141 lines
18 KiB
XML
141 lines
18 KiB
XML
<document id="5C3C10519FC31FF6A33C9E4976C14CCB" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFDEFFFFFCBEFF7A5D4F39AA" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) bicandida Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="15" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="15" updateTime="1700168056660" updateUser="ExternalLinkService">
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<mods:title id="7E63380ACEF52F75A878F0138DA344C8">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="6A3BAEB48B270EE0C607E33A130D9A99">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="C053ABB89F34929511C17E473B1C3977">text</mods:typeOfResource>
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<mods:title id="B3AFC6BE3CA0E455FDC9808AF766D2D2">Zootaxa</mods:title>
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<mods:part id="7FC1F85CB4F985EB85704A3EA10B18A3">
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<mods:date id="08BAFEDE83632EB2B4BC68A7CF5ACB23">2023</mods:date>
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<mods:number id="87F5AE4E7EA28F7957F0C9D6920DCD6A">2023-11-07</mods:number>
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<mods:number id="FA5E6A041CBFB383958FBD43478FAC3E">5367</mods:number>
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<mods:url id="F2702849B2483C71C31230D96BE542A9">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
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<mods:identifier id="4B9F5429A67709ABA1356256F0771C91" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
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<mods:identifier id="CE8A04B201C33878E347002EC4EE4453" type="ISSN">1175-5334</mods:identifier>
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<subSubSection id="C34E6516FFDEFFFFFCBEFF7A5B813CCE" box="[151,750,223,250]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BEB369DFFDEFFFFFCBEFF7A5B813CCE" blockId="14.[151,764,223,286]" box="[151,750,223,250]" pageId="14" pageNumber="15">
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<heading id="D0A381F1FFDEFFFFFCBEFF7A5B813CCE" bold="true" box="[151,750,223,250]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
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<emphasis id="B920EA8FFFDEFFFFFCBEFF7A5B813CCE" bold="true" box="[151,750,223,250]" pageId="14" pageNumber="15">
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<taxonomicName id="4C544D1EFFDEFFFFFCBEFF7A5B203CCD" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,591,223,250]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bicandida" status="sp. nov." subGenus="Hydraenopsis">
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<emphasis id="B920EA8FFFDEFFFFFCBEFF7A5B203CCD" bold="true" box="[151,591,223,250]" italics="true" pageId="14" pageNumber="15">Hydraena (Hydraenopsis) bicandida</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21357F4FFDEFFFFFE77FF7A5B813CCE" box="[606,750,223,249]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCBEFEA65B933D2A" box="[151,764,259,286]" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BEB369DFFDEFFFFFCBEFEA65B933D2A" blockId="14.[151,764,223,286]" box="[151,764,259,286]" pageId="14" pageNumber="15">
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<figureCitation id="136F2A18FFDEFFFFFCBEFEA6598C3D2A" box="[151,227,259,285]" captionStart="FIGURE 1" captionStartId="5.[151,250,1867,1891]" captionTargetBox="[221,1366,181,1844]" captionTargetId="figure-18@5.[221,1366,181,1844]" captionTargetPageId="5" captionText="FIGURE 1. Dorsal habitus of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.10145108" httpUri="https://zenodo.org/record/10145108/files/figure.png" pageId="14" pageNumber="15">Figs. 1</figureCitation>
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(habitus comparison), 7 (habitus and aedeagus)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCBEFEEE5B463DBE" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8BEB369DFFDEFFFFFCBEFEEE5B463DBE" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">
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<emphasis id="B920EA8FFFDEFFFFFCBEFEEE58273D52" bold="true" box="[151,328,331,357]" pageId="14" pageNumber="15">Type Material.</emphasis>
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<materialsCitation id="3B3C3CC0FFDEFFFFFD66FEEE5B463DBE" collectionCode="FMNH" collectorName="A. Newton" country="Mexico" location="Temascaltepec" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
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<emphasis id="B920EA8FFFDEFFFFFD66FEEE58D33D52" bold="true" box="[335,444,331,357]" pageId="14" pageNumber="15">
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<typeStatus id="54EF883FFFDEFFFFFD66FEEE58D33D52" box="[335,444,331,357]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male): “
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<collectingCountry id="F343760DFFDEFFFFFE0DFEEE5BF63D52" box="[548,665,331,357]" name="Mexico" pageId="14" pageNumber="15">MEXICO</collectingCountry>
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:
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<collectingRegion id="4990F87FFFDEFFFFFE8AFEEE5B963D52" box="[675,761,331,357]" country="Mexico" name="Mexico" pageId="14" pageNumber="15">Mexico</collectingRegion>
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<quantity id="4CAC9B78FFDEFFFFFF29FEE95A5C3D52" box="[768,819,331,358]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="14" pageNumber="15" unit="mi" value="7.0">7 mi</quantity>
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NE
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<location id="8E8B6046FFDEFFFFFF4CFEEE5D633D52" LSID="urn:lsid:plazi:treatment:03FD878BFFDEFFFFFCBEFF7A5D4F39AA:8E8B6046FFDEFFFFFF4CFEEE5D633D52" box="[869,1036,331,357]" country="Mexico" name="Temascaltepec" pageId="14" pageNumber="15" stateProvince="Mexico">Temascaltepec</location>
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, 7000’ ix.12–15.1973 // under moss on rock mass
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<collectorName id="26A1534BFFDEFFFFFD19FECA58C13DBE" box="[304,430,367,393]" pageId="14" pageNumber="15">A. Newton</collectorName>
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” (
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<collectionCode id="ED45AE58FFDEFFFFFDEFFECA5B763DBE" box="[454,537,367,393]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="14" pageNumber="15" type="Museum">FMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCEEFE3658633E56" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BEB369DFFDEFFFFFCEEFE3658633E56" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">
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<emphasis id="B920EA8FFFDEFFFFFCEEFE3658BD3D9A" bold="true" box="[199,466,403,429]" pageId="14" pageNumber="15">Differential Diagnosis.</emphasis>
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This new species is a member of the
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<taxonomicName id="4C544D1EFFDEFFFFFF59FE315D023D9A" authority="Subgroup" authorityName="Subgroup (Perkins" authorityYear="1980" box="[880,1133,403,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="argutipes">
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<emphasis id="B920EA8FFFDEFFFFFF59FE315A953D9A" box="[880,1018,404,429]" italics="true" pageId="14" pageNumber="15">H. argutipes</emphasis>
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Subgroup
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</taxonomicName>
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of the
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<taxonomicName id="4C544D1EFFDEFFFFF893FE315C713D9A" box="[1210,1310,403,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="leechi">
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<emphasis id="B920EA8FFFDEFFFFF893FE315C713D9A" box="[1210,1310,403,429]" italics="true" pageId="14" pageNumber="15">H. leechi</emphasis>
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</taxonomicName>
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Group; the subgroup is previously known from nine species (see
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<bibRefCitation id="EFC54B6CFFDEFFFFFED0FE125AD53DE6" author="Perkins, P. D." box="[761,954,439,465]" pageId="14" pageNumber="15" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 37</bibRefCitation>
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, 151). Based on the aedeagal morphology, this species is clearly related to
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<taxonomicName id="4C544D1EFFDEFFFFFDD1FE795A713DC2" authority="Perkins 1980" authorityName="Perkins" authorityYear="1980" box="[504,798,475,501]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="argutipes">
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<emphasis id="B920EA8FFFDEFFFFFDD1FE795BEC3DC2" box="[504,643,476,501]" italics="true" pageId="14" pageNumber="15">H. argutipes</emphasis>
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<bibRefCitation id="EFC54B6CFFDEFFFFFEA0FE7E5A713DC2" author="Perkins, P. D." box="[649,798,475,501]" pageId="14" pageNumber="15" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980</bibRefCitation>
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</taxonomicName>
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. It is recognized by the combination of the large, brightly shining metaventral plaques, the deep pronotal foveae, the lack of a pronotal scintilla, and the aedeagus (
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<figureCitation id="136F2A18FFDEFFFFF961FE5A5CE13E2E" box="[1352,1422,511,537]" captionStart="FIGURE 7" captionStartId="13.[151,250,1792,1816]" captionTargetBox="[320,1267,181,1768]" captionTargetId="figure-18@13.[320,1267,181,1768]" captionTargetPageId="13" captionText="FIGURE 7. Hydraena (Hydraenopsis) bicandida new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145120" httpUri="https://zenodo.org/record/10145120/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
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). Reliable determinations will require dissection of males to separate this species from others in the
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<taxonomicName id="4C544D1EFFDEFFFFF927FD8158633E56" authority="Subgroup." authorityName="Subgroup (Perkins" authorityYear="1980" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="argutipes">
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<emphasis id="B920EA8FFFDEFFFFF927FD815CF33E0A" box="[1294,1436,548,573]" italics="true" pageId="14" pageNumber="15">H. argutipes</emphasis>
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Subgroup.
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCEEFDCE5A993962" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BEB369DFFDEFFFFFCEEFDCE5AE13F6A" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">
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<emphasis id="B920EA8FFFDEFFFFFCEEFDCE583E3EB2" bold="true" box="[199,337,619,645]" pageId="14" pageNumber="15">Description</emphasis>
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. Size:
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<typeStatus id="54EF883FFFDEFFFFFD8DFDCE5B693EB2" box="[420,518,619,645]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.38/0.60; head width 0.36; pronotum 0.33/0.48, PA 0.38, PB 0.42; elytra 0.87/0.60. Habitus as illustrated (
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<figureCitation id="136F2A18FFDEFFFFF829FD2A5D2C3E9E" box="[1024,1091,655,681]" captionStart="FIGURE 7" captionStartId="13.[151,250,1792,1816]" captionTargetBox="[320,1267,181,1768]" captionTargetId="figure-18@13.[320,1267,181,1768]" captionTargetPageId="13" captionText="FIGURE 7. Hydraena (Hydraenopsis) bicandida new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145120" httpUri="https://zenodo.org/record/10145120/files/figure.png" pageId="14" pageNumber="15">Fig. 7</figureCitation>
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). Dorsum dark brown to black, palpi brown, tip of last palpomere darker. Dorsum of head and pronotum moderately coarsely moderately densely punctulate, interstices shining. Punctures of frons disc ca. 1xef, interstices 1–2xpd on disc, shining, lateral areas microreticulate, dull; punctures of clypeus very fine, lateral areas microreticulate. Punctures of pronotal disc deep, ca. 2–3xef, interstices on pronotal disc ca. 1–2xpd. All dorsal punctures with very indistince, very short recumbent seta. Labrum apicomedially excised, free margins weakly upturned.
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</paragraph>
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<paragraph id="8BEB369DFFDEFFFFFCEEFCCD5DD73F92" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">Pronotum with anterior margin very weakly emarginate over middle ½, without scintilla; anterolateral, posterolateral and posteromedial foveae deep; anterior part of disc with transverse impression.</paragraph>
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<paragraph id="8BEB369DFFDEFFFFFCEEFC0A5BE5386E" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">Elytra with summit of posterior declivity slightly past midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd of largest pronotal punctures, or slightly larger, punctures becoming gradually smaller toward posterior; most punctures with minute granule at anterior margin. Intervals not raised, width ca. 1xpd, as are interstices between punctures of a row.Apices in dorsal aspect conjointly, rather sharply rounded, in posterior aspect margins not forming angle with one another.</paragraph>
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<paragraph id="8BEB369DFFDEFFFFFCEEFBC15A993962" blockId="14.[151,1437,331,1438]" pageId="14" pageNumber="15">Venter: Mentum very sparsely very finely punctulate, shining; postmentum very finely densely micropunctulate in median concavity, surrounding areas smooth, shining. Genae raised, shining, without posterior ridge. Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/2/5/2. P1 laminate; median carina sinuate in profile. P2 l/w ca. 4/1, sides slightly converging toward blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques large, brightly shining, parallel, slightly raised, narrowly separated by slightly less than plaque width. AIS width at straight posterior margin ca. 2x P2. Protibia slightly arcuate. Mesotibia straight, slender. Metatibia slightly widened over distal 2/3. Abdominal apex symmetrical; last tergite with apicomedian notch.</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCEEFAFA5A32394E" box="[199,861,1375,1402]" pageId="14" pageNumber="15" type="distribution">
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<paragraph id="8BEB369DFFDEFFFFFCEEFAFA5A32394E" blockId="14.[151,1437,331,1438]" box="[199,861,1375,1402]" pageId="14" pageNumber="15">
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<emphasis id="B920EA8FFFDEFFFFFCEEFAFA580E394E" bold="true" box="[199,353,1375,1401]" pageId="14" pageNumber="15">Distribution.</emphasis>
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Currently known only from the
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<typeStatus id="54EF883FFFDEFFFFFEE5FAC55B93394D" box="[716,764,1376,1402]" pageId="14" pageNumber="15">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFDEFFFFFCEEFA265D4F39AA" box="[199,1056,1411,1438]" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="8BEB369DFFDEFFFFFCEEFA265D4F39AA" blockId="14.[151,1437,331,1438]" box="[199,1056,1411,1438]" pageId="14" pageNumber="15">
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<emphasis id="B920EA8FFFDEFFFFFCEEFA26582139AA" bold="true" box="[199,334,1411,1437]" pageId="14" pageNumber="15">Etymology.</emphasis>
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Named in reference to the brightly shining metaventral plaques.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |