365 lines
50 KiB
XML
365 lines
50 KiB
XML
<document id="571EE8D89BF33D0F96E8381709CDCBBD" ID-DOI="10.5281/zenodo.5341533" ID-ISSN="2345-7600" ID-Zenodo-Dep="5341533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190745178" checkinUser="diego" docAuthor="Cai, Yixiong & Husana, Daniel Edison M." docDate="2009" docId="03FD8790FC32FFAFFE8B3727B31E0AC3" docLanguage="en" docName="57rbz0051-063.pdf" docOrigin="Raffles Bulletin of Zoology 57 (1)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Edoneus marulas Cai & Husana 2009, new species" docType="treatment" docVersion="3" lastPageNumber="63" masterDocId="FFC4FFE8FC3BFFA3FF8B335BB2240E30" masterDocTitle="Cave Shrimps Of The Genus Edoneus Holthuis, 1978, From Luzon, The Philippines, With Descriptions Of Three New Species (Crustacea: Decapoda: Atyidae)" masterLastPageNumber="63" masterPageNumber="51" pageNumber="60" updateTime="1699181455100" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="4AA0FB7A4B0853E61FD1273909A7404D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="819CE94C81978FD193C318A39A9B2A90">
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<mods:title id="FA3A363F8FCC5FDF5201B41A08D973EB">Cave Shrimps Of The Genus Edoneus Holthuis, 1978, From Luzon, The Philippines, With Descriptions Of Three New Species (Crustacea: Decapoda: Atyidae)</mods:title>
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<mods:name id="B8CE41370D7313F1C4E0DE665E95C2A7" type="personal">
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<mods:roleTerm id="09E2F254C8E42F61A8AD478232440C51">Author</mods:roleTerm>
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<mods:namePart id="233BB1BE2302C5D09959D37D672A2715">Cai, Yixiong</mods:namePart>
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<mods:name id="EA0DA5751D403D6F14D0B2BC81B60113" type="personal">
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<mods:roleTerm id="255C678D8ED5B5CB6C6F33FC113736B8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="361FC54E37D1B3060757F30AD654C8BB">Husana, Daniel Edison M.</mods:namePart>
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<mods:typeOfResource id="6A1F09CBF8A35825283EF8049DF8254B">text</mods:typeOfResource>
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<mods:titleInfo id="BDE4EDAC5DBECD83CD6B50743BAD18EF">
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<mods:title id="EECAD053A2558A2C67CB79896FA852F1">Raffles Bulletin of Zoology</mods:title>
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</mods:titleInfo>
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<mods:part id="6905461A256F7024B05B010C941E90A2">
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<mods:date id="342B26B9EEC2054D21109243FF465FF3">2009</mods:date>
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<mods:detail id="25D01688D7002B18203CA7366DAC0ADB" type="pubDate">
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<mods:number id="FFEE7125102E952B79FFB04463E75FB5">2009-02-28</mods:number>
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</mods:detail>
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<mods:detail id="A039791BAA45A276D86454D21DCD8116" type="volume">
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<mods:number id="223D04C5ED58994C3D8D1650EF1103B3">57</mods:number>
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</mods:detail>
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<mods:detail id="9329F4CC81D11532D491B747CC74133C" type="issue">
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<mods:number id="FB871D05268ECB70A6D0A822AFECD153">1</mods:number>
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</mods:detail>
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<mods:extent id="1061EC9623E3A9A0E4F363D8AD79A3F4" unit="page">
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<mods:start id="D2840E98C1D4B7C37BD416C5D97FB304">51</mods:start>
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<mods:classification id="0057990D6D3EE12ECBC90FAF6E5A53C6">journal article</mods:classification>
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<mods:identifier id="2410FA8E4768D328885F4E798B4DA178" type="DOI">10.5281/zenodo.5341533</mods:identifier>
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<mods:identifier id="9FBFFFE34C81E2D0CD7FA79C76C3A60E" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="2F22DD7C99DF5324980DAB0846F42081" type="Zenodo-Dep">5341533</mods:identifier>
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<treatment id="03FD8790FC32FFAFFE8B3727B31E0AC3" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3" httpUri="http://treatment.plazi.org/id/03FD8790FC32FFAFFE8B3727B31E0AC3" lastPageId="12" lastPageNumber="63" pageId="9" pageNumber="60">
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<subSubSection id="C34E650DFC32FFAAFE8B3727B07D0AA3" box="[256,601,1147,1172]" pageId="9" pageNumber="60" type="nomenclature">
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<paragraph id="8BEB3686FC32FFAAFE8B3727B07D0AA3" blockId="9.[256,601,1147,1203]" box="[256,601,1147,1172]" pageId="9" pageNumber="60">
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<heading id="D0A381EAFC32FFAAFE8B3727B07D0AA3" bold="true" box="[256,601,1147,1172]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="60" reason="2">
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<emphasis id="B920EA94FC32FFAAFE8B3727B07D0AA3" bold="true" box="[256,601,1147,1172]" pageId="9" pageNumber="60">
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<taxonomicName id="4C544D05FC32FFAAFE8B3727B3E70AA4" ID-CoL="8VKJ6" authority="Cai & Husana, 2009" authorityName="Cai & Husana" authorityYear="2009" box="[256,451,1148,1172]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="60" phylum="Arthropoda" rank="species" species="marulas" status="sp. nov.">
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<emphasis id="B920EA94FC32FFAAFE8B3727B3E70AA4" bold="true" box="[256,451,1148,1172]" italics="true" pageId="9" pageNumber="60">Edoneus marulas</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21357EFFC32FFAAFE583720B07D0AA3" box="[467,601,1147,1171]" pageId="9" pageNumber="60" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E650DFC32FFAAFEFB37C7B3CC0A83" box="[368,488,1180,1203]" pageId="9" pageNumber="60" type="description">
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<paragraph id="8BEB3686FC32FFAAFEFB37C7B3CC0A83" blockId="9.[256,601,1147,1203]" box="[368,488,1180,1203]" pageId="9" pageNumber="60">
