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<document id="78390B4ADB5DCDDADBFF4C55A068B0EB" ID-CLB-Dataset="106789" ID-DOI="10.11646/zootaxa.5169.1.1" ID-GBIF-Dataset="b76236d0-0336-4d17-804a-80f0344b097e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911151" ID-ZooBank="3CB49021-90C5-46F0-AAAF-619EE8068F42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658900973851" checkinUser="plazi" docAuthor="Grall, Elena &amp; Jäger, Peter" docDate="2022" docId="03FD8792583DFFCAFF56FB9A82F64643" docLanguage="en" docName="zootaxa.5169.1.1.pdf" docOrigin="Zootaxa 5169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tiomaniella Grall &amp; Jäger 2022, gen. nov." docType="treatment" docVersion="6" lastPageNumber="16" masterDocId="FFC4FFEA5832FFC5FFC1FFA480644008" masterDocTitle="Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="16" updateTime="1699362128362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="DACA736E69497AE14CBF860A9AB9A3CA">Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)</mods:title>
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<mods:namePart id="B255BC559E778960B42E5323D17735D1">Grall, Elena</mods:namePart>
<mods:nameIdentifier id="0067569FB6198D1FD5347A02F116EA44" type="ORCID">0000-0002-8938-4105</mods:nameIdentifier>
<mods:affiliation id="97B388484C7DD53B72061CEFE7232E06">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; Institute of Ecology, Diversity and Evolution, Campus Riedberg, Goethe University, 60438 Frankfurt am Main, Germany; elena. grall @ senckenberg. de, https: // orcid. org / 0000 - 0002 - 8938 - 4105</mods:affiliation>
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<mods:namePart id="D7E8D86413BA1D3586A5CA536FA4655D">Jäger, Peter</mods:namePart>
<mods:nameIdentifier id="B2A90925F362A74E514A6211832E9AC5" type="ORCID">0000-0003-1563-0147</mods:nameIdentifier>
<mods:affiliation id="1B8DADB92B015DC7106DC331CDC0EF0E">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; peter. jaeger @ senckenberg. de, https: // orcid. org / 0000 - 0003 - 1563 - 0147</mods:affiliation>
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<mods:title id="4985296920426C82BFE1D4FE928B8C30">Zootaxa</mods:title>
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<mods:part id="DB61CB0D077D24EF0E782A43B06AEACE">
<mods:date id="CAF1EFDD844A573F6B6959007F86E8B2">2022</mods:date>
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<mods:number id="C51C727702A347192B0449E6F39A8EDA">2022-07-27</mods:number>
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<mods:number id="E7802792050CF58A891A0A56E0E915FC">5169</mods:number>
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<mods:identifier id="BEE53511526A1B7CF736913538BC2279" type="DOI">10.11646/zootaxa.5169.1.1</mods:identifier>
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<mods:identifier id="A1D5B0DDE7E0F4194F15B860820041E4" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03FD8792583DFFCAFF56FB9A82F64643" ID-DOI="http://doi.org/10.5281/zenodo.6911185" ID-GBIF-Taxon="197859751" ID-Zenodo-Dep="6911185" LSID="urn:lsid:plazi:treatment:03FD8792583DFFCAFF56FB9A82F64643" httpUri="http://treatment.plazi.org/id/03FD8792583DFFCAFF56FB9A82F64643" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C34E650F583DFFCAFF56FB9A81FA4450" box="[151,414,1086,1113]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BEB3684583DFFCAFF56FB9A81FA4450" blockId="15.[151,414,1086,1148]" box="[151,414,1086,1113]" pageId="15" pageNumber="16">
<heading id="D0A381E8583DFFCAFF56FB9A81FA4450" bold="true" box="[151,414,1086,1113]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B920EA96583DFFCAFF56FB9A81FA4450" bold="true" box="[151,414,1086,1113]" pageId="15" pageNumber="16">
<taxonomicName id="4C544D07583DFFCAFF56FB9A814D4451" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[151,297,1086,1113]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EA96583DFFCAFF56FB9A814D4451" bold="true" box="[151,297,1086,1113]" italics="true" pageId="15" pageNumber="16">Tiomaniella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357ED583DFFCAFEF1FB9A81FA4450" box="[304,414,1086,1112]" pageId="15" pageNumber="16" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF56FBC6810C4474" box="[151,360,1122,1148]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEB3684583DFFCAFF56FBC6810C4474" blockId="15.[151,414,1086,1148]" box="[151,360,1122,1148]" pageId="15" pageNumber="16">
<figureCitation id="136F2A01583DFFCAFF56FBC681734474" box="[151,279,1122,1148]" captionStart-0="FIGURES 66–69" captionStart-1="FIGURES 70–72" captionStart-2="FIGURES 73–81" captionStartId-0="16.[151,264,1679,1703]" captionStartId-1="17.[151,264,1831,1855]" captionStartId-2="18.[151,264,1842,1866]" captionTargetBox-0="[151,1420,464,1644]" captionTargetBox-1="[151,1436,309,1808]" captionTargetBox-2="[151,1436,536,1819]" captionTargetId-0="figure-135@16.[151,1436,452,1655]" captionTargetId-1="figure-101@17.[151,1436,309,1808]" captionTargetId-2="figure-169@18.[151,1436,525,1819]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionText-0="FIGURES 66–69. Tiomaniella ladam gen. &amp; spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis." captionText-1="FIGURES 70–72. Tiomaniella ladam gen. &amp; spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca." captionText-2="FIGURES 73–81. Tiomaniella ladam gen. &amp; spec. nov. holotype male (73, 75, 78, 80) and female paratype (74, 76–77, 79, 81) from Tioman Island. 73–74 Eyes, dorsal; 75–77 Claws (75 right leg I, retrolateral claw, retrolateral; 76 left palp, prolateral; 77 left leg I, retrolateral claw, retrolateral); 78–79 Left chelicera, ventral (79 escort setae not pictured); 80–81 trilobate membrane, distal metatarsus of leg I, dorsal (78 right, 79 left)." figureDoi-0="http://doi.org/10.5281/zenodo.6911193" figureDoi-1="http://doi.org/10.5281/zenodo.6911197" figureDoi-2="http://doi.org/10.5281/zenodo.6911201" httpUri-0="https://zenodo.org/record/6911193/files/figure.png" httpUri-1="https://zenodo.org/record/6911197/files/figure.png" httpUri-2="https://zenodo.org/record/6911201/files/figure.png" pageId="15" pageNumber="16">Figs 66–81</figureCitation>
,
<figureCitation id="136F2A01583DFFCAFEE3FBC6810C4474" box="[290,360,1122,1148]" captionStart-0="FIGURES 82–97" captionStart-1="FIGURE 98" captionStartId-0="19.[151,264,1899,1923]" captionStartId-1="20.[151,250,1436,1460]" captionTargetBox-0="[152,1435,182,1873]" captionTargetBox-1="[151,1436,956,1413]" captionTargetId-0="figure-19@19.[151,1436,181,1875]" captionTargetId-1="figure-372@20.[151,1436,956,1413]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionText-0="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." captionText-1="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi-0="http://doi.org/10.5281/zenodo.6911203" figureDoi-1="http://doi.org/10.5281/zenodo.6911205" httpUri-0="https://zenodo.org/record/6911203/files/figure.png" httpUri-1="https://zenodo.org/record/6911205/files/figure.png" pageId="15" pageNumber="16">94–98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF56FB03823244B7" box="[151,598,1190,1215]" pageId="15" pageNumber="16" type="type_taxon">
<paragraph id="8BEB3684583DFFCAFF56FB03823244B7" blockId="15.[151,598,1190,1215]" box="[151,598,1190,1215]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF56FB03814744B6" bold="true" box="[151,291,1190,1215]" pageId="15" pageNumber="16">
<typeStatus id="54EF8826583DFFCAFF56FB0380A844B7" box="[151,204,1191,1215]" pageId="15" pageNumber="16">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C544D07583DFFCAFEEBFB03818C44B7" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[298,488,1191,1215]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ladam" status="sp. nov.">
<emphasis id="B920EA96583DFFCAFEEBFB03818C44B7" box="[298,488,1191,1215]" italics="true" pageId="15" pageNumber="16">Tiomaniella ladam</emphasis>
</taxonomicName>
<emphasis id="B920EA96583DFFCAFE2EFB03823244B7" bold="true" box="[495,598,1191,1215]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A21357ED583DFFCAFE2EFB03823244B7" box="[495,598,1191,1215]" pageId="15" pageNumber="16" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF56FB49848F450F" box="[151,1259,1261,1287]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BEB3684583DFFCAFF56FB49848F450F" blockId="15.[151,1436,1261,1611]" box="[151,1259,1261,1287]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF56FB49817A450F" bold="true" box="[151,286,1261,1287]" pageId="15" pageNumber="16">Etymology.</emphasis>
The generic name is derived from the name of Tioman Island. The gender is feminine.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF06FAB5856045D7" pageId="15" pageNumber="16" type="diagnosis">
<paragraph id="8BEB3684583DFFCAFF06FAB5856045D7" blockId="15.[151,1436,1261,1611]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF06FAB581264523" bold="true" box="[199,322,1297,1323]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
Small
<taxonomicName id="4C544D07583DFFCAFE55FAB5827C4523" authorityName="Bertkau" authorityYear="1872" box="[404,536,1297,1323]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
with TL &lt;7 (
<figureCitation id="136F2A01583DFFCAFD75FAB583534523" box="[692,823,1297,1323]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="15" pageNumber="16">Figs 94–97</figureCitation>
) belonging to the subfamily
<taxonomicName id="4C544D07583DFFCAFB42FAB585484523" authorityName="Thorell" authorityYear="1873" box="[1155,1324,1297,1323]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">Heteropodinae</taxonomicName>
. Males of
<taxonomicName id="4C544D07583DFFCAFF56FA9181794547" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[151,285,1333,1359]" class="Arachnida" family="Sparassidae" genus="Tiomaniella" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B920EA96583DFFCAFF56FA9181794547" box="[151,285,1333,1359]" italics="true" pageId="15" pageNumber="16">Tiomaniella</emphasis>
</taxonomicName>
<emphasis id="B920EA96583DFFCAFEE7FA9181F64547" bold="true" box="[294,402,1333,1359]" pageId="15" pageNumber="16">
<taxonomicNameLabel id="A21357ED583DFFCAFEE7FA9181F64547" box="[294,402,1333,1359]" pageId="15" pageNumber="16" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all other
<taxonomicName id="4C544D07583DFFCAFCF4FA9183BE4547" authorityName="Thorell" authorityYear="1873" box="[821,986,1333,1359]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">Heteropodinae</taxonomicName>
by the subtegular functional conductor (
<figureCitation id="136F2A01583DFFCAFF5EFAFD8179457B" box="[159,285,1369,1395]" captionStart="FIGURES 66–69" captionStartId="16.[151,264,1679,1703]" captionTargetBox="[151,1420,464,1644]" captionTargetId="figure-135@16.[151,1436,452,1655]" captionTargetPageId="16" captionText="FIGURES 66–69. Tiomaniella ladam gen. &amp; spec. nov. holotype male from Tioman Island. 66–68 Left male palp (66 prolateral, 67 ventral, 68 retrolateral); 69 Embolus tip, ventral. C—conductor, E—embolus, rB—retrolateral bulge of cymbium, rpP—retro-proximal projection of cymbium, RTA—retrolateral tibial apophysis, S—spermophor, ST—subtegulum, acting as functional conductor, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911193" httpUri="https://zenodo.org/record/6911193/files/figure.png" pageId="15" pageNumber="16">Figs 66–67</figureCitation>
). Females may be recognised by the following combination of characters: 1. Copulatory organs situated laterally with epigynal pockets located posteriorly, with first winding of internal duct system postero-mediad, 2. Median septum not recognisable, “fusion bubbles” indicating fused seams between lateral lobes and median septum and 3. Internal duct system simple with very broad ducts leading to large spermathecae (
<figureCitation id="136F2A01583DFFCAFBB4FA61849D45D7" box="[1141,1273,1477,1503]" captionStart="FIGURES 70–72" captionStartId="17.[151,264,1831,1855]" captionTargetBox="[151,1436,309,1808]" captionTargetId="figure-101@17.[151,1436,309,1808]" captionTargetPageId="17" captionText="FIGURES 70–72. Tiomaniella ladam gen. &amp; spec. nov. female paratype from Tioman Island. 70 Epigyne, ventral; 71 Vulva, dorsal; 72 Schematic course of internal duct system, dorsal. CO—copulatory opening, EF—epigynal field, EP—epigynal pock- et, FB— fusion bubbles, FD—fertilization ducts, GA—glandular appendages, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911197" httpUri="https://zenodo.org/record/6911197/files/figure.png" pageId="15" pageNumber="16">Figs 70–72</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF06FA4D82A5460B" box="[199,705,1513,1539]" pageId="15" pageNumber="16" type="description">
<paragraph id="8BEB3684583DFFCAFF06FA4D82A5460B" blockId="15.[151,1436,1261,1611]" box="[199,705,1513,1539]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF06FA4D813D460B" bold="true" box="[199,345,1513,1539]" pageId="15" pageNumber="16">Description.</emphasis>
See description of
<typeStatus id="54EF8826583DFFCAFDF0FA4D8205460B" box="[561,609,1513,1539]" pageId="15" pageNumber="16">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F583DFFCAFF06F9A982F64643" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BEB3684583DFFCAFF06F9A9831A462F" blockId="15.[151,1436,1261,1611]" box="[199,894,1549,1575]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF06F9A98105462F" bold="true" box="[199,353,1549,1575]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="F3437614583DFFCAFEA6F9A981B6462F" box="[359,466,1549,1575]" name="Malaysia" pageId="15" pageNumber="16">Malaysia</collectingCountry>
: Tioman Island (
<figureCitation id="136F2A01583DFFCAFD4FF9A98286462F" box="[654,738,1549,1575]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="15" pageNumber="16">Fig. 98</figureCitation>
: red triangle).
</paragraph>
<paragraph id="8BEB3684583DFFCAFF06F99582F64643" blockId="15.[151,1436,1261,1611]" box="[199,658,1585,1611]" pageId="15" pageNumber="16">
<emphasis id="B920EA96583DFFCAFF06F99581F74643" bold="true" box="[199,403,1585,1611]" pageId="15" pageNumber="16">Species included.</emphasis>
Only the
<typeStatus id="54EF8826583DFFCAFDC3F99582564643" box="[514,562,1585,1611]" pageId="15" pageNumber="16">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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