amp; spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911193" httpUri="https://zenodo.org/record/6911193/files/figure.png" pageId="16" pageNumber="17">Figs 66–69</figureCitation>
): 1. Distinct, apically fanned conductor homologue arising in 1-o’clock-position from tegulum, 2. TA arising centrally, with broad base and retrolatero-distad acute tip, 3. Dorsal branch of RTA as long as ventral branch (best seen in retrolateral view). Females (
<figureCitation id="136F2A015822FFD5FDAAFEEE8295416C" box="[619,753,330,356]" captionStart="FIGURES 70–72" captionStartId="17.[151,264,1831,1855]" captionTargetBox="[151,1436,309,1808]" captionTargetId="figure-101@17.[151,1436,309,1808]" captionTargetPageId="17" captionText="FIGURES 70–72. Tiomaniella ladam gen. &amp; spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911197" httpUri="https://zenodo.org/record/6911197/files/figure.png" pageId="16" pageNumber="17">Figs 70–72</figureCitation>
): 1. Epigyne with epigynal pockets running from laterally to postero-medially, 2. Lateral lobes extending not in anterior half of epigyne and 3. Internal duct system only fused anteriorly at ventral part.
</paragraph>
</subSubSection>
<caption id="DF2B660C5822FFD5FF56F92B8203471B" ID-DOI="http://doi.org/10.5281/zenodo.6911193" ID-Zenodo-Dep="6911193" httpUri="https://zenodo.org/record/6911193/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,264,1679,1703]" targetBox="[151,1420,464,1644]" targetPageId="16">
<paragraph id="8BEB36845822FFD5FF56F92B8203471B" blockId="16.[151,1437,1679,1811]" pageId="16" pageNumber="17">
<emphasis id="B920EA965822FFD5FF56F92B813246AF" bold="true" box="[151,342,1679,1703]" pageId="16" pageNumber="17">FIGURES 66–69.</emphasis>
<taxonomicName id="4C544D075822FFD5FE9FF92B827A46AF" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[350,542,1679,1703]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ladam">
<emphasis id="B920EA965822FFD5FE9FF92B827A46AF" box="[350,542,1679,1703]" italics="true" pageId="16" pageNumber="17">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA965822FFD5FDE6F92B828646AF" bold="true" box="[551,738,1679,1703]" pageId="16" pageNumber="17">gen. &amp; spec. nov.</emphasis>
holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis.
</paragraph>
</caption>
<subSubSection id="C34E650F5822FFD1FF06F8E5840B4685" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEB36845822FFD4FF06F8E583364115" blockId="16.[151,1436,1857,2028]" lastBlockId="17.[151,1437,151,285]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B920EA965822FFD5FF06F8E582404753" bold="true" box="[199,548,1857,1883]" pageId="16" pageNumber="17">
Description. Male (
<typeStatus id="54EF88265822FFD5FE6BF8E5827C4753" box="[426,536,1857,1883]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
):
</emphasis>
TL 5.3; PL 2.7, PW 2.5, AW 1.3; OL 2.6, OW 1.6. Eye measurements (
<figureCitation id="136F2A015822FFD5FAFAF8E585EB4753" box="[1339,1423,1857,1883]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="16" pageNumber="17">Fig. 73</figureCitation>
): AME 0.13; ALE 0.20; PME 0.11; PLE 0.25; AME–AME 0.13; AME–ALE 0.05; PME–PME 0.21; PME–PLE 0.26; AME–PME 0.20; ALE–PLE 0.21; clypeus AME 0.16; clypeus ALE 0.14. Leg formula: 2431. Measurements of palp and legs: palp: 4.00 (1.10, 0.70, 0.80, 1.40); I: 8.65 (2.40, 1.10, 2.20, 2.10, 0.85); II: 10.70 (3.00, 1.30, 2.90, 2.60, 0.90); III: 8.95 (2.60, 1.15, 2.30, 2.10, 0.80); IV: 9.20 (2.60, 1.00, 2.40, 2.40, 0.80). Spination: palp: 131, 101, 2111; legs: Fe I 323, Fe II &amp; III 222, Fe IV 221; Pa I–IV 000; Ti I &amp;
<date id="FFEA10445823FFD4FC9AFF3383D440B9" box="[859,944,151,177]" pageId="17" pageNumber="18" value="1026-02">II 1026</date>
, Ti III &amp;
<date id="FFEA10445823FFD4FBDBFF33841E40B9" box="[1050,1146,151,177]" pageId="17" pageNumber="18" value="2126-04">IV 2126</date>
; Mt I &amp;
<date id="FFEA10445823FFD4FB1AFF33855440B9" box="[1243,1328,151,177]" pageId="17" pageNumber="18" value="1026-02">II 1026</date>
, Mt III &amp; Mt IV 3036. Cheliceral furrow with 3 promarginal, 5 retromarginal teeth, 16 intermarginal denticles and 1 escort seta (
<figureCitation id="136F2A015823FFD4FF10FF7B814140F1" box="[209,293,223,249]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="17" pageNumber="18">Fig. 78</figureCitation>
). Retrolateral claw of right leg I with 11 teeth (
<figureCitation id="136F2A015823FFD4FCECFF7B83E540F1" box="[813,897,222,249]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="17" pageNumber="18">Fig. 75</figureCitation>
). Trilobate membrane of left leg I with short and narrow median hook, shorter than lateral projections (
<figureCitation id="136F2A015823FFD4FD33FEA783224115" box="[754,838,259,285]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="17" pageNumber="18">Fig. 80</figureCitation>
).
</paragraph>
<caption id="DF2B660C5823FFD4FF56F8838415478F" ID-DOI="http://doi.org/10.5281/zenodo.6911197" ID-Zenodo-Dep="6911197" httpUri="https://zenodo.org/record/6911197/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1831,1855]" targetBox="[151,1436,309,1808]" targetPageId="17">
<paragraph id="8BEB36845823FFD4FF56F8838415478F" blockId="17.[151,1437,1831,1927]" pageId="17" pageNumber="18">
<emphasis id="B920EA965823FFD4FF56F88381314737" bold="true" box="[151,341,1831,1855]" pageId="17" pageNumber="18">FIGURES 70–72.</emphasis>
<taxonomicName id="4C544D075823FFD4FE9DF883827F4737" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[348,539,1831,1855]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ladam">
<emphasis id="B920EA965823FFD4FE9DF883827F4737" box="[348,539,1831,1855]" italics="true" pageId="17" pageNumber="18">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA965823FFD4FDE3F88382BD4737" bold="true" box="[546,729,1831,1855]" pageId="17" pageNumber="18">gen. &amp; spec. nov.</emphasis>
female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca.
</paragraph>
</caption>
<paragraph id="8BEB36845820FFD7FF06FF3384054149" blockId="18.[151,1437,151,501]" pageId="18" pageNumber="19">
Palp as in diagnosis (
<figureCitation id="136F2A015820FFD7FE75FF33825C40B9" box="[436,568,151,177]" captionStart="FIGURES 66–69" captionStartId="16.[151,264,1679,1703]" captionTargetBox="[151,1420,464,1644]" captionTargetId="figure-135@16.[151,1436,452,1655]" captionTargetPageId="16" captionText="FIGURES 66–69. Tiomaniella ladam gen. &amp; spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911193" httpUri="https://zenodo.org/record/6911193/files/figure.png" pageId="18" pageNumber="19">Figs 66–69</figureCitation>
). RTA arising subdistally from tibia, with two branches, both latter connected with a concave margin. Cymbium roughly twice as long as tibia, with enormous retrolateral bulge and small retroproximal projection. Subtegulum sail-shaped, covers half of the embolus length in ventral view, tip filiform elongated parallel to embolus tip. Median part of tegulum bulging prolaterally. Spermophor distinctly S-shaped. Embolus basally wide, then filiform, running a semicircle, its tip close to conductor homologue.
</paragraph>
<paragraph id="8BEB36845820FFD7FF06FEEF83AB41FD" blockId="18.[151,1437,151,501]" pageId="18" pageNumber="19">
Colouration in ethanol (
<figureCitation id="136F2A015820FFD7FE17FEEF8238416D" box="[470,604,330,357]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="18" pageNumber="19">Figs 94–95</figureCitation>
): Prosoma yellowish-brown, latero-posteriorly with brown U-shaped pattern, medially with brown longitudinal stripe, lateral margin with three dark brown dots. Sternum yellow. Opisthosoma grey with brown spots, medially with grey longitudinal stripe, posteriorly with grey dot, ventral part grey, laterally with brown stripes. Chelicerae yellowish-brown. Coxa dorsally with brown spots. Palps yellowish-brown. Legs yellowish-brown; Ti distally reddish-brow; Mt &amp; Ta reddish-brown.
</paragraph>
<caption id="DF2B660C5820FFD7FF56F89682F347BE" ID-DOI="http://doi.org/10.5281/zenodo.6911201" ID-Zenodo-Dep="6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,264,1842,1866]" targetBox="[151,1436,536,1819]" targetPageId="18">
<paragraph id="8BEB36845820FFD7FF56F89682F347BE" blockId="18.[151,1437,1842,1975]" pageId="18" pageNumber="19">
<emphasis id="B920EA965820FFD7FF56F89681374742" bold="true" box="[151,339,1842,1866]" pageId="18" pageNumber="19">FIGURES 73–81.</emphasis>
<taxonomicName id="4C544D075820FFD7FE98F89682724742" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[345,534,1842,1866]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ladam">
<emphasis id="B920EA965820FFD7FE98F89682724742" box="[345,534,1842,1866]" italics="true" pageId="18" pageNumber="19">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA965820FFD7FDDDF89782AB4743" bold="true" box="[540,719,1842,1867]" pageId="18" pageNumber="19">gen. &amp; spec. nov.</emphasis>
holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left).
</paragraph>
</caption>
<caption id="DF2B660C5821FFD6FF56F8CF84AF47C3" ID-DOI="http://doi.org/10.5281/zenodo.6911203" ID-Zenodo-Dep="6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,264,1899,1923]" targetBox="[152,1435,182,1873]" targetPageId="19">
<paragraph id="8BEB36845821FFD6FF56F8CF84AF47C3" blockId="19.[151,1437,1899,1995]" pageId="19" pageNumber="20">
<emphasis id="B920EA965821FFD6FF56F8CF8131478B" bold="true" box="[151,341,1899,1923]" pageId="19" pageNumber="20">FIGURES 82–97.</emphasis>
<taxonomicName id="4C544D075821FFD6FE9DF8CF8197478B" authorityName="Thorell" authorityYear="1873" box="[348,499,1899,1923]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">
<emphasis id="B920EA965821FFD6FE9DF8CF8197478B" box="[348,499,1899,1923]" italics="true" pageId="19" pageNumber="20">Heteropodinae</emphasis>
</taxonomicName>
<emphasis id="B920EA965821FFD6FE3BF8CF82CD478B" bold="true" box="[506,681,1899,1923]" pageId="19" pageNumber="20">gen. &amp; spp. nov.</emphasis>
, habitus (dorsal and ventral, respectively). 82–85
<taxonomicName id="4C544D075821FFD6FB6AF8CF8528478B" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[1195,1356,1899,1923]" class="Arachnida" family="Sparassidae" genus="Borniella" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="B920EA965821FFD6FB6AF8CF8528478B" box="[1195,1356,1899,1923]" italics="true" pageId="19" pageNumber="20">Borniella parva</emphasis>
</taxonomicName>
<emphasis id="B920EA965821FFD6FA95F8CF809947AF" bold="true" pageId="19" pageNumber="20">gen. &amp; spec. nov.</emphasis>
; 86–89
<taxonomicName id="4C544D075821FFD6FE8EF82B81BE47AF" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[335,474,1935,1959]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kecil">
<emphasis id="B920EA965821FFD6FE8EF82B81BE47AF" box="[335,474,1935,1959]" italics="true" pageId="19" pageNumber="20">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA965821FFD6FE1EF82B82F547AF" bold="true" box="[479,657,1935,1959]" pageId="19" pageNumber="20">gen. &amp; spec. nov.</emphasis>
; 90–93
<taxonomicName id="4C544D075821FFD6FD23F82B83C047AF" box="[738,932,1935,1959]" class="Arachnida" family="Sparassidae" genus="Micropoda" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="davisae">
<emphasis id="B920EA965821FFD6FD23F82B83C047AF" box="[738,932,1935,1959]" italics="true" pageId="19" pageNumber="20">Micropoda davisae</emphasis>
</taxonomicName>
<emphasis id="B920EA965821FFD6FC6BF82B843847AF" bold="true" box="[938,1116,1935,1959]" pageId="19" pageNumber="20">gen. &amp; spec. nov.</emphasis>
; 94–97
<taxonomicName id="4C544D075821FFD6FB6CF82B850E47AF" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[1197,1386,1935,1959]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ladam">
<emphasis id="B920EA965821FFD6FB6CF82B850E47AF" box="[1197,1386,1935,1959]" italics="true" pageId="19" pageNumber="20">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA965821FFD6FAB1F82B817E47C3" bold="true" pageId="19" pageNumber="20">gen. &amp; spec. nov.</emphasis>
(82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype).
</paragraph>
</caption>
<paragraph id="8BEB36845826FFD1FF06FF3281E041FC" blockId="20.[151,1437,150,933]" pageId="20" pageNumber="21">
<emphasis id="B920EA965826FFD1FF06FF3281CF40B8" bold="true" box="[199,427,150,176]" pageId="20" pageNumber="21">
Female (
<typeStatus id="54EF88265826FFD1FEEDFF3281FA40B8" box="[300,414,150,176]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
):
</emphasis>
TL 6.0; PL 2.6,
<specimenCode id="DBF29EFF5826FFD1FD9FFF3382D640B9" box="[606,690,150,177]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="20" pageNumber="21">PW 2.5</specimenCode>
,
<specimenCode id="DBF29EFF5826FFD1FD7AFF32837740B9" box="[699,787,150,177]" collectionCode="AW" pageId="20" pageNumber="21">AW 1.5</specimenCode>
;
<specimenCode id="DBF29EFF5826FFD1FCDFFF32830A40B8" box="[798,878,150,176]" collectionCode="OL" pageId="20" pageNumber="21">OL 3.4</specimenCode>
,
<specimenCode id="DBF29EFF5826FFD1FCB6FF3283B540B8" box="[887,977,150,176]" collectionCode="OW" pageId="20" pageNumber="21">OW 2.2</specimenCode>
. Eye measurements (
<figureCitation id="136F2A015826FFD1FB7AFF32856B40B8" box="[1211,1295,150,176]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="20" pageNumber="21">Fig. 74</figureCitation>
): AME 0.16; ALE 0.23; PME 0.14; PLE 0.27; AME–AME 0.12; AME–ALE 0.04; PME–PME 0.19; PME–PLE 0.32; AME– PME 0.21; ALE–PLE 0.23; clypeus AME 0.15; clypeus ALE 0.13.
<collectorName id="26A153525826FFD1FC53FF7B83DA40F1" box="[914,958,223,249]" pageId="20" pageNumber="21">Leg</collectorName>
formula: 2431; measurements of palp and legs: palp: 4.10 (1.10, 0.70, 1.00, 1.30); I: 8.55 (2.30, 1.20, 2.25, 2.00, 0.80); II: 10.20 (2.80, 1.30, 2.70, 2.50, 0.90); III: 9.20 (2.60, 1.20, 2.40, 2.20, 0.80); IV: 9.30 (2.50, 1.10, 2.30, 2.50, 0.90).
<collectorName id="26A153525826FFD1FC31FE8284044148" box="[1008,1120,294,320]" pageId="20" pageNumber="21">Spination</collectorName>
: palp: 131, 101, 2121, 1004; legs:
<collectorName id="26A153525826FFD1FF13FEEF809A416D" box="[210,254,331,357]" pageId="20" pageNumber="21">Fe I</collectorName>
323,
<collectorName id="26A153525826FFD1FEFDFEEF8115416D" box="[316,369,331,357]" pageId="20" pageNumber="21">Fe II</collectorName>
&amp; III 222,
<collectorName id="26A153525826FFD1FE2CFEEF8248416D" box="[493,556,331,357]" pageId="20" pageNumber="21">Fe IV</collectorName>
221;
<collectorName id="26A153525826FFD1FDAAFEEF82DC416D" box="[619,696,331,357]" pageId="20" pageNumber="21">Pa I–V</collectorName>
000;
<collectorName id="26A153525826FFD1FD37FEEE8379416D" box="[758,797,330,357]" pageId="20" pageNumber="21">
Ti
<date id="FFEA10445826FFD1FCD5FEEF8379416D" box="[788,797,331,357]" pageId="20" pageNumber="21" value="1004-01">I</date>
</collectorName>
1004,
<collectorName id="26A153525826FFD1FCA8FEEE83FE416D" box="[873,922,330,357]" pageId="20" pageNumber="21">Ti II</collectorName>
0004,
<collectorName id="26A153525826FFD1FC24FEEE8399416C" box="[997,1021,330,356]" pageId="20" pageNumber="21">Ti</collectorName>
<date id="FFEA10445826FFD1FBC2FEEF8404416C" box="[1027,1120,330,357]" pageId="20" pageNumber="21" value="2026-03">
<collectingDate id="EFAEE9AC5826FFD1FBC2FEEF8404416C" box="[1027,1120,330,357]" pageId="20" pageNumber="21" value="2026-03">III 2026</collectingDate>
</date>
,
<collectorName id="26A153525826FFD1FBABFEEE84E6416C" box="[1130,1154,330,356]" pageId="20" pageNumber="21">Ti</collectorName>
<date id="FFEA10445826FFD1FB49FEEF8483416D" box="[1160,1255,330,357]" pageId="20" pageNumber="21" value="2125-04">
<collectingDate id="EFAEE9AC5826FFD1FB49FEEF8483416D" box="[1160,1255,330,357]" pageId="20" pageNumber="21" value="2125-04">IV 2125</collectingDate>
</date>
;
<collectorName id="26A153525826FFD1FB33FEEF8546416D" box="[1266,1314,331,357]" pageId="20" pageNumber="21">Mt I</collectorName>
&amp; II 0004,
<collectorName id="26A153525826FFD1FF56FECB80DC4181" box="[151,184,367,393]" pageId="20" pageNumber="21">Mt</collectorName>
<date id="FFEA10445826FFD1FF01FECB817B4180" box="[192,287,366,393]" pageId="20" pageNumber="21" value="1006-03">III 1006</date>
, MT IV 3036.
<collectorName id="26A153525826FFD1FE0BFECA82264180" box="[458,578,366,392]" pageId="20" pageNumber="21">Chelicerae</collectorName>
with 3 promarginal, 4 retromarginal teeth, 22 intermarginal denticles and 1 escort seta (
<figureCitation id="136F2A015826FFD1FEDFFE36811041A4" box="[286,372,402,428]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="20" pageNumber="21">Fig. 79</figureCitation>
).
<collectorName id="26A153525826FFD1FE48FE3681AB41A4" box="[393,463,402,428]" pageId="20" pageNumber="21">Palpal</collectorName>
claw with 6 long and 1–2 short teeth (
<figureCitation id="136F2A015826FFD1FC48FE3683BB41A4" box="[905,991,402,428]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="20" pageNumber="21">Fig. 76</figureCitation>
).
<collectorName id="26A153525826FFD1FC32FE36841341A4" box="[1011,1143,402,428]" pageId="20" pageNumber="21">Retrolateral</collectorName>
claw of left leg I with 13 teeth (
<figureCitation id="136F2A015826FFD1FF1EFE12815141D8" box="[223,309,438,464]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="20" pageNumber="21">Fig. 77</figureCitation>
).
<collectorName id="26A153525826FFD1FE88FE1281CB41D8" box="[329,431,438,464]" pageId="20" pageNumber="21">Trilobate</collectorName>
membrane of left leg I with short and narrow median hook, slightly shorter than lateral projections (
<figureCitation id="136F2A015826FFD1FEE2FE7E811241FC" box="[291,374,474,500]" captionStart="FIGURES 73–81" captionStartId="18.[151,264,1842,1866]" captionTargetBox="[151,1436,536,1819]" captionTargetId="figure-169@18.[151,1436,525,1819]" captionTargetPageId="18" captionText="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi="http://doi.org/10.5281/zenodo.6911201" httpUri="https://zenodo.org/record/6911201/files/figure.png" pageId="20" pageNumber="21">Fig. 81</figureCitation>
).
</paragraph>
<paragraph id="8BEB36845826FFD1FF06FE5A81A5428C" blockId="20.[151,1437,150,933]" pageId="20" pageNumber="21">
Copulatory organ as in diagnosis (
<figureCitation id="136F2A015826FFD1FD8FFE5A82B04210" box="[590,724,510,536]" captionStart="FIGURES 70–72" captionStartId="17.[151,264,1831,1855]" captionTargetBox="[151,1436,309,1808]" captionTargetId="figure-101@17.[151,1436,309,1808]" captionTargetPageId="17" captionText="FIGURES 70–72. Tiomaniella ladam gen. &amp; spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911197" httpUri="https://zenodo.org/record/6911197/files/figure.png" pageId="20" pageNumber="21">Figs 70–72</figureCitation>
). Epigynal field slightly wider than long, with distinct anterior bands. Many “fusion bubbles” situated almost over entire epigyne. Lateral lobes posteriorly with median indentation. Internal duct system nearly as long as wide. Dorsal part of ducts S-shaped. Fertilization ducts arising posteromedially, long and curved.
</paragraph>
<paragraph id="8BEB36845826FFD1FF06FD2A81F043AC" blockId="20.[151,1437,150,933]" pageId="20" pageNumber="21">
Colouration in ethanol (
<figureCitation id="136F2A015826FFD1FE17FD2A823842A0" box="[470,604,654,680]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="20" pageNumber="21">Figs 96–97</figureCitation>
): Prosoma yellowish-brown, latero-posteriorly with brown U-shaped pattern, medially with brown longitudinal stripe, lateral margin with three dark brown spots. Sternum yellow, margin light brown. Opisthosoma brown with yellowish-white spots, medially with white longitudinal stripe, ventral part yellowish-brown, laterally with brown spots. Chelicerae yellowish-brown, right chelicera with three brown dots. Coxa dorsally with brown spots. Palps yellowish-brown; Fe prolaterally, retrolaterally and distally with brown dots; Mt distal part reddish-brown. Legs yellowish-brown; Fe medially and distally with brown band; Pa prolaterally and retrolaterally brown; Ti I–III distally and proximally with brown band, Ti IV only distal part with brown band; Mt and Ta reddish-brown.
</paragraph>
<caption id="DF2B660C5826FFD1FF56FA3884C545BC" ID-DOI="http://doi.org/10.5281/zenodo.6911205" ID-Zenodo-Dep="6911205" box="[151,1185,1436,1461]" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1436,1460]" targetBox="[151,1436,956,1413]" targetPageId="20">
<paragraph id="8BEB36845826FFD1FF56FA3884C545BC" blockId="20.[151,1185,1436,1461]" box="[151,1185,1436,1461]" pageId="20" pageNumber="21">
<emphasis id="B920EA965826FFD1FF56FA38814445BC" bold="true" box="[151,288,1436,1460]" pageId="20" pageNumber="21">FIGURE 98.</emphasis>
Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea.
</paragraph>
</caption>
<paragraph id="8BEB36845826FFD1FF06FA46840B4685" blockId="20.[151,1437,1506,1713]" pageId="20" pageNumber="21">
<emphasis id="B920EA965826FFD1FF06FA4681BB45F4" bold="true" box="[199,479,1506,1533]" pageId="20" pageNumber="21">Variation. Male (n = 1):</emphasis>
TL 4.85; PL 2.45, PW 2.4; OL 2.4, OW 1.4. Measurements leg I: total length 8.20, Fe 2.20, Pa 1.10, Ti 2.40, Mt 2.00, Ta 0.80. Spination: palp: Ti 1121; legs: Fe I 322, Ti
<date id="FFEA10445826FFD1FB8DF9A384FC4629" box="[1100,1176,1543,1569]" pageId="20" pageNumber="21" value="2026-01">I 2026</date>
, Ti
<date id="FFEA10445826FFD1FB03F9A3857D4629" box="[1218,1305,1543,1569]" pageId="20" pageNumber="21" value="2026-02">II 2026</date>
; Mt
<date id="FFEA10445826FFD1FA8DF9A385FC4629" box="[1356,1432,1543,1569]" pageId="20" pageNumber="21" value="2026-01">I 2026</date>
, Mt
<date id="FFEA10445826FFD1FF7FF98F8177464D" box="[190,275,1579,1605]" pageId="20" pageNumber="21" value="2026-02">II 2026</date>
.
<emphasis id="B920EA965826FFD1FEDDF98F81BD464C" bold="true" box="[284,473,1578,1605]" pageId="20" pageNumber="21">Females (n = 2):</emphasis>
TL 6.0–6.8; PL 2.6–3.1, PW 2.5–2.9, AW 1.5–1.7; OL 3.3–3.7, OW 2.2–2.5. Measurements leg I: total length 8.55–9.55, Fe 2.30–2.60, Pa 1.20–1.30, Ti 2.25–2.50, Mt 2.00–2.30, Ta 0.75–0.85. Spination: palp: Mt 1014; legs: Fe II 223; Ti
<date id="FFEA10445826FFD1FD8EF9D782FE4685" box="[591,666,1651,1677]" pageId="20" pageNumber="21" value="1006-01">I 1006</date>
, Ti II 006, Ti
<date id="FFEA10445826FFD1FCF2F9D783F64685" box="[819,914,1650,1677]" pageId="20" pageNumber="21" value="2135-04">IV 2135</date>
; Mt
<date id="FFEA10445826FFD1FC07F9D784404685" box="[966,1060,1651,1677]" pageId="20" pageNumber="21" value="2006-03">III 2006</date>
, 2016.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5826FFD1FF06F932838F46B9" box="[199,1003,1686,1713]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BEB36845826FFD1FF06F932838F46B9" blockId="20.[151,1437,1506,1713]" box="[199,1003,1686,1713]" pageId="20" pageNumber="21">
<emphasis id="B920EA965826FFD1FF06F932810546B8" bold="true" box="[199,353,1686,1712]" pageId="20" pageNumber="21">Distribution.</emphasis>
Known only from the
<typeStatus id="54EF88265826FFD1FDA1F93382F446B9" box="[608,656,1687,1713]" pageId="20" pageNumber="21">type</typeStatus>
locality (
<figureCitation id="136F2A015826FFD1FD3AF933832B46B9" box="[763,847,1687,1713]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="20" pageNumber="21">Fig. 98</figureCitation>
: red triangle).
</paragraph>
</subSubSection>
</treatment>
</document>
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