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(
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<figureCitation id="136F2A03FC32FFAAFEF337C7B3E10A83" box="[376,453,1180,1203]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Figs. 6</figureCitation>
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,
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<figureCitation id="136F2A03FC32FFAAFE5837C7B3C50A83" box="[467,481,1180,1203]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E650DFC32FFAAFFE63781B39208AF" pageId="9" pageNumber="60" type="materials_examined">
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<paragraph id="8BEB3686FC32FFAAFFE63781B36D0B19" blockId="9.[109,748,1242,1321]" pageId="9" pageNumber="60">
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<emphasis id="B920EA94FC32FFAAFFE63781B36A0ADF" bold="true" box="[109,334,1242,1263]" italics="true" pageId="9" pageNumber="60">Material examined. –</emphasis>
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<materialsCitation id="3B3C3CDBFC32FFAAFED33781B36D0B19" collectingDate="2002-11-11" collectionCode="NMCR" collectorName="D. E. Husana" country="Philippines" location="Marulas Cave" pageId="9" pageNumber="60" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8824FC32FFAAFED33781B39C0ADF" box="[344,440,1242,1263]" pageId="9" pageNumber="60" type="holotype">Holotype</typeStatus>
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: Male, cl
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<quantity id="4CAC9B63FC32FFAAFDA83781B0500ADF" box="[547,628,1242,1263]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="60" unit="mm" value="4.1">4.1 mm</quantity>
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,
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<collectionCode id="ED45AE43FC32FFAAFD093781B0E90ADF" box="[642,717,1242,1263]" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="9" pageNumber="60" type="Herbarium">NMCR</collectionCode>
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||
, at middle of
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<location id="8E8B605DFC32FFAAFF5637ACB3540B3C" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFF5637ACB3540B3C" box="[221,368,1271,1292]" country="Philippines" name="Marulas Cave" pageId="9" pageNumber="60">Marulas Cave</location>
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, Luzon, the
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<collectingCountry id="F3437616FC32FFAAFE7237ACB04B0B3C" box="[505,623,1271,1292]" name="Philippines" pageId="9" pageNumber="60">Philippines</collectingCountry>
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||
, coll.
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<collectorName id="26A15350FC32FFAAFD3F37ACB2930B19" pageId="9" pageNumber="60">D. E. Husana</collectorName>
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,
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<date id="FFEA1046FC32FFAAFF48364FB3600B19" box="[195,324,1300,1321]" pageId="9" pageNumber="60" value="2002-11-11">
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<collectingDate id="EFAEE9AEFC32FFAAFF48364FB3600B19" box="[195,324,1300,1321]" pageId="9" pageNumber="60" value="2002-11-11">11 Nov.2002</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BEB3686FC32FFAAFFE63616B39208AF" blockId="9.[109,749,1357,1695]" pageId="9" pageNumber="60">
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<materialsCitation id="3B3C3CDBFC32FFAAFFE63616B3C00B52" box="[109,484,1357,1378]" location="Seven" pageId="9" pageNumber="60" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="54EF8824FC32FFAAFFE63616B2EA0B52" box="[109,206,1357,1378]" pageId="9" pageNumber="60" type="paratype">Paratypes</typeStatus>
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||
:
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||
<location id="8E8B605DFC32FFAAFF5D3616B3350B52" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFF5D3616B3350B52" box="[214,273,1357,1378]" name="Seven" pageId="9" pageNumber="60">Seven</location>
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males, cl 3.4–4.0 mm
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||
</materialsCitation>
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||
,
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||
<materialsCitation id="3B3C3CDBFC32FFAAFE673616B3E70BAC" collectingDate="2002-11-11" collectionCode="ZRC" collectorName="D. E. Husana" country="Philippines" location="Marulas Cave" municipality="Luzon" pageId="9" pageNumber="60" specimenCode="ZRC 2007.0490" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
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||
<specimenCount id="9D52FD0FFC32FFAAFE673616B06E0B52" box="[492,586,1357,1378]" pageId="9" pageNumber="60" type="female">3 females</specimenCount>
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||
, cl
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||
<quantity id="4CAC9B63FC32FFAAFDE13616B0910B52" box="[618,693,1357,1378]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="60" unit="mm" value="4.5">4.5 mm</quantity>
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||
,
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||
<specimenCode id="DBF29EFDFC32FFAAFD363616B2F30B4F" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="60">ZRC 2007.0490</specimenCode>
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||
, at middle of
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||
<location id="8E8B605DFC32FFAAFEE23631B3D20B4F" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFEE23631B3D20B4F" box="[361,502,1386,1407]" country="Philippines" municipality="Luzon" name="Marulas Cave" pageId="9" pageNumber="60">Marulas Cave</location>
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||
,
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<collectingMunicipality id="6B8FACFCFC32FFAAFD883631B0600B4F" box="[515,580,1386,1407]" pageId="9" pageNumber="60">Luzon</collectingMunicipality>
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, the
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||
<collectingCountry id="F3437616FC32FFAAFDF33631B0CC0B4F" box="[632,744,1386,1407]" name="Philippines" pageId="9" pageNumber="60">Philippines</collectingCountry>
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||
, coll.
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||
<collectorName id="26A15350FC32FFAAFF2936DCB30A0BAC" box="[162,302,1415,1436]" pageId="9" pageNumber="60">D. E. Husana</collectorName>
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,
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<date id="FFEA1046FC32FFAAFEB036DCB3E70BAC" box="[315,451,1415,1436]" pageId="9" pageNumber="60" value="2002-11-11">
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<collectingDate id="EFAEE9AEFC32FFAAFEB036DCB3E70BAC" box="[315,451,1415,1436]" pageId="9" pageNumber="60" value="2002-11-11">11 Nov.2002</collectingDate>
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</date>
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</materialsCitation>
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;
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<specimenCount id="9D52FD0FFC32FFAAFE5B36DCB0050BAC" box="[464,545,1415,1436]" pageId="9" pageNumber="60" type="male">6 males</specimenCount>
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||
, cl 3.4–4.0 mm,
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<materialsCitation id="3B3C3CDBFC32FFAAFD5836DCB3510BC2" collectingDate="2002-11-11" collectionCode="ZRC" collectorName="D. E. Husana" country="Philippines" location="Marulas Cave" municipality="Luzon" pageId="9" pageNumber="60" specimenCode="ZRC 2007.0491" specimenCount="11" specimenCount-female="11" typeStatus="paratype">
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||
<specimenCount id="9D52FD0FFC32FFAAFD5836DCB29E0B89" pageId="9" pageNumber="60" type="female">11 females</specimenCount>
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, cl
|
||
<quantity id="4CAC9B63FC32FFAAFF5036FFB3740B89" box="[219,336,1444,1465]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.3" metricValueMin="3.7" pageId="9" pageNumber="60" unit="mm" value="4.0" valueMax="4.3" valueMin="3.7">3.7–4.3 mm</quantity>
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||
, 3 ovigerous females, cl 4.0–
|
||
<quantity id="4CAC9B63FC32FFAAFDE336FFB0900B89" box="[616,692,1444,1465]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="60" unit="mm" value="4.4">4.4 mm</quantity>
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||
,
|
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<specimenCode id="DBF29EFDFC32FFAAFD3636FFB2F10BE6" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="60">ZRC 2007.0491</specimenCode>
|
||
,
|
||
<location id="8E8B605DFC32FFAAFF55369AB3410BE6" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFF55369AB3410BE6" box="[222,357,1473,1494]" country="Philippines" municipality="Luzon" name="Marulas Cave" pageId="9" pageNumber="60">Marulas Cave</location>
|
||
, lowest part,
|
||
<collectingMunicipality id="6B8FACFCFC32FFAAFE6A369AB0050BE6" box="[481,545,1473,1494]" pageId="9" pageNumber="60">Luzon</collectingMunicipality>
|
||
, the
|
||
<collectingCountry id="F3437616FC32FFAAFDC0369AB09D0BE6" box="[587,697,1473,1494]" name="Philippines" pageId="9" pageNumber="60">Philippines</collectingCountry>
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, coll.
|
||
<collectorName id="26A15350FC32FFAAFFE63686B2C90BC2" box="[109,237,1501,1522]" pageId="9" pageNumber="60">D. E. Husana</collectorName>
|
||
,
|
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<date id="FFEA1046FC32FFAAFF7D3686B3510BC2" box="[246,373,1501,1522]" pageId="9" pageNumber="60" value="2002-11-11">
|
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<collectingDate id="EFAEE9AEFC32FFAAFF7D3686B3510BC2" box="[246,373,1501,1522]" pageId="9" pageNumber="60" value="2002-11-11">11 Nov.2002</collectingDate>
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</date>
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||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D52FD0FFC32FFAAFEF43686B3F20BC2" box="[383,470,1501,1522]" pageId="9" pageNumber="60" type="male">12 males</specimenCount>
|
||
, cl 3.4–4.0 mm,
|
||
<materialsCitation id="3B3C3CDBFC32FFAAFDF83686B3870879" collectingDate="2002-11-11" collectionCode="ZRC" collectorName="D. E. Husana" country="Philippines" location="Marulas Cave" municipality="Luzon" pageId="9" pageNumber="60" specimenCode="ZRC 2007.0492" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
|
||
<specimenCount id="9D52FD0FFC32FFAAFDF83686B0F50BC2" box="[627,721,1501,1522]" pageId="9" pageNumber="60" type="female">6 females</specimenCount>
|
||
, cl
|
||
<quantity id="4CAC9B63FC32FFAAFFE636A1B2C7083F" box="[109,227,1530,1551]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.5" metricValueMin="4.1" pageId="9" pageNumber="60" unit="mm" value="4.3" valueMax="4.5" valueMin="4.1">4.1–4.5 mm</quantity>
|
||
, 1 ovigerous female, cl
|
||
<quantity id="4CAC9B63FC32FFAAFE4D36A1B036083F" box="[454,530,1530,1551]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="60" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, eggs 0.90 ×
|
||
<quantity id="4CAC9B63FC32FFAAFD1B36A1B0CC083F" box="[656,744,1530,1551]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="60" unit="mm" value="0.55">0.55 mm</quantity>
|
||
,
|
||
<specimenCode id="DBF29EFDFC32FFAAFFE6354CB32F081C" box="[109,267,1559,1580]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="9" pageNumber="60">ZRC 2007.0492</specimenCode>
|
||
, middle of
|
||
<location id="8E8B605DFC32FFAAFEF2354CB026081C" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFEF2354CB026081C" box="[377,514,1559,1580]" country="Philippines" municipality="Luzon" name="Marulas Cave" pageId="9" pageNumber="60">Marulas Cave</location>
|
||
,
|
||
<collectingMunicipality id="6B8FACFCFC32FFAAFD87354CB068081C" box="[524,588,1559,1580]" pageId="9" pageNumber="60">Luzon</collectingMunicipality>
|
||
, the
|
||
<collectingCountry id="F3437616FC32FFAAFDF1354CB0CD081C" box="[634,745,1559,1580]" name="Philippines" pageId="9" pageNumber="60">Philippines</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26A15350FC32FFAAFF17356FB3380879" box="[156,284,1588,1609]" pageId="9" pageNumber="60">D. E. Husana</collectorName>
|
||
,
|
||
<date id="FFEA1046FC32FFAAFEAF356FB3870879" box="[292,419,1588,1609]" pageId="9" pageNumber="60" value="2002-11-11">
|
||
<collectingDate id="EFAEE9AEFC32FFAAFEAF356FB3870879" box="[292,419,1588,1609]" pageId="9" pageNumber="60" value="2002-11-11">11 Nov.2002</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount id="9D52FD0FFC32FFAAFE27356FB3D30879" box="[428,503,1588,1609]" pageId="9" pageNumber="60" type="male">8 males</specimenCount>
|
||
, cl
|
||
<quantity id="4CAC9B63FC32FFAAFD9D356FB0AF0879" box="[534,651,1588,1609]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="9" pageNumber="60" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.3–3.8 mm</quantity>
|
||
,
|
||
<materialsCitation id="3B3C3CDBFC32FFAAFD18356FB39608AF" collectingDate="2002-11-11" collectionCode="NMCR" collectorName="D. E. Husana" country="Philippines" county="Luzon" location="Marulas Well" municipality="Marulas Cave" pageId="9" pageNumber="60" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D52FD0FFC32FFAAFD18356FB0CC0879" box="[659,744,1588,1609]" pageId="9" pageNumber="60" type="female">1 female</specimenCount>
|
||
, cl
|
||
<quantity id="4CAC9B63FC32FFAAFF0F350AB2EB0856" box="[132,207,1617,1638]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="9" pageNumber="60" unit="mm" value="4.1">4.1 mm</quantity>
|
||
, 1 ovigerous female, cl
|
||
<quantity id="4CAC9B63FC32FFAAFE39350AB3DA0856" box="[434,510,1617,1638]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="60" unit="mm" value="4.5">4.5 mm</quantity>
|
||
, with hatchings,
|
||
<collectionCode id="ED45AE43FC32FFAAFD14350AB0CC0856" box="[671,744,1617,1638]" country="USA" lsid="urn:lsid:biocol.org:col:14936" name="New Mexico State University" pageId="9" pageNumber="60" type="Herbarium">NMCR</collectionCode>
|
||
,
|
||
<location id="8E8B605DFC32FFAAFFE63536B2D108B2" LSID="urn:lsid:plazi:treatment:03FD8790FC32FFAFFE8B3727B31E0AC3:8E8B605DFC32FFAAFFE63536B2D108B2" box="[109,245,1645,1666]" country="Philippines" county="Luzon" municipality="Marulas Cave" name="Marulas Well" pageId="9" pageNumber="60">Marulas Well</location>
|
||
, outside the
|
||
<collectingMunicipality id="6B8FACFCFC32FFAAFEF83536B3DA08B2" box="[371,510,1645,1666]" pageId="9" pageNumber="60">Marulas Cave</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="628A4E0AFC32FFAAFD823536B06D08B2" box="[521,585,1645,1666]" pageId="9" pageNumber="60">Luzon</collectingCounty>
|
||
, the
|
||
<collectingCountry id="F3437616FC32FFAAFDF13536B0CD08B2" box="[634,745,1645,1666]" name="Philippines" pageId="9" pageNumber="60">Philippines</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26A15350FC32FFAAFF1435D1B30008AF" box="[159,292,1674,1695]" pageId="9" pageNumber="60">D. E. Husana</collectorName>
|
||
,
|
||
<date id="FFEA1046FC32FFAAFEBB35D1B39608AF" box="[304,434,1674,1695]" pageId="9" pageNumber="60" value="2002-11-11">
|
||
<collectingDate id="EFAEE9AEFC32FFAAFEBB35D1B39608AF" box="[304,434,1674,1695]" pageId="9" pageNumber="60" value="2002-11-11">11 Nov.2002</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E650DFC32FFAFFFE63593B07C0D03" lastPageId="12" lastPageNumber="63" pageId="9" pageNumber="60" type="description">
|
||
<paragraph id="8BEB3686FC32FFAAFFE63593B33E0950" blockId="9.[109,748,1736,1888]" pageId="9" pageNumber="60">
|
||
<emphasis id="B920EA94FC32FFAAFFE63593B32F08D0" bold="true" box="[109,267,1736,1760]" italics="true" pageId="9" pageNumber="60">Description. –</emphasis>
|
||
Rostrum (
|
||
<figureCitation id="136F2A03FC32FFAAFE0F3592B3FF08D0" box="[388,475,1737,1760]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6A</figureCitation>
|
||
) short, straight, pointed, unarmed, reaching near to or slightly beyond end of basal segment of antennular peduncle. Inferior orbital angle acute, fused with antennal spine; pterygostomian margin sub-rectangular.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFFE634D2B68A0EC3" blockId="9.[109,748,1929,2112]" lastBlockId="9.[808,1449,188,243]" pageId="9" pageNumber="60">
|
||
Sixth abdominal somite 0.54 times length of carapace, 1.3 times as long as fifth somite, as long as or slightly longer than telson. Telson (
|
||
<figureCitation id="136F2A03FC32FFAAFEC33492B3E209D0" box="[328,454,1993,2016]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Figs. 6B, C</figureCitation>
|
||
) 2.8 times as long as wide, distal margin rounded, with a very tiny median projection, with 4 pairs of dorsal spinules, and one pair of dorsolateral spinules; distal end with 3–4 pairs of spines, lateral pair longer than intermediate pairs; sublateral pair shortest. Preanal carina (
|
||
<figureCitation id="136F2A03FC32FFAAFCF23387B1EB0EC3" box="[889,975,220,243]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6D</figureCitation>
|
||
) low, lacking spine.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFCA33247B63F0C63" blockId="9.[808,1448,284,595]" pageId="9" pageNumber="60">
|
||
Eyes strongly reduced, cornea absent, unpigmented or occasionally with very tiny dot; anterior end reaching to half length of basal segment of antennular peduncle. Antennular peduncle 0.5 times as long as carapace; basal segment of antennular peduncle as long as combined lengths of second and third segments, anterolateral angle 0.25 times length of second segment, second segment distinctly longer than third segment. Stylocerite reaching 0.7 times length of basal segment of antennular peduncle. Scaphocerite (
|
||
<figureCitation id="136F2A03FC32FFAAFAAD3147B75C0C03" box="[1318,1400,540,563]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6E</figureCitation>
|
||
) 3.0 times as long as wide.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFCA33127B62B0A03" blockId="9.[808,1448,636,1075]" pageId="9" pageNumber="60">
|
||
Incisor process of mandible (
|
||
<figureCitation id="136F2A03FC32FFAAFBDF3127B6850CA3" box="[1108,1185,636,659]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6F</figureCitation>
|
||
) ending in a row of small teeth, molar process truncated. Lower lacinia of maxillula (
|
||
<figureCitation id="136F2A03FC32FFAAFCBB31E7B1AD0CE3" box="[816,905,700,723]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6G</figureCitation>
|
||
) broadly rounded, upper lacinia elongated, with numerous distinct teeth on inner margin, palp slender. Upper endites of maxilla (
|
||
<figureCitation id="136F2A03FC32FFAAFB8D31A7B6440D23" box="[1030,1120,764,787]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6H</figureCitation>
|
||
) subdivided, palp elongated, scaphognathite tapering posteriorly with numerous long, curved setae at posterior end. Distal end of palp of first maxilliped (
|
||
<figureCitation id="136F2A03FC32FFAAFC223007B1D50D43" box="[937,1009,860,883]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6I</figureCitation>
|
||
) triangular, with a finger-like projection; flagellum of the exopod very short, endopod high, as long as flagellum of exopod in length. Second maxilliped (
|
||
<figureCitation id="136F2A03FC32FFAAFAF630C7B1640DE3" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6J</figureCitation>
|
||
) typical, except the slightly reduced podobranch. Third maxilliped (
|
||
<figureCitation id="136F2A03FC32FFAAFC253087B6220DC3" box="[942,1030,988,1011]" captionStart="Fig" captionStartId="10.[132,167,2028,2049]" captionTargetBox="[194,1406,193,1966]" captionTargetId="figure-8@10.[168,1436,140,2005]" captionTargetPageId="10" captionText="Fig. 6. Edoneus marulas, new species: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="9" pageNumber="60">Fig. 6K</figureCitation>
|
||
) reaching to end of third segment of antennular peduncle, with ultimate segment as long as penultimate segment.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFCA33707B64308C3" blockId="9.[808,1448,1116,1779]" pageId="9" pageNumber="60">
|
||
Epipods well developed on first four pereiopods. First pereiopod (
|
||
<figureCitation id="136F2A03FC32FFAAFC2C3727B1DB0AA3" box="[935,1023,1148,1171]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Fig. 7A</figureCitation>
|
||
) reaching to end of basal segment of antennular peduncle; merus 2.1 times as long as broad, as long as carpus; carpus excavated anteriorly, shorter than chela, 1.6 times as long as high; chela 1.8 times as long as broad; fingers slightly shorter than palm. Second pereiopod (
|
||
<figureCitation id="136F2A03FC32FFAAFCBB3647B1A60B03" box="[816,898,1308,1331]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Fig. 7B</figureCitation>
|
||
) reaching to middle of third segment of antennular peduncle; merus shorter than carpus, 5.3 times as long as broad; carpus 1.4 times as long as chela, 4.8 times as long as high; chela 2.4 times as long as broad; fingers1.6 times as long as palm. Third pereiopod (
|
||
<figureCitation id="136F2A03FC32FFAAFBFA36C7B6CF0B83" box="[1137,1259,1436,1459]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Figs. 7C, D</figureCitation>
|
||
) reaching beyond scaphocerite by length of dactylus, propodus 10.4 times as long as broad, 4.1 times as long as dactylus; dactylus 3.3 times as long as wide (spines included), terminating in two claws, 3 or 4 accessory spines on its flexor margin. Fourth pereiopod reaching to end of scaphocerite, similar in form as third one. Fifth pereiopod (
|
||
<figureCitation id="136F2A03FC32FFAAFBF93507B6CA0843" box="[1138,1262,1628,1651]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Figs. 7E, F</figureCitation>
|
||
) reaching to end of scaphocerite, propodus 14 times as long as broad, 3.2 times as long as dactylus; dactylus 4.8 times as long as wide (spinules included), terminating in a single claw, with 42–49 spinules on its flexor margin.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFCA33447B1D10983" blockId="9.[808,1448,1820,1971]" pageId="9" pageNumber="60">
|
||
Endopod of male first pleopod (
|
||
<figureCitation id="136F2A03FC32FFAAFB143447B6DD0903" box="[1183,1273,1820,1843]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Fig. 7G</figureCitation>
|
||
) subtriangular, 2.1 times as long as wide, 0.4 times length of exopod, no appendix interna. Appendix masculina of male second pleopod (
|
||
<figureCitation id="136F2A03FC32FFAAFC053427B1C009A3" box="[910,996,1916,1939]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Fig. 7H</figureCitation>
|
||
) sub-cylindrical, spinose, reaching to 2/3 length of endopod.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC32FFAAFCA33487B76B0623" blockId="9.[808,1448,2012,2067]" pageId="9" pageNumber="60">
|
||
Uropodal diaeresis (
|
||
<figureCitation id="136F2A03FC32FFAAFB8F3487B66B09C3" box="[1028,1103,2012,2035]" captionStart="Fig" captionStartId="11.[109,144,2030,2051]" captionTargetBox="[182,1407,194,1985]" captionTargetId="figure-75@11.[140,1422,174,2016]" captionTargetPageId="11" captionText="Fig. 7. Edoneus marulas, new species: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC)." figureDoi="http://doi.org/10.5281/zenodo.5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="9" pageNumber="60">Fig. 7I</figureCitation>
|
||
) with 14–17 movable spinules, outer most one distinctly longer than lateral angle.
|
||
</paragraph>
|
||
<caption id="DF2B660EFC31FFA9FF0F34B7B654060B" ID-DOI="http://doi.org/10.5281/zenodo.5341545" ID-Zenodo-Dep="5341545" httpUri="https://zenodo.org/record/5341545/files/figure.png" pageId="10" pageNumber="61" startId="10.[132,167,2028,2049]" targetBox="[194,1406,193,1966]" targetPageId="10">
|
||
<paragraph id="8BEB3686FC31FFA9FF0F34B7B654060B" blockId="10.[132,1469,2028,2107]" pageId="10" pageNumber="61">
|
||
Fig. 6.
|
||
<taxonomicName id="4C544D05FC31FFA9FF4734B7B3520631" authority="Cai & Husana, 2009" authorityName="Cai & Husana" authorityYear="2009" box="[204,374,2028,2049]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="61" phylum="Arthropoda" rank="species" species="marulas" status="sp. nov.">
|
||
<emphasis id="B920EA94FC31FFA9FF4734B7B3520631" box="[204,374,2028,2049]" italics="true" pageId="10" pageNumber="61">Edoneus marulas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21357EFFC31FFA9FE0834B7B3D80631" box="[387,508,2028,2049]" pageId="10" pageNumber="61" rank="species">new species</taxonomicNameLabel>
|
||
: A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, preanal carina; E, scaphocerite; F, mandible; G, maxillula; H, maxilla; I, first maxilliped; J, second maxilliped; K, third maxilliped. Scale bars: A = 1 mm; B, E = 0.5 mm; C = 0.1 mm; D, F–K = 0.3 mm. (male, cl 3.9 mm, paratype, ZRC).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2B660EFC30FFA8FFE634B5B171060C" ID-DOI="http://doi.org/10.5281/zenodo.5341547" ID-Zenodo-Dep="5341547" httpUri="https://zenodo.org/record/5341547/files/figure.png" pageId="11" pageNumber="62" startId="11.[109,144,2030,2051]" targetBox="[182,1407,194,1985]" targetPageId="11">
|
||
<paragraph id="8BEB3686FC30FFA8FFE634B5B171060C" blockId="11.[109,1448,2030,2108]" pageId="11" pageNumber="62">
|
||
Fig. 7.
|
||
<taxonomicName id="4C544D05FC30FFA8FF3934B5B37F0633" authority="Cai & Husana, 2009" authorityName="Cai & Husana" authorityYear="2009" box="[178,347,2030,2051]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="62" phylum="Arthropoda" rank="species" species="marulas" status="sp. nov.">
|
||
<emphasis id="B920EA94FC30FFA8FF3934B5B37F0633" box="[178,347,2030,2051]" italics="true" pageId="11" pageNumber="62">Edoneus marulas</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A21357EFFC30FFA8FEEC34B5B3FB0633" box="[359,479,2030,2051]" pageId="11" pageNumber="62" rank="species">new species</taxonomicNameLabel>
|
||
: A, first pereiopod; B, second pereiopod; C, third pereiopod; D, dactylus of third pereiopod; E, fifth pereiopod; F, dactylus of fifth pereiopod; G, male first pleopod; H, male second pleopod; I, uropodal diaeresis. Scale bars: A–C, E = 0.5 mm; D, F, I = 0.1 mm; G, H = 0.2 mm; (male, cl 3.9 mm, paratype, ZRC).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEB3686FC37FFAFFF0F33E7B0910EE3" blockId="12.[132,693,188,211]" box="[132,693,188,211]" pageId="12" pageNumber="63">
|
||
Ovigerous female with eggs sized 0.90 ×
|
||
<quantity id="4CAC9B63FC37FFAFFDC633E7B0950EE3" box="[589,689,188,211]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="63" unit="mm" value="0.55">0.55 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB3686FC37FFAFFF0F33A7B07C0D03" blockId="12.[132,771,252,819]" pageId="12" pageNumber="63">
|
||
<emphasis id="B920EA94FC37FFAFFF0F33A7B2D70F24" bold="true" box="[132,243,252,276]" italics="true" pageId="12" pageNumber="63">Habitat. –</emphasis>
|
||
Marulas Cave is located in Barangay Tinandog, same area as the previous two caves visited. Of the three caves studied, this is the shortest one. Its coordinates are
|
||
<geoCoordinate id="EE605041FC37FFAFFF0F3207B33E0F43" box="[132,282,348,371]" degrees="13" direction="north" minutes="59" orientation="latitude" pageId="12" pageNumber="63" precision="1" seconds="40.6" value="13.994612">13º59'40.6"N</geoCoordinate>
|
||
<geoCoordinate id="EE605041FC37FFAFFEAF3207B3E30F43" box="[292,455,348,371]" degrees="121" direction="east" minutes="52" orientation="longitude" pageId="12" pageNumber="63" precision="1" seconds="23.2" value="121.873116">121º52'23.2"E</geoCoordinate>
|
||
,
|
||
<quantity id="4CAC9B63FC37FFAFFE5C3207B0370F43" box="[471,531,348,371]" metricMagnitude="1" metricUnit="m" metricValue="7.8" pageId="12" pageNumber="63" unit="m" value="78.0">78 m</quantity>
|
||
above sea level and about
|
||
<quantity id="4CAC9B63FC37FFAFFF433227B3260FA3" box="[200,258,380,403]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="63" unit="km" value="4.0">4 km</quantity>
|
||
away from the sea at its northeast. There is a dry creek on the surface that sinks to the cave entrance. Water is not present at the cave entrance until about
|
||
<quantity id="4CAC9B63FC37FFAFFD4132E7B1270FE3" box="[714,771,444,467]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="63" unit="m" value="36.0">36 m</quantity>
|
||
inside where it leaks (spring) from the rock crevices. The stairway-like morphology of the underground stream where pools could be found in each step provides refuge for the aquatic animals. There was also one ovigerous female of
|
||
<emphasis id="B920EA94FC37FFAFFF0F3107B3080C43" box="[132,300,604,627]" italics="true" pageId="12" pageNumber="63">
|
||
<taxonomicName id="4C544D05FC37FFAFFF0F3107B2CC0C43" box="[132,232,604,627]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Caridina</taxonomicName>
|
||
<typeStatus id="54EF8824FC37FFAFFF793107B3080C43" box="[242,300,604,627]" pageId="12" pageNumber="63">typus</typeStatus>
|
||
</emphasis>
|
||
H.
|
||
<bibRefCitation id="EFC54B77FC37FFAFFED63107B0720C43" author="Milne Edwards, H" box="[349,598,604,627]" pageId="12" pageNumber="63" pagination="1 - 532" refId="ref7736" refString="Milne Edwards, H., 1837, Histoire naturelle des Crustaces, comprenant l'anatomie, la physiologie et la classification de ces animaux, 2, 1 - 532, atlas, Pls. 1 - 14." type="journal article" year="1837">Milne Edwards, 1837</bibRefCitation>
|
||
, with carapace length
|
||
<quantity id="4CAC9B63FC37FFAFFF5B3127B30C0CA3" box="[208,296,636,659]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="12" pageNumber="63" unit="mm" value="8.3">8.3 mm</quantity>
|
||
, collected from lowest part of this cave in the present collection.
|
||
<emphasis id="B920EA94FC37FFAFFEFD31C7B0310C83" box="[374,533,668,691]" italics="true" pageId="12" pageNumber="63">
|
||
<taxonomicName id="4C544D05FC37FFAFFEFD31C7B3F20C83" box="[374,470,668,691]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="genus">Caridina</taxonomicName>
|
||
<typeStatus id="54EF8824FC37FFAFFE5631C7B0310C83" box="[477,533,668,691]" pageId="12" pageNumber="63">typus</typeStatus>
|
||
</emphasis>
|
||
is one of most widely distributed species in islands of the Indo-West Pacific and requires seawater to complete its life cycle (Cai, pers. obs.). The presence of
|
||
<emphasis id="B920EA94FC37FFAFFEB531A7B3BC0D23" box="[318,408,764,787]" italics="true" pageId="12" pageNumber="63">
|
||
C.
|
||
<typeStatus id="54EF8824FC37FFAFFEEB31A7B3BC0D23" box="[352,408,764,787]" pageId="12" pageNumber="63">typus</typeStatus>
|
||
</emphasis>
|
||
thus indicates that the stream in Marulas Cave has connection with the sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E650DFC37FFAFFF0F3007B2DE0D83" pageId="12" pageNumber="63" type="etymology">
|
||
<paragraph id="8BEB3686FC37FFAFFF0F3007B2DE0D83" blockId="12.[132,770,860,947]" pageId="12" pageNumber="63">
|
||
<emphasis id="B920EA94FC37FFAFFF0F3007B3300D44" bold="true" box="[132,276,860,884]" italics="true" pageId="12" pageNumber="63">Etymology. –</emphasis>
|
||
The species is named after the
|
||
<typeStatus id="54EF8824FC37FFAFFDF83007B0850D43" box="[627,673,860,883]" pageId="12" pageNumber="63">type</typeStatus>
|
||
locality- Marulas Cave in Luzon, the
|
||
<collectingCountry id="F3437616FC37FFAFFE343027B01F0DA3" box="[447,571,892,915]" name="Philippines" pageId="12" pageNumber="63">Philippines</collectingCountry>
|
||
. Name used as an apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E650DFC37FFAFFF0F3087B31E0AC3" pageId="12" pageNumber="63" type="discussion">
|
||
<paragraph id="8BEB3686FC37FFAFFF0F3087B31E0AC3" blockId="12.[132,771,988,1267]" pageId="12" pageNumber="63">
|
||
<emphasis id="B920EA94FC37FFAFFF0F3087B2D80DC4" bold="true" box="[132,252,988,1012]" italics="true" pageId="12" pageNumber="63">Remarks. –</emphasis>
|
||
With respect to the rostrum and the eyes,
|
||
<taxonomicName id="4C544D05FC37FFAFFD233087B2FF0A23" authorityName="Cai & Husana" authorityYear="2009" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="marulas">
|
||
<emphasis id="B920EA94FC37FFAFFD233087B2FF0A23" italics="true" pageId="12" pageNumber="63">Edoneus marulas</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C544D05FC37FFAFFEE030A7B3C70A23" authorityName="Holthuis" authorityYear="1978" box="[363,483,1020,1043]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="atheatus">
|
||
<emphasis id="B920EA94FC37FFAFFEE030A7B3C70A23" box="[363,483,1020,1043]" italics="true" pageId="12" pageNumber="63">E. atheatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D05FC37FFAFFD9130A7B04D0A23" authorityName="Cai & Husana" authorityYear="2009" box="[538,617,1020,1043]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sketi">
|
||
<emphasis id="B920EA94FC37FFAFFD9130A7B04D0A23" box="[538,617,1020,1043]" italics="true" pageId="12" pageNumber="63">E. sketi</emphasis>
|
||
</taxonomicName>
|
||
. But the stout carpus of the first pereiopod (carpus 1.6 times as long as high vs. 2.0 times in
|
||
<taxonomicName id="4C544D05FC37FFAFFEE13767B3C70A63" authorityName="Holthuis" authorityYear="1978" box="[362,483,1084,1107]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="atheatus">
|
||
<emphasis id="B920EA94FC37FFAFFEE13767B3C70A63" box="[362,483,1084,1107]" italics="true" pageId="12" pageNumber="63">E. atheatus</emphasis>
|
||
</taxonomicName>
|
||
, and 2.4 times in
|
||
<taxonomicName id="4C544D05FC37FFAFFD223767B0DE0A63" authorityName="Cai & Husana" authorityYear="2009" box="[681,762,1084,1107]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="sketi">
|
||
<emphasis id="B920EA94FC37FFAFFD223767B0DE0A63" box="[681,762,1084,1107]" italics="true" pageId="12" pageNumber="63">E. sketi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EA94FC37FFAFFD723767B1260A64" bold="true" box="[761,770,1084,1108]" pageId="12" pageNumber="63">)</emphasis>
|
||
can easily differentiate it from the latter two species. With regard to the form of the second pereiopod,
|
||
<taxonomicName id="4C544D05FC37FFAFFDE43727B0C30AA3" authorityName="Cai & Husana" authorityYear="2009" box="[623,743,1148,1171]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="marulas">
|
||
<emphasis id="B920EA94FC37FFAFFDE43727B05B0AA3" box="[623,639,1148,1171]" italics="true" pageId="12" pageNumber="63">E</emphasis>
|
||
.
|
||
<emphasis id="B920EA94FC37FFAFFD1B3727B0C30AA3" box="[656,743,1148,1171]" italics="true" pageId="12" pageNumber="63">marulas</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C544D05FC37FFAFFF7C37C7B3780A83" authorityName="Cai & Husana" authorityYear="2009" box="[247,348,1180,1203]" class="Malacostraca" family="Atyidae" genus="Edoneus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="63" phylum="Arthropoda" rank="species" species="erwini">
|
||
<emphasis id="B920EA94FC37FFAFFF7C37C7B3230A83" box="[247,263,1180,1203]" italics="true" pageId="12" pageNumber="63">E</emphasis>
|
||
.
|
||
<emphasis id="B920EA94FC37FFAFFE9337C7B3780A83" box="[280,348,1180,1203]" italics="true" pageId="12" pageNumber="63">erwini</emphasis>
|
||
</taxonomicName>
|
||
, but can be distinguished by its short and unarmed rostrum, and the strongly reduced eyes (vs. eyes developed).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |